1926 lines
432 KiB
XML
1926 lines
432 KiB
XML
<document id="77F2CAA5E4AFCF3BCFECA88F6D962CEF" ID-DOI="10.11646/zootaxa.4027.2.1" ID-GBIF-Dataset="058b8bab-084b-4a45-8d74-29446fed1877" ID-ISSN="1175-5326" ID-Zenodo-Dep="238764" ID-ZooBank="B2D0A15A-16BE-4311-B9F1-ADC27341D40C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461281236055" checkinUser="plazi" docAuthor="Norton, Roy A. & Fuangarworn, Marut" docDate="2015" docId="03B987B9FFE3FFC4FF37FB3A3088F368" docLanguage="en" docName="zt04027p204.pdf" docOrigin="Zootaxa 4027 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nanohystrix hammerae Norton & Fuangarworn, 2015, n. sp." docType="treatment" docUuid="86B17F71-3C6C-4829-9243-E31B65849341" docUuidSource="ZooBank" docVersion="9" lastPageNumber="180" masterDocId="FF80FFC1FFE0FFD9FFA0FFBC3175F778" masterDocTitle="Nanohystricidae n. fam., an unusual, plesiomorphic enarthronote mite family endemic to New Zealand (Acari, Oribatida)" masterLastPageNumber="204" masterPageNumber="151" pageNumber="154" updateTime="1698614142856" updateUser="plazi">
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<mods:title id="70D19BAE8EEF73390B1B57C6AB93648A">Nanohystricidae n. fam., an unusual, plesiomorphic enarthronote mite family endemic to New Zealand (Acari, Oribatida)</mods:title>
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<mods:namePart id="9AEBD9DF46486F85AFB1B9B524B1F2BA">Fuangarworn, Marut</mods:namePart>
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<mods:date id="A1570197996E2C6E1B0B22996EDD909B">2015</mods:date>
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<mods:number id="8B74AD5979C800F52FB15E0EB2E65A3E">4027</mods:number>
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<taxonomicName id="4C104D2CFFE3FFDAFF37FB3A30C1F3D8" box="[151,436,1158,1184]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
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<emphasis id="B964EABDFFE3FFDAFF37FB3A30C1F3D8" bold="true" box="[151,436,1158,1184]" italics="true" pageId="3" pageNumber="154">Nanohystrix hammerae</emphasis>
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Adult (
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<figureCitation id="132B2A2AFFE3FFDAFF49FB71302BF39E" box="[233,350,1229,1254]" captionStart-0="FIGURE 1. A – H" captionStart-1="FIGURE 2. A – E" captionStart-10="FIGURE 11. A – G" captionStart-11="FIGURE 12. A – F" captionStart-12="FIGURE 13. A – E" captionStart-13="FIGURE 14. A – I" captionStart-14="FIGURE 15. A – I" captionStart-15="FIGURE 16. A – G" captionStart-16="FIGURE 17. A – C" captionStart-2="FIGURE 3. A – D" captionStart-3="FIGURE 4. A – G" captionStart-4="FIGURE 5. A – G" captionStart-5="FIGURE 6. A – G" captionStart-6="FIGURE 7. A – J" captionStart-7="FIGURE 8. A – E" captionStart-8="FIGURE 9. A – B" captionStart-9="FIGURE 10. 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A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." captionText-1="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." captionText-10="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." captionText-11="FIGURE 12. A – F Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ 1 and coupled seta d; E, dorsolateral aspect of tibia I seta d (coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5 – 20 layers. Scale bars: 25 Μm (A – C, F); 10 Μm (D, E)." captionText-12="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." captionText-13="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." captionText-14="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." captionText-15="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." captionText-16="FIGURE 17. A – C Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, left leg III, tarsus partially shown; B, left tarsus III; C, left leg IV (slightly twisted at base). Labels all indicate setiform organs (see text). Scale bars: 100 Μm." captionText-2="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." captionText-3="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." captionText-4="FIGURE 5. A – G Nanohystrix