457 lines
65 KiB
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457 lines
65 KiB
XML
<document id="A681EA22B0EF2897F616C52781181890" ID-CLB-Dataset="5669" ID-DOI="10.5852/ejt.2021.758.1423" ID-GBIF-Dataset="de9c0c06-0e7d-44a6-a582-d41a346233f6" ID-ISSN="2118-9773" ID-Zenodo-Dep="5075824" ID-ZooBank="01BBC12C-E715-4393-A9F6-6EA85CB1289F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625579995989" checkinUser="felipe" docAuthor="Wesener, Thomas & Sagorny, Christina" docDate="2021" docId="03BF4B62FFB6E0391C66FDA586A5F862" docLanguage="en" docName="ejt-758_1-48.pdf" docOrigin="European Journal of Taxonomy 758 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.758.1423" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zoosphaerium nigrum Wesener & Sagorny 2021, sp. nov." docType="treatment" docUuid="4F170F43-C7A9-44A4-A004-47FD58FDBA2F" docUuidSource="ZooBank" docVersion="7" lastPageNumber="14" masterDocId="FF86331AFFBEE0341E06FFFB8261FFB5" masterDocTitle="Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="9" updateTime="1699025899143" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9DF8C246806C9D79C0F7F7226570E01D">Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)</mods:title>
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<mods:namePart id="8E855D4E212255FC3790E44D0DF2AC55">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="E77C4856724EDB93A194CAE29246CA2A">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="B029251BDC9C17CE21D89845C4670750" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:namePart id="9057843388343EE291E79B745E328D29">Sagorny, Christina</mods:namePart>
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<mods:affiliation id="03B4009A2B12DF8AD81ED3A040E4CC64">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. & University of Bonn, Institute of Evolutionary Biology and Ecology, D- 53121 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="69C972E85573A2AF978AD51CCCA6C035" type="email">csagorny@evolution.uni-bonn.de</mods:nameIdentifier>
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<mods:title id="C6896827524B4E1F429B3F42DE20EB63">European Journal of Taxonomy</mods:title>
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<mods:date id="AC00E2E3B47C3167884A7EF6AC6857AA">2021</mods:date>
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<treatment id="03BF4B62FFB6E0391C66FDA586A5F862" ID-DOI="http://doi.org/10.5281/zenodo.5076194" ID-GBIF-Taxon="183392507" ID-Zenodo-Dep="5076194" LSID="urn:lsid:zoobank.org:act:4F170F43-C7A9-44A4-A004-47FD58FDBA2F" httpUri="http://treatment.plazi.org/id/03BF4B62FFB6E0391C66FDA586A5F862" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9">
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<subSubSection id="C30CA9FFFFB6E03C1C66FDA581B2FDCF" box="[608,979,606,634]" pageId="8" pageNumber="9" type="nomenclature">
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<heading id="D0E14D18FFB6E03C1C66FDA581B2FDCF" box="[608,979,606,634]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
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<taxonomicName id="4C1681F7FFB6E03C1C66FDA58113FDCC" ID-CoL="9BDGN" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[608,882,606,633]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
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<emphasis id="B9622666FFB6E03C1C66FDA58113FDCC" bold="true" box="[608,882,606,633]" italics="true" pageId="8" pageNumber="9">Zoosphaerium nigrum</emphasis>
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<taxonomicNameLabel id="A2519B1DFFB6E03C1D7FFD9B81B2FDCF" box="[889,979,608,634]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C30CA9FFFFB6E03C1F6AFD7981E1FD75" pageId="8" pageNumber="9" type="description">
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<uri id="FF87F676FFB6E03C1F6AFD7986A6FD28" box="[364,1223,642,669]" pageId="8" pageNumber="9">
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urn:lsid:zoobank.org:act:
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<uuid id="FFB0C0A1FFB6E03C1C8DFD7986A6FD28" box="[651,1223,642,669]" pageId="8" pageNumber="9">4F170F43-C7A9-44A4-A004-47FD58FDBA2F</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BA9FA74FFB6E03C1CB3FD5E81E1FD75" blockId="8.[364,1223,606,704]" box="[693,896,677,704]" pageId="8" pageNumber="9">
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<figureCitation id="132DE6F1FFB6E03C1CB3FD5E8174FD75" box="[693,789,677,704]" captionStart="Fig" captionStartId="9.[189,232,1764,1790]" captionTargetBox="[198,1390,265,1713]" captionTargetId="figure-92@9.[189,1398,265,1719]" captionTargetPageId="9" captionText="Fig. 2. Habitus photographs of the seven new species of Zoosphaerium, lateral view. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1).D. Z. beanka sp. nov., holotype (FMNH-INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). F. Z. masoala sp. nov., holotype (FMNH-INS 3195651). G. Z. spinopiligerum sp. nov., holotype (FMNH-INS 3119885). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5075828" httpUri="https://zenodo.org/record/5075828/files/figure.png" pageId="8" pageNumber="9">Figs 2A</figureCitation>
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,
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<figureCitation id="132DE6F1FFB6E03C1D27FD5D8131FD75" box="[801,848,678,704]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[189,232,1891,1917]" captionStartId-1="11.[189,232,1750,1776]" captionTargetBox-0="[228,1372,266,1870]" captionTargetBox-1="[251,1332,267,1720]" captionTargetId-0="figure-87@10.[202,1385,239,1881]" captionTargetId-1="figure-150@11.[244,1343,244,1730]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Fig. 3. Distribution map of the newly described species of Zoosphaerium. Vegetation color marks modified from Moat & Smith 2007. 1 = Z. nigrum sp. nov.; 2 = Z. silens sp. nov.; 3 = Z. ambatovaky sp. nov.; 4 = Z. beanka sp. nov.; 5 = Z. voahangy sp. nov.; 6 = Z. masoala sp. nov.; 7 = Z. spinopiligerum sp. nov. Numbers followed by ‘a’ mark additional, non-type localities." captionText-1="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5075830" figureDoi-1="http://doi.org/10.5281/zenodo.5075832" httpUri-0="https://zenodo.org/record/5075830/files/figure.png" httpUri-1="https://zenodo.org/record/5075832/files/figure.png" pageId="8" pageNumber="9">3–4</figureCitation>
