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<document id="D403869189E31B387B746709E6410812" ID="10.11646/zootaxa.4263.3.6SLASH11002" ID-CLB-Dataset="33071" ID-DOI="10.11646/zootaxa.4263.3.6" ID-GBIF-Dataset="892554a9-032d-4aee-a5d1-da1a46043dfa" ID-ISSN="1175-5326" ID-Zenodo-Dep="573667" ID-ZooBank="A08F6DE4-9A1D-4687-B84B-57E1E67C4720" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494405758088" checkinUser="plazi" docAuthor="Wang, Zongqing &amp; Che, Yanli" docDate="2017" docId="03B987DDDB47FFD9F487E758D812FF62" docLanguage="en" docName="zootaxa.4263.3.6SLASH11002.pdf" docOrigin="Zootaxa 4263 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemithyrsocera forcipata Wang &amp; Che, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="552" masterDocId="FF80FFA5DB40FFD0F410E539DA70FF9A" masterDocTitle="Three new species of cockroach genus Hemithyrsocera Saussure, 1893 (Blattodea: Ectobiidae: Blattellinae) with redescriptions of two known species from China" masterLastPageNumber="556" masterPageNumber="543" pageNumber="550" updateTime="1698439448157" updateUser="ExternalLinkService">
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<mods:title id="87140EE252C4B5304830C11F44EC115A">Three new species of cockroach genus Hemithyrsocera Saussure, 1893 (Blattodea: Ectobiidae: Blattellinae) with redescriptions of two known species from China</mods:title>
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<mods:namePart id="9288A42799E2258C11CFBFC045B9CCB9">Wang, Zongqing</mods:namePart>
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<mods:namePart id="3E08B709CDB3CDF50EBD15CC390FAE5D">Che, Yanli</mods:namePart>
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<mods:date id="DDDE6F36C165E992EF289C4BDA9F8443">2017</mods:date>
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<treatment id="03B987DDDB47FFD9F487E758D812FF62" ID-DOI="http://doi.org/10.5281/zenodo.6025513" ID-GBIF-Taxon="130509356" ID-Zenodo-Dep="6025513" LSID="urn:lsid:plazi:treatment:03B987DDDB47FFD9F487E758D812FF62" httpUri="http://treatment.plazi.org/id/03B987DDDB47FFD9F487E758D812FF62" lastPageId="9" lastPageNumber="552" pageId="7" pageNumber="550">
<subSubSection id="C30A6540DB47FFD7F487E758D83FFDE1" box="[151,591,608,635]" pageId="7" pageNumber="550" type="nomenclature">
<paragraph id="8BAF36CBDB47FFD7F487E758D83FFDE1" blockId="7.[151,591,608,669]" box="[151,591,608,635]" pageId="7" pageNumber="550">
<heading id="D0E781A7DB47FFD7F487E758D83FFDE1" bold="true" box="[151,591,608,635]" fontSize="11" level="1" pageId="7" pageNumber="550" reason="1">
<emphasis id="B964EAD9DB47FFD7F487E758D83FFDE1" bold="true" box="[151,591,608,635]" pageId="7" pageNumber="550">
3.
