treatments-xml/data/03/BF/A7/03BFA7558B6EFF30FEA6FEA88C1F427B.xml
2024-06-21 12:22:17 +02:00

168 lines
19 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D1FA726C73D6F312C0CFE382CD448CD5" ID-DOI="10.5281/zenodo.170622" ID-GBIF-Dataset="ae589893-7091-4d5e-a8a3-7d42291d490e" ID-ISSN="1175­5326" ID-Zenodo-Dep="170622" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459714173681" checkinUser="plazi" docAuthor="Campos, Luiz Alexandre" docDate="2005" docId="03BFA7558B6EFF30FEA6FEA88C1F427B" docLanguage="en" docName="zt00820.pdf" docOrigin="Zootaxa 820" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cromata graziae Campos, 2005, sp. nov." docType="treatment" docVersion="10" lastPageNumber="5" masterDocId="FF86DF2D8B6CFF34FFAEFFB48D02444D" masterDocTitle="A new species of Cromata Rolston from Brazilian and Venezuelan Amazon Forest (Hemiptera: Pentatomidae: Discocephalinae)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="3" updateTime="1698207528039" updateUser="plazi">
<mods:mods id="6B344CD6951C54F0570A8A0D49D22B5C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="168DECB8B90B3A4AC716253F78DEF2CB">
<mods:title id="9A930324321B9B4B91FB0C6C8222279C">A new species of Cromata Rolston from Brazilian and Venezuelan Amazon Forest (Hemiptera: Pentatomidae: Discocephalinae)</mods:title>
</mods:titleInfo>
<mods:name id="522E2FA089201C0C010E209709990727" type="personal">
<mods:role id="1B5DBC02ABCA8573197FA5CEF64FD9F9">
<mods:roleTerm id="5AE23AD3783788195AE6A3DC87410525">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6780262E99DB2D0E876DC1825FB12AF1">Campos, Luiz Alexandre</mods:namePart>
</mods:name>
<mods:typeOfResource id="9C26545E37B9A705108632C2DF6655A8">text</mods:typeOfResource>
<mods:relatedItem id="21A9AFDD470628E168772EDD62B1FD81" type="host">
<mods:titleInfo id="39E0006CD0AB60AE862D86FBF53518EB">
<mods:title id="7735BB781B4E387E5E0D62AF99CA4CBD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D51CC1FC76C688BCEFC6DBCA88F5C004">
<mods:date id="16EDAA90593279DCCE592F2EAEC90D81">2005</mods:date>
<mods:detail id="2C13B65E99076D8BB4D07C92403E4B07" type="volume">
<mods:number id="999CD81ED2DA69C7CF5A9B6D773A2548">820</mods:number>
</mods:detail>
<mods:extent id="128B446C74B32A6EDDD95F7163E39CD0" unit="page">
<mods:start id="797CDAFF7C3230403A00C35749060729">1</mods:start>
<mods:end id="7F895896AFD05E5F1890D4DB28899575">6</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F56DA9BCFEFFF9BB126AE92DFC4FC7F8">journal article</mods:classification>
<mods:identifier id="AD59E69EC00B28101F66953996FDE455" type="DOI">10.5281/zenodo.170622</mods:identifier>
<mods:identifier id="7D087893B5DE5E43B23521854F79BA88" type="GBIF-Dataset">ae589893-7091-4d5e-a8a3-7d42291d490e</mods:identifier>
<mods:identifier id="88E9AA29231D886091C6B58D6932C2CE" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="82866AC0B343177C61DC42D2A265E515" type="Zenodo-Dep">170622</mods:identifier>
</mods:mods>
<treatment id="03BFA7558B6EFF30FEA6FEA88C1F427B" ID-DOI="http://doi.org/10.5281/zenodo.6266661" ID-GBIF-Taxon="119334179" ID-Zenodo-Dep="6266661" LSID="urn:lsid:zoobank.org:act:7D2F5EA2-57D1-4AE5-B3BF-CC487EEFAEA6" httpUri="http://treatment.plazi.org/id/03BFA7558B6EFF30FEA6FEA88C1F427B" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="C30C45C88B6EFF36FEA6FEA88C8A4514" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BA916438B6EFF36FEA6FEA88F3E4578" blockId="2.[264,572,283,345]" box="[264,572,283,310]" pageId="2" pageNumber="3">