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,
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<figureCitation id="132DE6F1FFB6E03C1D5AFD5E81E1FD75" box="[860,896,677,704]" captionStart="Fig" captionStartId="12.[189,232,1891,1917]" captionTargetBox="[270,1317,266,1874]" captionTargetId="figure-78@12.[270,1317,266,1875]" captionTargetPageId="12" captionText="Fig. 5. Scanning electron micrographs of the endoterga of midbody rings. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1). D. Z. beanka sp. nov., holotype (FMNH- INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5075834" httpUri="https://zenodo.org/record/5075834/files/figure.png" pageId="8" pageNumber="9">5A</figureCitation>
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</paragraph>
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<subSubSection id="C30CA9FFFFB6E03C1EBBFD158716FBBF" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BA9FA74FFB6E03C1EBBFD158357FCBD" blockId="8.[189,310,750,776]" box="[189,310,750,776]" pageId="8" pageNumber="9">
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<heading id="D0E14D18FFB6E03C1EBBFD158357FCBD" bold="true" box="[189,310,750,776]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
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<emphasis id="B9622666FFB6E03C1EBBFD158357FCBD" bold="true" box="[189,310,750,776]" pageId="8" pageNumber="9">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9FA74FFB6E03C1EBBFCE68716FBBF" blockId="8.[189,1400,797,1035]" pageId="8" pageNumber="9">
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Posterior telopods (
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<figureCitation id="132DE6F1FFB6E03C1FA0FCE6807CFC82" box="[422,541,797,823]" captionStart="Fig" captionStartId="11.[189,232,1750,1776]" captionTargetBox="[251,1332,267,1720]" captionTargetId="figure-150@11.[244,1343,244,1730]" captionTargetPageId="11" captionText="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="8" pageNumber="9">Fig. 4H–I</figureCitation>
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) identify this species as a member of the
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<taxonomicName id="4C1681F7FFB6E03C1A1FFCE586A4FC82" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[1049,1221,797,823]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="libidinosum">
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<emphasis id="B9622666FFB6E03C1A1FFCE586A4FC82" box="[1049,1221,797,823]" italics="true" pageId="8" pageNumber="9">Z. libidinosum</emphasis>
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</taxonomicName>
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species-group. Within this group,
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<taxonomicName id="4C1681F7FFB6E03C1F9EFCBB80FCFCEF" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[408,669,832,858]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
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<emphasis id="B9622666FFB6E03C1F9EFCBB80FCFCEF" box="[408,669,832,858]" italics="true" pageId="8" pageNumber="9">Zoosphaerium nigrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFB6E03C1CA2FCBA809FFCEE" box="[676,766,833,859]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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shares a number of characters with only
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<taxonomicName id="4C1681F7FFB6E03C1ADAFCBA8058FCC8" authority="(de Saussure & Zehntner, 1902)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1902" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="anomalum">
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<emphasis id="B9622666FFB6E03C1ADAFCBA8717FCEF" box="[1244,1398,832,858]" italics="true" pageId="8" pageNumber="9">Z. anomalum</emphasis>
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(de
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<bibRefCitation id="EF878785FFB6E03C1EEDFC988050FCC8" author="Saussure H. de & Zehntner L." box="[235,561,867,893]" pageId="8" pageNumber="9" pagination="1 - 356" refId="ref27698" refString="Saussure H. de & Zehntner L. 1902. Myriapodes de Madagascar. In: Grandidier A. (ed.) Histoire physique, naturelle et politique de Madagascar 27 (53): i - viii, 1 - 356." type="journal article" year="1902">Saussure & Zehntner, 1902</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="4C1681F7FFB6E03C1C71FC9F81B3FCC8" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[631,978,867,894]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pulchellum">
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<emphasis id="B9622666FFB6E03C1C71FC9F817BFCC8" box="[631,794,867,893]" italics="true" pageId="8" pageNumber="9">Z. pulchellum</emphasis>
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Wesener, 2009
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</taxonomicName>
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. These include the presence of two locking carinae, four apical cones on each antenna, and shiny, glossy tergites. Moreover, the second and third podomeres of the posterior telopods are covered with numerous minute hairs.
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<taxonomicName id="4C1681F7FFB6E03C1AC8FC518373FC52" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
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<emphasis id="B9622666FFB6E03C1AC8FC518373FC52" italics="true" pageId="8" pageNumber="9">Zoosphaerium nigrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFB6E03C1F1FFC368312FC52" box="[281,371,973,999]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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differs from these two species in the black coloration (jade-like green in the others), the presence of only a single weak stridulation rib on the anterior telopod and differently shaped podomeres.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30CA9FFFFB6E03C1EBBFBC28131FB37" box="[189,848,1081,1154]" pageId="8" pageNumber="9" type="etymology">