<taxonomicName id="4C104D48DB47FFD7F4A4E759DB9AFDE0" ID-CoL="8N9KY" box="[180,490,608,634]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="forcipata" status="sp. nov.">
<emphasis id="B964EAD9DB47FFD7F4A4E759DB9AFDE0" bold="true" box="[180,490,608,634]" italics="true" pageId="7" pageNumber="550">Hemithyrsocera forcipata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757A2DB47FFD7F5E1E758D83FFDE1" box="[497,591,609,635]" pageId="7" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A6540DB47FFD7F487E7BDD846F917" pageId="7" pageNumber="550" type="description">
<paragraph id="8BAF36CBDB47FFD7F487E7BDDB1FFD07" blockId="7.[151,591,608,669]" box="[151,367,644,669]" pageId="7" pageNumber="550">
<figureCitation id="132B2A4EDB47FFD7F487E7BDDB6DFD07" box="[151,285,644,669]" captionStart="FIGURES 10 19. 10 11" captionStartId="3.[151,264,1880,1903]" captionTargetBox="[159,1431,530,1833]" captionTargetId="figure@3.[151,1436,519,1859]" captionTargetPageId="3" captionText="FIGURES 10 19. 10 11 Hemithyrsocera simulans (Bey-Bienko, 1969), male: 10 dorsal view; 11 ventral view. 12 13 Hemithyrsocera banvaneuensis Roth, 1985, male: 12 dorsal view; 13 ventral view. 14 15 Hemithyrsocera forcipata sp. nov., holotype: 14 dorsal view; 15 ventral view. 16 17 Hemithyrsocera spinibarbis sp. nov., holotype: 16 dorsal view; 17 ventral view. 18 19 Hemithyrsocera longiseta sp. nov., holotype: 18 dorsal view; 19 ventral view. Scale bars = 1.0 cm." httpUri="https://zenodo.org/record/573669/files/figure.png" pageId="7" pageNumber="550">Figs. 1415</figureCitation>
,
<figureCitation id="132B2A4EDB47FFD7F539E7BDDB1FFD07" box="[297,367,644,669]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">4455</figureCitation>
</paragraph>
<paragraph id="8BAF36CBDB47FFD7F487E7FFD8FBFCBD" blockId="7.[151,1437,710,2037]" pageId="7" pageNumber="550">
<emphasis id="B964EAD9DB47FFD7F487E7FFDB36FD45" bold="true" box="[151,326,710,735]" pageId="7" pageNumber="550">Measurements</emphasis>
(mm).
<emphasis id="B964EAD9DB47FFD7F58AE7FFDBA7FD45" bold="true" box="[410,471,710,735]" pageId="7" pageNumber="550">Male</emphasis>
, overall length including tegmina: 17.518.5; pronotum length × width: 3.13.4×4.1 4.4; tegmina length: 15.015.3.
<emphasis id="B964EAD9DB47FFD7F617E7D3D82EFC99" bold="true" box="[519,606,746,771]" pageId="7" pageNumber="550">Female</emphasis>
, overall length including tegmina: 16.017.1; pronotum length × width: 3.13.5×4.04.3; tegmina length: 14.015.0.
</paragraph>
<paragraph id="8BAF36CBDB47FFD7F4D7E60BDB2BFBBC" blockId="7.[151,1437,710,2037]" pageId="7" pageNumber="550">
Body dark brown (
<figureCitation id="132B2A4EDB47FFD7F5B1E60BDB87FCD1" box="[417,503,818,843]" captionStart="FIGURES 10 19. 10 11" captionStartId="3.[151,264,1880,1903]" captionTargetBox="[159,1431,530,1833]" captionTargetId="figure@3.[151,1436,519,1859]" captionTargetPageId="3" captionText="FIGURES 10 19. 10 11 Hemithyrsocera simulans (Bey-Bienko, 1969), male: 10 dorsal view; 11 ventral view. 12 13 Hemithyrsocera banvaneuensis Roth, 1985, male: 12 dorsal view; 13 ventral view. 14 15 Hemithyrsocera forcipata sp. nov., holotype: 14 dorsal view; 15 ventral view. 16 17 Hemithyrsocera spinibarbis sp. nov., holotype: 16 dorsal view; 17 ventral view. 18 19 Hemithyrsocera longiseta sp. nov., holotype: 18 dorsal view; 19 ventral view. Scale bars = 1.0 cm." httpUri="https://zenodo.org/record/573669/files/figure.png" pageId="7" pageNumber="550">Fig. 14</figureCitation>