<heading id="D0E1A12F8B6EFF36FEA6FEA88F3E4578" bold="true" box="[264,572,283,310]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B962CA518B6EFF36FEA6FEA88F3E4578" bold="true" box="[264,572,283,310]" pageId="2" pageNumber="3">
<taxonomicName id="4C166DC08B6EFF36FEA6FEA88CD3457B" box="[264,465,284,310]" class="Insecta" family="Pentatomidae" genus="Cromata" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B962CA518B6EFF36FEA6FEA88CD3457B" bold="true" box="[264,465,284,310]" italics="true" pageId="2" pageNumber="3">Cromata graziae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A251772A8B6EFF36FE71FEAF8F3E4578" box="[479,572,283,309]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA916438B6EFF36FEA6FE8B8C8A4514" blockId="2.[264,572,283,345]" box="[264,392,319,345]" pageId="2" pageNumber="3">
(
<figureCitation id="132D0AC68B6EFF36FEBFFE8B8C7D4514" box="[273,383,319,345]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 6" captionStartId-0="2.[264,368,1882,1906]" captionStartId-1="3.[264,383,1738,1762]" captionTargetBox-1="[297,1287,589,1710]" captionTargetId-1="figure@3.[296,1291,575,1718]" captionTargetPageId-1="3" captionText-0="FIGURE 1. Cromata graziae, sp. nov., holotype male, dorsal." captionText-1="FIGURES 2 6. Cromata graziae, sp. nov., female and male genitalia. Fig. 2, female genital plates, ventroposterior view. (gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; la 8 = laterotergites 8; la 9 = laterotergites 9; sp = spiracle). Figs. 3 6, pygophore, ventral, dorsal, posterior and lateral view respectively. (c = carina; dp = dorsal process over pygophore; fr = hair fringes; p = posterolateral angle; pr = proctiger; vr = ventral rim)." httpUri-1="https://zenodo.org/record/170623/files/figure.png" pageId="2" pageNumber="3">Figs. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C45C88B6EFF36FEA6FE3B8E8245B4" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BA916438B6EFF36FEA6FE3B8C8745E5" blockId="2.[264,1323,399,505]" box="[264,389,399,424]" pageId="2" pageNumber="3">
<emphasis id="B962CA518B6EFF36FEA6FE3B8C8745E5" box="[264,389,399,424]" italics="true" pageId="2" pageNumber="3">Etymology</emphasis>
</paragraph>
<paragraph id="8BA916438B6EFF36FE96FE038E8245B4" blockId="2.[264,1323,399,505]" pageId="2" pageNumber="3">The species is named in honor to Dr. Jocélia Grazia, for her great and valuable contributions to the knowledge of Neotropical pentatomids.</paragraph>
</subSubSection>
<subSubSection id="C30C45C88B6EFF36FEA6FDE38FEF47C4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BA916438B6EFF36FEA6FDE38C6D463D" blockId="2.[264,1325,599,905]" box="[264,367,599,624]" pageId="2" pageNumber="3">
<emphasis id="B962CA518B6EFF36FEA6FDE38C6D463D" box="[264,367,599,624]" italics="true" pageId="2" pageNumber="3">Material</emphasis>
</paragraph>
<paragraph id="8BA916438B6EFF36FE96FDCB8FEF47C4" blockId="2.[264,1325,599,905]" pageId="2" pageNumber="3">
<typeStatus id="54ADA8E18B6EFF36FE96FDCB8CA746D4" box="[312,421,639,665]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
male,
<emphasis id="B962CA518B6EFF36FE58FDCA8E1846D5" bold="true" box="[502,794,638,664]" pageId="2" pageNumber="3">
<collectingCountry id="F30156D38B6EFF36FE58FDCA8FAC46D5" box="[502,686,638,664]" name="Venezuela" pageId="2" pageNumber="3">VENEZUELA</collectingCountry>
, Bolivar
</emphasis>
: El Bochinche, with the following labels: a)
<collectingCountry id="F30156D38B6EFF36FEA6FD138C86468C" box="[264,388,679,705]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
, Bolivar, El Bochinche, Res. Forestal Imataca.