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<paragraph id="8BA9FA74FFB6E03C1EBBFBC28325FBE6" blockId="8.[189,324,1081,1107]" box="[189,324,1081,1107]" pageId="8" pageNumber="9">
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|
||
<emphasis id="B9622666FFB6E03C1EBBFBC28325FBE6" bold="true" box="[189,324,1081,1107]" pageId="8" pageNumber="9">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB6E03C1EBBFB938131FB37" blockId="8.[189,848,1128,1154]" box="[189,848,1128,1154]" pageId="8" pageNumber="9">‘Nigrum’, adjective, after the dark colour of the species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFB6E03C1EBBFB4B818CFAD6" pageId="8" pageNumber="9" type="materials_examined">
|
||
<paragraph id="8BA9FA74FFB6E03C1EBBFB4B83CBFB7F" blockId="8.[189,426,1200,1226]" box="[189,426,1200,1226]" pageId="8" pageNumber="9">
|
||
<heading id="D0E14D18FFB6E03C1EBBFB4B83CBFB7F" bold="true" box="[189,426,1200,1226]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="B9622666FFB6E03C1EBBFB4B83CBFB7F" bold="true" box="[189,426,1200,1226]" pageId="8" pageNumber="9">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB6E03C1EECFB24818CFAD6" blockId="8.[189,1399,1247,1380]" pageId="8" pageNumber="9">
|
||
<materialsCitation id="3B7EF029FFB6E03C1EECFB24818CFAD6" ID-GBIF-Occurrence="3314642328" accessionNumber="BLF14945" collectingDate="2006-11-17" collectionCode="CASENT" collectorName="B. L. Fisher" country="Madagascar" county="Mount Papango" elevation="1250" latitude="-23.840834" location="Fianarantosa" longLatPrecision="21" longitude="46.9575" municipality="Papango" pageId="8" pageNumber="9" specimenCode="CASENT 9068289" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B9622666FFB6E03C1EECFB24833DFB4C" bold="true" box="[234,348,1247,1273]" pageId="8" pageNumber="9">
|
||
<typeStatus id="54AD44D6FFB6E03C1EECFB24833DFB4C" box="[234,348,1247,1273]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F301BAE4FFB6E03C1EBBFAF883EFFAA8" box="[189,398,1283,1309]" name="Madagascar" pageId="8" pageNumber="9">MADAGASCAR</collectingCountry>
|
||
–
|
||
<emphasis id="B9622666FFB6E03C1FB1FAF9803CFAA9" bold="true" box="[439,605,1282,1308]" pageId="8" pageNumber="9">
|
||
<location id="8EC9ACAFFFB6E03C1FB1FAF9803CFAA9" LSID="urn:lsid:plazi:treatment:03BF4B62FFB6E0391C66FDA586A5F862:8EC9ACAFFFB6E03C1FB1FAF9803CFAA9" box="[439,605,1282,1308]" country="Madagascar" county="Mount Papango" latitude="-23.840834" longLatPrecision="21" longitude="46.9575" municipality="Papango" name="Fianarantosa" pageId="8" pageNumber="9">Fianarantosa</location>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D1031FDFFB6E03C1C84FAF880FAFAA8" box="[642,667,1283,1309]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
|
||
;
|
||
<location id="8EC9ACAFFFB6E03C1CA8FAF88620FAA9" LSID="urn:lsid:plazi:treatment:03BF4B62FFB6E0391C66FDA586A5F862:8EC9ACAFFFB6E03C1CA8FAF88620FAA9" box="[686,1089,1282,1309]" country="Madagascar" county="Mount Papango" latitude="-23.840834" longLatPrecision="21" longitude="46.9575" municipality="Papango" name="Parc National Befotaka-Midongy" pageId="8" pageNumber="9">Parc National Befotaka-Midongy</location>
|
||
,
|
||
<collectingMunicipality id="6BCD600EFFB6E03C1A55FAF886DAFAA8" box="[1107,1211,1283,1309]" pageId="8" pageNumber="9">Papango</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="9A1BE356FFB6E03C1ACBFAF98302FAF5" location="Midongy-Sud" pageId="8" pageNumber="9">
|
||
<quantity id="4CEE5791FFB6E03C1ACBFAF98752FAA9" box="[1229,1331,1282,1309]" metricMagnitude="4" metricUnit="m" metricValue="2.85" pageId="8" pageNumber="9" unit="km" value="28.5">28.5 km</quantity>
|
||
S of Midongy-Sud
|
||
</locationDeviation>
|
||
,
|
||
<collectingCounty id="62C882F8FFB6E03C1F77FADD8052FAF5" box="[369,563,1318,1344]" pageId="8" pageNumber="9">Mount Papango</collectingCounty>
|
||
;
|
||
<geoCoordinate id="EE229CB3FFB6E03C1C47FADF80B5FAF5" box="[577,724,1316,1344]" degrees="23" direction="south" minutes="50" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="27" value="-23.840834">23°50′27″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE229CB3FFB6E03C1CE2FADF8118FAF5" box="[740,889,1316,1344]" degrees="46" direction="east" minutes="57" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="27" value="46.9575">46°57′27″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CEE5791FFB6E03C1DB1FADE8677FAF5" box="[951,1046,1317,1344]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="9" unit="m" value="1250.0">
|
||
<elevation id="003B1D47FFB6E03C1DB1FADE8677FAF5" box="[951,1046,1317,1344]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="9" unit="m" value="1250.0">1250 m</elevation>
|
||
</quantity>
|
||
; montane rainforest;
|
||
<date id="FFA8DCB4FFB6E03C1B11FADD829AFAD6" pageId="8" pageNumber="9" value="2006-11-17">
|
||
<collectingDate id="EFEC255CFFB6E03C1B11FADD829AFAD6" pageId="8" pageNumber="9" value="2006-11-17">17 Nov. 2006</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26E39FA2FFB6E03C1F01FAB283EDFAD6" box="[263,396,1353,1379]" pageId="8" pageNumber="9">B.L. Fisher</collectorName>
|
||
<emphasis id="B9622666FFB6E03C1F95FAB183AFFAD6" box="[403,462,1353,1379]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
leg.; coll. no. BLF14945;
|
||
<specimenCode id="DBB0520FFFB6E03C1D03FAB28188FAD6" box="[773,1001,1353,1379]" collectionCode="CASENT" pageId="8" pageNumber="9">CASENT 9068289</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFB6E0391EBBFA698358F894" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="8BA9FA74FFB6E03C1EBBFA69832FFA19" blockId="8.[189,334,1426,1452]" box="[189,334,1426,1452]" pageId="8" pageNumber="9">
|
||
<heading id="D0E14D18FFB6E03C1EBBFA69832FFA19" bold="true" box="[189,334,1426,1452]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="B9622666FFB6E03C1EBBFA69832FFA19" bold="true" box="[189,334,1426,1452]" pageId="8" pageNumber="9">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB6E03C1EECFA3B834BFA6F" blockId="8.[188,1399,1472,1570]" box="[234,298,1472,1498]" pageId="8" pageNumber="9">
|
||
<emphasis id="B9622666FFB6E03C1EECFA3B834BFA6F" bold="true" box="[234,298,1472,1498]" pageId="8" pageNumber="9">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB6E03C1EBBFA1F866EF997" blockId="8.[188,1399,1472,1570]" pageId="8" pageNumber="9">
|
||
BODY LENGTH.