). Vertex yellowish brown. Face brown with wide yellowish brown band between eyes (
<figureCitation id="132B2A4EDB47FFD7F4CEE66FDB49FCF5" box="[222,313,854,879]" captionStart="FIGURES 10 19. 10 11" captionStartId="3.[151,264,1880,1903]" captionTargetBox="[159,1431,530,1833]" captionTargetId="figure@3.[151,1436,519,1859]" captionTargetPageId="3" captionText="FIGURES 10 19. 10 11 Hemithyrsocera simulans (Bey-Bienko, 1969), male: 10 dorsal view; 11 ventral view. 12 13 Hemithyrsocera banvaneuensis Roth, 1985, male: 12 dorsal view; 13 ventral view. 14 15 Hemithyrsocera forcipata sp. nov., holotype: 14 dorsal view; 15 ventral view. 16 17 Hemithyrsocera spinibarbis sp. nov., holotype: 16 dorsal view; 17 ventral view. 18 19 Hemithyrsocera longiseta sp. nov., holotype: 18 dorsal view; 19 ventral view. Scale bars = 1.0 cm." httpUri="https://zenodo.org/record/573669/files/figure.png" pageId="7" pageNumber="550">Fig. 15</figureCitation>
). Ocellar spot pale yellow. Third maxillary palpomere yellowish brown with fourth and fifth maxillary palpomeres brown (
<figureCitation id="132B2A4EDB47FFD7F5E3E643D83BFC09" box="[499,587,890,915]" captionStart="FIGURES 10 19. 10 11" captionStartId="3.[151,264,1880,1903]" captionTargetBox="[159,1431,530,1833]" captionTargetId="figure@3.[151,1436,519,1859]" captionTargetPageId="3" captionText="FIGURES 10 19. 10 11 Hemithyrsocera simulans (Bey-Bienko, 1969), male: 10 dorsal view; 11 ventral view. 12 13 Hemithyrsocera banvaneuensis Roth, 1985, male: 12 dorsal view; 13 ventral view. 14 15 Hemithyrsocera forcipata sp. nov., holotype: 14 dorsal view; 15 ventral view. 16 17 Hemithyrsocera spinibarbis sp. nov., holotype: 16 dorsal view; 17 ventral view. 18 19 Hemithyrsocera longiseta sp. nov., holotype: 18 dorsal view; 19 ventral view. Scale bars = 1.0 cm." httpUri="https://zenodo.org/record/573669/files/figure.png" pageId="7" pageNumber="550">Fig. 15</figureCitation>
). Pronotum dark brown with yellowish brown stripe, lateral and anterior borders yellowish brown (
<figureCitation id="132B2A4EDB47FFD7F5D0E6A7D86FFC2D" box="[448,543,926,951]" captionStart="FIGURES 10 19. 10 11" captionStartId="3.[151,264,1880,1903]" captionTargetBox="[159,1431,530,1833]" captionTargetId="figure@3.[151,1436,519,1859]" captionTargetPageId="3" captionText="FIGURES 10 19. 10 11 Hemithyrsocera simulans (Bey-Bienko, 1969), male: 10 dorsal view; 11 ventral view. 12 13 Hemithyrsocera banvaneuensis Roth, 1985, male: 12 dorsal view; 13 ventral view. 14 15 Hemithyrsocera forcipata sp. nov., holotype: 14 dorsal view; 15 ventral view. 16 17 Hemithyrsocera spinibarbis sp. nov., holotype: 16 dorsal view; 17 ventral view. 18 19 Hemithyrsocera longiseta sp. nov., holotype: 18 dorsal view; 19 ventral view. Scale bars = 1.0 cm." httpUri="https://zenodo.org/record/573669/files/figure.png" pageId="7" pageNumber="550">Figs. 14</figureCitation>
,
<figureCitation id="132B2A4EDB47FFD7F63CE6A7D83CFC2D" box="[556,588,926,951]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">46</figureCitation>
). Tegmina brown with discoidal area yellowish brown; hind wings hyaline and pale yellowish brown. Fore legs pale brown with apices of all tibiae dark brown, femora pale brown with dark brown stripes. 6th8th sternites yellowish brown with dark brown irregular maculae on the middle and lateral borders (
