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<document id="E1838177A42A1B19B786E231AA26A87B" ID-CLB-Dataset="31250" ID-DOI="10.11646/zootaxa.4362.1.9" ID-GBIF-Dataset="9434ce3f-b243-4379-9137-90c77646a55b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1298337" ID-ZooBank="A0509E0B-A6F0-4F96-8D7C-F46095990D1F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512393502497" checkinUser="plazi" docAuthor="Hardy, Martin" docDate="2017" docId="03B987D35871997BFF71F976FDD3FE55" docLanguage="en" docName="zootaxa.4362.1.9.pdf" docOrigin="Zootaxa 4362 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bothynus thrix Hardy 2017, new species" docType="treatment" docVersion="10" lastPageNumber="145" masterDocId="FF80FFAB5871997FFFE6FFB5FFDDFFF4" masterDocTitle="Description of a new species of Bothynus Hope from Argentina and Bolivia (Coleoptera: Scarabaeidae: Dynastinae)" masterLastPageNumber="145" masterPageNumber="141" pageNumber="141" updateTime="1698507580620" updateUser="ExternalLinkService">
<mods:mods id="263BC3F1F70753E4302E218906A2DB12" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A495A86A7829E9EA8E834FA1E120D02A">
<mods:title id="3D9D505B03A2A7E21AB4A4D24B9F2701">Description of a new species of Bothynus Hope from Argentina and Bolivia (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
</mods:titleInfo>
<mods:name id="FCA97970EAA43F8688F1A690BD887BD2" type="personal">
<mods:role id="323E326EC8B9B2FEEE723FE2879A4CE0">
<mods:roleTerm id="C65C92B85F1B222EE4ECBB06245BE385">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F42DF12113416E5E031900FC979D28F9">Hardy, Martin</mods:namePart>
</mods:name>
<mods:typeOfResource id="342B48D892B942E457FF9A0908FBEE82">text</mods:typeOfResource>
<mods:relatedItem id="D539B5A56BE675CDC61F885EA661993D" type="host">
<mods:titleInfo id="A84C32CB204E5FAEF2BC3C7EA4F951B8">
<mods:title id="2D9B1235526EBE15FCD09111E820D008">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E56EF3D4365FF04E5682F22F6F2F9D14">
<mods:date id="C2125F4B59B0E263EB0E92642C3B2A37">2017</mods:date>
<mods:detail id="3B7484492C45C6923282D5EF4F2A708C" type="pubDate">
<mods:number id="307B103C7122D1AFF3CDEAFF61ADE16D">2017-12-04</mods:number>
</mods:detail>
<mods:detail id="95227A37E3668B163CEBDBEAC9F651F7" type="volume">
<mods:number id="E7D7A031E8F6E52A883E46A96B5E33B1">4362</mods:number>
</mods:detail>
<mods:detail id="8BBE12F8217D3F941D554B13252CDC9C" type="issue">
<mods:number id="70C3F88AFDD2A8A97AB12FA2E142D941">1</mods:number>
</mods:detail>
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<mods:start id="0B6D71A4F913DE1C0A27FCB09B4CE1B3">141</mods:start>
<mods:end id="24DC2E5C5460F96EB459AD3A0CE85FE4">145</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E7F2CF0232AC85063B9A8CC423395151">journal article</mods:classification>
<mods:identifier id="0A7C26A259FB95C7C39500EB1FDAC361" type="CLB-Dataset">31250</mods:identifier>
<mods:identifier id="7E38D7D88195EEF355FC814A6A2C17BF" type="DOI">10.11646/zootaxa.4362.1.9</mods:identifier>
<mods:identifier id="3433965BDC478B4E974DF00C728A8956" type="GBIF-Dataset">9434ce3f-b243-4379-9137-90c77646a55b</mods:identifier>
<mods:identifier id="CA3A6CFCE7CF2A0195E6E81D5F1E38E6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="38A533E03CB5B2EBC234C8F07736DDEF" type="Zenodo-Dep">1298337</mods:identifier>
<mods:identifier id="71C63D3E86FB4A5A6DFB861A4325411D" type="ZooBank">A0509E0B-A6F0-4F96-8D7C-F46095990D1F</mods:identifier>
</mods:mods>
<treatment id="03B987D35871997BFF71F976FDD3FE55" ID-DOI="http://doi.org/10.5281/zenodo.6053159" ID-GBIF-Taxon="137584815" ID-Zenodo-Dep="6053159" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55" httpUri="http://treatment.plazi.org/id/03B987D35871997BFF71F976FDD3FE55" lastPageId="4" lastPageNumber="145" pageId="0" pageNumber="141">
<subSubSection id="C30A654E5871997FFF71F976FD11F92A" box="[151,716,1731,1758]" pageId="0" pageNumber="141" type="nomenclature">
<paragraph id="8BAF36C55871997FFF71F976FD11F92A" blockId="0.[151,1436,1731,2032]" box="[151,716,1731,1758]" pageId="0" pageNumber="141">
<heading id="D0E781A95871997FFF71F976FD11F92A" bold="true" box="[151,716,1731,1758]" fontSize="11" level="1" pageId="0" pageNumber="141" reason="1">
<emphasis id="B964EAD75871997FFF71F976FD11F92A" bold="true" box="[151,716,1731,1758]" pageId="0" pageNumber="141">
<taxonomicName id="4C104D465871997FFF71F976FE7BF92A" ID-CoL="5WQKN" authority="Hardy" authorityName="Hardy" authorityYear="2017" box="[151,422,1731,1758]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="thrix" status="sp. nov.">
<emphasis id="B964EAD75871997FFF71F976FE90F929" bold="true" box="[151,333,1731,1757]" italics="true" pageId="0" pageNumber="141">Bothynus thrix</emphasis>
Hardy
</taxonomicName>
,
<taxonomicNameLabel id="A25757AC5871997FFE54F971FD9FF92A" box="[434,578,1732,1758]" pageId="0" pageNumber="141" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="132B2A405871997FFDB4F971FD1EF92A" box="[594,707,1732,1758]" captionStart-0="FIGURES 1" captionStart-1="FIGURES 2" captionStart-2="FIGURES3" captionStart-3="FIGURES 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,258,1301,1323]" captionStartId-1="2.[151,258,1139,1161]" captionStartId-2="2.[151,258,1666,1688]" captionStartId-3="3.[151,258,1373,1395]" captionStartId-4="3.[151,244,1850,1872]" captionTargetBox-0="[447,1139,368,1277]" captionTargetBox-1="[444,1142,180,1115]" captionTargetBox-2="[478,1105,1229,1644]" captionTargetBox-3="[470,1121,969,1340]" captionTargetBox-4="[446,1141,1478,1826]" captionTargetId-0="figure@1.[445,1142,365,1281]" captionTargetId-1="figure@2.[444,1143,179,1118]" captionTargetId-2="figure@2.[477,1109,1224,1645]" captionTargetId-3="figure@3.[453,1133,959,1351]" captionTargetId-4="figure@3.[441,1146,1472,1830]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionTargetPageId-4="3" captionText-0="FIGURES 1A–C. Bothynus thrix new species, holotype male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view." captionText-1="FIGURES 2A–C. Bothynus thrix new species, allotype female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view." captionText-2="FIGURES3A–C. Bothynus thrix new species,male parameres.A,caudal view;B, lateral view;C, oblique view." captionText-3="FIGURES 4A–B. Bothynus striatellus (Fairmaire, 1878), holotype male. A, habitus, dorsal view; B, labels. Photographs courtesy of A.Mantilleri, Muséum national d’Histoire naturelle, Paris, France." captionText-4="FIGURE 5. Distribution map showing currently known localities in South America for Bothynus thrix (black dots)." httpUri-0="https://zenodo.org/record/1298339/files/figure.png" httpUri-1="https://zenodo.org/record/1298341/files/figure.png" httpUri-2="https://zenodo.org/record/1298343/files/figure.png" httpUri-3="https://zenodo.org/record/1298345/files/figure.png" httpUri-4="https://zenodo.org/record/1298347/files/figure.png" pageId="0" pageNumber="141">Figs. 1–5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A654E5871997EFF71F953FB77FF00" lastPageId="1" lastPageNumber="142" pageId="0" pageNumber="141" type="materials_examined">
<paragraph id="8BAF36C55871997EFF71F953FB77FF00" blockId="0.[151,1436,1731,2032]" lastBlockId="1.[151,1436,152,313]" lastPageId="1" lastPageNumber="142" pageId="0" pageNumber="141">
<emphasis id="B964EAD75871997FFF71F953FEE3F909" bold="true" box="[151,318,1766,1789]" pageId="0" pageNumber="141">Type material.</emphasis>
<typeStatus id="54AB88675871997FFEA1F953FE77F909" box="[327,426,1766,1789]" pageId="0" pageNumber="141" type="holotype">Holotype</typeStatus>
male,
<typeStatus id="54AB88675871997FFE13F953FD96F909" box="[501,587,1766,1789]" pageId="0" pageNumber="141" type="allotype">allotype</typeStatus>
female,
<specimenCount id="9D16FD4C5871997FFD4DF953FD22F909" box="[683,767,1766,1789]" pageId="0" pageNumber="141" type="male">5 males</specimenCount>
<typeStatus id="54AB88675871997FFCEEF953FCACF909" box="[776,881,1766,1789]" pageId="0" pageNumber="141" type="paratype">paratypes</typeStatus>
, and
<specimenCount id="9D16FD4C5871997FFC4BF953FBC8F909" box="[941,1045,1766,1789]" pageId="0" pageNumber="141" type="female">4 females</specimenCount>
<typeStatus id="54AB88675871997FFBF8F953FB55F909" box="[1054,1160,1766,1789]" pageId="0" pageNumber="141" type="paratype">paratypes</typeStatus>
.