hammerae n. sp., transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (pr. mu; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (po. mu) and deeper plane (right) with cuticular disk cd 5 (with underlying fibers, fib) and seta h 1 (with depressor muscle, h 1. mu); F, section of middle scissure (more lateral than indicated in A) showing base of seta f 1 and associated depressor muscle (f 1. mu); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations: bng (anterior border of notogaster), cr. mu (cheliceral retractor muscles); cw (caecum wall), ep 1 – 3 (plates of epimeres 1 – 3), nc (nuchal constriction), t. gr (transverse groove), sej (sejugal articulation), t. sul (transverse sulcus). All composites of 5 – 20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C – G)." captionText-5="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." captionText-6="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." captionText-7="FIGURE 8. A – E Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (A, C uncleared). A, Anterior portion of notogaster dissected from body (midline indicated by thin black line; prodorsal retractor muscles torn anteriorly) under strong DIC illumination to show muscles; B, left cuticular disk cd 4 and pale extension from middle scissure (posterodorsal view, anterior to right); C, region of cuticular disk cd 3 showing tendon insertion files and proximal parts of thin tendons (lateral aspect, anterior to right); D, portion of seta f 1, transmitted bright-field illumination; E, same, but with polarized light. A – C composites of 5 - 15 layers. Abbreviations: cd 1, cd 3 (cuticular disks), e 1. mu, f 1. mu (muscles acting on respective setae), pr. mu (dorsal components of prodorsal retractor muscles), ti (tendon insertions), t. sul (transverse sulcus, with concave wall appearing brown in transmitted light; other labels indicate setae (see text). Scale bars 20 Μm (A); 10 Μm (B – E)." captionText-8="FIGURE 9. A – B Nanohystrix hammerae n. sp., adult female. A, several of the large setae that are abbreviated in B and in Fig. 2 B; B, ventral aspect, legs and some setae only partially shown. Abbreviations: ips (lyrifissure); other labels indicate setae (see text). Scale bar: 100 Μm." captionText-9="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri-0="https://zenodo.org/record/238765/files/figure.png" httpUri-1="https://zenodo.org/record/238766/files/figure.png" httpUri-10="https://zenodo.org/record/238775/files/figure.png" httpUri-11="https://zenodo.org/record/238776/files/figure.png" httpUri-12="https://zenodo.org/record/238777/files/figure.png" httpUri-13="https://zenodo.org/record/238779/files/figure.png" httpUri-14="https://zenodo.org/record/238780/files/figure.png" httpUri-15="https://zenodo.org/record/238781/files/figure.png" httpUri-16="https://zenodo.org/record/238782/files/figure.png" httpUri-2="https://zenodo.org/record/238767/files/figure.png" httpUri-3="https://zenodo.org/record/238768/files/figure.png" httpUri-4="https://zenodo.org/record/238769/files/figure.png" httpUri-5="https://zenodo.org/record/238770/files/figure.png" httpUri-6="https://zenodo.org/record/238771/files/figure.png" httpUri-7="https://zenodo.org/record/238772/files/figure.png" httpUri-8="https://zenodo.org/record/238773/files/figure.png" httpUri-9="https://zenodo.org/record/238774/files/figure.png" pageId="3" pageNumber="154">Figs 1–17</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<emphasis id="B964EABDFFE3FFDAFED3FB71333DF39E" box="[371,584,1229,1254]" italics="true" pageId="3" pageNumber="154">General attributes.</emphasis>
|
||
Unusually large mites: total length 968–1213 Μm (smallest contracted male, largest distended female, respectively, not including setae). Average total length of females (n = 10) 1095 Μm (range
|
||
<date id="FFAE106FFFE3FFDAFF47FAAA301CF256" box="[231,361,1302,1326]" pageId="3" pageNumber="154" value="1029" valueMax="1213">1029–1213</date>
|
||