<specimenCount id="9D10DDCA8B6EFF36FC6CFD13890E468C" box="[962,1036,679,705]" pageId="2" pageNumber="3" type="generic">
<quantity id="4CEEBBA68B6EFF36FC6CFD13890A468C" box="[962,1032,679,705]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="m" value="200.0">200m</quantity>
.
</specimenCount>
<date id="FFA830838B6EFF36FBB8FD1389A8468C" box="[1046,1194,679,705]" pageId="2" pageNumber="3" value="1974-12-06" valueMax="1974-12-13" valueMin="1974-12-06">613­XII­74</date>
[1974]; b) Expedicion Instituto Zoologia Agrícola Fac. Agronomia. U.C.V. Deposited in the Museo del Instituto de Zoología Agrícola Francisco Fernández Yépez.
<typeStatus id="54ADA8E18B6EFF36FBBEFD438974475C" box="[1040,1142,759,785]" pageId="2" pageNumber="3" type="paratype">Paratype</typeStatus>
female,
<emphasis id="B962CA518B6EFF36FB4DFD428CD74775" bold="true" pageId="2" pageNumber="3">
<collectionCode id="ED078E868B6EFF36FB4DFD428829475D" box="[1251,1323,758,784]" pageId="2" pageNumber="3">BRA­</collectionCode>
<collectionCode id="ED078E868B6EFF36FEA6FCAA8C3E4775" box="[264,316,798,824]" httpUri="http://grbio.org/cool/74rs-wjen" name="Academy of Sciences, Zoological Institute" pageId="2" pageNumber="3">ZIL</collectionCode>
, Amazonas
</emphasis>
: Coari, with the following labels: a) BR, AM, Coari, rio Urucu
<geoCoordinate id="EE2270848B6EFF36FEA6FCF38C9A472C" box="[264,408,839,865]" direction="south" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="-4.8231664">4°4923.4”S</geoCoordinate>
/
<geoCoordinate id="EE2270848B6EFF36FE0CFCF38F3D472C" box="[418,575,839,865]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="-64.917305">64°552.3”W</geoCoordinate>
<date id="FFA830838B6EFF36FDE6FCF38FFC472C" box="[584,766,839,865]" pageId="2" pageNumber="3" value="1991-04-02" valueMax="1991-04-08" valueMin="1991-04-02">0208/IV/1991</date>
, N.O. Aguiar
<emphasis id="B962CA518B6EFF36FC06FCF38EDF472D" box="[936,989,839,864]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. col. Deposited in the Instituto Nacional de Pesquisas da Amazônia.
</paragraph>
</subSubSection>
<subSubSection id="C30C45C88B6EFF30FEA6FC0B8CF640BB" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="8BA916438B6EFF36FEA6FC0B8CB04795" blockId="2.[264,1322,959,1105]" box="[264,434,959,984]" pageId="2" pageNumber="3">
<emphasis id="B962CA518B6EFF36FEA6FC0B8CB04795" box="[264,434,959,984]" italics="true" pageId="2" pageNumber="3">Measurements</emphasis>
</paragraph>
<paragraph id="8BA916438B6EFF36FE96FC528EDA401C" blockId="2.[264,1322,959,1105]" pageId="2" pageNumber="3">
<emphasis id="B962CA518B6EFF36FE96FC528C98404D" bold="true" box="[312,410,998,1024]" pageId="2" pageNumber="3">Female.</emphasis>
Body length 9.12; abdominal width 5.44; head length 1.44; head width 1.84; pronotum length 2.00; pronotum width 5.04; scutellar length 3.68; scutellar width 2.88; length of antennal segments: I 0.64; II 0.56; III 0.88; IV 1.12.
</paragraph>
<caption id="DF6946CB8B6EFF36FEA6F8EE8EA2433C" box="[264,928,1882,1906]" pageId="2" pageNumber="3">
<paragraph id="8BA916438B6EFF36FEA6F8EE8EA2433C" blockId="2.[264,928,1882,1906]" box="[264,928,1882,1906]" pageId="2" pageNumber="3">
<emphasis id="B962CA518B6EFF36FEA6F8EE8C88433F" bold="true" box="[264,394,1882,1906]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
<taxonomicName id="4C166DC08B6EFF36FE3FF8EE8F47433C" box="[401,581,1882,1905]" class="Insecta" family="Pentatomidae" genus="Cromata" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B962CA518B6EFF36FE3FF8EE8F47433C" box="[401,581,1882,1905]" italics="true" pageId="2" pageNumber="3">Cromata graziae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A251772A8B6EFF36FDFDF8EE8FAA433C" box="[595,680,1882,1905]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male, dorsal.