|
||
<typeStatus id="54AD44D6FFB6E03C1F74FA1883BEFA48" box="[370,479,1507,1533]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
|
||
male: length ca
|
||
<quantity id="4CEE5791FFB6E03C1C9BFA1F8093FA4B" box="[669,754,1508,1534]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="9" unit="mm" value="28.0">28 mm</quantity>
|
||
, width of thoracic shield
|
||
<quantity id="4CEE5791FFB6E03C1A13FA1F86E1FA4B" box="[1045,1152,1508,1534]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="8" pageNumber="9" unit="mm" value="13.1">13.1 mm</quantity>
|
||
, of tergite 7
|
||
<quantity id="4CEE5791FFB6E03C1B09FA188716FA4B" box="[1295,1399,1507,1534]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="9" unit="mm" value="13.5">13.5 mm</quantity>
|
||
(widest), height of thoracic shield
|
||
<quantity id="4CEE5791FFB6E03C1C48F9FC80CAF994" box="[590,683,1543,1569]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="8" pageNumber="9" unit="mm" value="7.1">7.1 mm</quantity>
|
||
, of tergite 7
|
||
<quantity id="4CEE5791FFB6E03C1D38F9FC81F9F994" box="[830,920,1543,1569]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="9" unit="mm" value="7.4">7.4 mm</quantity>
|
||
(highest).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB6E03C1EBBF9AB8376F905" blockId="8.[188,1398,1616,1712]" pageId="8" pageNumber="9">
|
||
COLORATION. Some discoloration may have occurred because of preservation in ethanol. Tergites, head, anal shield and appendages dark green to black (
|
||
<figureCitation id="132DE6F1FFB6E03C1CF7F988812EF938" box="[753,847,1651,1677]" captionStart="Fig" captionStartId="9.[189,232,1764,1790]" captionTargetBox="[198,1390,265,1713]" captionTargetId="figure-92@9.[189,1398,265,1719]" captionTargetPageId="9" captionText="Fig. 2. Habitus photographs of the seven new species of Zoosphaerium, lateral view. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1).D. Z. beanka sp. nov., holotype (FMNH-INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). F. Z. masoala sp. nov., holotype (FMNH-INS 3195651). G. Z. spinopiligerum sp. nov., holotype (FMNH-INS 3119885). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5075828" httpUri="https://zenodo.org/record/5075828/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
|
||
). Posterior margin of tergites with a thin brown margin.
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB6E03C1EBBF9248126F8D6" blockId="8.[187,1399,1759,1891]" pageId="8" pageNumber="9">HEAD. Wide and short. Eyes with>120 ommatidia. Aberrant ocellus located in antennal groove. Antennae short, with cylindrical joints, reaching posteriorly to leg pair 3. Length of antennomeres 1= 2=3= 4=5<6. A shallow groove present on antennomere 1. Male apical disc with four apical sensory cones. Organ of Tömösváry located in antennal groove.</paragraph>
|
||
<paragraph id="8BA9FA74FFB6E03C1EBBF8698169F819" blockId="8.[189,776,1938,1964]" box="[189,776,1938,1964]" pageId="8" pageNumber="9">GNATHOCHILARIUM AND MANDIBLE. Not dissected.</paragraph>
|
||
<caption id="DF69AAFCFFB7E03D1EBBF91F8701F83E" ID-DOI="http://doi.org/10.5281/zenodo.5075828" ID-Zenodo-Dep="5075828" httpUri="https://zenodo.org/record/5075828/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1764,1790]" targetBox="[198,1390,265,1713]" targetPageId="9">
|
||
<paragraph id="8BA9FA74FFB7E03D1EBBF91F8701F83E" blockId="9.[187,1399,1764,1932]" pageId="9" pageNumber="10">
|
||
<emphasis id="B9622666FFB7E03D1EBBF91F836AF94B" bold="true" box="[189,267,1764,1790]" pageId="9" pageNumber="10">Fig. 2.</emphasis>
|
||
Habitus photographs of the seven new species of
|
||
<taxonomicName id="4C1681F7FFB7E03D1D5CF91F8662F94B" authorityName="Pocock" authorityYear="1895" box="[858,1027,1764,1790]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9622666FFB7E03D1D5CF91F8662F94B" box="[858,1027,1764,1790]" italics="true" pageId="9" pageNumber="10">Zoosphaerium</emphasis>
|
||
</taxonomicName>
|
||
, lateral view.
|
||
<emphasis id="B9622666FFB7E03D1AAFF91F86DFF94B" bold="true" box="[1193,1214,1764,1790]" pageId="9" pageNumber="10">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB7E03D1ACBF91F8373F894" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB7E03D1ACBF91F8373F894" italics="true" pageId="9" pageNumber="10">Zoosphaerium nigrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB7E03D1F1CF8F38319F897" box="[282,376,1800,1826]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (CASENT 9068289).
|
||
<emphasis id="B9622666FFB7E03D1CFFF8F3816DF897" bold="true" box="[761,780,1800,1826]" pageId="9" pageNumber="10">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB7E03D1D1DF8F3811FF894" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[795,894,1799,1825]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="silens" status="sp. nov.">
|
||
<emphasis id="B9622666FFB7E03D1D1DF8F3811FF894" box="[795,894,1799,1825]" italics="true" pageId="9" pageNumber="10">Z. silens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB7E03D1D80F8F38185F897" box="[902,996,1800,1826]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119887).
|
||
<emphasis id="B9622666FFB7E03D1EBBF8D182B3F8F1" bold="true" box="[189,210,1834,1860]" pageId="9" pageNumber="10">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB7E03D1ED9F8D083EBF8F0" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[223,394,1835,1861]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ambatovaky" status="sp. nov.">
|
||
<emphasis id="B9622666FFB7E03D1ED9F8D083EBF8F0" box="[223,394,1835,1861]" italics="true" pageId="9" pageNumber="10">Z. ambatovaky</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB7E03D1F89F8D0838BF8F0" box="[399,490,1835,1861]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (CASENT 9068294/1).
