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<document id="6AAA522A71C2038FF456890F1FDF5023" ID-DOI="10.11646/zootaxa.3652.2.2" ID-GBIF-Dataset="0d490ce9-0872-4a4a-8721-ca1a10de1879" ID-ISSN="1175-5326" ID-Zenodo-Dep="219927" ID-ZooBank="B721A1E5-707A-476F-A3E7-E1B0D9559706" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460609319723" checkinUser="plazi" docAuthor="Chipman, Ariel D., Dor, Neta &amp; Bonato, Lucio" docDate="2013" docId="03B987F9143DFFE2CDB7D1FEF0DFB252" docLanguage="en" docName="zt03652p248.pdf" docOrigin="Zootaxa 3652 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachymerium ferrugineum Koch 1835" docType="treatment" docVersion="9" lastPageNumber="238" masterDocId="FF80FF811438FFE4CD20D717F30BB12D" masterDocTitle="Diversity and biogeography of Israeli geophilomorph centipedes (Chilopoda: Geophilomorpha)" masterLastPageNumber="248" masterPageNumber="232" pageNumber="237" updateTime="1698328166891" updateUser="plazi">
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<mods:title id="6DDCB2A14A1926BB06C9390FB6E84761">Diversity and biogeography of Israeli geophilomorph centipedes (Chilopoda: Geophilomorpha)</mods:title>
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<mods:namePart id="526A6FD44C77C2FB5FBA1CCDF18E7498">Chipman, Ariel D.</mods:namePart>
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<mods:namePart id="44425452771488456D73BC731AB4D106">Dor, Neta</mods:namePart>
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<mods:namePart id="2A8A670A9DB2F634890E4155CE518851">Bonato, Lucio</mods:namePart>
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<mods:date id="44AB7752523089AC9E56F04B60A577CD">2013</mods:date>
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<treatment id="03B987F9143DFFE2CDB7D1FEF0DFB252" ID-DOI="http://doi.org/10.5281/zenodo.5678043" ID-GBIF-Taxon="119568184" ID-Zenodo-Dep="5678043" LSID="urn:lsid:plazi:treatment:03B987F9143DFFE2CDB7D1FEF0DFB252" httpUri="http://treatment.plazi.org/id/03B987F9143DFFE2CDB7D1FEF0DFB252" lastPageId="6" lastPageNumber="238" pageId="5" pageNumber="237">
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<paragraph id="8BAF36EF143DFFE1CDB7D1FEF18CB62F" blockId="5.[151,647,1768,1795]" box="[151,647,1768,1795]" pageId="5" pageNumber="237">
<heading id="D0E78183143DFFE1CDB7D1FEF18CB62F" bold="true" box="[151,647,1768,1795]" fontSize="11" level="1" pageId="5" pageNumber="237" reason="1">
<taxonomicName id="4C104D6C143DFFE1CDB7D1FEF18CB62F" ID-CoL="4BSVF" authority="Koch, 1835" authorityName="Koch" authorityYear="1835" box="[151,647,1768,1795]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="237" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="B964EAFD143DFFE1CDB7D1FEF18CB62F" bold="true" box="[151,647,1768,1795]" pageId="5" pageNumber="237">
<emphasis id="B964EAFD143DFFE1CDB7D1FEF2EAB62E" bold="true" box="[151,481,1769,1795]" italics="true" pageId="5" pageNumber="237">Pachymerium ferrugineum</emphasis>
(Koch, 1835)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF36EF143DFFE1CDB7D038F3D4B6F5" blockId="5.[151,1436,1839,2008]" pageId="5" pageNumber="237">
Published records: “Jerusalem” (Verhoeff 1902, sub
<taxonomicName id="4C104D6C143DFFE1CFD4D026F700B665" box="[756,1035,1840,1864]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="237" phylum="Arthropoda" rank="subSpecies" species="ferrugineum" subSpecies="vosseleri">
<emphasis id="B964EAFD143DFFE1CFD4D026F700B665" box="[756,1035,1840,1864]" italics="true" pageId="5" pageNumber="237">P. ferrugineum vosseleri</emphasis>
</taxonomicName>
); “Chuldah” [Hulda], “El Muraw” [possibly near Jericho?], “Nahr Rubin” [Nahal Soreq], “Rehobot” [Rehovot], “Ranletz” [Ramleh?], “See Genezareth” [Sea of Galilee] (Verhoeff 1925, sub.