<materialsCitation id="3B783C985871997FFB75F953FEE7F8B7" ID-GBIF-Occurrence="2430682494" collectingDate="1964-02" collectionCode="CMNC" collectorName="Martinez &amp; Howden &amp; A. Martinez" country="Argentina" location="Pocitos" municipality="Dpto. San Martin" pageId="0" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Salta" typeStatus="holotype">
<typeStatus id="54AB88675871997FFB75F953FB2BF909" box="[1171,1270,1766,1789]" pageId="0" pageNumber="141" type="holotype">Holotype</typeStatus>
<specimenCount id="9D16FD4C5871997FFB19F953FAEEF909" box="[1279,1331,1766,1789]" pageId="0" pageNumber="141" type="male">male</specimenCount>
(
<collectionCode id="ED01AE005871997FFAA4F953FA49F909" box="[1346,1428,1766,1789]" pageId="0" pageNumber="141">CMNC</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555871997FFF10F8BCFE49F8D4" box="[246,404,1801,1824]" name="Argentina" pageId="0" pageNumber="141">ARGENTINA</collectingCountry>
/
<collectingRegion id="49D4F8275871997FFE49F8BCFE39F8D4" box="[431,484,1801,1824]" country="Argentina" name="Salta" pageId="0" pageNumber="141">Salta</collectingRegion>
/
<collectingMunicipality id="6BCBACBF5871997FFDE6F8BCFD1DF8D4" box="[512,704,1801,1824]" pageId="0" pageNumber="141">Dpto. San Martìn</collectingMunicipality>
/
<location id="8ECF601E5871997FFD3DF8BCFCF4F8D4" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5871997FFD3DF8BCFCF4F8D4" box="[731,809,1801,1824]" country="Argentina" municipality="Dpto. San Martin" name="Pocitos" pageId="0" pageNumber="141" stateProvince="Salta">Pocitos</location>
/ coll.
<collectorName id="26E553135871997FFC9BF8BCFC00F8D4" box="[893,989,1801,1824]" pageId="0" pageNumber="141">Martinez</collectorName>
/
<collectingDate id="EFEAE9ED5871997FFC1FF8BCFBB6F8D4" box="[1017,1131,1801,1824]" pageId="0" pageNumber="141" value="1964-02">
<date id="FFAE10055871997FFC1FF8BCFBB6F8D4" box="[1017,1131,1801,1824]" pageId="0" pageNumber="141" value="1964-02">Feb. 1964</date>
</collectingDate>
&quot;, // &quot;
<collectorName id="26E553135871997FFB45F8BCFADDF8D4" box="[1187,1280,1801,1824]" pageId="0" pageNumber="141">Howden</collectorName>
coll. Ex. /
<collectorName id="26E553135871997FFA64F8BCFF2AF8B7" pageId="0" pageNumber="141">A. Martinez</collectorName>
coll.&quot;
</materialsCitation>
.
<materialsCitation id="3B783C985871997FFEA5F899FD38F891" ID-GBIF-Occurrence="2430682493" collectingDate="1950-02" collectionCode="CMNC" collectorName="Martinez &amp; Howden &amp; A. Martinez" country="Bolivia" location="Buena Vista" municipality="Pua. Ichilo" pageId="0" pageNumber="141" specimenCount="1" specimenCount-female="1" stateProvince="Dpto. Sta Cruz" typeStatus="allotype">
<typeStatus id="54AB88675871997FFEA5F899FE7DF8B7" box="[323,416,1836,1859]" pageId="0" pageNumber="141" type="allotype">Allotype</typeStatus>
<specimenCount id="9D16FD4C5871997FFE41F899FE2DF8B7" box="[423,496,1836,1859]" pageId="0" pageNumber="141" type="female">female</specimenCount>
(
<collectionCode id="ED01AE005871997FFE1BF899FD92F8B7" box="[509,591,1836,1859]" pageId="0" pageNumber="141">CMNC</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555871997FFD50F899FCFBF8B7" box="[694,806,1836,1859]" name="Bolivia" pageId="0" pageNumber="141">BOLIVIA</collectingCountry>
/
<collectingRegion id="49D4F8275871997FFCDDF899FC05F8B7" box="[827,984,1836,1859]" inferredBySuffix="true" pageId="0" pageNumber="141">Dpto. Sta Cruz</collectingRegion>
/
<collectingMunicipality id="6BCBACBF5871997FFC0DF899FB83F8B7" box="[1003,1118,1836,1859]" pageId="0" pageNumber="141">Pua. Ichilo</collectingMunicipality>
/
<location id="8ECF601E5871997FFB94F899FB2FF8B7" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5871997FFB94F899FB2FF8B7" box="[1138,1266,1836,1859]" country="Bolivia" municipality="Pua. Ichilo" name="Buena Vista" pageId="0" pageNumber="141" stateProvince="Dpto. Sta Cruz">Buena Vista</location>
/ coll.
<collectorName id="26E553135871997FFADDF899FA46F8B7" box="[1339,1435,1836,1859]" pageId="0" pageNumber="141">Martinez</collectorName>
/
<collectingDate id="EFEAE9ED5871997FFF40F8FBFECBF891" box="[166,278,1870,1893]" pageId="0" pageNumber="141" value="1950-02">
<date id="FFAE10055871997FFF40F8FBFECBF891" box="[166,278,1870,1893]" pageId="0" pageNumber="141" value="1950-02">Feb. 1950</date>
</collectingDate>
&quot;, // &quot;
<collectorName id="26E553135871997FFEAFF8FBFE7BF891" box="[329,422,1870,1893]" pageId="0" pageNumber="141">Howden</collectorName>
coll. Ex. /
<collectorName id="26E553135871997FFDF9F8FBFD7DF891" box="[543,672,1870,1893]" pageId="0" pageNumber="141">A. Martinez</collectorName>
coll.&quot;
</materialsCitation>
.
<materialsCitation id="3B783C985871997FFD16F8FBFB9AF87C" ID-GBIF-Occurrence="2430682495" collectingDate="2000-10-17" collectingDateMax="2000-10-20" collectingDateMin="2000-10-17" collectionCode="MAHC" collectorName="R. Morris" country="Bolivia" location="Flora &amp; Fauna Hotel" municipality="Buena Vista" pageId="0" pageNumber="141" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="54AB88675871997FFD16F8FBFC81F891" box="[752,860,1870,1893]" pageId="0" pageNumber="141" type="paratype">Paratypes</typeStatus>
: one
<specimenCount id="9D16FD4C5871997FFC72F8FBFC15F891" box="[916,968,1870,1893]" pageId="0" pageNumber="141" type="male">male</specimenCount>
(
<collectionCode id="ED01AE005871997FFC31F8FBFBF7F891" box="[983,1066,1870,1893]" pageId="0" pageNumber="141">MAHC</collectionCode>
) and one
<specimenCount id="9D16FD4C5871997FFB73F8FBFB03F891" box="[1173,1246,1870,1893]" pageId="0" pageNumber="141" type="female">female</specimenCount>
(
<collectionCode id="ED01AE005871997FFB0AF8FBFAE2F891" box="[1260,1343,1870,1893]" pageId="0" pageNumber="141">MAHC</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555871997FFF79F8C4FECEF87C" box="[159,275,1905,1928]" name="Bolivia" pageId="0" pageNumber="141">BOLIVIA</collectingCountry>
:
<collectingRegion id="49D4F8275871997FFEFBF8C4FE4FF87C" box="[285,402,1905,1928]" country="Bolivia" name="Santa Cruz" pageId="0" pageNumber="141">Santa Cruz</collectingRegion>
/
<collectingMunicipality id="6BCBACBF5871997FFE43F8C4FDFAF87C" box="[421,551,1905,1928]" country="Bolivia" name="Buena Vista" pageId="0" pageNumber="141" stateProvince="Santa Cruz">Buena Vista</collectingMunicipality>
vic. /
<location id="8ECF601E5871997FFD81F8C4FCEAF87C" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5871997FFD81F8C4FCEAF87C" box="[615,823,1905,1928]" country="Bolivia" municipality="Buena Vista" name="Flora &amp; Fauna Hotel" pageId="0" pageNumber="141" stateProvince="Santa Cruz">Flora&amp;Fauna Hotel</location>
/
<collectingDate id="EFEAE9ED5871997FFCACF8C4FC15F87C" box="[842,968,1905,1928]" pageId="0" pageNumber="141" value="2000-10-17" valueMax="2000-10-20" valueMin="2000-10-17">
<date id="FFAE10055871997FFCACF8C4FC15F87C" box="[842,968,1905,1928]" pageId="0" pageNumber="141" value="2000-10-17" valueMax="2000-10-20" valueMin="2000-10-17">17-20/X/00</date>
</collectingDate>
.