<figureCitation id="132B2A4EDB47FFD7F4EAE134DB3DFBBC" box="[250,333,1037,1062]" captionStart="FIGURES 10 19. 10 11" captionStartId="3.[151,264,1880,1903]" captionTargetBox="[159,1431,530,1833]" captionTargetId="figure@3.[151,1436,519,1859]" captionTargetPageId="3" captionText="FIGURES 10 19. 10 11 Hemithyrsocera simulans (Bey-Bienko, 1969), male: 10 dorsal view; 11 ventral view. 12 13 Hemithyrsocera banvaneuensis Roth, 1985, male: 12 dorsal view; 13 ventral view. 14 15 Hemithyrsocera forcipata sp. nov., holotype: 14 dorsal view; 15 ventral view. 16 17 Hemithyrsocera spinibarbis sp. nov., holotype: 16 dorsal view; 17 ventral view. 18 19 Hemithyrsocera longiseta sp. nov., holotype: 18 dorsal view; 19 ventral view. Scale bars = 1.0 cm." httpUri="https://zenodo.org/record/573669/files/figure.png" pageId="7" pageNumber="550">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BAF36CBDB47FFD7F4D7E10BD9E2FA67" blockId="7.[151,1437,710,2037]" pageId="7" pageNumber="550">
Vertex with interocular space distinctly narrower than the distance between antennal sockets (
<figureCitation id="132B2A4EDB47FFD7F0E3E108DF39FBD0" box="[1267,1353,1073,1098]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 44</figureCitation>
). Third and fourth maxillary palpomeres approximately same length, and both slightly longer than the fifth (
<figureCitation id="132B2A4EDB47FFD7F127E16CDFE0FBF4" box="[1335,1424,1109,1134]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 45</figureCitation>
). Pronotum subelliptical, the middle of hind margin more or less produced (
<figureCitation id="132B2A4EDB47FFD7F7FCE140DE33FB08" box="[1004,1091,1145,1170]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 46</figureCitation>
). Hind wings with radius area narrow, R1 with 4 branches, posterior branches of R extended behind the base of anterior rami of radius and bifurcated at apical; M simple, Cu with two branches; apical triangle distinct. Front femur
<typeStatus id="54AB8869DB47FFD7F084E1FBDEBCFB40" box="[1172,1228,1218,1242]" pageId="7" pageNumber="550">Type</typeStatus>
A2 or A3 (
<figureCitation id="132B2A4EDB47FFD7F12BE1F8DFE0FB40" box="[1339,1424,1217,1242]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 47</figureCitation>
). Seventh abdominal tergum specialized but without setae; laterally on each side, a rounded lobe, covered with minute setae, directed laterad and with base robust (
<figureCitation id="132B2A4EDB47FFD7F6C9E035D95CFABF" box="[729,812,1292,1317]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 50</figureCitation>
). Supra-anal plate (
<figureCitation id="132B2A4EDB47FFD7F014E035DE25FABF" box="[1028,1109,1292,1317]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 51</figureCitation>
) in ventral view symmetrical with hind margin rounded, right and left intercercal processes absent; right and left paraprocts dissimilar (
<figureCitation id="132B2A4EDB47FFD7F12BE009DFFEFAD3" box="[1339,1422,1328,1353]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 51</figureCitation>
), right one bifurcated with apices rounded or tapering, one larger spine arising from the middle; left one similarly bifurcated with apices truncate, one small spine arising from the middle. Subgenital plate in dorsal view strongly asymmetrical (
<figureCitation id="132B2A4EDB47FFD7F52DE0A5DBE1FA2F" box="[317,401,1436,1461]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 52</figureCitation>