<taxonomicName id="4C104D6C143DFFE1CFD4D06EF714B6BD" box="[756,1055,1912,1936]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="237" phylum="Arthropoda" rank="subSpecies" species="ferrugineum" subSpecies="insulanum">
<emphasis id="B964EAFD143DFFE1CFD4D06EF714B6BD" box="[756,1055,1912,1936]" italics="true" pageId="5" pageNumber="237">P. ferrugineum insulanum</emphasis>
</taxonomicName>
); “Ein Fashka-Wadi Dara” [Ein Fashkha], “En Gedi” [Ein Gedi], “Nahal Bezet” [Nahal Betzet], “Wadi Shumriya” [Wadi Shomriya] (Zapparoli 1995).
</paragraph>
<paragraph id="8BAF36EF143EFFE2CDE9D78FF0C2B049" blockId="6.[151,1436,151,896]" pageId="6" pageNumber="238">
New records: Arad (
<specimenCount id="9D16FD66143EFFE2CC8FD780F130B182" box="[431,571,151,175]" pageId="6" pageNumber="238" type="generic">4 specimens</specimenCount>
1964);
<collectingCountry id="F307767F143EFFE2CFB3D78FF1C3B182" box="[659,712,152,175]" name="Senegal" pageId="6" pageNumber="238">Beer</collectingCountry>
Sheva (
<specimenCount id="9D16FD66143EFFE2CE07D780F0AEB182" box="[807,933,151,175]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2011); Bteha (
<specimenCount id="9D16FD66143EFFE2C970D780F7C5B182" box="[1104,1230,151,175]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
1934); Ein Tamar (
<specimenCount id="9D16FD66143EFFE2CD82D7ABF22AB1F9" box="[162,289,188,212]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
1963); Ezuz (
<specimenCount id="9D16FD66143EFFE2CCE8D7ABF14CB1F9" box="[456,583,188,212]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2011); Hermon, Emek HaMan (
<specimenCount id="9D16FD66143EFFE2CE9FD7ABF734B1F9" box="[959,1087,188,212]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2011); Jerusalem (
<specimenCount id="9D16FD66143EFFE2C83DD7ABF697B1F9" box="[1309,1436,188,212]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
1934); Jerusalem - Deir Es Sheikh (
<specimenCount id="9D16FD66143EFFE2CF6ED7C8F1D4B1DA" box="[590,735,223,247]" pageId="6" pageNumber="238" type="generic">3 specimens</specimenCount>
1944); Jerusalem - Talpiot (
<specimenCount id="9D16FD66143EFFE2C91ED7C8F7CAB1DA" box="[1086,1217,223,247]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
1957); Kalanit (
<specimenCount id="9D16FD66143EFFE2C8AED7C8F207B031" pageId="6" pageNumber="238" type="generic">3 specimens</specimenCount>
2011); Maale Adumim (
<specimenCount id="9D16FD66143EFFE2CF19D613F1CCB031" box="[569,711,260,284]" pageId="6" pageNumber="238" type="generic">2 specimens</specimenCount>
2011); Maale Efraim (
<specimenCount id="9D16FD66143EFFE2CEC1D613F769B031" box="[993,1122,260,284]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2011); Magenim Forest (