<collectorName id="26E553135871997FFC34F8C4FBE2F87C" box="[978,1087,1905,1928]" pageId="0" pageNumber="141">R. Morris</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="3B783C985871997FFBB7F8C4FB23F85F" ID-GBIF-Occurrence="1703235515" collectingDate="2000-10-14" collectionCode="CERPE" collectorName="F &amp; F Hotel" country="Bolivia" location="Oct." municipality="Buena Vista" pageId="0" pageNumber="141" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="54AB88675871997FFBB7F8C4FB6EF87C" box="[1105,1203,1905,1928]" pageId="0" pageNumber="141" type="paratype">Paratype</typeStatus>
: one female (
<collectionCode id="ED01AE005871997FFAD8F8C4FA49F87C" box="[1342,1428,1905,1928]" pageId="0" pageNumber="141">CERPE</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555871997FFF14F821FEBEF85F" box="[242,355,1940,1963]" name="Bolivia" pageId="0" pageNumber="141">BOLIVIA</collectingCountry>
<collectingRegion id="49D4F8275871997FFE8FF821FE02F85F" box="[361,479,1940,1963]" country="Bolivia" name="Santa Cruz" pageId="0" pageNumber="141">Santa Cruz</collectingRegion>
/ 4-6k SSE
<collectingMunicipality id="6BCBACBF5871997FFDB9F821FD3DF85F" box="[607,736,1940,1963]" pageId="0" pageNumber="141">Buena Vista</collectingMunicipality>
/
<collectorName id="26E553135871997FFD13F821FCB4F85F" box="[757,873,1940,1963]" pageId="0" pageNumber="141">F&amp;F Hotel</collectorName>
<date id="FFAE10055871997FFC96F821FBF6F85F" box="[880,1067,1940,1963]" pageId="0" pageNumber="141" value="2000-10-14" valueMax="2000-10-16" valueMin="2000-10-14">
14-16
<location id="8ECF601E5871997FFC53F821FC3FF85F" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5871997FFC53F821FC3FF85F" box="[949,994,1940,1963]" country="Bolivia" municipality="Buena Vista" name="Oct." pageId="0" pageNumber="141" stateProvince="Santa Cruz">Oct.</location>
/ 2000
</date>
Wappes &amp; Morris&quot;
</materialsCitation>
.
<materialsCitation id="3B783C985871997FFAEEF821FA45F839" ID-GBIF-Occurrence="1703235520" collectingDate="2002-11-01" collectingDateMax="2002-11-08" collectingDateMin="2002-11-01" collectionCode="ACM" collectorName="J. E. Wappes" country="Bolivia" location="Buena Vista" pageId="0" pageNumber="141" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="54AB88675871997FFAEEF821FAB7F85F" box="[1288,1386,1940,1963]" pageId="0" pageNumber="141" type="paratype">Paratype</typeStatus>
: two males (
<collectionCode id="ED01AE005871997FFF00F803FEFBF839" box="[230,294,1974,1997]" country="Australia" httpUri="http://grbio.org/cool/12ic-3ajm" name="Australian Collection of Microorganisms" pageId="0" pageNumber="141">ACM</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555871997FFE70F803FDDAF839" box="[406,519,1974,1997]" name="Bolivia" pageId="0" pageNumber="141">BOLIVIA</collectingCountry>
<collectingRegion id="49D4F8275871997FFDF7F803FD57F839" box="[529,650,1974,1997]" country="Bolivia" name="Santa Cruz" pageId="0" pageNumber="141">Santa Cruz</collectingRegion>
/ 4-6k SSE
<location id="8ECF601E5871997FFCFEF803FC40F839" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5871997FFCFEF803FC40F839" box="[792,925,1974,1997]" country="Bolivia" name="Buena Vista" pageId="0" pageNumber="141" stateProvince="Santa Cruz">Buena Vista</location>
/ F &amp; F Hotel
<collectingDate id="EFEAE9ED5871997FFBA8F803FB27F839" box="[1102,1274,1974,1997]" pageId="0" pageNumber="141" value="2002-11-01" valueMax="2002-11-08" valueMin="2002-11-01">
<date id="FFAE10055871997FFBA8F803FB27F839" box="[1102,1274,1974,1997]" pageId="0" pageNumber="141" value="2002-11-01" valueMax="2002-11-08" valueMin="2002-11-01">Nov 1-8 / 2002</date>
</collectingDate>
<collectorName id="26E553135871997FFAE2F803FA4DF839" box="[1284,1424,1974,1997]" pageId="0" pageNumber="141">J.E. Wappes</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="3B783C985871997EFF71F86CFDE9FF5B" ID-GBIF-Occurrence="1703235521" collectingDate="2002-10-22" collectingDateMax="2002-10-26" collectingDateMin="2002-10-22" collectionCode="RFMC" country="Bolivia" county="Buena Vista" lastPageId="1" lastPageNumber="142" location="Wappes" municipality="Morris" pageId="0" pageNumber="141" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="54AB88675871997FFF71F86CFF24F804" box="[151,249,2009,2032]" pageId="0" pageNumber="141" type="paratype">Paratype</typeStatus>
: one male (
<collectionCode id="ED01AE005871997FFE95F86CFE1DF804" box="[371,448,2009,2032]" pageId="0" pageNumber="141">RFMC</collectionCode>
) and one female (
<collectionCode id="ED01AE005871997FFD64F86CFD12F804" box="[642,719,2009,2032]" pageId="0" pageNumber="141">RFMC</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555871997FFCDDF86CFC72F804" box="[827,943,2009,2032]" name="Bolivia" pageId="0" pageNumber="141">BOLIVIA</collectingCountry>
:
<collectingRegion id="49D4F8275871997FFC5CF86CFBEDF804" box="[954,1072,2009,2032]" country="Bolivia" name="Santa Cruz" pageId="0" pageNumber="141">Santa Cruz</collectingRegion>
/
<collectingCounty id="62CE4E495871997FFBA0F86CFB14F804" box="[1094,1225,2009,2032]" pageId="0" pageNumber="141">Buena Vista</collectingCounty>
vic. / Flora&amp;Fauna Hotel /
<collectingDate id="EFEAE9ED5870997EFF0FFF2DFEBBFF5B" box="[233,358,152,175]" pageId="1" pageNumber="142" value="2002-10-22" valueMax="2002-10-26" valueMin="2002-10-22">
<date id="FFAE10055870997EFF0FFF2DFEBBFF5B" box="[233,358,152,175]" pageId="1" pageNumber="142" value="2002-10-22" valueMax="2002-10-26" valueMin="2002-10-22">22-26/X/02</date>
</collectingDate>
,
<collectingMunicipality id="6BCBACBF5870997EFE94FF2DFE67FF5B" box="[370,442,152,175]" pageId="1" pageNumber="142">Morris</collectingMunicipality>
/
<location id="8ECF601E5870997EFE34FF2DFDF1FF5B" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5870997EFE34FF2DFDF1FF5B" box="[466,556,152,175]" country="Bolivia" county="Buena Vista" municipality="Morris" name="Wappes" pageId="1" pageNumber="142" stateProvince="Santa Cruz">Wappes</location>
&quot;
</materialsCitation>
.