</paragraph>
</caption>
<paragraph id="8BA916438B6FFF37FE96FEAF893A45CC" blockId="3.[264,1323,283,385]" pageId="3" pageNumber="4">
<emphasis id="B962CA518B6FFF37FE96FEAF8C7C4578" bold="true" box="[312,382,283,309]" pageId="3" pageNumber="4">Male.</emphasis>
Body length 8.88; abdominal width 5.12; head length 1.60; head width 2.00; pronotum length 1.92; pronotum width 4.72; scutellar length 3.60; scutellar width 3.04; length of antennal segments: I 0.64; II 0.60; III 0.88; IV 1.04; V 1.52.
</paragraph>
<paragraph id="8BA916438B6FFF37FEA6FE1B8C904585" blockId="3.[264,1320,431,537]" box="[264,402,431,456]" pageId="3" pageNumber="4">
<emphasis id="B962CA518B6FFF37FEA6FE1B8C904585" box="[264,402,431,456]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
</paragraph>
<paragraph id="8BA916438B6FFF37FE96FE638ED94654" blockId="3.[264,1320,431,537]" pageId="3" pageNumber="4">Dorsal surface brown to light brown densely punctured in darker punctures (fig. 1). Ventral surface punctured in concolorous or darker punctures.</paragraph>
<caption id="DF6946CB8B6FFF37FEA6F97E8F464326" httpUri="https://zenodo.org/record/170623/files/figure.png" pageId="3" pageNumber="4" targetBox="[297,1287,589,1710]" targetPageId="3">
<paragraph id="8BA916438B6FFF37FEA6F97E8F464326" blockId="3.[264,1323,1738,1899]" pageId="3" pageNumber="4">
<emphasis id="B962CA518B6FFF37FEA6F97E8CB642AF" bold="true" box="[264,436,1738,1762]" pageId="3" pageNumber="4">FIGURES 26.</emphasis>
<taxonomicName id="4C166DC08B6FFF37FE15F97E8F6D42AC" box="[443,623,1738,1761]" class="Insecta" family="Pentatomidae" genus="Cromata" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B962CA518B6FFF37FE15F97E8F6D42AC" box="[443,623,1738,1761]" italics="true" pageId="3" pageNumber="4">Cromata graziae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A251772A8B6FFF37FDD2F97E8FD342AC" box="[636,721,1738,1761]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
, female and male genitalia. Fig. 2, female genital plates, ventroposterior view. (gc8 = gonocoxites 8; gc9 = gonocoxites 9; la8 = laterotergites 8; la9 = laterotergites 9; sp = spiracle). Figs. 36, pygophore, ventral, dorsal, posterior and lateral view respectively. (c = carina; dp = dorsal process over pygophore; fr = hair fringes; p = posterolateral angle; pr = proctiger; vr = ventral rim).