), right side distinctly produced into one rounded lobe, whose apex is upturned and scattered with apical spines; two styli dissimilar with apical long hairs, arising from the middle of hind margin; hind margin with 3 processes, whose apices scattered with lots of teeth (
<figureCitation id="132B2A4EDB47FFD7F720E0DDD9F5FA67" box="[816,901,1508,1533]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="8BAF36CBDB47FFD7F4D7E331D846F917" blockId="7.[151,1437,710,2037]" pageId="7" pageNumber="550">
Male genitalia. L3 (
<figureCitation id="132B2A4EDB47FFD7F5B8E331DB8AF9BB" box="[424,506,1544,1569]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 53</figureCitation>
) long and slender, sclerotized portion small and without preapical incision; L2vm (
<figureCitation id="132B2A4EDB47FFD7F48FE315DA82F9DF" box="[159,242,1580,1605]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 54</figureCitation>
) long, curved and slender with base pointed, one short and irregular accessory structure originated in the middle; R2 (
<figureCitation id="132B2A4EDB47FFD7F536E369DB09F9F3" box="[294,377,1616,1641]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 55</figureCitation>
) simple and irregular, R3 (
<figureCitation id="132B2A4EDB47FFD7F6BEE369D971F9F3" box="[686,769,1616,1641]" captionStart="FIGURES 44 55" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[217,1382,194,1868]" captionTargetId="figure@8.[196,1385,193,1878]" captionTargetPageId="8" captionText="FIGURES 44 55. Hemithyrsocera forcipata sp. nov. male: 44 head; 45 maxillary palp segments 3 5; 46 pronotum; 47 front femur; 48 tegmen; 49 hind wing; 50 seventh abdominal tergum; 51 supra-anal plate and paraprocts, ventral view; 52 subgenital plate, dorsal view; 53 hook-like phallomere (L 3); 54 median phallomere (L 2 vm); 55 right phallomere (R 2 and R 3). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="7" pageNumber="550">Fig. 55</figureCitation>
) with 2 obvious sclerites, one of which nearly inversed Yshaped, the other small and irregular.
</paragraph>
</subSubSection>
<subSubSection id="C30A6540DB47FFD7F4D7E3A1DB70F887" pageId="7" pageNumber="550" type="materials_examined">
<paragraph id="8BAF36CBDB47FFD7F4D7E3A1D940F94F" blockId="7.[151,1437,710,2037]" pageId="7" pageNumber="550">
<materialsCitation id="3B783C96DB47FFD7F4D7E3A1D940F94F" ID-GBIF-Occurrence="1502353026" collectingDate="2014-05-28" collectorName="Li Xinran." country="China" county="Material" elevation="1299" latitude="22.369833" location="Jinghong" longLatPrecision="1" longitude="100.949615" municipality="Xishuangbanna" pageId="7" pageNumber="550" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B964EAD9DB47FFD7F4D7E3A1D843F92B" bold="true" box="[199,563,1688,1713]" pageId="7" pageNumber="550">
<collectingCounty id="62CE4E47DB47FFD7F4D7E3A1DB5FF92B" box="[199,303,1688,1713]" pageId="7" pageNumber="550">Material</collectingCounty>
examined.
<typeStatus id="54AB8869DB47FFD7F5D6E3A1D843F92B" box="[454,563,1688,1713]" pageId="7" pageNumber="550" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D16FD42DB47FFD7F659E3A0D810F92A" box="[585,608,1689,1712]" pageId="7" pageNumber="550" type="male"></specimenCount>
,
<collectingCountry id="F307765BDB47FFD7F663E3A1D8C9F92B" box="[627,697,1688,1713]" name="China" pageId="7" pageNumber="550">China</collectingCountry>