<materialsCitation id="3B783C985870997EFDD9FF2DFD79FF25" ID-GBIF-Occurrence="1703235519" collectingDate="2000-10-20" collectingDateMax="2000-10-21" collectingDateMin="2000-10-20" collectionCode="CERPE" country="Bolivia" county="Rio Selva" location="Wappes" municipality="Morris" pageId="1" pageNumber="142" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="54AB88675870997EFDD9FF2DFD7CFF5B" box="[575,673,152,175]" pageId="1" pageNumber="142" type="paratype">Paratype</typeStatus>
: one male (
<collectionCode id="ED01AE005870997EFCFAFF2DFCAFFF5B" box="[796,882,152,175]" pageId="1" pageNumber="142">CERPE</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555870997EFC39FF2DFB8EFF5B" box="[991,1107,152,175]" name="Bolivia" pageId="1" pageNumber="142">BOLIVIA</collectingCountry>
:
<collectingRegion id="49D4F8275870997EFBB8FF2DFB08FF5B" box="[1118,1237,152,175]" country="Bolivia" name="Santa Cruz" pageId="1" pageNumber="142">Santa Cruz</collectingRegion>
/ 5k ESE Warnes, Hotel /
<collectingCounty id="62CE4E495870997EFF0FFF0FFE88FF25" box="[233,341,186,209]" pageId="1" pageNumber="142">Rio Selva</collectingCounty>
,
<collectingDate id="EFEAE9ED5870997EFE87FF0FFE06FF25" box="[353,475,186,209]" pageId="1" pageNumber="142" value="2000-10-20" valueMax="2000-10-21" valueMin="2000-10-20">
<date id="FFAE10055870997EFE87FF0FFE06FF25" box="[353,475,186,209]" pageId="1" pageNumber="142" value="2000-10-20" valueMax="2000-10-21" valueMin="2000-10-20">20-21/X/00</date>
</collectingDate>
/
<collectingMunicipality id="6BCBACBF5870997EFE14FF0FFDE1FF25" box="[498,572,186,209]" pageId="1" pageNumber="142">Morris</collectingMunicipality>
/
<location id="8ECF601E5870997EFDA2FF0FFD41FF25" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5870997EFDA2FF0FFD41FF25" box="[580,668,186,209]" country="Bolivia" county="Rio Selva" municipality="Morris" name="Wappes" pageId="1" pageNumber="142" stateProvince="Santa Cruz">Wappes</location>
&quot;
</materialsCitation>
.
<materialsCitation id="3B783C985870997EFD49FF0FFB7AFF00" ID-GBIF-Occurrence="2430682492" collectingDate="1970-12" collectionCode="CMNC" collectorName="Martinez &amp; Howden &amp; A. Martinez" country="Argentina" location="Dique Itiyuro" municipality="Dpto. San Martin" pageId="1" pageNumber="142" specimenCount="1" specimenCount-female="1" stateProvince="Salta" typeStatus="paratype">
<typeStatus id="54AB88675870997EFD49FF0FFCCDFF25" box="[687,784,186,209]" pageId="1" pageNumber="142" type="paratype">Paratype</typeStatus>
: one
<specimenCount id="9D16FD4C5870997EFCACFF0FFC4FFF25" box="[842,914,186,209]" pageId="1" pageNumber="142" type="female">female</specimenCount>
(
<collectionCode id="ED01AE005870997EFC47FF0FFC2EFF25" box="[929,1011,186,209]" pageId="1" pageNumber="142">CMNC</collectionCode>
) labeled &quot;
<collectingCountry id="F30776555870997EFBB8FF0FFB20FF25" box="[1118,1277,186,209]" name="Argentina" pageId="1" pageNumber="142">ARGENTINA</collectingCountry>
/
<collectingRegion id="49D4F8275870997EFAF5FF0FFA95FF25" box="[1299,1352,186,209]" country="Argentina" name="Salta" pageId="1" pageNumber="142">Salta</collectingRegion>
/
<collectingMunicipality id="6BCBACBF5870997EFA86FF0FFED3FF00" pageId="1" pageNumber="142">Dpto. San Martìn</collectingMunicipality>
/
<location id="8ECF601E5870997EFEC5FF68FE6EFF00" LSID="urn:lsid:plazi:treatment:03B987D35871997BFF71F976FDD3FE55:8ECF601E5870997EFEC5FF68FE6EFF00" box="[291,435,221,244]" country="Argentina" municipality="Dpto. San Martin" name="Dique Itiyuro" pageId="1" pageNumber="142" stateProvince="Salta">Dique Itiyuro</location>
/ coll.
<collectorName id="26E553135870997EFE1BFF68FD80FF00" box="[509,605,221,244]" pageId="1" pageNumber="142">Martinez</collectorName>
/
<collectingDate id="EFEAE9ED5870997EFD94FF68FD3DFF00" box="[626,736,221,244]" pageId="1" pageNumber="142" value="1970-12">
<date id="FFAE10055870997EFD94FF68FD3DFF00" box="[626,736,221,244]" pageId="1" pageNumber="142" value="1970-12">Dic. 1970</date>
</collectingDate>
&quot;, // &quot;
<collectorName id="26E553135870997EFCF7FF68FCB3FF00" box="[785,878,221,244]" pageId="1" pageNumber="142">Howden</collectorName>
coll. Ex. /
<collectorName id="26E553135870997EFC05FF68FBBEFF00" box="[995,1123,221,244]" pageId="1" pageNumber="142">A. Martinez</collectorName>
coll.&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A654E5870997DFF21FEB5FABAF81E" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142" type="description">
<paragraph id="8BAF36C55870997EFF21FEB5FD58FECD" blockId="1.[151,1436,152,313]" pageId="1" pageNumber="142">
<emphasis id="B964EAD75870997EFF21FEB5FE0AFEE3" bold="true" box="[199,471,256,279]" pageId="1" pageNumber="142">
Description of
<typeStatus id="54AB88675870997EFE93FEB5FE0AFEE3" box="[373,471,256,279]" pageId="1" pageNumber="142" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="132B2A405870997EFE0FFEB5FDB1FEE3" box="[489,620,256,279]" captionStart="FIGURES 1" captionStartId="1.[151,258,1301,1323]" captionTargetBox="[447,1139,368,1277]" captionTargetId="figure@1.[445,1142,365,1281]" captionTargetPageId="1" captionText="FIGURES 1A–C. Bothynus thrix new species, holotype male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view." httpUri="https://zenodo.org/record/1298339/files/figure.png" pageId="1" pageNumber="142">Figs. 1A–C</figureCitation>
). Male. Length
<quantity id="4CE89B205870997EFCF9FEB5FC58FEE3" box="[799,901,256,279]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="142" unit="mm" value="19.5">19.5 mm</quantity>
; width at elytra humeri
<quantity id="4CE89B205870997EFB75FEB5FB24FEE3" box="[1171,1273,256,279]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="142" unit="mm" value="10.5">10.5 mm</quantity>
. Color reddish brown, the pronotum and head slightly darker.
</paragraph>
<caption id="DF6F664D5870997EFF71FAA0FF16FABC" httpUri="https://zenodo.org/record/1298339/files/figure.png" pageId="1" pageNumber="142" targetBox="[447,1139,368,1277]" targetPageId="1">
<paragraph id="8BAF36C55870997EFF71FAA0FF16FABC" blockId="1.[151,1436,1301,1352]" pageId="1" pageNumber="142">
<emphasis id="B964EAD75870997EFF71FAA0FE99FADF" bold="true" box="[151,324,1301,1323]" pageId="1" pageNumber="142">FIGURES 1A–C</emphasis>
.
<taxonomicName id="4C104D465870997EFEB4FAA3FE3DFADF" authority="Hardy, 2017" authorityName="Hardy" authorityYear="2017" box="[338,480,1302,1323]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="thrix" status="sp. nov.">
<emphasis id="B964EAD75870997EFEB4FAA3FE3DFADF" box="[338,480,1302,1323]" italics="true" pageId="1" pageNumber="142">Bothynus thrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757AC5870997EFE0EFAA3FDBDFADF" box="[488,608,1302,1323]" pageId="1" pageNumber="142" rank="species">new species</taxonomicNameLabel>
,
<typeStatus id="54AB88675870997EFD8DFAA3FD62FADF" box="[619,703,1302,1323]" pageId="1" pageNumber="142" type="holotype">holotype</typeStatus>
male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view.
</paragraph>
</caption>
<paragraph id="8BAF36C55870997EFF21FADBFE3BF9C7" blockId="1.[151,1436,1390,2037]" pageId="1" pageNumber="142">
<emphasis id="B964EAD75870997EFF21FADBFED6FA72" bold="true" box="[199,267,1390,1414]" italics="true" pageId="1" pageNumber="142">Head:</emphasis>
Clypeus triangular, sides straight and weakly carinate, apex with anterior margin slightly emarginate, anterior angles bidentate with both teeth broadly separated and weakly reflexed. Clypeus and frons coarsely rugopunctate, punctures on frons setose. Frontoclypeal ridge weakly marked, feebly elevated not reaching sides of eyes. Interocular width equals 4 transverse eye diameters. Ocular canthus small, rounded. Antenna with 10 antennomeres, club slightly longer than antennomeres 2–7 (funicle without scape), club round. Mandibles tridentate, the 2 anterior teeth pointed, third tooth rounded on the side.