|
||
<emphasis id="B9622666FFB7E03D1D75F8D081E9F8F0" bold="true" box="[883,904,1835,1861]" pageId="9" pageNumber="10">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB7E03D1D91F8D08668F8F0" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[919,1033,1835,1861]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
|
||
<emphasis id="B9622666FFB7E03D1D91F8D08668F8F0" box="[919,1033,1835,1861]" italics="true" pageId="9" pageNumber="10">Z. beanka</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB7E03D1A08F8D08608F8F0" box="[1038,1129,1835,1861]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119897).
|
||
<emphasis id="B9622666FFB7E03D1F3AF8B58331F8DD" bold="true" box="[316,336,1870,1896]" pageId="9" pageNumber="10">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB7E03D1F66F8B48393F8DD" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[352,498,1870,1896]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
|
||
<emphasis id="B9622666FFB7E03D1F66F8B48393F8DD" box="[352,498,1870,1896]" italics="true" pageId="9" pageNumber="10">Z. voahangy</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB7E03D1FFDF8B5803BF8DD" box="[507,602,1870,1896]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119921).
|
||
<emphasis id="B9622666FFB7E03D1DF1F8B58669F8DD" bold="true" box="[1015,1032,1870,1896]" pageId="9" pageNumber="10">F</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB7E03D1A1EF8B486FAF8DD" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[1048,1179,1870,1896]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="masoala" status="sp. nov.">
|
||
<emphasis id="B9622666FFB7E03D1A1EF8B486FAF8DD" box="[1048,1179,1870,1896]" italics="true" pageId="9" pageNumber="10">Z. masoala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB7E03D1AA2F8B58762F8DD" box="[1188,1283,1870,1896]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3195651).
|
||
<emphasis id="B9622666FFB7E03D1FDDF88A8393F83E" bold="true" box="[475,498,1905,1931]" pageId="9" pageNumber="10">G</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB7E03D1C06F88980B0F83E" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[512,721,1905,1931]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spinopiligerum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB7E03D1C06F88980B0F83E" box="[512,721,1905,1931]" italics="true" pageId="9" pageNumber="10">Z. spinopiligerum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB7E03D1CDEF8898154F839" box="[728,821,1906,1932]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119885). Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF69AAFCFFB4E03E1EBBF8988195F853" ID-DOI="http://doi.org/10.5281/zenodo.5075830" ID-Zenodo-Dep="5075830" httpUri="https://zenodo.org/record/5075830/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1891,1917]" targetBox="[228,1372,266,1870]" targetPageId="10">
|
||
<paragraph id="8BA9FA74FFB4E03E1EBBF8988195F853" blockId="10.[189,1400,1891,2023]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9622666FFB4E03E1EBBF898836EF8C8" bold="true" box="[189,271,1891,1917]" pageId="10" pageNumber="11">Fig. 3.</emphasis>
|
||
Distribution map of the newly described species of
|
||
<taxonomicName id="4C1681F7FFB4E03E1DA6F8988628F8C8" authorityName="Pocock" authorityYear="1895" box="[928,1097,1891,1917]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9622666FFB4E03E1DA6F8988628F8C8" box="[928,1097,1891,1917]" italics="true" pageId="10" pageNumber="11">Zoosphaerium</emphasis>
|
||
</taxonomicName>
|
||
. Vegetation color marks modified from
|
||
<bibRefCitation id="EF878785FFB4E03E1F76F87C8005F815" author="Moat J. & Smith P." box="[368,612,1926,1953]" pageId="10" pageNumber="11" refId="ref27053" refString="Moat J. & Smith P. 2007. Atlas of the Vegetation of Madagascar Vegetation / Atlas de la Vegetation de Madagascar. Royal Botanic Gardens, Kew (bilingual edition)." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
. 1 =
|
||
<taxonomicName id="4C1681F7FFB4E03E1CA4F87C8176F815" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[674,791,1927,1952]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB4E03E1CA4F87C8176F815" box="[674,791,1927,1952]" italics="true" pageId="10" pageNumber="11">Z. nigrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB4E03E1D26F87C811EF814" box="[800,895,1927,1953]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 2 =
|
||
<taxonomicName id="4C1681F7FFB4E03E1DB9F87C8642F815" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[959,1059,1926,1952]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="silens" status="sp. nov.">
|
||
<emphasis id="B9622666FFB4E03E1DB9F87C8642F815" box="[959,1059,1926,1952]" italics="true" pageId="10" pageNumber="11">Z. silens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB4E03E1A2CF87C86EBF814" box="[1066,1162,1927,1953]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 3 =
|
||
<taxonomicName id="4C1681F7FFB4E03E1ACCF87C8719F815" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[1226,1400,1926,1952]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ambatovaky" status="sp. nov.">
|
||
<emphasis id="B9622666FFB4E03E1ACCF87C8719F815" box="[1226,1400,1926,1952]" italics="true" pageId="10" pageNumber="11">Z. ambatovaky</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB4E03E1EBBF851837BF871" box="[189,282,1962,1988]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 4 =
|
||
<taxonomicName id="4C1681F7FFB4E03E1F57F85183A5F876" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[337,452,1961,1987]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
|
||
<emphasis id="B9622666FFB4E03E1F57F85183A5F876" box="[337,452,1961,1987]" italics="true" pageId="10" pageNumber="11">Z. beanka</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB4E03E1FCCF8518046F871" box="[458,551,1962,1988]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 5 =
|
||
<taxonomicName id="4C1681F7FFB4E03E1C58F851808FF876" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[606,750,1961,1987]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
|
||
<emphasis id="B9622666FFB4E03E1C58F851808FF876" box="[606,750,1961,1987]" italics="true" pageId="10" pageNumber="11">Z. voahangy</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB4E03E1CF0F8518132F871" box="[758,851,1962,1988]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 6 =
|
||
<taxonomicName id="4C1681F7FFB4E03E1D8FF851866AF876" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[905,1035,1961,1987]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="masoala" status="sp. nov.">
|
||
<emphasis id="B9622666FFB4E03E1D8FF851866AF876" box="[905,1035,1961,1987]" italics="true" pageId="10" pageNumber="11">Z. masoala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB4E03E1A14F851860EF871" box="[1042,1135,1962,1988]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 7 =
|
||
<taxonomicName id="4C1681F7FFB4E03E1AA0F8518717F876" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[1190,1398,1961,1987]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinopiligerum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB4E03E1AA0F8518717F876" box="[1190,1398,1961,1987]" italics="true" pageId="10" pageNumber="11">Z. spinopiligerum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB4E03E1EBBF8368379F852" box="[189,280,1997,2023]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Numbers followed by ‘a’ mark additional, non-type localities.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF69AAFCFFB5E03F1EBBF92D81FFF852" ID-DOI="http://doi.org/10.5281/zenodo.5075832" ID-Zenodo-Dep="5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1750,1776]" targetBox="[251,1332,267,1720]" targetPageId="11">
|
||
<paragraph id="8BA9FA74FFB5E03F1EBBF92D81FFF852" blockId="11.[188,1400,1750,2023]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9622666FFB5E03F1EBBF92D836FF945" bold="true" box="[189,270,1750,1776]" pageId="11" pageNumber="12">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C1681F7FFB5E03F1F1EF92D8040F945" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[280,545,1750,1776]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB5E03F1F1EF92D8040F945" box="[280,545,1750,1776]" italics="true" pageId="11" pageNumber="12">Zoosphaerium nigrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB5E03F1C2DF92C80EDF944" box="[555,652,1751,1777]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (CASENT 9068289).