,
<collectingRegion id="49D4F829DB47FFD7F6DBE3A0D951F92A" box="[715,801,1689,1712]" country="China" name="Yunnan" pageId="7" pageNumber="550">Yunnan</collectingRegion>
Prov.,
<collectingMunicipality id="6BCBACB1DB47FFD7F76FE3A1DE40F92B" box="[895,1072,1688,1713]" pageId="7" pageNumber="550">Xishuangbanna</collectingMunicipality>
,
<location id="8ECF6010DB47FFD7F052E3A1DEDCF92B" LSID="urn:lsid:plazi:treatment:03B987DDDB47FFD9F487E758D812FF62:8ECF6010DB47FFD7F052E3A1DEDCF92B" box="[1090,1196,1688,1713]" country="China" county="Material" latitude="22.369833" longLatPrecision="1" longitude="100.949615" municipality="Xishuangbanna" name="Jinghong" pageId="7" pageNumber="550" stateProvince="Yunnan">Jinghong</location>
,
<location id="8ECF6010DB47FFD7F0AFE3A1DF45F92B" LSID="urn:lsid:plazi:treatment:03B987DDDB47FFD9F487E758D812FF62:8ECF6010DB47FFD7F0AFE3A1DF45F92B" box="[1215,1333,1688,1713]" country="China" county="Material" latitude="22.369833" longLatPrecision="1" longitude="100.949615" municipality="Xishuangbanna" name="Dadugang" pageId="7" pageNumber="550" stateProvince="Yunnan">Dadugang</location>
,
<elevation id="003DD1F8DB47FFD7F157E3A1DFE8F92A" box="[1351,1432,1688,1712]" metricMagnitude="3" metricUnit="m" metricValue="1.299" pageId="7" pageNumber="550" unit="m" value="1299.0">
<quantity id="4CE89B2EDB47FFD7F157E3A1DFE8F92A" box="[1351,1432,1688,1712]" metricMagnitude="3" metricUnit="m" metricValue="1.299" pageId="7" pageNumber="550" unit="m" value="1299.0">1299m</quantity>
</elevation>
,
<geoCoordinate id="EE24500CDB47FFD7F487E384DB5EF94E" box="[151,302,1725,1749]" direction="north" orientation="latitude" pageId="7" pageNumber="550" precision="1" value="22.369833">22°22.190' N</geoCoordinate>
,
<geoCoordinate id="EE24500CDB47FFD7F529E385DBABF94E" box="[313,475,1724,1749]" direction="east" orientation="longitude" pageId="7" pageNumber="550" precision="1" value="100.949615">100°56.977' E</geoCoordinate>
,
<collectingDate id="EFEAE9E3DB47FFD7F5F6E384D80FF94F" box="[486,639,1725,1749]" pageId="7" pageNumber="550" value="2014-05-28">
<date id="FFAE100BDB47FFD7F5F6E384D80FF94F" box="[486,639,1725,1749]" pageId="7" pageNumber="550" value="2014-05-28">28 May 2014</date>
</collectingDate>
, leg.
<collectorName id="26E5531DDB47FFD7F6AAE385D940F94F" box="[698,816,1724,1749]" pageId="7" pageNumber="550">Li Xinran.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BAF36CBDB47FFD7F4D7E3D9DB70F887" blockId="7.[151,1437,710,2037]" pageId="7" pageNumber="550">
<emphasis id="B964EAD9DB47FFD7F4D7E3D9DB4FF963" bold="true" box="[199,319,1760,1785]" pageId="7" pageNumber="550">Paratypes</emphasis>
: 1♂, Xishuangbanna, Jinghong, Dadugang,
<date id="FFAE100BDB47FFD7F727E3D9D9A2F962" box="[823,978,1760,1785]" pageId="7" pageNumber="550" value="2014-05-29">29 May 2014</date>
, leg. Liu Hongguang;
<materialsCitation id="3B783C96DB47FFD7F0C2E3D9DB70F887" ID-GBIF-Occurrence="1502353024" collectingDate="2014-05-28" collectorName="Li Xinran." country="China" county="Material" elevation="1299" latitude="22.369833" location="Jinghong" longLatPrecision="1" longitude="100.949615" municipality="Xishuangbanna" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D16FD42DB47FFD7F0C2E3D9DE8DF962" box="[1234,1277,1760,1784]" pageId="7" pageNumber="550" type="female">1 ♀</specimenCount>