</paragraph>
</caption>
<paragraph id="8BA916438B68FF30FE96FEA88CDE45B3" blockId="4.[264,1323,284,1270]" pageId="4" pageNumber="5">Head with few punctures on vertex and juga; ventral surface black, except apex of juga and anterior margin of buccula yellowish. Central disk of tylus swollen. Lateral margins of juga sinuous and emarginated; juga slightly surpassing tylus. Ocelli near posterolateral margin of head, yellowish. Antennal segments light brown, except basal third of segment V yellowish. Labrum rufous, labium yellowish; apical segment of rostrum attaining base of urosternite VII.</paragraph>
<paragraph id="8BA916438B68FF30FE96FDB889A94773" blockId="4.[264,1323,284,1270]" pageId="4" pageNumber="5">Pronotum with punctures two to four times larger than on head; disc bearing a large callose yellow area with few punctures; posterior to this area punctures dense and confluent; cicatrices inconspicuous. Anterolateral margins slightly sinuous; humeral angles rufous. Legs dark brown, except tarsi and basal half of femora yellowish, distal fourth of femora and base of tibia rufous. Scutellum with dense and confluent punctures, these decreasing in size toward apex; shallow, black fovea at basal angles of scutellum. Hemelytra finely, densely punctured, with 1+1 yellowish spots near the apex of radial vein; membranes dark brown. Connexival segments bearing a mesial orange macula.</paragraph>
<paragraph id="8BA916438B68FF30FE96FCF88821472B" blockId="4.[264,1323,284,1270]" box="[312,1315,844,870]" pageId="4" pageNumber="5">Abdominal venter dark brown, densely punctured laterad to unpunctured central disc.</paragraph>
<paragraph id="8BA916438B68FF30FE96FCC08CBC404B" blockId="4.[264,1323,284,1270]" pageId="4" pageNumber="5">
<emphasis id="B962CA518B68FF30FE96FCC08C9847C3" bold="true" box="[312,410,884,910]" pageId="4" pageNumber="5">Female.</emphasis>
Genital plates (fig. 2). Laterotergites 8 (la8) trapezoidal, posterior margin slightly convex; posterior margin of gonocoxites 8 (gc8) strongly convex, produced caudad over laterotergites 9 (la9). Laterotergites 9 not attaining the medial band uniting laterotergites 8.
</paragraph>
<paragraph id="8BA916438B68FF30FE96FBA08CF640BB" blockId="4.[264,1323,284,1270]" pageId="4" pageNumber="5">
<emphasis id="B962CA518B68FF30FE96FBA08C7C4063" bold="true" box="[312,382,1044,1070]" pageId="4" pageNumber="5">Male.</emphasis>
Pygophore (figs. 36). Ventral rim (vr) tumescent, u­shaped from ventral view and v­shaped from caudal view. With slight, mesial carina (c) projecting downwards from ventral rim forming a “y” mark over posterior surface of pygophore. Posterolateral angles (p) rounded, excavated, posteriorly projected, bearing subapical hair fringes (fr). Proctiger (pr) large, truncate at apex, covering most of pygophore; 1+1 hair fringes over dorsolateral surface of proctiger.
</paragraph>
</subSubSection>
<subSubSection id="C30C45C88B68FF30FEA6FA988C1F427B" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BA916438B68FF30FEA6FA988C814108" blockId="4.[264,1323,1324,1590]" box="[264,387,1324,1349]" pageId="4" pageNumber="5">
<emphasis id="B962CA518B68FF30FEA6FA988C814108" box="[264,387,1324,1349]" italics="true" pageId="4" pageNumber="5">Comments</emphasis>
</paragraph>
<paragraph id="8BA916438B68FF30FE96FAE08C1F427B" blockId="4.[264,1323,1324,1590]" pageId="4" pageNumber="5">
<taxonomicName id="4C166DC08B68FF30FE96FAE08CB14120" box="[312,435,1364,1389]" class="Insecta" family="Pentatomidae" genus="Cromata" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B962CA518B68FF30FE96FAE08CB14120" box="[312,435,1364,1389]" italics="true" pageId="4" pageNumber="5">C. graziae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A251772A8B68FF30FE6CFAE08F1F4123" box="[450,541,1364,1390]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C166DC08B68FF30FD0CFAE08E114120" box="[674,787,1364,1389]" class="Insecta" family="Pentatomidae" genus="Cromata" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="B962CA518B68FF30FD0CFAE08E114120" box="[674,787,1364,1389]" italics="true" pageId="4" pageNumber="5">C. ornata</emphasis>
</taxonomicName>
in the proportions of the antennal segments, color pattern of legs, shape and proportions of the head, length of rostrum, presence of mesial carina on gonocoxites 9, and laterotergites 9 touching mesially and covering proctiger. The new species can be distinguished from
<taxonomicName id="4C166DC08B68FF30FC9BFA788EA641A8" box="[821,932,1484,1509]" class="Insecta" family="Pentatomidae" genus="Cromata" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="B962CA518B68FF30FC9BFA788EA641A8" box="[821,932,1484,1509]" italics="true" pageId="4" pageNumber="5">C. ornata</emphasis>
</taxonomicName>
by the former's smaller size, dorsal color pattern, contour of the posterior rim of gonocoxites 8, and shape of laterotergites 9.
</paragraph>
</subSubSection>
</treatment>
</document>