), average maximum width 697 Μm (range 653–774), average maximum height 636 Μm (range 605–677); average total length of males (n = 10) 1036 Μm (range 968–1137), average maximum width 622 Μm (range 593–653), average maximum height 584 Μm (range 532–665). Sejugal band of soft cuticle ca. 100 Μm wide dorsally, ca. 20 Μm ventrally (exposed or hidden according to degree of distention and prodorsal flexing).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE3FFDAFF67FA193244F1C2" blockId="3.[151,1437,1229,2010]" pageId="3" pageNumber="154">
|
||
Body shiny in reflected light. Cuticle mostly of three
|
||
<typeStatus id="54AB880DFFE3FFDAFC85FA1A3211F2C6" box="[805,868,1446,1470]" pageId="3" pageNumber="154">types</typeStatus>
|
||
. (a)
|
||
<emphasis id="B964EABDFFE3FFDAFC36FA193564F2C6" box="[918,1041,1445,1470]" italics="true" pageId="3" pageNumber="154">Sclerotized</emphasis>
|
||
cuticle conspicuously porose (
|
||
<figureCitation id="132B2A2AFFE3FFDAFACFFA1931ABF29A" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="3" pageNumber="154">Fig. 7A, B</figureCitation>
|
||
), brownish-orange in recently collected mature adults (
|
||
<figureCitation id="132B2A2AFFE3FFDAFCFAFA7532C6F29A" box="[858,947,1481,1506]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="3" pageNumber="154">Fig. 1D</figureCitation>
|
||
) but pale, brownish-yellow and somewhat translucent (eggs and food boluses visible within uncleared females) after long preservation (
|
||
<figureCitation id="132B2A2AFFE3FFDAFB49FA5134E5F17E" box="[1257,1424,1517,1542]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="3" pageNumber="154">Fig. 1A, E–H</figureCitation>
|
||
); tubercles of notogastral setae appearing slightly darker than surrounding cuticle; legs often slightly paler than body. (b)
|
||
<emphasis id="B964EABDFFE3FFDAFF64F98A3185F136" box="[196,240,1590,1614]" italics="true" pageId="3" pageNumber="154">Soft</emphasis>
|
||
cuticle of most articulations: colorless, thin, highly flexible, usually with surface granules (probably cerotegument) of various density and shape, in places forming thick vermiform pattern (
|
||
<figureCitation id="132B2A2AFFE3FFDAFB21F9E53459F10A" box="[1153,1324,1625,1650]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="3" pageNumber="154">Fig. 11A, D, E</figureCitation>
|
||
). (c)
|
||
<emphasis id="B964EABDFFE3FFDAFAC9F9E534E9F10A" box="[1385,1436,1625,1650]" italics="true" pageId="3" pageNumber="154">Pale</emphasis>
|
||
cuticle of apparently intermediate flexibility: with very faint pigmentation, fine and inconspicuous porosity, and in places with fine, dense surface striation (
|
||
<figureCitation id="132B2A2AFFE3FFDAFDFFF91D33BEF1C2" box="[607,715,1697,1722]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="3" pageNumber="154">Figs 10D</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE3FFDAFD76F91E3250F1C2" box="[726,805,1698,1722]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="3" pageNumber="154">11F, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE3FFDAFF67F97A3195F016" blockId="3.[151,1437,1229,2010]" pageId="3" pageNumber="154">
|
||
Setae hyaline, lustrous and glassy in reflected light (
|
||
<figureCitation id="132B2A2AFFE3FFDAFCB0F9793224F1A6" box="[784,849,1733,1758]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
|
||
), strongly birefringent in polarized transmitted light (
|
||
<figureCitation id="132B2A2AFFE3FFDAFF3FF955306DF07A" box="[159,280,1769,1794]" captionStart="FIGURE 8. A – E" captionStartId="13.[151,250,1220,1242]" captionTargetBox="[151,1436,193,1198]" captionTargetId="figure@13.[151,1436,193,1198]" captionTargetPageId="13" captionText="FIGURE 8. A – E Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (A, C uncleared). A, Anterior portion of notogaster dissected from body (midline indicated by thin black line; prodorsal retractor muscles torn anteriorly) under strong DIC illumination to show muscles; B, left cuticular disk cd 4 and pale extension from middle scissure (posterodorsal view, anterior to right); C, region of cuticular disk cd 3 showing tendon insertion files and proximal parts of thin tendons (lateral aspect, anterior to right); D, portion of seta f 1, transmitted bright-field illumination; E, same, but with polarized light. A – C composites of 5 - 15 layers. Abbreviations: cd 1, cd 3 (cuticular disks), e 1. mu, f 1. mu (muscles acting on respective setae), pr. mu (dorsal components of prodorsal retractor muscles), ti (tendon insertions), t. sul (transverse sulcus, with concave wall appearing brown in transmitted light; other labels indicate setae (see text). Scale bars 20 Μm (A); 10 Μm (B – E)." httpUri="https://zenodo.org/record/238772/files/figure.png" pageId="3" pageNumber="154">Fig. 8A, E</figureCitation>