, same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A6540DB47FFD7F4D7E211DE68F86F" pageId="7" pageNumber="550" type="discussion">
<paragraph id="8BAF36CBDB47FFD7F4D7E211DE68F86F" blockId="7.[151,1437,710,2037]" pageId="7" pageNumber="550">
<emphasis id="B964EAD9DB47FFD7F4D7E211DB4BF8DB" bold="true" box="[199,315,1832,1857]" pageId="7" pageNumber="550">Remarks.</emphasis>
<taxonomicName id="4C104D48DB47FFD7F555E210DBA1F8DA" box="[325,465,1832,1856]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis id="B964EAD9DB47FFD7F555E210DBA1F8DA" box="[325,465,1832,1856]" italics="true" pageId="7" pageNumber="550">H. forcipata</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C104D48DB47FFD7F6D8E210DE4FF8DB" authority="Bey-Bienko, 1969" authorityName="Bey-Bienko" authorityYear="1969" box="[712,1087,1832,1857]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B964EAD9DB47FFD7F6D8E210D922F8DA" box="[712,850,1832,1856]" italics="true" pageId="7" pageNumber="550">H. simulans</emphasis>
(
<bibRefCitation id="EF814B3ADB47FFD7F773E211DE47F8DB" author="Bey-Bienko" box="[867,1079,1832,1857]" pageId="7" pageNumber="550" refString="Bey-Bienko, G. Y. (1969) New genera and species of cockroaches (Blattoptera) from tropical and subtropical Asia. Entomologicheskoe Obozrenie, 48, 831 - 862." type="journal article" year="1969">Bey-Bienko, 1969</bibRefCitation>
)
</taxonomicName>
based on the body shape and color. It differs from
<taxonomicName id="4C104D48DB47FFD7F59CE274D865F8FE" box="[396,533,1868,1892]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B964EAD9DB47FFD7F59CE274D865F8FE" box="[396,533,1868,1892]" italics="true" pageId="7" pageNumber="550">H. simulans</emphasis>
</taxonomicName>
in the following: 1) right side of subgenital plate distinctly produced into one rounded lobe, two styli arising from the middle of hind margin; hind margin with 3 processes (in
<taxonomicName id="4C104D48DB47FFD7F0C4E248DF29F812" box="[1236,1369,1904,1928]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B964EAD9DB47FFD7F0C4E248DF29F812" box="[1236,1369,1904,1928]" italics="true" pageId="7" pageNumber="550">H. simulans</emphasis>
</taxonomicName>
, right posterolateral corners rounded, two styli arising from the middle and left corner of hind margin, hind margin uneven with 2 robust fingerlike processes); 2) apex of L2vm rounded (tapering in
<taxonomicName id="4C104D48DB47FFD7F02DE280DEB4F84A" box="[1085,1220,1976,2000]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B964EAD9DB47FFD7F02DE280DEB4F84A" box="[1085,1220,1976,2000]" italics="true" pageId="7" pageNumber="550">H. simulans</emphasis>
</taxonomicName>
); 3) both right and left paraprocts with small spines arising from the middle (large in
<taxonomicName id="4C104D48DB47FFD7F76BE2E4DE71F86E" box="[891,1025,2012,2036]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B964EAD9DB47FFD7F76BE2E4DE71F86E" box="[891,1025,2012,2036]" italics="true" pageId="7" pageNumber="550">H. simulans</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF6F6643DB48FFD8F487E249DB70F879" httpUri="https://zenodo.org/record/573672/files/figure.png" pageId="8" pageNumber="551" targetBox="[217,1382,194,1868]" targetPageId="8">
<paragraph id="8BAF36CBDB48FFD8F487E249DB70F879" blockId="8.[151,1436,1904,2019]" pageId="8" pageNumber="551">
<emphasis id="B964EAD9DB48FFD8F487E249DB3EF81C" bold="true" box="[151,334,1904,1927]" pageId="8" pageNumber="551">FIGURES 4455</emphasis>
.