<specimenCount id="9D16FD66143EFFE2C8B0D613F20AB012" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2011); Nahal Afik (
<specimenCount id="9D16FD66143EFFE2CCCFD630F164B012" box="[495,623,295,319]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2010); Nahal Oren (
<specimenCount id="9D16FD66143EFFE2CE41D630F0E6B012" box="[865,1005,295,319]" pageId="6" pageNumber="238" type="generic">3 specimens</specimenCount>
2011); Tel Gezer (
<specimenCount id="9D16FD66143EFFE2C9EBD630F640B012" box="[1227,1355,295,319]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2011); Upper Nahal Deragot (
<specimenCount id="9D16FD66143EFFE2CCBAD65BF11CB049" box="[410,535,332,356]" pageId="6" pageNumber="238" type="generic">1 specimen</specimenCount>
2011); Yerucham (
<specimenCount id="9D16FD66143EFFE2CFCFD65BF072B049" box="[751,889,332,356]" pageId="6" pageNumber="238" type="generic">3 specimens</specimenCount>
1963);
</paragraph>
<paragraph id="8BAF36EF143EFFE2CDE9D678F0DFB0E2" blockId="6.[151,1436,151,896]" pageId="6" pageNumber="238">
Distribution in
<collectingCountry id="F307767F143EFFE2CC55D678F2BCB0AA" box="[373,439,367,391]" name="Israel" pageId="6" pageNumber="238">Israel</collectingCountry>
: the species has been found in a wide range of habitats, throughout the country, from the Dead Sea and the Negev desert up to the Upper Galilee and Mt. Hermon. Average annual temperatures are between 1624˚C and annual precipitations are between
<quantity id="4CE89B0A143EFFE2CF8ED6A0F048B0E2" box="[686,835,439,463]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="12.5" metricValueMin="0.5" pageId="6" pageNumber="238" unit="mm" value="650.0" valueMax="1250.0" valueMin="50.0">501250 mm</quantity>
(
<figureCitation id="132B2A6A143EFFE2CE71D6A0F0BAB0E2" box="[849,945,439,463]" captionStart="FIGURE 3" captionStartId="9.[151,250,1941,1964]" captionTargetBox="[380,1139,222,1903]" captionTargetId="figure@9.[363,1178,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Sampling locations and presumptive distribution sites for members of the Geophilidae and Mecistocephalidae. A) Clinopodes escherichii. B) Geophilus cf. carpophagus. C) Pachymerium ferrugineum. D) Stenotaenia naxia. E) Stenotaenia palaestina. F) Mecistocephalus evansi." httpUri="https://zenodo.org/record/219930/files/figure.png" pageId="6" pageNumber="238">Figure 3</figureCitation>
C).
</paragraph>
<paragraph id="8BAF36EF143EFFE2CDE9D6CBF686B0D9" blockId="6.[151,1436,151,896]" box="[201,1421,476,500]" pageId="6" pageNumber="238">Global distribution: the species is widespread in the western Palearctic, but it has been introduced elsewhere.</paragraph>
<paragraph id="8BAF36EF143EFFE2CDE9D6E8F0DFB252" blockId="6.[151,1436,151,896]" pageId="6" pageNumber="238">
Taxonomic and nomenclatural notes.