|
||
). Most setae characterized below as:
|
||
<emphasis id="B964EABDFFE3FFDAFD18F9553260F07A" box="[696,789,1769,1794]" italics="true" pageId="3" pageNumber="154">acicular</emphasis>
|
||
if stiff, straight or slightly curved, tapering to thick point, heavily barbed (unless noted otherwise) almost to end; and
|
||
<emphasis id="B964EABDFFE3FFDAFC93F8B232EEF05D" box="[819,923,1806,1829]" italics="true" pageId="3" pageNumber="154">attenuate</emphasis>
|
||
if with long, taper to thin, smooth tip, with at least modest distal curvature and apparent flexibility. Barbs usually strongly developed and birefringent, like setal shafts.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE3FFDDFF67F8C53363F46D" blockId="3.[151,1437,1229,2010]" lastBlockId="4.[151,1437,151,1977]" lastPageId="4" lastPageNumber="155" pageId="3" pageNumber="154">
|
||
<emphasis id="B964EABDFFE3FFDAFF67F8C53031F0EA" box="[199,324,1913,1938]" italics="true" pageId="3" pageNumber="154">Prodorsum</emphasis>
|
||
. Outline of main aspis semi-elliptical in dorsal aspect (
|
||
<figureCitation id="132B2A2AFFE3FFDAFC66F8C53555F0EA" box="[966,1056,1913,1938]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="3" pageNumber="154">Fig. 2B</figureCitation>
|
||
), seen flat about 1.2 times wider than long (e.g. 430 x 370 Μm). In lateral view, plane of aspis usually at 50–70° angle from horizontal (
|
||
<figureCitation id="132B2A2AFFE3FFDAFAAAF8213412F0CE" box="[1290,1383,1949,1974]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="3" pageNumber="154">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE3FFDAFAD3F82131D9F0A2" captionStart="FIGURE 5. A – G" captionStartId="10.[151,250,1619,1641]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. A – G Nanohystrix hammerae n. sp., transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (pr. mu; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (po. mu) and deeper plane (right) with cuticular disk cd 5 (with underlying fibers, fib) and seta h 1 (with depressor muscle, h 1. mu); F, section of middle scissure (more lateral than indicated in A) showing base of seta f 1 and associated depressor muscle (f 1. mu); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations: bng (anterior border of notogaster), cr. mu (cheliceral retractor muscles); cw (caecum wall), ep 1 – 3 (plates of epimeres 1 – 3), nc (nuchal constriction), t. gr (transverse groove), sej (sejugal articulation), t. sul (transverse sulcus). All composites of 5 – 20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C – G)." httpUri="https://zenodo.org/record/238769/files/figure.png" pageId="3" pageNumber="154">5A, B</figureCitation>
|
||
), maximum seen ca. 80°, minimum ca. 10° (great rotation allowed by ventrosejugal articulation and broad soft cuticle of dorsosejugal, disjugal and pleural regions). Aspis cuticle finely porose throughout (
|
||
<figureCitation id="132B2A2AFFE4FFDDFB7FFF2B342AF7C8" box="[1247,1375,151,176]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">Fig. 7A, B</figureCitation>
|
||
) and uniformly colored except for eyes (see below); sigilla of cheliceral retractor and subcapitular levator muscles in several pairs of patches occupying most of posterior half (
|
||
<figureCitation id="132B2A2AFFE4FFDDFC99FF6332E3F780" box="[825,918,223,248]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">Fig. 7A</figureCitation>
|
||
). Aspis dorsally convex (
|
||
<figureCitation id="132B2A2AFFE4FFDDFB1DFF633462F780" box="[1213,1303,223,248]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="4" pageNumber="155">Fig. 1C</figureCitation>