</paragraph>
<paragraph id="8BAF36C55870997EFF21F98BFD5BF914" blockId="1.[151,1436,1390,2037]" pageId="1" pageNumber="142">
<emphasis id="B964EAD75870997EFF21F98BFEE2F9A2" bold="true" box="[199,319,1598,1622]" italics="true" pageId="1" pageNumber="142">Pronotum:</emphasis>
Anterior angles acute, posterior angles very obtuse, rounded. Marginal bead completes anteriorly and laterally, incomplete basally. Anterior two thirds of pronotum with a transversally declivous area, almost flat, not delimited on side; behind anterior margin with a strong, pointed tubercle. Surface covered with large rounded punctures, dense and equally spaced, smaller on posterior third; punctures behind tubercle, in anterior angles and on sides setose. Row of setae also present on the basal margin.
</paragraph>
<paragraph id="8BAF36C55870997EFF21F95EFC31F8D1" blockId="1.[151,1436,1390,2037]" pageId="1" pageNumber="142">
<emphasis id="B964EAD75870997EFF21F95EFEC9F8F7" bold="true" box="[199,276,1771,1795]" italics="true" pageId="1" pageNumber="142">Elytra:</emphasis>
Surface shiny, with indistinct row of small to moderate punctures, ocellate; covered with long, pale setae. Sutural stria indistinct, marked with small punctures equal in size. Intervals flat.
</paragraph>
<paragraph id="8BAF36C55870997EFF21F885FE78F89F" blockId="1.[151,1436,1390,2037]" pageId="1" pageNumber="142">
<taxonomicName id="4C104D465870997EFF21F885FE68F8BC" authority=": Surface" authorityName="Surface" box="[199,437,1840,1864]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="142" phylum="Chordata" rank="genus">
<emphasis id="B964EAD75870997EFF21F885FE83F8BC" bold="true" box="[199,350,1840,1864]" italics="true" pageId="1" pageNumber="142">Propygidium:</emphasis>
Surface
</taxonomicName>
finely and transversely rugose on sides, with a large stridulary band in the middle, completely covered with small setae.
</paragraph>
<paragraph id="8BAF36C55870997EFF21F8C3FEE8F844" blockId="1.[151,1436,1390,2037]" pageId="1" pageNumber="142">
<taxonomicName id="4C104D465870997EFF21F8C3FE4CF879" authority=": Surface" authorityName="Surface" box="[199,401,1910,1934]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="142" phylum="Chordata" rank="genus">
<emphasis id="B964EAD75870997EFF21F8C3FEE4F87A" bold="true" box="[199,313,1910,1934]" italics="true" pageId="1" pageNumber="142">Pygidium:</emphasis>
Surface
</taxonomicName>
completely finely rugopunctate, less on disc; covered with small setae, longer on sides. Convex in lateral view.
</paragraph>
<paragraph id="8BAF36C55870997EFF21F80EFC9EF801" blockId="1.[151,1436,1390,2037]" pageId="1" pageNumber="142">
<emphasis id="B964EAD75870997EFF21F80EFED9F827" bold="true" box="[199,260,1979,2003]" italics="true" pageId="1" pageNumber="142">Legs:</emphasis>
Protibia tridentate, teeth equally spaced; protarsis not thickened, claws elongate and equal in size; inner claw not incised. Metatibia with apex truncate, with 32 small bristles.
</paragraph>
<caption id="DF6F664D5873997DFF71FBC6FF16FB52" httpUri="https://zenodo.org/record/1298341/files/figure.png" pageId="2" pageNumber="143" targetBox="[444,1142,180,1115]" targetPageId="2">
<paragraph id="8BAF36C55873997DFF71FBC6FF16FB52" blockId="2.[151,1435,1139,1190]" pageId="2" pageNumber="143">
<emphasis id="B964EAD75873997DFF71FBC6FE9EFB7D" bold="true" box="[151,323,1139,1161]" pageId="2" pageNumber="143">FIGURES 2A–C</emphasis>
.
<taxonomicName id="4C104D465873997DFEB6FBC1FE03FB7D" authority="Hardy, 2017" authorityName="Hardy" authorityYear="2017" box="[336,478,1140,1161]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="thrix" status="sp. nov.">
<emphasis id="B964EAD75873997DFEB6FBC1FE03FB7D" box="[336,478,1140,1161]" italics="true" pageId="2" pageNumber="143">Bothynus thrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757AC5873997DFE03FBC1FD81FB7D" box="[485,604,1140,1161]" pageId="2" pageNumber="143" rank="species">new species</taxonomicNameLabel>
,
<typeStatus id="54AB88675873997DFD81FBC1FD69FB7D" box="[615,692,1140,1161]" pageId="2" pageNumber="143" type="allotype">allotype</typeStatus>
female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view.
</paragraph>
</caption>
<caption id="DF6F664D5873997DFF71F937FB0BF96C" box="[151,1238,1666,1688]" httpUri="https://zenodo.org/record/1298343/files/figure.png" pageId="2" pageNumber="143" targetBox="[478,1105,1229,1644]" targetPageId="2">
<paragraph id="8BAF36C55873997DFF71F937FB0BF96C" blockId="2.[151,1238,1666,1688]" box="[151,1238,1666,1688]" pageId="2" pageNumber="143">
<emphasis id="B964EAD75873997DFF71F937FE9CF96C" bold="true" box="[151,321,1666,1688]" pageId="2" pageNumber="143">FIGURES 3A–C</emphasis>
.
<taxonomicName id="4C104D465873997DFEA8F936FE07F96C" authority="Hardy, 2017" authorityName="Hardy" authorityYear="2017" box="[334,474,1667,1688]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="thrix" status="sp. nov.">
<emphasis id="B964EAD75873997DFEA8F936FE07F96C" box="[334,474,1667,1688]" italics="true" pageId="2" pageNumber="143">Bothynus thrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757AC5873997DFE06F936FD8BF96C" box="[480,598,1667,1688]" pageId="2" pageNumber="143" rank="species">new species</taxonomicNameLabel>
, male parameres. A, caudal view; B, lateral view; C, oblique view.
</paragraph>
</caption>
<paragraph id="8BAF36C55873997DFF21F908FE09F903" blockId="2.[151,1436,1725,2026]" pageId="2" pageNumber="143">
<emphasis id="B964EAD75873997DFF21F908FECAF921" bold="true" box="[199,279,1725,1749]" italics="true" pageId="2" pageNumber="143">Venter:</emphasis>
Metasternum rugopunctate and densely setigerous, setae long. Prosternal process short, rounded, densely covered with numerous setae.
</paragraph>
<paragraph id="8BAF36C55873997DFF21F8B6FD26F8C8" blockId="2.[151,1436,1725,2026]" pageId="2" pageNumber="143">
<emphasis id="B964EAD75873997DFF21F8B6FE9BF8EF" bold="true" box="[199,326,1795,1819]" italics="true" pageId="2" pageNumber="143">Parameres:</emphasis>
In caudal view (
<figureCitation id="132B2A405873997DFDE3F8B6FD80F8EE" box="[517,605,1795,1818]" captionStart="FIGURES3" captionStartId="2.[151,258,1666,1688]" captionTargetBox="[478,1105,1229,1644]" captionTargetId="figure@2.[477,1109,1224,1645]" captionTargetPageId="2" captionText="FIGURES3A–C. Bothynus thrix new species,male parameres.A,caudal view;B, lateral view;C, oblique view." httpUri="https://zenodo.org/record/1298343/files/figure.png" pageId="2" pageNumber="143">Fig. 3A</figureCitation>
), enlarge at base then slightly tapering, apex dilated; in lateral and oblique views (
<figureCitation id="132B2A405873997DFF05F890FE82F8C8" box="[227,351,1829,1852]" captionStart="FIGURES3" captionStartId="2.[151,258,1666,1688]" captionTargetBox="[478,1105,1229,1644]" captionTargetId="figure@2.[477,1109,1224,1645]" captionTargetPageId="2" captionText="FIGURES3A–C. Bothynus thrix new species,male parameres.A,caudal view;B, lateral view;C, oblique view." httpUri="https://zenodo.org/record/1298343/files/figure.png" pageId="2" pageNumber="143">Figs. 3B–C</figureCitation>
) with a distinct, small tooth at middle.