<taxonomicName id="4C104D48DB48FFD8F54CE248D812F81D" box="[348,610,1905,1927]" class="Insecta" family="Ectobiidae" genus="Hemithyrsocera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="551" phylum="Arthropoda" rank="species" species="forcipata" status="sp. nov.">
<emphasis id="B964EAD9DB48FFD8F54CE248D812F81D" box="[348,610,1905,1927]" italics="true" pageId="8" pageNumber="551">Hemithyrsocera forcipata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757A2DB48FFD8F67AE248D8CBF81C" box="[618,699,1905,1926]" pageId="8" pageNumber="551" rank="species">
<emphasis id="B964EAD9DB48FFD8F67AE248D8CBF81C" bold="true" box="[618,699,1905,1926]" pageId="8" pageNumber="551">sp. nov.</emphasis>
</taxonomicNameLabel>
male:
<emphasis id="B964EAD9DB48FFD8F713E248D96CF81C" bold="true" box="[771,796,1905,1926]" pageId="8" pageNumber="551">44</emphasis>
head;
<emphasis id="B964EAD9DB48FFD8F772E248D90BF81C" bold="true" box="[866,891,1905,1926]" pageId="8" pageNumber="551">45</emphasis>
maxillary palp segments 35;
<emphasis id="B964EAD9DB48FFD8F0A8E248DEA1F81C" bold="true" box="[1208,1233,1905,1926]" pageId="8" pageNumber="551">46</emphasis>
pronotum;
<emphasis id="B964EAD9DB48FFD8F15AE248DF14F81C" bold="true" box="[1354,1380,1905,1926]" pageId="8" pageNumber="551">47</emphasis>
front femur;
<emphasis id="B964EAD9DB48FFD8F4F0E2A9DA8AF83F" bold="true" box="[224,250,1936,1957]" pageId="8" pageNumber="551">48</emphasis>
tegmen;
<emphasis id="B964EAD9DB48FFD8F548E2A9DB01F83F" bold="true" box="[344,369,1936,1957]" pageId="8" pageNumber="551">49</emphasis>
hind wing;
<emphasis id="B964EAD9DB48FFD8F5FAE2A9D874F83F" bold="true" box="[490,516,1936,1957]" pageId="8" pageNumber="551">50</emphasis>
seventh abdominal tergum;
<emphasis id="B964EAD9DB48FFD8F733E2A9D94CF83F" bold="true" box="[803,828,1936,1957]" pageId="8" pageNumber="551">51</emphasis>
supra-anal plate and paraprocts, ventral view;
<emphasis id="B964EAD9DB48FFD8F104E2A9DF5DF83F" bold="true" box="[1300,1325,1936,1957]" pageId="8" pageNumber="551">52</emphasis>
subgenital plate, dorsal view;
<emphasis id="B964EAD9DB48FFD8F549E296DB02F85E" bold="true" box="[345,370,1967,1988]" pageId="8" pageNumber="551">53</emphasis>
hook-like phallomere (L3);
<emphasis id="B964EAD9DB48FFD8F684E296D8DEF85E" bold="true" box="[660,686,1967,1988]" pageId="8" pageNumber="551">54</emphasis>
median phallomere (L2vm);
<emphasis id="B964EAD9DB48FFD8F7C9E296D982F85E" bold="true" box="[985,1010,1967,1988]" pageId="8" pageNumber="551">55</emphasis>
right phallomere (R2 and R3). Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C30A6540DB49FFD9F4D7E5AEDBAFFF4F" pageId="9" pageNumber="552" type="etymology">
<paragraph id="8BAF36CBDB49FFD9F4D7E5AEDBAFFF4F" blockId="9.[151,1436,151,248]" pageId="9" pageNumber="552">
<emphasis id="B964EAD9DB49FFD9F4D7E5AEDB3EFF2A" bold="true" box="[199,334,151,176]" pageId="9" pageNumber="552">Etymology.</emphasis>
The specific epithet “forcipatus” is derived from Latin, referring to the pronotum with pincersshaped, dark brown maculae.
</paragraph>
</subSubSection>
<subSubSection id="C30A6540DB49FFD9F4D7E5E6D812FF62" box="[199,610,223,248]" pageId="9" pageNumber="552" type="distribution">
<paragraph id="8BAF36CBDB49FFD9F4D7E5E6D812FF62" blockId="9.[151,1436,151,248]" box="[199,610,223,248]" pageId="9" pageNumber="552">
<emphasis id="B964EAD9DB49FFD9F4D7E5E6DB10FF62" bold="true" box="[199,352,223,248]" pageId="9" pageNumber="552">Distribution.</emphasis>
<collectingCountry id="F307765BDB49FFD9F577E5E6DBDAFF62" box="[359,426,223,248]" name="China" pageId="9" pageNumber="552">China</collectingCountry>
(
<collectingRegion id="49D4F829DB49FFD9F5A8E5D9D87FFF62" box="[440,527,224,248]" country="China" name="Yunnan" pageId="9" pageNumber="552">Yunnan</collectingRegion>
Prov.).
</paragraph>
</subSubSection>
</treatment>
</document>