|
||
); anteriorly with slightly recurved rostral margin (
|
||
<emphasis id="B964EABDFFE4FFDDFDE7FEB93313F664" box="[583,614,261,284]" italics="true" pageId="4" pageNumber="155">rm</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFE4FFDDFDD6FEB833BEF665" box="[630,715,260,285]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">Fig. 6F</figureCitation>
|
||
); margin continuing posteriorly on each side as sharply defined carina (
|
||
<emphasis id="B964EABDFFE4FFDDFF4FFE94306DF638" box="[239,280,296,320]" italics="true" pageId="4" pageNumber="155">ca1</emphasis>
|
||
), ending at lateral eye, thereby forming outline of aspis in dorsal aspect; with second carina (
|
||
<emphasis id="B964EABDFFE4FFDDFACAFE9434E7F638" box="[1386,1426,296,320]" italics="true" pageId="4" pageNumber="155">ca2</emphasis>
|
||
) descending from eye at right angle (
|
||
<figureCitation id="132B2A2AFFE4FFDDFD95FEF033E6F61C" box="[565,659,332,357]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="4" pageNumber="155">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFD01FEF033B2F61D" box="[673,711,332,357]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">6A</figureCitation>
|
||
). Lateral region between carinae concave (
|
||
<figureCitation id="132B2A2AFFE4FFDDFB12FEF0347EF61C" box="[1202,1291,332,357]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="4" pageNumber="155">Fig. 1C</figureCitation>
|
||
; presumably relating to motion of leg I), subtriangular in lateral view, with irregular ventral margin merging with soft supracoxal region (i.e., no ‘prodorsal tectum’ present); with long, narrow extension forming supportive strut (
|
||
<emphasis id="B964EABDFFE4FFDDFB56FE293466F6D4" box="[1270,1299,405,428]" italics="true" pageId="4" pageNumber="155">str</emphasis>
|
||
) with distal condyle that articulates with subcapitulum (
|
||
<figureCitation id="132B2A2AFFE4FFDDFD04FE0B327DF6A8" box="[676,776,439,464]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="4" pageNumber="155">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFCBBFE04321EF6A8" box="[795,875,440,464]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">7D, G</figureCitation>
|
||
). Rostral tectum very narrow, crescent-shaped; undulating vertically to accommodate chelicerae (
|
||
<figureCitation id="132B2A2AFFE4FFDDFD55FE603223F68D" box="[757,854,476,501]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">Fig. 6B</figureCitation>
|
||
); reflected ventral part (rostrophragma) without noticeable solid limb; about 60 Μm deep medially, effacing laterally and ending at level midway between rostral and lamellar setae. Tectum anteromedially with low, well-circumscribed, pale rostral bulge precisely dorsal to median eye, ca. 50 µm wide (
|
||
<emphasis id="B964EABDFFE4FFDDFE42FDFB3371F518" box="[482,516,583,608]" italics="true" pageId="4" pageNumber="155">rb;</emphasis>
|
||
<figureCitation id="132B2A2AFFE4FFDDFDABFDFB3310F518" box="[523,613,583,608]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">Figs 6C</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFDD0FDF433D9F518" box="[624,684,584,608]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">7H, I</figureCitation>
|
||
). Prodorsum abruptly constricted posteriorly, forming near-vertical ‘occipital’ wall (
|
||
<emphasis id="B964EABDFFE4FFDDFEF3FDD13001F5FC" box="[339,372,621,644]" italics="true" pageId="4" pageNumber="155">ow</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFE4FFDDFE23FDD030ABF5FC" box="[387,478,620,645]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="4" pageNumber="155">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFE4AFDD0337BF5FC" box="[490,526,620,644]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="4" pageNumber="155">3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFDBBFDD0334BF5FD" box="[539,574,620,645]" captionStart="FIGURE 5. A – G" captionStartId="10.