|
||
<emphasis id="B9622666FFB5E03F1A12F92D8648F945" bold="true" box="[1044,1065,1750,1776]" pageId="11" pageNumber="12">A</emphasis>
|
||
. Left leg 9, posterior view.
|
||
<emphasis id="B9622666FFB5E03F1EBBF90182B0F8A1" bold="true" box="[189,209,1786,1812]" pageId="11" pageNumber="12">B</emphasis>
|
||
. Left coxa 2 with gonopore, posterior view.
|
||
<emphasis id="B9622666FFB5E03F1CF8F9028172F8A6" bold="true" box="[766,787,1785,1811]" pageId="11" pageNumber="12">C</emphasis>
|
||
. Left coxa 1 with stigmatic plate, posterior view.
|
||
<emphasis id="B9622666FFB5E03F1EBBF8E682B3F882" bold="true" box="[189,210,1821,1847]" pageId="11" pageNumber="12">D</emphasis>
|
||
. Left anterior gonopod, mesal view.
|
||
<emphasis id="B9622666FFB5E03F1C7DF8E680EEF882" bold="true" box="[635,655,1821,1847]" pageId="11" pageNumber="12">E</emphasis>
|
||
. Left anterior gonopod, anterior view.
|
||
<emphasis id="B9622666FFB5E03F1A49F8E68600F882" bold="true" box="[1103,1121,1821,1847]" pageId="11" pageNumber="12">F</emphasis>
|
||
. Left anterior gonopod, posterior view.
|
||
<emphasis id="B9622666FFB5E03F1F7BF8BB83F5F8EF" bold="true" box="[381,404,1856,1882]" pageId="11" pageNumber="12">G</emphasis>
|
||
. Left anterior gonopod, lateral view.
|
||
<emphasis id="B9622666FFB5E03F1D6FF8BB81E1F8EF" bold="true" box="[873,896,1856,1882]" pageId="11" pageNumber="12">H</emphasis>
|
||
. Right posterior telopod, posterior view.
|
||
<emphasis id="B9622666FFB5E03F1EBBF89882A9F8C8" bold="true" box="[189,200,1891,1917]" pageId="11" pageNumber="12">I</emphasis>
|
||
. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF69AAFCFFB2E0381EBBF8988778F85D" ID-DOI="http://doi.org/10.5281/zenodo.5075834" ID-Zenodo-Dep="5075834" httpUri="https://zenodo.org/record/5075834/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1891,1917]" targetBox="[270,1317,266,1874]" targetPageId="12">
|
||
<paragraph id="8BA9FA74FFB2E0381EBBF8988778F85D" blockId="12.[189,1399,1891,2024]" pageId="12" pageNumber="13">
|
||
<emphasis id="B9622666FFB2E0381EBBF898836FF8CB" bold="true" box="[189,270,1891,1918]" pageId="12" pageNumber="13">Fig. 5.</emphasis>
|
||
Scanning electron micrographs of the endoterga of midbody rings.
|
||
<emphasis id="B9622666FFB2E0381A41F898863DF8C8" bold="true" box="[1095,1116,1891,1917]" pageId="12" pageNumber="13">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB2E0381A68F8988716F8C8" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[1134,1399,1891,1917]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB2E0381A68F8988716F8C8" box="[1134,1399,1891,1917]" italics="true" pageId="12" pageNumber="13">Zoosphaerium nigrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB2E0381EBBF87C8342F814" box="[189,291,1927,1953]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (CASENT 9068289).
|
||
<emphasis id="B9622666FFB2E0381CC7F87C80B4F814" bold="true" box="[705,725,1927,1953]" pageId="12" pageNumber="13">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB2E0381CEAF87C8136F814" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[748,855,1927,1953]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="silens" status="sp. nov.">
|
||
<emphasis id="B9622666FFB2E0381CEAF87C8136F814" box="[748,855,1927,1953]" italics="true" pageId="12" pageNumber="13">Z. silens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB2E0381D61F87C81ACF814" box="[871,973,1927,1953]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119887).
|
||
<emphasis id="B9622666FFB2E0381EBBF85182B3F871" bold="true" box="[189,210,1962,1988]" pageId="12" pageNumber="13">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB2E0381EE4F85083F0F871" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[226,401,1962,1988]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ambatovaky" status="sp. nov.">
|
||
<emphasis id="B9622666FFB2E0381EE4F85083F0F871" box="[226,401,1962,1988]" italics="true" pageId="12" pageNumber="13">Z. ambatovaky</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB2E0381F9FF8518399F871" box="[409,504,1962,1988]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (CASENT 9068294/1).
