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<document id="F398B3F1871A4741B43460D95ADEDCFD" ID-DOI="10.1111/j.1096-3642.2008.00496.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5444719" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630458066632" checkinUser="felipe" docAuthor="Schmidt, B. Christian" docDate="2009" docId="03B987FAFFB9F72FFC5269D8FD7AE768" docLanguage="en" docName="j.1096-3642.2008.00496.x.pdf" docOrigin="Zoological Journal of the Linnean Society 156 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00496.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Grammia Rambur 1866" docType="treatment" docVersion="3" lastPageNumber="520" masterDocId="FF80FF82FFB4F721FF926F2DFFC1E46B" masterDocTitle="Taxonomic revision of the genus Grammia Rambur (Lepidoptera: Noctuidae: Arctiinae)" masterLastPageNumber="597" masterPageNumber="507" pageNumber="520" updateTime="1699278284593" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="02C70D82A684834F35E22904E7E815BE">Taxonomic revision of the genus Grammia Rambur (Lepidoptera: Noctuidae: Arctiinae)</mods:title>
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<mods:namePart id="153AB61E3EA6C48A3C30E0E839228B7A">Schmidt, B. Christian</mods:namePart>
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<mods:date id="0948D1777D5A3598CBCFD67C85A63158">2009</mods:date>
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<treatment id="03B987FAFFB9F72FFC5269D8FD7AE768" LSID="urn:lsid:plazi:treatment:03B987FAFFB9F72FFC5269D8FD7AE768" httpUri="http://treatment.plazi.org/id/03B987FAFFB9F72FFC5269D8FD7AE768" lastPageId="14" lastPageNumber="520" pageId="13" pageNumber="520">
<subSubSection id="C30A6567FFB9F72CFC5269D8FADAE361" box="[960,1307,1781,1802]" pageId="13" pageNumber="520" type="nomenclature">
<paragraph id="8BAF36ECFFB9F72CFC5269D8FADAE361" blockId="13.[960,1307,1781,1802]" box="[960,1307,1781,1802]" pageId="13" pageNumber="520">
<heading id="D0E78180FFB9F72CFC5269D8FADAE361" box="[960,1307,1781,1802]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="520" reason="2">
SUBGENUS
<taxonomicName id="4C104D6FFFB9F72CFBD669D8FADAE361" authority="RAMBUR" authorityName="Rambur" authorityYear="1866" box="[1092,1307,1781,1802]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="571" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAFEFFB9F72CFBD669D8FB70E362" box="[1092,1201,1781,1802]" italics="true" pageId="13" pageNumber="520">GRAMMIA</emphasis>
RAMBUR
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A6567FFB9F72FFCA86831FD7AE768" lastPageId="14" lastPageNumber="521" pageId="13" pageNumber="520" type="diagnosis">
<paragraph id="8BAF36ECFFB9F72FFCA86831FD7AE768" blockId="13.[826,1441,1820,1903]" lastBlockId="14.[144,759,198,771]" lastPageId="14" lastPageNumber="521" pageId="13" pageNumber="520">
<emphasis id="B964EAFEFFB9F72CFCA86831FC70E35A" box="[826,945,1820,1841]" italics="true" pageId="13" pageNumber="520">Diagnosis:</emphasis>
All but two of the 36 species of the genus are included in the subgenus
<taxonomicName id="4C104D6FFFB9F72CFB1D6816FB3CE33B" authorityName="Rambur" authorityYear="1866" box="[1167,1277,1851,1872]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="520" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAFEFFB9F72CFB1D6816FB3CE33B" box="[1167,1277,1851,1872]" italics="true" pageId="13" pageNumber="520">Grammia</emphasis>
</taxonomicName>
. Although the wing pattern is superficially like
<taxonomicName id="4C104D6FFFB9F72CFB796874FAA2E305" box="[1259,1379,1881,1902]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFB9F72CFB796874FAA2E305" box="[1259,1379,1881,1902]" italics="true" pageId="13" pageNumber="520">Mimarctia</emphasis>
</taxonomicName>
, the forewing costal cell is never entirely pale-scaled, and the costal portion of the postmedial band is not angled sharply basad. Structurally, the dorsal margin of the male juxta is broadly concave and u-shaped (somewhat v-shaped in
<taxonomicName id="4C104D6FFFBAF72FFEE76E6CFE2CE53D" box="[373,493,321,342]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFBAF72FFEE76E6CFE2CE53D" box="[373,493,321,342]" italics="true" pageId="14" pageNumber="521">Mimarctia</emphasis>