[151,250,1619,1641]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. A – G Nanohystrix hammerae n. sp., transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (pr. mu; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (po. mu) and deeper plane (right) with cuticular disk cd 5 (with underlying fibers, fib) and seta h 1 (with depressor muscle, h 1. mu); F, section of middle scissure (more lateral than indicated in A) showing base of seta f 1 and associated depressor muscle (f 1. mu); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations: bng (anterior border of notogaster), cr. mu (cheliceral retractor muscles); cw (caecum wall), ep 1 – 3 (plates of epimeres 1 – 3), nc (nuchal constriction), t. gr (transverse groove), sej (sejugal articulation), t. sul (transverse sulcus). All composites of 5 – 20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C – G)." httpUri="https://zenodo.org/record/238769/files/figure.png" pageId="4" pageNumber="155">5C</figureCitation>
|
||
), ca. 50 Μm high, edge of constriction bordered by carina
|
||
<emphasis id="B964EABDFFE4FFDDFB6DFDD03583F5FC" box="[1229,1270,620,644]" italics="true" pageId="4" pageNumber="155">ca2</emphasis>
|
||
(laterally) and setal pairs
|
||
<emphasis id="B964EABDFFE4FFDDFEB4FD2D305FF5D0" box="[276,298,657,680]" italics="true" pageId="4" pageNumber="155">in</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE4FFDDFE9AFD333023F5D0" box="[314,342,655,680]" italics="true" pageId="4" pageNumber="155">bo</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE4FFDDFE32FD2D30DCF5D0" box="[402,425,657,680]" italics="true" pageId="4" pageNumber="155">xs</emphasis>
|
||
(dorsally); wall with sharp posterior bend to form nuchal constriction (
|
||
<emphasis id="B964EABDFFE4FFDDFB45FD2D3475F5D0" box="[1253,1280,657,680]" italics="true" pageId="4" pageNumber="155">nc</emphasis>
|
||
) that extends posteriorly for ca. 30 Μm before joining soft sejugal cuticle (
|
||
<figureCitation id="132B2A2AFFE4FFDDFCF8FD0832CDF5B4" box="[856,952,692,717]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="4" pageNumber="155">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFC67FD08329FF5B5" box="[967,1002,692,717]" captionStart="FIGURE 5. A – G" captionStartId="10.[151,250,1619,1641]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. A – G Nanohystrix hammerae n. sp., transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (pr. mu; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (po. mu) and deeper plane (right) with cuticular disk cd 5 (with underlying fibers, fib) and seta h 1 (with depressor muscle, h 1. mu); F, section of middle scissure (more lateral than indicated in A) showing base of seta f 1 and associated depressor muscle (f 1. mu); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations: bng (anterior border of notogaster), cr. mu (cheliceral retractor muscles); cw (caecum wall), ep 1 – 3 (plates of epimeres 1 – 3), nc (nuchal constriction), t. gr (transverse groove), sej (sejugal articulation), t. sul (transverse sulcus). All composites of 5 – 20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C – G)." httpUri="https://zenodo.org/record/238769/files/figure.png" pageId="4" pageNumber="155">5C</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFC58FD08356BF5B5" box="[1016,1054,692,717]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">6A</figureCitation>
|
||
). Nuchal constriction comprising posterior fifth of aspis; hidden in fully contracted specimens, in which occipital wall abuts notogaster, protecting sejugal articulation (
|
||
<figureCitation id="132B2A2AFFE4FFDDFEDBFD4030A2F46C" box="[379,471,764,789]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="4" pageNumber="155">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFE43FD40337DF46D" box="[483,520,764,789]" captionStart="FIGURE 5. A – G" captionStartId="10.