<taxonomicName id="4C104D6C143EFFE2CF4AD517F7A6B33A" authority="Verhoeff, 1902" authorityName="Verhoeff" authorityYear="1902" box="[618,1197,511,536]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="6" pageNumber="238" phylum="Arthropoda" rank="subSpecies" species="ferrugineum" subSpecies="vosseleri">
<emphasis id="B964EAFD143EFFE2CF4AD517F70AB335" box="[618,1025,512,536]" italics="true" pageId="6" pageNumber="238">Pachymerium ferrugineum vosseleri</emphasis>
Verhoeff, 1902
</taxonomicName>
is here recognized as a junior synonym of
<taxonomicName id="4C104D6C143EFFE2CCA8D533F05DB311" authority="Koch, 1835" authorityName="Koch" authorityYear="1835" box="[392,854,548,572]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="B964EAFD143EFFE2CCA8D533F1BCB311" box="[392,695,548,572]" italics="true" pageId="6" pageNumber="238">Pachymerium ferrugineum</emphasis>
(Koch, 1835)
</taxonomicName>
(
<emphasis id="B964EAFD143EFFE2CE49D533F0BBB311" bold="true" box="[873,944,548,572]" pageId="6" pageNumber="238">n. syn</emphasis>
.). Doubts on the validity of the subspecies
<emphasis id="B964EAFD143EFFE2CDB7D55FF3F0B34D" box="[151,251,584,608]" italics="true" pageId="6" pageNumber="238">vosseleri</emphasis>
have been already expressed by Zapparoli (1991). After re-assessing all information provided by Verhoeff (1902 1925 1934) and after examining specimens of
<taxonomicName id="4C104D6C143EFFE2CE2CD57AF0BCB3A9" box="[780,951,620,644]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="B964EAFD143EFFE2CE2CD57AF0BCB3A9" box="[780,951,620,644]" italics="true" pageId="6" pageNumber="238">P. ferrugineum</emphasis>
</taxonomicName>
from the
<typeStatus id="54AB884D143EFFE2C913D57AF768B3A8" box="[1075,1123,621,645]" pageId="6" pageNumber="238">type</typeStatus>
locality of
<taxonomicName id="4C104D6C143EFFE2C9D1D57AF28EB38A" authority="(Jerusalem)" baseAuthorityName="Jerusalem" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="6" pageNumber="238" phylum="Arthropoda" rank="subSpecies" species="ferrugineum" subSpecies="vosseleri">
<emphasis id="B964EAFD143EFFE2C9D1D57AF3F0B385" italics="true" pageId="6" pageNumber="238">P. ferrugineum vosseleri</emphasis>
(Jerusalem)
</taxonomicName>
and other sites in
<collectingCountry id="F307767F143EFFE2CF75D598F19DB38A" box="[597,662,655,679]" name="Israel" pageId="6" pageNumber="238">Israel</collectingCountry>
, we found no evidence for any significant difference between
<collectingCountry id="F307767F143EFFE2C876D598F697B38A" box="[1366,1436,655,679]" name="Israel" pageId="6" pageNumber="238">Israeli</collectingCountry>
populations of
<taxonomicName id="4C104D6C143EFFE2CC6AD5A2F2FEB3E1" box="[330,501,692,716]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="B964EAFD143EFFE2CC6AD5A2F2FEB3E1" box="[330,501,692,716]" italics="true" pageId="6" pageNumber="238">P. ferrugineum</emphasis>
</taxonomicName>
and populations from other parts of the species range. All the few characters described by Verhoeff (1902) for the two
<typeStatus id="54AB884D143EFFE2CFA5D5CFF1ECB3DD" box="[645,743,728,752]" pageId="6" pageNumber="238" type="syntype">syntypes</typeStatus>
of the subspecies
<emphasis id="B964EAFD143EFFE2CEE6D5CFF722B3DD" box="[966,1065,728,752]" italics="true" pageId="6" pageNumber="238">vosseleri</emphasis>
(almost exclusively: body size, number of denticles and setae on the mouth parts, and number of legs) are within the range of variation reported in typical
<taxonomicName id="4C104D6C143EFFE2CDCDD436F29DB215" box="[237,406,800,824]" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="B964EAFD143EFFE2CDCDD436F29DB215" box="[237,406,800,824]" italics="true" pageId="6" pageNumber="238">P. ferrugineum</emphasis>
</taxonomicName>
by other authors. With respect to the number of legs, we found that it is highly variable between specimens from different sites in
<collectingCountry id="F307767F143EFFE2CF8FD453F1E6B271" box="[687,749,836,860]" name="Israel" pageId="6" pageNumber="238">Israel</collectingCountry>
(4769 pairs), but without any obvious geographic or environmental pattern, and with all possible intermediate values recorded.
</paragraph>
</subSubSection>
</treatment>
</document>