</taxonomicName>
). The male vesica is moderately developed with poorly to moderately differentiated chambers and diverticula (
<figureCitation id="132B2A69FFBAF72FFDF96E50FD2AE5FF" box="[619,747,381,404]" captionStart-0="Figures 40–45" captionStart-1="Figures 46–51" captionStart-2="Figures 52–57" captionStart-3="Figures 58–63" captionStart-4="Figures 64–68" captionStart-5="Figures 69–71" captionStartId-0="73.[164,254,1744,1763]" captionStartId-1="74.[144,234,1816,1835]" captionStartId-2="75.[164,254,1755,1774]" captionStartId-3="76.[144,234,1779,1798]" captionStartId-4="77.[164,254,1667,1686]" captionStartId-5="78.[144,234,1560,1579]" captionTargetBox-0="[226,1383,197,1711]" captionTargetBox-1="[221,1347,196,1783]" captionTargetBox-2="[263,1345,198,1722]" captionTargetBox-3="[216,1352,196,1746]" captionTargetBox-4="[273,1335,198,1635]" captionTargetBox-5="[254,1311,196,1527]" captionTargetId-0="figure-0@73.[224,1386,194,1714]" captionTargetId-1="figure-0@74.[219,1349,194,1786]" captionTargetId-2="figure-0@75.[260,1348,195,1725]" captionTargetId-3="figure-0@76.[213,1355,193,1749]" captionTargetId-4="figure-0@77.[270,1337,195,1637]" captionTargetId-5="figure-0@78.[254,1313,194,1530]" captionTargetPageId-0="73" captionTargetPageId-1="74" captionTargetPageId-2="75" captionTargetPageId-3="76" captionTargetPageId-4="77" captionTargetPageId-5="78" captionText-0="Figures 40–45. Genitalia of male Holarctia and Grammia, ventral aspect of valve (a) and right lateral aspect of aedeagus (b) (scale bar = 1 mm). 40, Holarctia obliterata (Stretch). 41, Grammia arge (Drury). 42, Grammia doris doris (Bdv.). 43, Grammia phyllira (Drury). 44, Grammia placentia (J. E. Smith). 45, Grammia virgo virgo (L.)." captionText-1="Figures 46–51. Genitalia of male Grammia, ventral aspect of valve and right lateral aspect of aedeagus (scale bar = 1 mm). 46, Grammia anna (Grt.). 47, Grammia parthenice (W. Kby.). 48, Grammia quenseli (Paykull). 49, Grammia virguncula (W. Kby.). 50, Grammia margo Schmidt. 51, Grammia franconia (H. Edw.)." captionText-2="Figures 52–57. Genitalia of male Grammia, ventral aspect of valve and right lateral aspect of aedeagus (scale bar = 1 mm). 52, Grammia figurata (Drury). 53, Grammia f-pallida (Stkr.). 54, Grammia incorrupta (Hy. Edwards). 55, Grammia allectans Ferguson. 56, Grammia ursina Schmidt. 57, Grammia nevadensis (G. &amp; R.)." captionText-3="Figures 58–63. Genitalia of male Grammia, ventral aspect of valve and right lateral aspect of aedeagus (scale bar = 1 mm). 58, Grammia behrii (Stretch). 59, Grammia eureka Ferguson &amp; Schmidt. 60, Grammia brillians Schmidt. 61, Grammia bowmani Ferguson &amp; Schmidt. 62, Grammia williamsii (Dodge). 63, Grammia elongata (Stretch)." captionText-4="Figures 64–68. Genitalia of male Grammia, ventral aspect of valve and right lateral aspect of aedeagus (scale bar = 1 mm). 64, Grammia yavapai Schmidt. 65, Grammia favorita (Neum.). 66, Grammia blakei (Grt.). 67, Grammia cervinoides (Stkr.). 68, Grammia fergusoni Schmidt." captionText-5="Figures 69–71. Genitalia of male Grammia, ventral aspect of valve and right lateral aspect of aedeagus (scale bar = 1 mm). 69, Grammia ornata (Pack.). 70, Grammia complicata (Walker). 71, Grammia hewletti (B. &amp; McD.)." figureDoi-0="http://doi.org/10.5281/zenodo.5444725" figureDoi-1="http://doi.org/10.5281/zenodo.5444727" figureDoi-2="http://doi.org/10.5281/zenodo.5444729" figureDoi-3="http://doi.org/10.5281/zenodo.5444731" figureDoi-4="http://doi.org/10.5281/zenodo.5444733" figureDoi-5="http://doi.org/10.5281/zenodo.5444735" httpUri-0="https://zenodo.org/record/5444725/files/figure.png" httpUri-1="https://zenodo.org/record/5444727/files/figure.png" httpUri-2="https://zenodo.org/record/5444729/files/figure.png" httpUri-3="https://zenodo.org/record/5444731/files/figure.png" httpUri-4="https://zenodo.org/record/5444733/files/figure.png" httpUri-5="https://zenodo.org/record/5444735/files/figure.png" pageId="14" pageNumber="521">Figs 44–71</figureCitation>