|
||
<emphasis id="B9622666FFB2E0381D97F85181C4F871" bold="true" box="[913,933,1962,1988]" pageId="12" pageNumber="13">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB2E0381DB3F850864BF871" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[949,1066,1962,1988]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
|
||
<emphasis id="B9622666FFB2E0381DB3F850864BF871" box="[949,1066,1962,1988]" italics="true" pageId="12" pageNumber="13">Z. beanka</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB2E0381A35F85186F0F871" box="[1075,1169,1962,1988]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH- INS 3119897).
|
||
<emphasis id="B9622666FFB2E0381F74F83683E7F852" bold="true" box="[370,390,1997,2023]" pageId="12" pageNumber="13">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C1681F7FFB2E0381F92F8358044F852" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[404,549,1997,2023]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
|
||
<emphasis id="B9622666FFB2E0381F92F8358044F852" box="[404,549,1997,2023]" italics="true" pageId="12" pageNumber="13">Z. voahangy</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB2E0381C2AF83580E8F85D" box="[556,649,1998,2024]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119921). Scale bars = 100 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFEF28076FEDC" blockId="13.[189,1399,265,361]" pageId="13" pageNumber="14">
|
||
STIGMATIC PLATES. First stigmatic plate triangular, apex well rounded, nearly as wide as at base. Mesal margin straight, not curved towards coxa. Hair most abundant on apical margin, very few hairs on basal and lateral margins (
|
||
<figureCitation id="132DE6F1FFB3E0391FABFEB4806BFEDC" box="[429,522,335,361]" captionStart="Fig" captionStartId="11.[189,232,1750,1776]" captionTargetBox="[251,1332,267,1720]" captionTargetId="figure-150@11.[244,1343,244,1730]" captionTargetPageId="11" captionText="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="13" pageNumber="14">Fig. 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFE76801FFE7F" blockId="13.[189,1398,397,459]" pageId="13" pageNumber="14">PLEURITES. First pleurite laterally modified and extending backwards with a broad, apically rounded process. Other pleurites well-rounded.</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFE15829FFD9E" blockId="13.[189,1398,494,555]" pageId="13" pageNumber="14">COLLUM. Glabrous with very sparse setae around margins, mostly located at corners on either side of head.</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFDB481D0FDDC" blockId="13.[189,945,591,617]" box="[189,945,591,617]" pageId="13" pageNumber="14">THORACIC SHIELD. Smooth and glabrous, without protuberances.</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFD7783A1FD7C" blockId="13.[189,1399,652,713]" pageId="13" pageNumber="14">TERGITES. Surface glabrous and shiny, lacking small pits. Tips of paratergites of midbody tergites projecting posteriorly.</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFD168025FCC7" blockId="13.[189,1400,749,882]" pageId="13" pageNumber="14">
|
||
ENDOTERGUM. Inner section with numerous short triangular spines and very few longer setae. Middle area above spines with single row of sparse, circular cuticular impressions slightly varying in size. Apically a single row of short marginal bristles, protruding to ⅔ distance to tergite margin. Bristles with numerous small spines (
|
||
<figureCitation id="132DE6F1FFB3E0391FDEFCAC8056FCC7" box="[472,567,855,882]" captionStart="Fig" captionStartId="12.[189,232,1891,1917]" captionTargetBox="[270,1317,266,1874]" captionTargetId="figure-78@12.[270,1317,266,1875]" captionTargetPageId="12" captionText="Fig. 5. Scanning electron micrographs of the endoterga of midbody rings. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1). D. Z. beanka sp. nov., holotype (FMNH- INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5075834" httpUri="https://zenodo.org/record/5075834/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFC6E8376FC4D" blockId="13.[187,1398,916,1016]" pageId="13" pageNumber="14">ANAL SHIELD. Large and slightly bell-shaped. Several tiny setae present in posterior part. Underside with two black locking carinae, posterior one 2 times as long as anterior one, both located close to anal shield margin.</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFBE78677FBC9" blockId="13.[187,1398,1052,1149]" pageId="13" pageNumber="14">
|
||
LEGS. Leg 1 with 5 ventral spines, leg 2 with 3, leg 3 with 3–5. First two leg pairs without an apical spine, spine present on leg 3. Leg pairs 4–21 with 5–7 ventral spines and an apical spine. On leg 9, femur 1.8 and tarsus 4.6 times as long as wide. All podomeres with setae (
|
||
<figureCitation id="132DE6F1FFB3E0391DACFB998668FBC9" box="[938,1033,1122,1148]" captionStart="Fig" captionStartId="11.[189,232,1750,1776]" captionTargetBox="[251,1332,267,1720]" captionTargetId="figure-150@11.[244,1343,244,1730]" captionTargetPageId="11" captionText="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="13" pageNumber="14">Fig. 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EECFB5F806AFB0B" blockId="13.[189,1170,1188,1249]" box="[234,523,1188,1214]" pageId="13" pageNumber="14">
|
||
<emphasis id="B9622666FFB3E0391EECFB5F806AFB0B" bold="true" box="[234,523,1188,1214]" pageId="13" pageNumber="14">Male sexual characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFB3C86F3FB54" blockId="13.[189,1170,1188,1249]" box="[189,1170,1223,1249]" pageId="13" pageNumber="14">
|
||
GONOPORE. Covered with a single undivided, rounded membranous plate (
|
||
<figureCitation id="132DE6F1FFB3E0391A2EFB3C86E4FB54" box="[1064,1157,1223,1249]" captionStart="Fig" captionStartId="11.[189,232,1750,1776]" captionTargetBox="[251,1332,267,1720]" captionTargetId="figure-150@11.[244,1343,244,1730]" captionTargetPageId="11" captionText="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="13" pageNumber="14">Fig. 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFAFE805AFA7A" blockId="13.[188,1399,1284,1487]" pageId="13" pageNumber="14">
|
||
ANTERIOR TELOPODS. Harp with a single weak stridulation rib (
|
||
<figureCitation id="132DE6F1FFB3E0391D9CFAFF8197FAAA" box="[922,1014,1284,1311]" captionStart="Fig" captionStartId="11.[189,232,1750,1776]" captionTargetBox="[251,1332,267,1720]" captionTargetId="figure-150@11.[244,1343,244,1730]" captionTargetPageId="11" captionText="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="13" pageNumber="14">Fig. 4E</figureCitation>
|
||
). First podomere with only a few setae, restricted to apical parts of lateral margins, 1.5 times wider than long. Process of second podomere as long as ⅔ of third podomere, not visible in anterior view, with apical mound of a few sclerotized spots juxtaposed to third podomere. Third podomere without sclerotized teeth, but with a conspicuous process carrying sclerotized spots (
|
||
<figureCitation id="132DE6F1FFB3E0391C5DFA6A80BBFA1E" box="[603,730,1425,1451]" captionStart="Fig" captionStartId="11.[189,232,1750,1776]" captionTargetBox="[251,1332,267,1720]" captionTargetId="figure-150@11.[244,1343,244,1730]" captionTargetPageId="11" captionText="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="13" pageNumber="14">Fig. 4F–G</figureCitation>
|
||
); base of process with four spines (
|
||
<figureCitation id="132DE6F1FFB3E0391A7DFA6A86BBFA1E" box="[1147,1242,1425,1451]" captionStart="Fig" captionStartId="11.[189,232,1750,1776]" captionTargetBox="[251,1332,267,1720]" captionTargetId="figure-150@11.[244,1343,244,1730]" captionTargetPageId="11" captionText="Fig. 4. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). A. Left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, mesal view. E. Left anterior gonopod, anterior view. F. Left anterior gonopod, posterior view. G. Left anterior gonopod, lateral view. H. Right posterior telopod, posterior view. I. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075832" httpUri="https://zenodo.org/record/5075832/files/figure.png" pageId="13" pageNumber="14">Fig. 4G</figureCitation>
|
||
). Second and third podomeres sparsely setose.