</paragraph>
<paragraph id="8BAF36C55873997DFF21F8FDFABAF81E" blockId="2.[151,1436,1725,2026]" pageId="2" pageNumber="143">
<emphasis id="B964EAD75873997DFF21F8FDFEE4F8AB" bold="true" box="[199,313,1864,1887]" pageId="2" pageNumber="143">Variation.</emphasis>
<typeStatus id="54AB88675873997DFEA2F8FDFE7CF8AB" box="[324,417,1864,1887]" pageId="2" pageNumber="143" type="allotype">Allotype</typeStatus>
female (
<figureCitation id="132B2A405873997DFDE1F8FDFD50F8AB" box="[519,653,1864,1887]" captionStart="FIGURES 2" captionStartId="2.[151,258,1139,1161]" captionTargetBox="[444,1142,180,1115]" captionTargetId="figure@2.[444,1143,179,1118]" captionTargetPageId="2" captionText="FIGURES 2A–C. Bothynus thrix new species, allotype female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus, lateral view." httpUri="https://zenodo.org/record/1298341/files/figure.png" pageId="2" pageNumber="143">Figs. 2A–C</figureCitation>
): length
<quantity id="4CE89B205873997DFD17F8FDFC8AF8AB" box="[753,855,1864,1887]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="2" pageNumber="143" unit="mm" value="20.5">20.5 mm</quantity>
; width at elytra humeri
<quantity id="4CE89B205873997DFB8CF8FDFB60F8AB" box="[1130,1213,1864,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="143" unit="mm" value="10.0">10 mm</quantity>
. Female
<typeStatus id="54AB88675873997DFAC2F8FDFAA7F8AB" box="[1316,1402,1864,1887]" pageId="2" pageNumber="143" type="allotype">allotype</typeStatus>
as
<typeStatus id="54AB88675873997DFF71F8DEFF29F876" box="[151,244,1899,1922]" pageId="2" pageNumber="143" type="holotype">holotype</typeStatus>
except in the followings characteristics.
<emphasis id="B964EAD75873997DFD54F8DEFD2AF877" bold="true" box="[690,759,1899,1923]" italics="true" pageId="2" pageNumber="143">Head:</emphasis>
Antenna club the same size as antennomeres 2–7.
<emphasis id="B964EAD75873997DFAC2F8DEFA41F877" bold="true" box="[1316,1436,1899,1923]" italics="true" pageId="2" pageNumber="143">Pronotum:</emphasis>
Tubercle behind anterior margin absent, reduce to a low gibbosity. Surface covered with larger and denser punctures, becoming confluent on disc.
<emphasis id="B964EAD75873997DFE36F805FDC3F83C" bold="true" box="[464,542,1968,1992]" italics="true" pageId="2" pageNumber="143">Elytra:</emphasis>
Setae more abundant over entire surface.
<taxonomicName id="4C104D465873997DFC00F805FB6FF833" authority=": Pilosity" authorityName="Pilosity" box="[998,1202,1968,1992]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="143" phylum="Chordata" rank="genus">
<emphasis id="B964EAD75873997DFC00F805FB84F83C" bold="true" box="[998,1113,1968,1992]" italics="true" pageId="2" pageNumber="143">Pygidium:</emphasis>
Pilosity
</taxonomicName>
longer and denser on disc. Flat in lateral view.
<typeStatus id="54AB88675873997DFE45F866FDD2F81E" box="[419,527,2003,2026]" pageId="2" pageNumber="143" type="paratype">Paratypes</typeStatus>
: length 19.5–22.0 mm. Characters similar to
<typeStatus id="54AB88675873997DFC09F866FB91F81E" box="[1007,1100,2003,2026]" pageId="2" pageNumber="143" type="holotype">holotype</typeStatus>
male and
<typeStatus id="54AB88675873997DFB5CF866FACDF81E" box="[1210,1296,2003,2026]" pageId="2" pageNumber="143" type="allotype">allotype</typeStatus>
female.
</paragraph>
</subSubSection>
<subSubSection id="C30A654E5872997CFF21FF2DFD2AFF25" pageId="3" pageNumber="144" type="etymology">
<paragraph id="8BAF36C55872997CFF21FF2DFD2AFF25" blockId="3.[151,1436,152,417]" pageId="3" pageNumber="144">
<emphasis id="B964EAD75872997CFF21FF2DFE9AFF5B" bold="true" box="[199,327,152,175]" pageId="3" pageNumber="144">Etymology.</emphasis>
This species name is based after the Greek adjective
<taxonomicName id="4C104D465872997CFC78FF2DFC0DFF5B" authorityName="Hardy" authorityYear="2017" box="[926,976,152,175]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75872997CFC78FF2DFC0DFF5B" box="[926,976,152,175]" italics="true" pageId="3" pageNumber="144">thríx</emphasis>
</taxonomicName>
, meaning hairy or pilose, referring to the pubescence covering the elytra and part of the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C30A654E5872997CFF21FF68FE1DFEA8" pageId="3" pageNumber="144" type="distribution">
<paragraph id="8BAF36C55872997CFF21FF68FDFBFEE3" blockId="3.[151,1436,152,417]" pageId="3" pageNumber="144">
<emphasis id="B964EAD75872997CFF21FF68FE8FFF00" bold="true" box="[199,338,221,244]" pageId="3" pageNumber="144">Distribution</emphasis>
(
<figureCitation id="132B2A405872997CFE84FF68FE79FF00" box="[354,420,221,244]" captionStart="FIGURE 5" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[446,1141,1478,1826]" captionTargetId="figure@3.[441,1146,1472,1830]" captionTargetPageId="3" captionText="FIGURE 5. Distribution map showing currently known localities in South America for Bothynus thrix (black dots)." httpUri="https://zenodo.org/record/1298347/files/figure.png" pageId="3" pageNumber="144">Fig. 5</figureCitation>
).
<taxonomicName id="4C104D465872997CFE5CFF68FD8AFF00" authorityName="Hardy" authorityYear="2017" box="[442,599,221,244]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75872997CFE5CFF68FD8AFF00" box="[442,599,221,244]" italics="true" pageId="3" pageNumber="144">Bothynus thrix</emphasis>
</taxonomicName>
is known from the northern part of
<collectingCountry id="F30776555872997CFC39FF68FB94FF00" box="[991,1097,221,244]" name="Argentina" pageId="3" pageNumber="144">Argentina</collectingCountry>
in
<collectingRegion id="49D4F8275872997CFB8BFF68FAD4FF00" box="[1133,1289,221,244]" country="Argentina" name="Salta" pageId="3" pageNumber="144">Salta Province</collectingRegion>
and from the Department of
<collectingRegion id="49D4F8275872997CFEDEFEB5FE73FEE3" box="[312,430,256,279]" country="Bolivia" name="Santa Cruz" pageId="3" pageNumber="144">Santa Cruz</collectingRegion>
in
<collectingCountry id="F30776555872997CFE36FEB5FDFFFEE3" box="[464,546,256,279]" name="Bolivia" pageId="3" pageNumber="144">Bolivia</collectingCountry>
.
</paragraph>
<paragraph id="8BAF36C55872997CFF21FE97FE1DFEA8" blockId="3.[151,1436,152,417]" pageId="3" pageNumber="144">
<emphasis id="B964EAD75872997CFF21FE97FEB0FECD" bold="true" box="[199,365,290,313]" pageId="3" pageNumber="144">Temporal data</emphasis>
. February (2), October (6), November (2), December (1); these dates coincide with the beginning of the rainy season in October.
</paragraph>
</subSubSection>
<subSubSection id="C30A654E5872997BFF21FEDDFDD3FE55" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144" type="diagnosis">
<paragraph id="8BAF36C55872997CFF21FEDDFB76FE55" blockId="3.[151,1436,152,417]" pageId="3" pageNumber="144">
<emphasis id="B964EAD75872997CFF21FEDDFEE1FE8B" bold="true" box="[199,316,360,383]" pageId="3" pageNumber="144">Diagnosis.</emphasis>
<taxonomicName id="4C104D465872997CFEA3FEDDFE38FE8B" authorityName="Hardy" authorityYear="2017" box="[325,485,360,383]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75872997CFEA3FEDDFE38FE8B" box="[325,485,360,383]" italics="true" pageId="3" pageNumber="144">Bothynus thrix</emphasis>
</taxonomicName>
will key to the couplet 37(36) for
<taxonomicName id="4C104D465872997CFC96FEDDFA4AFE8B" authority="(Fairmaire, 1878) in Endrodi (1985)" authorityName="in Endrodi" authorityYear="1985" baseAuthorityName="Fairmaire" baseAuthorityYear="1878" box="[880,1431,360,383]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="striatellus">
<emphasis id="B964EAD75872997CFC96FEDDFC21FE8B" box="[880,1020,360,383]" italics="true" pageId="3" pageNumber="144">B. striatellus</emphasis>
(Fairmaire, 1878) in
<bibRefCitation id="EF814B345872997CFB08FEDDFA4AFE8B" author="Endrodi" box="[1262,1431,360,383]" firstAuthor="Endrodi" pageId="3" pageNumber="144" refId="ref2175" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
</taxonomicName>
, although it is easily distinguished from it based on the combination of characters listed in
<tableCitation id="C692037E5872997CFBB3FE3FFB78FE55" box="[1109,1189,394,417]" captionStart="TABLE 1" captionStartId="3.[151,239,464,487]" captionText="TABLE 1. Characters to differentiate Bothynus thrix and Bothynus striatellus." pageId="3" pageNumber="144">Table 1</tableCitation>
.
</paragraph>
<caption id="DF6F664D5872997CFF71FE65FC05FE13" box="[151,984,464,487]" pageId="3" pageNumber="144">
<paragraph id="8BAF36C55872997CFF71FE65FC05FE13" blockId="3.[151,984,464,487]" box="[151,984,464,487]" pageId="3" pageNumber="144">
<emphasis id="B964EAD75872997CFF71FE65FED7FE13" bold="true" box="[151,266,464,487]" pageId="3" pageNumber="144">TABLE 1.</emphasis>
Characters to differentiate
<taxonomicName id="4C104D465872997CFDCBFE65FD17FE13" authorityName="Hardy" authorityYear="2017" box="[557,714,464,487]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75872997CFDCBFE65FD17FE13" box="[557,714,464,487]" italics="true" pageId="3" pageNumber="144">Bothynus thrix</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D465872997CFD1BFE65FC0DFE13" authorityName="in Endrodi" authorityYear="1985" baseAuthorityName="Fairmaire" baseAuthorityYear="1878" box="[765,976,464,487]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="striatellus">
<emphasis id="B964EAD75872997CFD1BFE65FC0DFE13" box="[765,976,464,487]" italics="true" pageId="3" pageNumber="144">Bothynus striatellus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BAF36C55872997CFF79FDB7FB12FC80" pageId="3" pageNumber="144">
<table id="F910C46558726680FF79FDB7FAA0FC80" box="[159,1405,514,884]" gridcols="3" gridrows="8" pageId="3" pageNumber="144">
<tr id="3520348758726680FF79FDB7FAA0FDEC" box="[159,1405,514,536]" gridrow="0" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FDB7FE87FDEC" box="[159,346,514,536]" gridcol="0" gridrow="0" pageId="3" pageNumber="144">Character (male)</th>
<th id="76F15DFB58726680FE6CFDB7FC96FDEC" box="[394,843,514,536]" gridcol="1" gridrow="0" pageId="3" pageNumber="144">
<taxonomicName id="4C104D465872997CFE6CFDB7FE0FFDE3" authorityName="Hardy" authorityYear="2017" box="[394,466,514,535]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75872997CFE6CFDB7FE0FFDE3" box="[394,466,514,535]" italics="true" pageId="3" pageNumber="144">B. thrix</emphasis>
</taxonomicName>
</th>
<th id="76F15DFB58726680FC44FDB7FAA0FDEC" box="[930,1405,514,536]" gridcol="2" gridrow="0" pageId="3" pageNumber="144">
<taxonomicName id="4C104D465872997CFC44FDB7FBC2FDEC" authorityName="in Endrodi" authorityYear="1985" baseAuthorityName="Fairmaire" baseAuthorityYear="1878" box="[930,1055,514,536]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="striatellus">
<emphasis id="B964EAD75872997CFC44FDB7FBC2FDEC" bold="true" box="[930,1055,514,536]" italics="true" pageId="3" pageNumber="144">B. striatellus</emphasis>
</taxonomicName>
(Figs. 4A–B)
</th>
</tr>
<tr id="3520348758726680FF79FD9FFAA0FDCB" box="[159,1405,554,575]" gridrow="1" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FD9FFE87FDCB" box="[159,346,554,575]" gridcol="0" gridrow="1" pageId="3" pageNumber="144">Body size and form</th>
<td id="76F15DFB58726680FE6CFD9FFC96FDCB" box="[394,843,554,575]" gridcol="1" gridrow="1" pageId="3" pageNumber="144">
Length
<quantity id="4CE89B205872997CFE32FD9FFDE3FDCB" box="[468,574,554,575]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" pageNumber="144" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">19–22 mm</quantity>
, body more slender
</td>
<td id="76F15DFB58726680FC44FD9FFAA0FDCB" box="[930,1405,554,575]" gridcol="2" gridrow="1" pageId="3" pageNumber="144">
Length
<quantity id="4CE89B205872997CFC0AFD9FFB8BFDCB" box="[1004,1110,554,575]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="3" pageNumber="144" unit="mm" value="22.0" valueMax="25.0" valueMin="19.0">19–25 mm</quantity>
, body broader
</td>
</tr>
<tr id="3520348758726680FF79FDCFFAA0FD7B" box="[159,1405,634,655]" gridrow="2" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FDCFFE87FD7B" box="[159,346,634,655]" gridcol="0" gridrow="2" pageId="3" pageNumber="144">Clypeus</th>
<td id="76F15DFB58726680FE6CFDCFFC96FD7B" box="[394,843,634,655]" gridcol="1" gridrow="2" pageId="3" pageNumber="144">Apical teeth broadly spaced, feebly upturned</td>
<td id="76F15DFB58726680FC44FDCFFAA0FD7B" box="[930,1405,634,655]" gridcol="2" gridrow="2" pageId="3" pageNumber="144">Apical teeth closer, strongly upturned</td>
</tr>
<tr id="3520348758726680FF79FD17FAA0FD20" box="[159,1405,674,724]" gridrow="3" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FD17FE87FD20" box="[159,346,674,724]" gridcol="0" gridrow="3" pageId="3" pageNumber="144">Pronotum</th>
<td id="76F15DFB58726680FE6CFD17FC96FD20" box="[394,843,674,724]" gridcol="1" gridrow="3" pageId="3" pageNumber="144">No distinct fovea, only declivous, punctate in center</td>
<td id="76F15DFB58726680FC44FD17FAA0FD20" box="[930,1405,674,724]" gridcol="2" gridrow="3" pageId="3" pageNumber="144">Distinct fovae, surface wrinkled</td>
</tr>
<tr id="3520348758726680FF79FD52FAA0FD08" box="[159,1405,743,764]" gridrow="4" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FD52FE87FD08" box="[159,346,743,764]" gridcol="0" gridrow="4" pageId="3" pageNumber="144">Elytra</th>
<td id="76F15DFB58726680FE6CFD52FC96FD08" box="[394,843,743,764]" gridcol="1" gridrow="4" pageId="3" pageNumber="144">Indistinct striae, not furrowed; surface setose</td>
<td id="76F15DFB58726680FC44FD52FAA0FD08" box="[930,1405,743,764]" gridcol="2" gridrow="4" pageId="3" pageNumber="144">Striae punctate, finely furrowed; surface glabrous</td>
</tr>
<tr id="3520348758726680FF79FCBAFAA0FCD0" box="[159,1405,783,804]" gridrow="5" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FCBAFE87FCD0" box="[159,346,783,804]" gridcol="0" gridrow="5" pageId="3" pageNumber="144">
<taxonomicName id="4C104D465872997CFF79FCBAFF21FCD0" box="[159,252,783,804]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="144" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
</th>
<td id="76F15DFB58726680FE6CFCBAFC96FCD0" box="[394,843,783,804]" gridcol="1" gridrow="5" pageId="3" pageNumber="144">Finely reticulate, setose</td>
<td id="76F15DFB58726680FC44FCBAFAA0FCD0" box="[930,1405,783,804]" gridcol="2" gridrow="5" pageId="3" pageNumber="144">Finely punctate, glabrous</td>
</tr>
<tr id="3520348758726680FF79FC82FAA0FCB8" box="[159,1405,823,844]" gridrow="6" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FC82FE87FCB8" box="[159,346,823,844]" gridcol="0" gridrow="6" pageId="3" pageNumber="144">Metatibiae</th>
<td id="76F15DFB58726680FE6CFC82FC96FCB8" box="[394,843,823,844]" gridcol="1" gridrow="6" pageId="3" pageNumber="144">Apex with 32 spinules</td>
<td id="76F15DFB58726680FC44FC82FAA0FCB8" box="[930,1405,823,844]" gridcol="2" gridrow="6" pageId="3" pageNumber="144">Apex with 22–25 spinules</td>
</tr>
<tr id="3520348758726680FF79FCEAFAA0FC80" box="[159,1405,863,884]" gridrow="7" pageId="3" pageNumber="144">
<th id="76F15DFB58726680FF79FCEAFE87FC80" box="[159,346,863,884]" gridcol="0" gridrow="7" pageId="3" pageNumber="144">Parameres</th>
<td id="76F15DFB58726680FE6CFCEAFC96FC80" box="[394,843,863,884]" gridcol="1" gridrow="7" pageId="3" pageNumber="144">Longer, with small tooth visible in lateral view</td>
<td id="76F15DFB58726680FC44FCEAFAA0FC80" box="[930,1405,863,884]" gridcol="2" gridrow="7" pageId="3" pageNumber="144">Shorter, no tooth on lateral side</td>
</tr>
</table>
</paragraph>
<caption id="DF6F664D5872997CFF71FAE8FCC5FA7B" httpUri="https://zenodo.org/record/1298345/files/figure.png" pageId="3" pageNumber="144" targetBox="[470,1121,969,1340]" targetPageId="3">
<paragraph id="8BAF36C55872997CFF71FAE8FCC5FA7B" blockId="3.[151,1436,1373,1423]" pageId="3" pageNumber="144">
<emphasis id="B964EAD75872997CFF71FAE8FE9CFA87" bold="true" box="[151,321,1373,1395]" pageId="3" pageNumber="144">FIGURES 4A–B</emphasis>
.
<taxonomicName id="4C104D465872997CFEA8FAE8FD1CFA86" authority="(Fairmaire, 1878)" authorityName="in Endrodi" authorityYear="1985" baseAuthorityName="Fairmaire" baseAuthorityYear="1878" box="[334,705,1373,1394]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="striatellus">
<emphasis id="B964EAD75872997CFEA8FAE8FDD3FA86" box="[334,526,1373,1394]" italics="true" pageId="3" pageNumber="144">Bothynus striatellus</emphasis>
(Fairmaire, 1878)
</taxonomicName>
,
<typeStatus id="54AB88675872997CFD2AFAE8FCFDFA86" box="[716,800,1373,1394]" pageId="3" pageNumber="144" type="holotype">holotype</typeStatus>
male. A, habitus, dorsal view; B, labels. Photographs courtesy of A. Mantilleri, Muséum national d’Histoire naturelle, Paris,
<collectingCountry id="F30776555872997CFD36FACFFCC9FA7B" box="[720,788,1402,1423]" name="France" pageId="3" pageNumber="144">France</collectingCountry>
.
</paragraph>
</caption>
<caption id="DF6F664D5872997CFF71F88FFB32F8A4" box="[151,1263,1850,1872]" httpUri="https://zenodo.org/record/1298347/files/figure.png" pageId="3" pageNumber="144" targetBox="[446,1141,1478,1826]" targetPageId="3">
<paragraph id="8BAF36C55872997CFF71F88FFB32F8A4" blockId="3.[151,1263,1850,1872]" box="[151,1263,1850,1872]" pageId="3" pageNumber="144">
<emphasis id="B964EAD75872997CFF71F88FFED1F8A4" bold="true" box="[151,268,1850,1872]" pageId="3" pageNumber="144">FIGURE 5.</emphasis>
Distribution map showing currently known localities in South America for
<taxonomicName id="4C104D465872997CFC02F88EFBADF8A4" authorityName="Hardy" authorityYear="2017" box="[996,1136,1851,1872]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75872997CFC02F88EFBADF8A4" box="[996,1136,1851,1872]" italics="true" pageId="3" pageNumber="144">Bothynus thrix</emphasis>
</taxonomicName>
(black dots).
</paragraph>
</caption>
<paragraph id="8BAF36C55875997BFF21FF2DFCBAFEE3" blockId="4.[151,1436,152,417]" pageId="4" pageNumber="145">
The genital structure of males of
<taxonomicName id="4C104D465875997BFDADFF2DFD73FF5B" authorityName="Hope" authorityYear="1837" box="[587,686,152,175]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAD75875997BFDADFF2DFD73FF5B" box="[587,686,152,175]" italics="true" pageId="4" pageNumber="145">Bothynus</emphasis>
</taxonomicName>
species is reliable for species identification even if some general similarities are sometimes observed between some species. The parameres of the new species are similar to those of
<taxonomicName id="4C104D465875997BFF71FF68FDECFF00" authority="(Fairmaire, 1878)" authorityName="in Endrodi" authorityYear="1985" baseAuthorityName="Fairmaire" baseAuthorityYear="1878" box="[151,561,221,244]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="striatellus">
<emphasis id="B964EAD75875997BFF71FF68FEB1FF00" box="[151,364,221,244]" italics="true" pageId="4" pageNumber="145">Bothynus striatellus</emphasis>
(Fairmaire, 1878)
</taxonomicName>
, but they are shorter and have no visible tooth on the side, whereas they are more elongate and with a distinctive lateral tooth in
<taxonomicName id="4C104D465875997BFD63FEB5FD08FEE3" authorityName="Hardy" authorityYear="2017" box="[645,725,256,279]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75875997BFD63FEB5FD08FEE3" box="[645,725,256,279]" italics="true" pageId="4" pageNumber="145">B. thrix</emphasis>
</taxonomicName>
(
<figureCitation id="132B2A405875997BFD05FEB5FC86FEE3" box="[739,859,256,279]" captionStart="FIGURES3" captionStartId="2.[151,258,1666,1688]" captionTargetBox="[478,1105,1229,1644]" captionTargetId="figure@2.[477,1109,1224,1645]" captionTargetPageId="2" captionText="FIGURES3A–C. Bothynus thrix new species,male parameres.A,caudal view;B, lateral view;C, oblique view." httpUri="https://zenodo.org/record/1298343/files/figure.png" pageId="4" pageNumber="145">Figs 3B–C</figureCitation>
).
</paragraph>
<paragraph id="8BAF36C55875997BFF21FE97FDD3FE55" blockId="4.[151,1436,152,417]" pageId="4" pageNumber="145">
Given the particular appearance of this new species of
<taxonomicName id="4C104D465875997BFCF0FE97FCA4FECD" authorityName="Hope" authorityYear="1837" box="[790,889,290,313]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAD75875997BFCF0FE97FCA4FECD" box="[790,889,290,313]" italics="true" pageId="4" pageNumber="145">Bothynus</emphasis>
</taxonomicName>
, (the only species with such abundant setae on the elytra and part of the pronotum), it is surprising that
<taxonomicName id="4C104D465875997BFD35FEF0FCF9FEA8" authorityName="Hardy" authorityYear="2017" box="[723,804,325,348]" class="Insecta" family="Scarabaeidae" genus="Bothynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="thrix">
<emphasis id="B964EAD75875997BFD35FEF0FCF9FEA8" box="[723,804,325,348]" italics="true" pageId="4" pageNumber="145">B. thrix</emphasis>
</taxonomicName>
has remained unknown until now. This eastern part of the Bolivian and Argentinean Andes from which the specimens originate are known for their high endemism and biodiversity (
<bibRefCitation id="EF814B345875997BFEC2FE3FFDDEFE55" author="Swenson, J. J. &amp; Young, B. E. &amp; Beck, S. &amp; Comer, P. &amp; Cordova, J. H. &amp; Dyson, J. &amp; Embert, D. &amp; Encarnacion, F. &amp; Ferreira, W. &amp; Franke, I &amp; Grossman, D. &amp; Hernandez, P. &amp; Herzog, S. K. &amp; Josse, C. &amp; Navarro, G. &amp; Pacheco, V &amp; Stein, B. A. &amp; Timana, M. &amp; Tovar, A. &amp; Tovar, C. &amp; Vargas, J. &amp; Zambrana-Torrelio, C. M." box="[292,515,394,417]" etAl="et al." firstAuthor="Swenson" journalOrPublisher="BMC Ecology" pageId="4" pageNumber="145" pagination="1 - 18" part="12" refId="ref2404" refString="Swenson, J. J., Young, B. E., Beck, S., Comer, P., Cordova, J. H., Dyson, J., Embert, D., Encarnacion, F., Ferreira, W., Franke, I., Grossman, D., Hernandez, P., Herzog, S. K., Josse, C., Navarro, G., Pacheco, V., Stein, B. A., Timana, M., Tovar, A., Tovar, C., Vargas, J. &amp; Zambrana-Torrelio, C. M. (2012) Plant and animal endemism in the eastern Andean slope: challenges to conservation. BMC Ecology, 12 (1), 1 - 18. https: // doi. org / 10.1186 / 1472 - 6785 - 12 - 1" title="Plant and animal endemism in the eastern Andean slope: challenges to conservation" type="journal article" year="2012">
Swenson
<emphasis id="B964EAD75875997BFE6AFE3FFE1FFE55" box="[396,450,394,417]" italics="true" pageId="4" pageNumber="145">et al.</emphasis>
2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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