);
<taxonomicName id="4C104D6FFFBAF72FFF026EB0FEC9E5D9" box="[144,264,413,434]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFBAF72FFF026EB0FEC9E5D9" box="[144,264,413,434]" italics="true" pageId="14" pageNumber="521">Mimarctia</emphasis>
</taxonomicName>
exhibits large, well-defined chambers and diverticula (
<figureCitation id="132B2A69FFBAF72FFE8D6E96FE62E5BA" box="[287,419,443,465]" captionStart="Figures 40–45" captionStartId="73.[164,254,1744,1763]" captionTargetBox="[226,1383,197,1711]" captionTargetId="figure-0@73.[224,1386,194,1714]" captionTargetPageId="73" captionText="Figures 40–45. Genitalia of male Holarctia and Grammia, ventral aspect of valve (a) and right lateral aspect of aedeagus (b) (scale bar = 1 mm). 40, Holarctia obliterata (Stretch). 41, Grammia arge (Drury). 42, Grammia doris doris (Bdv.). 43, Grammia phyllira (Drury). 44, Grammia placentia (J. E. Smith). 45, Grammia virgo virgo (L.)." figureDoi="http://doi.org/10.5281/zenodo.5444725" httpUri="https://zenodo.org/record/5444725/files/figure.png" pageId="14" pageNumber="521">Figs 40, 41</figureCitation>
). The male distal portion of valve is well developed, simple, and rounded to slightly pointed (rounded or sickle-shaped in
<taxonomicName id="4C104D6FFFBAF72FFD396ED5FF04E647" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFBAF72FFD396ED5FF04E647" italics="true" pageId="14" pageNumber="521">Mimarctia</emphasis>
</taxonomicName>
). The female ductus bursae has two sclerotized lateral plates near the ostium, absent in
<taxonomicName id="4C104D6FFFBAF72FFDE56D1BFD2EE620" box="[631,751,566,587]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFBAF72FFDE56D1BFD2EE620" box="[631,751,566,587]" italics="true" pageId="14" pageNumber="521">Mimarctia</emphasis>
</taxonomicName>
. Where known, the cremaster of the pupa is short and broad, not elongate as in
<taxonomicName id="4C104D6FFFBAF72FFE5B6D5EFD80E6E3" box="[457,577,627,648]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFBAF72FFE5B6D5EFD80E6E3" box="[457,577,627,648]" italics="true" pageId="14" pageNumber="521">Mimarctia</emphasis>
</taxonomicName>
, and lacks the basal folds or grooves (except for
<taxonomicName id="4C104D6FFFBAF72FFD8E6DBCFDB8E6CD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[540,633,657,678]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="521" phylum="Arthropoda" rank="species" species="virgo">
<emphasis id="B964EAFEFFBAF72FFD8E6DBCFDB8E6CD" box="[540,633,657,678]" italics="true" pageId="14" pageNumber="521">G. virgo</emphasis>
</taxonomicName>
) of
<taxonomicName id="4C104D6FFFBAF72FFD396DBCFF04E6AE" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFBAF72FFD396DBCFF04E6AE" italics="true" pageId="14" pageNumber="521">Mimarctia</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF814B1DFFBAF72FFF466D82FEAEE6AE" author="Smith ME" box="[212,367,687,709]" pageId="14" pageNumber="521" refId="ref64404" refString="Smith ME. 1938 a. A revision of the genus Apantesis Walker (Lepidoptera, Arctiidae). Unpublished Ph. D. thesis. Urbana: University of Illinois." type="book" year="1938">Smith, 1938a</bibRefCitation>
, b). The larval tubercles are shiny silvery-white not dull as in
<taxonomicName id="4C104D6FFFBAF72FFE476DE2FD8CE68F" box="[469,589,719,740]" class="Insecta" family="Arctiidae" genus="Grammia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="568" phylum="Arthropoda" rank="subGenus" subGenus="Mimarctia">
<emphasis id="B964EAFEFFBAF72FFE476DE2FD8CE68F" box="[469,589,719,740]" italics="true" pageId="14" pageNumber="521">Mimarctia</emphasis>
</taxonomicName>
, with coarsely barbed (vs. finely barbed) setae (
<bibRefCitation id="EF814B1DFFBAF72FFD826DC0FD6FE768" author="Smith ME" box="[528,686,749,771]" pageId="14" pageNumber="521" refId="ref64404" refString="Smith ME. 1938 a. A revision of the genus Apantesis Walker (Lepidoptera, Arctiidae). Unpublished Ph. D. thesis. Urbana: University of Illinois." type="book" year="1938">Smith, 1938a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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