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBFA098649F909" blockId="13.[189,1401,1522,1725]" pageId="13" pageNumber="14">POSTERIOR TELOPODS.Movable finger weakly curved; hollowed-out inner margin with two non-sclerotized lobes and ca three smaller sclerotized spines, posterior aspect with ca 29 crenulated teeth. Fixed finger thinner than movable finger, with slightly curved tip; basally with single membranous lobe, on inner margin covered with numerous sclerotized spots. Second and third podomeres without any long hair, but on both sides densely covered with very small pits, one very small seta standing in each pit. First podomere without those pits but with some isolated, standard longer hairs.</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EECF91F8324F94B" blockId="13.[189,325,1764,1825]" box="[234,325,1764,1790]" pageId="13" pageNumber="14">
|
||
<emphasis id="B9622666FFB3E0391EECF91F8324F94B" bold="true" box="[234,325,1764,1790]" pageId="13" pageNumber="14">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBF8FC8358F894" blockId="13.[189,325,1764,1825]" box="[189,313,1799,1825]" pageId="13" pageNumber="14">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFB3E0391EBBF8B386A5F862" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="8BA9FA74FFB3E0391EBBF8B38390F8D7" blockId="13.[189,497,1864,1890]" box="[189,497,1864,1890]" pageId="13" pageNumber="14">
|
||
<heading id="D0E14D18FFB3E0391EBBF8B38390F8D7" bold="true" box="[189,497,1864,1890]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B9622666FFB3E0391EBBF8B38390F8D7" bold="true" box="[189,497,1864,1890]" pageId="13" pageNumber="14">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFB3E0391EBBF88C86A5F862" blockId="13.[189,1399,1911,2008]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C1681F7FFB3E0391EBBF88C83A6F825" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[189,455,1911,1937]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB3E0391EBBF88C83A6F825" box="[189,455,1911,1937]" italics="true" pageId="13" pageNumber="14">Zoosphaerium nigrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB3E0391FD5F88C8053F824" box="[467,562,1911,1937]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is currently known only from the montane rainforests of Midongy- Befotaka (
|
||
<figureCitation id="132DE6F1FFB3E0391F3FF86183E0F801" box="[313,385,1946,1972]" captionStart="Fig" captionStartId="10.[189,232,1891,1917]" captionTargetBox="[228,1372,266,1870]" captionTargetId="figure-87@10.[202,1385,239,1881]" captionTargetPageId="10" captionText="Fig. 3. Distribution map of the newly described species of Zoosphaerium. Vegetation color marks modified from Moat & Smith 2007. 1 = Z. nigrum sp. nov.; 2 = Z. silens sp. nov.; 3 = Z. ambatovaky sp. nov.; 4 = Z. beanka sp. nov.; 5 = Z. voahangy sp. nov.; 6 = Z. masoala sp. nov.; 7 = Z. spinopiligerum sp. nov. Numbers followed by ‘a’ mark additional, non-type localities." figureDoi="http://doi.org/10.5281/zenodo.5075830" httpUri="https://zenodo.org/record/5075830/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
|
||
) where it coexists with another species of the genus,
|
||
<taxonomicName id="4C1681F7FFB3E0391A0EF86086BAF801" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[1032,1243,1946,1972]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="spinopiligerum" status="sp. nov.">
|
||
<emphasis id="B9622666FFB3E0391A0EF86086BAF801" box="[1032,1243,1946,1972]" italics="true" pageId="13" pageNumber="14">Z. spinopiligerum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFB3E0391AE4F861875FF801" box="[1250,1342,1946,1972]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(see below), as well as with a species of the genus
|
||
<taxonomicName id="4C1681F7FFB3E0391CDFF84686DEF862" authority="(Moritz & Wesener 2017)" baseAuthorityName="Moritz & Wesener" baseAuthorityYear="2017" box="[729,1215,1981,2008]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9622666FFB3E0391CDFF84681E7F862" box="[729,902,1981,2007]" italics="true" pageId="13" pageNumber="14">Sphaeromimus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF878785FFB3E0391D90F84686D9F862" author="Moritz L. & Wesener T." box="[918,1208,1981,2008]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref27087" refString="Moritz L. & Wesener T. 2017. Integrative description of two new species of Malagasy chirping giant pillmillipedes, genus Sphaeromimus (Diplopoda, Sphaerotheriida, Arthrosphaeridae). European Journal of Taxonomy 381: 1 - 25. https: // doi. org / 10.5852 / ejt. 2017.381" type="journal article" year="2017">Moritz & Wesener 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |