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<document id="563C821CD98CB2638BE9FDF70739D380" ID-CLB-Dataset="56108" ID-DOI="10.6620/ZS.2019.58-14" ID-GBIF-Dataset="de0647eb-9b73-4bac-9ff7-f7bff7518dc0" ID-PMC="PMC6759935" ID-PubMed="31966315" ID-Zenodo-Dep="8065869" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687182873363" checkinUser="valdenar" docAuthor="Nguyen, Anh D., Nguyen, Manh-Ha, Nguyen, Thu-Anh T. &amp; Phung, Hong-Luong T." docDate="2019" docId="03B987C4FFFBAE4AFEE3FE58FBB7AAF4" docLanguage="en" docName="ZoolStud.58.14.1-31.pdf" docOrigin="Zoological Studies 58 (14)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Hylomus saiyans Nguyen &amp; Nguyen &amp; Nguyen &amp; Phung 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="22" masterDocId="FF80FFBCFFEAAE5FFFF0FFA5FF8AAB21" masterDocTitle="Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="18" updateTime="1687361907046" updateUser="ExternalLinkService">
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<mods:title id="8C25793BB8815BD33CF23FE291ACEE91">Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species</mods:title>
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<mods:namePart id="5F5AB86D6C5EAAEE934D2DDB3C41EB9C">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="4DA64B6AF7ACE7D063BE116BEE59138A">Nguyen, Manh-Ha</mods:namePart>
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<mods:namePart id="A6F49E3753283CCF8A5C75C08110FD9E">Nguyen, Thu-Anh T.</mods:namePart>
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<mods:namePart id="8DC3CB192E0759090B269AADCE0B1B71">Phung, Hong-Luong T.</mods:namePart>
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<treatment id="03B987C4FFFBAE4AFEE3FE58FBB7AAF4" ID-DOI="http://doi.org/10.5281/zenodo.8055838" ID-GBIF-Taxon="209694530" ID-Zenodo-Dep="8055838" LSID="urn:lsid:plazi:treatment:03B987C4FFFBAE4AFEE3FE58FBB7AAF4" httpUri="http://treatment.plazi.org/id/03B987C4FFFBAE4AFEE3FE58FBB7AAF4" lastPageId="21" lastPageNumber="22" pageId="17" pageNumber="18">
<subSubSection id="C30A6559FFFBAE4EFEE3FE58FDDCA937" box="[275,598,509,534]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BAF36D2FFFBAE4EFEE3FE58FDDCA937" blockId="17.[173,700,509,618]" box="[275,598,509,534]" pageId="17" pageNumber="18">
<heading id="D0E781BEFFFBAE4EFEE3FE58FDDCA937" bold="true" box="[275,598,509,534]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="18" reason="2">
<emphasis id="B964EAC0FFFBAE4EFEE3FE58FDDCA937" bold="true" box="[275,598,509,534]" pageId="17" pageNumber="18">
<taxonomicName id="4C104D51FFFBAE4EFEE3FE58FE67A937" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Phung, 2019" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Phung" authorityYear="2019" box="[275,493,509,534]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="saiyans" status="sp. nov.">
<emphasis id="B964EAC0FFFBAE4EFEE3FE58FE67A937" bold="true" box="[275,493,509,534]" italics="true" pageId="17" pageNumber="18">Hylomus saiyans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757BBFFFBAE4EFE04FE58FDDCA937" box="[500,598,509,534]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C30A6559FFFBAE4AFEAEFDB8FBB7AAF4" lastPageId="21" lastPageNumber="22" pageId="17" pageNumber="18" type="description">
<paragraph id="8BAF36D2FFFBAE4EFEAEFDB8FD83A94B" blockId="17.[173,700,509,618]" pageId="17" pageNumber="18">
(
<figureCitation id="132B2A57FFFBAE4EFE96FDB8FE3AA914" box="[358,432,541,565]" captionStart="Fig" captionStartId="2.[175,206,1813,1832]" captionTargetBox="[440,1185,515,1783]" captionTargetId="figure-96@2.[440,1185,486,1804]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Hylomus species in mainland Vietnam. 1: Hoang Lien National Park (Sa Pa, Lao Cai Province); 2: Nam Xay Commune (Van Ban District, Lao Cai Province); 3: Duc Xuan Commune (Bac Quang District, Ha Giang Province); 4: Xuan Son National Park (Phu Tho Province); 5: Tam Dao (Vinh Phuc Province); 6: Cuc Phuong National Park (Ninh Binh Province); 7: Phong Nha Ke Bang National Park (Quang Binh Province); 8: Lien Chieu District (Da Nang Province); 9: Ba Na National Park (Da Nang Province); 10: Lo Xo Pass (Kon Tum Province); 11: Ngoc Linh Mt. (Kon Tum Province); 12: Kon Chu Rang Natural Reserve (Gia Lai Province); 13: Buon Ho District (Dak Lak Province); 14: Hon Ba Mt. (Khanh Hoa Province); 15: Phan Rang (Ninh Thuan Province); 16: Cat Tien National Park (Dong Nai Province)." figureDoi="http://doi.org/10.5281/zenodo.8065873" httpUri="https://zenodo.org/record/8065873/files/figure.png" pageId="17" pageNumber="18">Figs. 1</figureCitation>
,
<figureCitation id="132B2A57FFFBAE4EFE4DFDB8FD89A914" box="[445,515,541,565]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="19.[136,167,1926,1945]" captionStartId-1="20.[174,205,1271,1290]" captionStartId-2="20.[174,205,1926,1945]" captionStartId-3="21.[136,167,1954,1973]" captionStartId-4="22.[174,205,1954,1973]" captionTargetBox-0="[136,1413,925,1898]" captionTargetBox-1="[174,1451,282,1245]" captionTargetBox-2="[174,1451,1395,1899]" captionTargetBox-3="[205,1345,749,1927]" captionTargetBox-4="[297,1328,945,1927]" captionTargetId-0="figure-339@19.[136,1413,925,1898]" captionTargetId-1="figure-80@20.[174,1451,282,1245]" captionTargetId-2="figure-86@20.[174,1451,1395,1899]" captionTargetId-3="figure-223@21.[205,1345,749,1927]" captionTargetId-4="figure-344@22.[297,1328,945,1927]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="20" captionTargetPageId-3="21" captionTargetPageId-4="22" captionText-0="Fig. 21. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Anterior part, dorsal view (A), lateral view (B), ventral view (C). Body rings 1011, dorsal view (C). Scale bar = 1 mm." captionText-1="Fig. 22. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Posterior part, dorsal view (A), ventral view (B). Telson and epiproct, dorsal view (C). Hypoproct and epiproct, ventral view (D). Scale bars:AB = 1 mm; CD = 0.5 mm." captionText-2="Fig. 23. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Sternal process between coxae 4, subposterior view (A). Legs 567 (B). Scale bars: A = 0.5 mm; B = 1 mm." captionText-3="Fig. 24. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Right gonopod, lateral view (A), mesal view (B). Scale bar = 1 mm." captionText-4="Fig. 25. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Right gonopod, lateral view (A), mesal view (B)." figureDoi-0="http://doi.org/10.5281/zenodo.8065928" figureDoi-1="http://doi.org/10.5281/zenodo.8065932" figureDoi-2="http://doi.org/10.5281/zenodo.8065934" figureDoi-3="http://doi.org/10.5281/zenodo.8065936" figureDoi-4="http://doi.org/10.5281/zenodo.8065938" httpUri-0="https://zenodo.org/record/8065928/files/figure.png" httpUri-1="https://zenodo.org/record/8065932/files/figure.png" httpUri-2="https://zenodo.org/record/8065934/files/figure.png" httpUri-3="https://zenodo.org/record/8065936/files/figure.png" httpUri-4="https://zenodo.org/record/8065938/files/figure.png" pageId="17" pageNumber="18">2125</figureCitation>
)
<uri id="FF813AD0FFFBAE4EFF5DFD9DFD83A94B" pageId="17" pageNumber="18">
urn:lsid:zoobank.org:act:
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<paragraph id="8BAF36D2FFFBAE4EFF35FD35FE6CA809" blockId="17.[136,737,656,873]" pageId="17" pageNumber="18">
<emphasis id="B964EAC0FFFBAE4EFF35FD35FE2BA989" box="[197,417,656,680]" italics="true" pageId="17" pageNumber="18">Material examined</emphasis>
:
<materialsCitation id="3B783C8FFFFBAE4EFE45FD35FE80A828" collectingDate="2012-04-11" collectingDateMax="2012-04-13" collectingDateMin="2012-04-11" collectorName="Anh D. Nguyen." location="Cuc Phuong National Park" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh" typeStatus="holotype">
<typeStatus id="54AB8870FFFBAE4EFE45FD35FDAFA989" box="[437,549,656,680]" pageId="17" pageNumber="18">Holotype</typeStatus>
:
<specimenCount id="9D16FD5BFFFBAE4EFDC3FD34FD08A989" box="[563,642,656,681]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
(IEBR- Myr 219H)
<collectingRegion id="49D4F830FFFBAE4EFEFCFD15FE63A9E9" box="[268,489,688,712]" country="Vietnam" name="Ninh Binh" pageId="17" pageNumber="18">Ninh Binh Province</collectingRegion>
,
<location id="8ECF6009FFFBAE4EFE06FD14FF35A9C9" LSID="urn:lsid:plazi:treatment:03B987C4FFFBAE4AFEE3FE58FBB7AAF4:8ECF6009FFFBAE4EFE06FD14FF35A9C9" name="Cuc Phuong National Park" pageId="17" pageNumber="18" stateProvince="Ninh Binh">Cuc Phuong National Park</location>
, limestone forest,
<date id="FFAE1012FFFBAE4EFE68FD74FDE2A9C8" box="[408,616,720,745]" pageId="17" pageNumber="18" value="2012-04-11" valueMax="2012-04-13" valueMin="2012-04-11">
<collectingDate id="EFEAE9FAFFFBAE4EFE68FD74FDE2A9C8" box="[408,616,720,745]" pageId="17" pageNumber="18" value="2012-04-11" valueMax="2012-04-13" valueMin="2012-04-11">1113 April 2012</collectingDate>
</date>
, coll.
<collectorName id="26E55304FFFBAE4EFD41FD75FE80A828" pageId="17" pageNumber="18">Anh D. Nguyen.</collectorName>
</materialsCitation>
<typeStatus id="54AB8870FFFBAE4EFEE3FD54FE08A828" box="[275,386,753,777]" pageId="17" pageNumber="18">Paratypes</typeStatus>
:
<specimenCount id="9D16FD5BFFFBAE4EFE7FFD54FE54A829" box="[399,478,752,777]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<specimenCount id="9D16FD5BFFFBAE4EFE1CFD55FDD2A829" box="[492,600,752,776]" pageId="17" pageNumber="18" type="female">5 females</specimenCount>
(IEBR-Myr 219P) same data as for
<typeStatus id="54AB8870FFFBAE4EFE72FCB5FE69A809" box="[386,483,784,808]" pageId="17" pageNumber="18">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BAF36D2FFFBAE4EFF35FC94FCE2AA54" blockId="17.[136,737,656,873]" lastBlockId="17.[813,1414,253,885]" pageId="17" pageNumber="18">
<emphasis id="B964EAC0FFFBAE4EFF35FC94FEB3A869" box="[197,313,817,840]" italics="true" pageId="17" pageNumber="18">Non-types</emphasis>
:
<specimenCount id="9D16FD5BFFFBAE4EFEBCFC94FE16A869" box="[332,412,816,841]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
(IEBR-Myr 44)
<collectingRegion id="49D4F830FFFBAE4EFD97FC95FF60A849" country="Vietnam" name="Ninh Binh" pageId="17" pageNumber="18">Ninh Binh Province</collectingRegion>
, Cuc Phuong National Park, limestone forest,
<date id="FFAE1012FFFBAE4EFCDDFF58FC2FAA34" box="[813,933,253,277]" pageId="17" pageNumber="18" value="2006-04">April 2006</date>
, coll. Thu-Anh T. Nguyen;
<specimenCount id="9D16FD5BFFFBAE4EFB21FF5BFAA5AA34" box="[1233,1327,253,278]" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(IEBR- Myr 63)
<collectingRegion id="49D4F830FFFBAE4EFC66FEB8FBF5AA14" box="[918,1151,285,309]" country="Vietnam" name="Ninh Binh" pageId="17" pageNumber="18">Ninh Binh Province</collectingRegion>
, Cuc Phuong National Park, limestone forest,
<date id="FFAE1012FFFBAE4EFBDCFE9BFB42AA74" box="[1068,1224,317,342]" pageId="17" pageNumber="18" value="2006-04-30">30 April 2006</date>
, coll. Luong Van Hien.
</paragraph>
<paragraph id="8BAF36D2FFFBAE4EFC99FED8FB1BAAF7" blockId="17.[813,1414,253,885]" pageId="17" pageNumber="18">
<emphasis id="B964EAC0FFFBAE4EFC99FED8FC7AAAB4" box="[873,1008,381,405]" italics="true" pageId="17" pageNumber="18">Etymology</emphasis>
: The name refers to naturally aggressive warriors in the Japanese manga “Dragon Ball” by Toriyama Akira (
<collectingCountry id="F3077642FFFBAE4EFBB2FE1BFB0FAAF7" box="[1090,1157,446,470]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
).
</paragraph>
<paragraph id="8BAF36D2FFFBAE4EFC99FE7BFC5AA9F4" blockId="17.[813,1414,253,885]" pageId="17" pageNumber="18">
<emphasis id="B964EAC0FFFBAE4EFC99FE7BFC5FAAD4" box="[873,981,478,501]" italics="true" pageId="17" pageNumber="18">Diagnosis</emphasis>
: The species differs from its congeners by the second metatergal row having 1 + 1 setiferous spines, others being setiferous knobs. Femora 6 and 7 with a ventral hump, sometimes femur 5 inflated. Gonopod femorite slightly curverd ventrad, enlarged distad; distal part with mesal shelf. Demarcation between femorite and postfemoral region absent. Tip of gonopod acute.
</paragraph>
<paragraph id="8BAF36D2FFFBAE4CFC99FD7BFE85AA14" blockId="17.[813,1414,253,885]" lastBlockId="19.[136,737,253,885]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<emphasis id="B964EAC0FFFBAE4EFC99FD7BFC64A9D4" box="[873,1006,734,757]" italics="true" pageId="17" pageNumber="18">Description</emphasis>
: Length ca.
<quantity id="4CE89B37FFFBAE4EFB74FD78FAA1A9D4" box="[1156,1323,733,757]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.58" metricValueMin="1.42" pageId="17" pageNumber="18" unit="mm" value="15.0" valueMax="15.8" valueMin="14.2">14.215.8 mm</quantity>
(male),
<quantity id="4CE89B37FFFBAE4EFCDDFD5BFC58A834" box="[813,978,766,790]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.42" metricValueMin="1.38" pageId="17" pageNumber="18" unit="mm" value="14.0" valueMax="14.2" valueMin="13.8">13.814.2 mm</quantity>
(female); width of midbody pro- and metazonae
<quantity id="4CE89B37FFFBAE4EFC5CFCBBFBBEA814" box="[940,1076,798,822]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="17" pageNumber="18" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 mm</quantity>
(male),
<quantity id="4CE89B37FFFBAE4EFB66FCBBFA95A814" box="[1174,1311,798,822]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="17" pageNumber="18" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
(female) and 1.0
<quantity id="4CE89B37FFFBAE4EFC6EFC9BFC76A874" box="[926,1020,830,854]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="18" unit="mm" value="1.1">1.1 mm</quantity>
(male),
<quantity id="4CE89B37FFFBAE4EFB9FFC9BFA8DA874" box="[1135,1287,830,854]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="17" pageNumber="18" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
(female), respectively.
<typeStatus id="54AB8870FFFBAE4EFC34FCF8FBA6A854" box="[964,1068,861,885]" pageId="17" pageNumber="18">Holotype</typeStatus>
length ca.
<quantity id="4CE89B37FFFBAE4EFB5FFCF8FA9DA854" box="[1199,1303,861,885]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="17" pageNumber="18" unit="mm" value="15.8">15.8 mm</quantity>
, width of midbody pro- and metazonae
<quantity id="4CE89B37FFF9AE4CFE1AFF5BFDC8AA34" box="[490,578,254,278]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="mm" value="0.8">0.8 mm</quantity>
and 1.0 mm, respectively.
</paragraph>
<paragraph id="8BAF36D2FFF9AE4CFF35FE98FECAAAB4" blockId="19.[136,737,253,885]" pageId="19" pageNumber="20">
Colouration (
<figureCitation id="132B2A57FFF9AE4CFEAAFE98FE6DAA77" box="[346,487,317,342]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="19.[136,167,1926,1945]" captionStartId-1="20.[174,205,1271,1290]" captionStartId-2="20.[174,205,1926,1945]" captionTargetBox-0="[136,1413,925,1898]" captionTargetBox-1="[174,1451,282,1245]" captionTargetBox-2="[174,1451,1395,1899]" captionTargetId-0="figure-339@19.[136,1413,925,1898]" captionTargetId-1="figure-80@20.[174,1451,282,1245]" captionTargetId-2="figure-86@20.[174,1451,1395,1899]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="20" captionText-0="Fig. 21. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Anterior part, dorsal view (A), lateral view (B), ventral view (C). Body rings 1011, dorsal view (C). Scale bar = 1 mm." captionText-1="Fig. 22. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Posterior part, dorsal view (A), ventral view (B). Telson and epiproct, dorsal view (C). Hypoproct and epiproct, ventral view (D). Scale bars:AB = 1 mm; CD = 0.5 mm." captionText-2="Fig. 23. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Sternal process between coxae 4, subposterior view (A). Legs 567 (B). Scale bars: A = 0.5 mm; B = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8065928" figureDoi-1="http://doi.org/10.5281/zenodo.8065932" figureDoi-2="http://doi.org/10.5281/zenodo.8065934" httpUri-0="https://zenodo.org/record/8065928/files/figure.png" httpUri-1="https://zenodo.org/record/8065932/files/figure.png" httpUri-2="https://zenodo.org/record/8065934/files/figure.png" pageId="19" pageNumber="20">Figs. 2123</figureCitation>
): body generally light brown in male, but darker in female. Legs and sterna brownish yellow.
</paragraph>
<paragraph id="8BAF36D2FFF9AE4CFF35FE38FD5FA954" blockId="19.[136,737,253,885]" pageId="19" pageNumber="20">
Head (
<figureCitation id="132B2A57FFF9AE4CFEFFFE38FED7AA97" box="[271,349,413,438]" captionStart="Fig" captionStartId="19.[136,167,1926,1945]" captionTargetBox="[136,1413,925,1898]" captionTargetId="figure-339@19.[136,1413,925,1898]" captionTargetPageId="19" captionText="Fig. 21. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Anterior part, dorsal view (A), lateral view (B), ventral view (C). Body rings 1011, dorsal view (C). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8065928" httpUri="https://zenodo.org/record/8065928/files/figure.png" pageId="19" pageNumber="20">Fig. 21</figureCitation>
AC) slightly broader than collum in width; surface with dense tiny setiferous spines. Clypeolabral region densely setose. Epicranial suture clearly distinct. Antennae (
<figureCitation id="132B2A57FFF9AE4CFE30FE58FD98A937" box="[448,530,509,534]" captionStart="Fig" captionStartId="19.[136,167,1926,1945]" captionTargetBox="[136,1413,925,1898]" captionTargetId="figure-339@19.[136,1413,925,1898]" captionTargetPageId="19" captionText="Fig. 21. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Anterior part, dorsal view (A), lateral view (B), ventral view (C). Body rings 1011, dorsal view (C). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8065928" httpUri="https://zenodo.org/record/8065928/files/figure.png" pageId="19" pageNumber="20">Fig. 21</figureCitation>
AC) slender and long, reaching segment 6 if stretched posteriorly; antennomere 2&gt; 3&gt; 4 = 5 = 6&gt; 1&gt;
<quantity id="4CE89B37FFF9AE4CFDA7FD9BFD0FA974" box="[599,645,573,597]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="19" pageNumber="20" unit="in" value="7.0">7 in</quantity>
length; antennomere 7 blackish, with four sensory cones at tip.
</paragraph>
<paragraph id="8BAF36D2FFF9AE4CFF35FDD8FF5DA814" blockId="19.[136,737,253,885]" pageId="19" pageNumber="20">
Collum slightly broader than segment
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width, subtrapeziform; surface with dense microgranulation, and two rows of well-developed setiferous spines: 3 + 3 anterior and 1 + 1 posterior. Paraterga spiniform, slightly longer than posterior metatergal spines, directed dorsad.
</paragraph>
<paragraph id="8BAF36D2FFF9AE4CFF35FC98FB71A937" blockId="19.[136,737,253,885]" lastBlockId="19.[813,1414,253,885]" pageId="19" pageNumber="20">
Width of body segment 3 &lt;4 &lt;2 =
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width, thereafter gradually tapering towards telson. Prozonae shining and shagreened. Metazonae (terga and pleura) with dense microgranulations. Metaterga with two rows of 1 + 1 anterior small setiferous knobs and 2 + 2 posterior setiferous knobs/spines including well developed, median setiferous spines (
<figureCitation id="132B2A57FFF9AE4CFB21FED8FA0BAAB7" box="[1233,1409,381,406]" captionStart="Fig" captionStartId="19.[136,167,1926,1945]" captionTargetBox="[136,1413,925,1898]" captionTargetId="figure-339@19.[136,1413,925,1898]" captionTargetPageId="19" captionText="Fig. 21. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Anterior part, dorsal view (A), lateral view (B), ventral view (C). Body rings 1011, dorsal view (C). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8065928" httpUri="https://zenodo.org/record/8065928/files/figure.png" pageId="19" pageNumber="20">Figs. 21A, 21D</figureCitation>
,
<figureCitation id="132B2A57FFF9AE4CFCDDFE3BFC10AA97" box="[813,922,414,438]" captionStart="Fig" captionStartId="20.[174,205,1271,1290]" captionTargetBox="[174,1451,282,1245]" captionTargetId="figure-80@20.[174,1451,282,1245]" captionTargetPageId="20" captionText="Fig. 22. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Posterior part, dorsal view (A), ventral view (B). Telson and epiproct, dorsal view (C). Hypoproct and epiproct, ventral view (D). Scale bars:AB = 1 mm; CD = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065932" httpUri="https://zenodo.org/record/8065932/files/figure.png" pageId="19" pageNumber="20">22A, 22C</figureCitation>
). Pleurosternal carinae absent. Waist between pro- and metazonae broad, not deep, neither striolate nor beaded at bottom. Transverse sulcus incomplete, starting on metatergum 4. Axial line vague.
</paragraph>
<paragraph id="8BAF36D2FFF9AE4CFC99FDBBFB2BA814" blockId="19.[813,1414,253,885]" pageId="19" pageNumber="20">
Paraterga (
<figureCitation id="132B2A57FFF9AE4CFC14FDB8FBCEA917" box="[996,1092,541,566]" captionStart="Fig" captionStartId="19.[136,167,1926,1945]" captionTargetBox="[136,1413,925,1898]" captionTargetId="figure-339@19.[136,1413,925,1898]" captionTargetPageId="19" captionText="Fig. 21. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Anterior part, dorsal view (A), lateral view (B), ventral view (C). Body rings 1011, dorsal view (C). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8065928" httpUri="https://zenodo.org/record/8065928/files/figure.png" pageId="19" pageNumber="20">Figs. 21</figureCitation>
,
<figureCitation id="132B2A57FFF9AE4CFBA2FDBBFB4BA917" box="[1106,1217,542,566]" captionStart="Fig" captionStartId="20.[174,205,1271,1290]" captionTargetBox="[174,1451,282,1245]" captionTargetId="figure-80@20.[174,1451,282,1245]" captionTargetPageId="20" captionText="Fig. 22. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Posterior part, dorsal view (A), ventral view (B). Telson and epiproct, dorsal view (C). Hypoproct and epiproct, ventral view (D). Scale bars:AB = 1 mm; CD = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065932" httpUri="https://zenodo.org/record/8065932/files/figure.png" pageId="19" pageNumber="20">22A, 22C</figureCitation>
) well developed, antler-shaped, highly elevated from metatergal surface, directed dorsad on segments 218, and directed caudad on segment 19. Paraterga with 2 branches on segment 2, 3 branches on segments 38, 11, 14, and 18, four branches on segments 910, 1213, 1517, and spiniform on segment 19. Ozopore on segments 5, 7, 910, 1213, and 1519, lying at the base of the second branch, invisible from dorsal view.
</paragraph>
<paragraph id="8BAF36D2FFF9AE4AFC99FC98FD1DAA14" blockId="19.[813,1414,253,885]" lastBlockId="21.[136,738,253,693]" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
Epiproct (
<figureCitation id="132B2A57FFF9AE4CFC24FC98FBA8A877" box="[980,1058,829,854]" captionStart="Fig" captionStartId="20.[174,205,1271,1290]" captionTargetBox="[174,1451,282,1245]" captionTargetId="figure-80@20.[174,1451,282,1245]" captionTargetPageId="20" captionText="Fig. 22. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Posterior part, dorsal view (A), ventral view (B). Telson and epiproct, dorsal view (C). Hypoproct and epiproct, ventral view (D). Scale bars:AB = 1 mm; CD = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065932" httpUri="https://zenodo.org/record/8065932/files/figure.png" pageId="19" pageNumber="20">Fig. 22</figureCitation>
BD) broadly truncated, dorsally inflated, with two lateral setiferous tubercles; tip with four spinnerets. Hypoproct (
<figureCitation id="132B2A57FFFFAE4AFE47FF58FD93AA37" box="[439,537,253,278]" captionStart="Fig" captionStartId="20.[174,205,1271,1290]" captionTargetBox="[174,1451,282,1245]" captionTargetId="figure-80@20.[174,1451,282,1245]" captionTargetPageId="20" captionText="Fig. 22. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Posterior part, dorsal view (A), ventral view (B). Telson and epiproct, dorsal view (C). Hypoproct and epiproct, ventral view (D). Scale bars:AB = 1 mm; CD = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065932" httpUri="https://zenodo.org/record/8065932/files/figure.png" pageId="21" pageNumber="22">Fig. 22D</figureCitation>
) trapeziform, with two well separated, distolateral, setiferous knobs.
</paragraph>
<paragraph id="8BAF36D2FFFFAE4AFF35FE9BFD18AAB7" blockId="21.[136,738,253,693]" pageId="21" pageNumber="22">
Sterna (
<figureCitation id="132B2A57FFFFAE4AFEE8FE98FEFDAA77" box="[280,375,317,342]" captionStart="Fig" captionStartId="20.[174,205,1271,1290]" captionTargetBox="[174,1451,282,1245]" captionTargetId="figure-80@20.[174,1451,282,1245]" captionTargetPageId="20" captionText="Fig. 22. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Posterior part, dorsal view (A), ventral view (B). Telson and epiproct, dorsal view (C). Hypoproct and epiproct, ventral view (D). Scale bars:AB = 1 mm; CD = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065932" httpUri="https://zenodo.org/record/8065932/files/figure.png" pageId="21" pageNumber="22">Fig. 22B</figureCitation>
) densely setose, cross impression distinct, without modifications except a setiferous trapeziform process between coxae 4 (
<figureCitation id="132B2A57FFFFAE4AFDD2FED8FD0CAAB7" box="[546,646,381,406]" captionStart="Fig" captionStartId="20.[174,205,1926,1945]" captionTargetBox="[174,1451,1395,1899]" captionTargetId="figure-86@20.[174,1451,1395,1899]" captionTargetPageId="20" captionText="Fig. 23. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Sternal process between coxae 4, subposterior view (A). Legs 567 (B). Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8065934" httpUri="https://zenodo.org/record/8065934/files/figure.png" pageId="21" pageNumber="22">Fig. 23A</figureCitation>
).
</paragraph>
<paragraph id="8BAF36D2FFFFAE4AFF35FE3BFE65A934" blockId="21.[136,738,253,693]" pageId="21" pageNumber="22">
Legs slender and long, ca. 23 times as long as midbody height. Prefemora not swollen. Femora 6 and 7 with a ventral hump, femur 5 sometimes inflated (
<figureCitation id="132B2A57FFFFAE4AFD5CFE78FF3DA937" captionStart="Fig" captionStartId="19.[136,167,1926,1945]" captionTargetBox="[136,1413,925,1898]" captionTargetId="figure-339@19.[136,1413,925,1898]" captionTargetPageId="19" captionText="Fig. 21. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Anterior part, dorsal view (A), lateral view (B), ventral view (C). Body rings 1011, dorsal view (C). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8065928" httpUri="https://zenodo.org/record/8065928/files/figure.png" pageId="21" pageNumber="22">Figs. 21B</figureCitation>
,
<figureCitation id="132B2A57FFFFAE4AFF32FE5BFF78A937" box="[194,242,510,534]" captionStart="Fig" captionStartId="20.[174,205,1926,1945]" captionTargetBox="[174,1451,1395,1899]" captionTargetId="figure-86@20.[174,1451,1395,1899]" captionTargetPageId="20" captionText="Fig. 23. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Sternal process between coxae 4, subposterior view (A). Legs 567 (B). Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8065934" httpUri="https://zenodo.org/record/8065934/files/figure.png" pageId="21" pageNumber="22">23B</figureCitation>
). Tarsal brushes absent.
</paragraph>
<paragraph id="8BAF36D2FFFFAE4AFF35FDB8FBB7AAF4" blockId="21.[136,738,253,693]" lastBlockId="21.[813,1414,253,469]" pageId="21" pageNumber="22">
Gonopod (
<figureCitation id="132B2A57FFFFAE4AFEB3FDB8FE5EA914" box="[323,468,541,565]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="21.[136,167,1954,1973]" captionStartId-1="22.[174,205,1954,1973]" captionTargetBox-0="[205,1345,749,1927]" captionTargetBox-1="[297,1328,945,1927]" captionTargetId-0="figure-223@21.[205,1345,749,1927]" captionTargetId-1="figure-344@22.[297,1328,945,1927]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Fig. 24. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Right gonopod, lateral view (A), mesal view (B). Scale bar = 1 mm." captionText-1="Fig. 25. Hylomus saiyans sp. nov. from Cuc Phuong National Park. Holotype. Right gonopod, lateral view (A), mesal view (B)." figureDoi-0="http://doi.org/10.5281/zenodo.8065936" figureDoi-1="http://doi.org/10.5281/zenodo.8065938" httpUri-0="https://zenodo.org/record/8065936/files/figure.png" httpUri-1="https://zenodo.org/record/8065938/files/figure.png" pageId="21" pageNumber="22">Figs. 2425</figureCitation>
) simple. Coxite short, ca. 1/3 as long as femorite length; distoventral part sparsely setose. Prefemorite densely setose, separated laterally from femorite by a transverse sulcus. Femorite slightly curved ventrad, enlarged distad; distal part with mesal shelf (ms). No demarcation between femorite and postfemoral region. The postfemoral region short, consisting only of solenophore and solenomere. S o l e n o m e r e f l a g e l l i f o r m, p a r t l y s h e a t h e d b y solenophore. Seminal groove running entirely mesally, directed distodorsad before entering the solenophore. Tip of solenophore acute.
</paragraph>
</subSubSection>
</treatment>
</document>