[151,250,1619,1641]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. A – G Nanohystrix hammerae n. sp., transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (pr. mu; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (po. mu) and deeper plane (right) with cuticular disk cd 5 (with underlying fibers, fib) and seta h 1 (with depressor muscle, h 1. mu); F, section of middle scissure (more lateral than indicated in A) showing base of seta f 1 and associated depressor muscle (f 1. mu); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations: bng (anterior border of notogaster), cr. mu (cheliceral retractor muscles); cw (caecum wall), ep 1 – 3 (plates of epimeres 1 – 3), nc (nuchal constriction), t. gr (transverse groove), sej (sejugal articulation), t. sul (transverse sulcus). All composites of 5 – 20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C – G)." httpUri="https://zenodo.org/record/238769/files/figure.png" pageId="4" pageNumber="155">5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE4FFDDFF67FCA33079F494" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="155">
|
||
Median eye (
|
||
<emphasis id="B964EABDFFE4FFDDFEFBFC9D30E5F440" box="[347,400,801,824]" italics="true" pageId="4" pageNumber="155">oc.m</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFE4FFDDFE00FCA33365F440" box="[416,528,799,824]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">Fig. 7H, J</figureCitation>
|
||
) about 50 µm diameter, located on rostrophragma directly under pale, dome-like rostral bulge (
|
||
<emphasis id="B964EABDFFE4FFDDFE99FCF83027F425" box="[313,338,836,861]" italics="true" pageId="4" pageNumber="155">rb</emphasis>
|
||
); with strongly convex but rather thin-walled cornea. No pigment observed in median eye, but sclerotized cuticle of rostrophragma thick and pigmented posterior to eye (in region of constriction
|
||
<emphasis id="B964EABDFFE4FFDDFA86FCD5344CF4F8" box="[1318,1337,873,896]" italics="true" pageId="4" pageNumber="155">st</emphasis>
|
||
). Paired lateral eye (
|
||
<emphasis id="B964EABDFFE4FFDDFEBBFC303031F4DD" box="[283,324,908,933]" italics="true" pageId="4" pageNumber="155">oc.l</emphasis>
|
||
) smaller, about 25 µm diameter, each immediately ventral to seta
|
||
<emphasis id="B964EABDFFE4FFDDFB87FC31354EF4DC" box="[1063,1083,909,932]" italics="true" pageId="4" pageNumber="155">xi</emphasis>
|
||
at junction of carinae
|
||
<emphasis id="B964EABDFFE4FFDDFA95FC30342BF4DC" box="[1333,1374,908,932]" italics="true" pageId="4" pageNumber="155">ca1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE4FFDDFACCFC3034E1F4DC" box="[1388,1428,908,932]" italics="true" pageId="4" pageNumber="155">ca2</emphasis>
|
||
; dark pigmentation visible beneath hemispherical cuticular cornea (
|
||
<figureCitation id="132B2A2AFFE4FFDDFC2BFC133293F4B0" box="[907,998,943,968]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">Fig. 7D</figureCitation>
|
||
), lost with clearing of specimens (
|
||
<figureCitation id="132B2A2AFFE4FFDDFACFFC133075F494" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">Fig. 7A, C, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE4FFDDFF67FC4B3393F170" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="155">
|
||
Prodorsal setae of various forms (
|
||
<figureCitation id="132B2A2AFFE4FFDDFD9EFC4B33ECF368" box="[574,665,1015,1040]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="4" pageNumber="155">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFD04FC4433BDF368" box="[676,712,1016,1040]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="4" pageNumber="155">3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFD74FC443383F368" box="[724,758,1016,1040]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="4" pageNumber="155">4B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFCA2FC4B325DF368" box="[770,808,1015,1040]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">6A</figureCitation>
|
||
), larger ones inserting on conspicuous tubercles. Rostral seta (
|
||
<emphasis id="B964EABDFFE4FFDDFF74FBA1319EF34C" box="[212,235,1053,1076]" italics="true" pageId="4" pageNumber="155">ro</emphasis>
|
||
) very thin, attenuate, ca. 170 Μm long, straight except for fine tip, projecting anteriorly, with only several inconspicuous barbs; pair slightly divergent, inserted close together (ca. 25–30 Μm mutual distance) on rostral bulge. Lamellar seta (
|
||
<emphasis id="B964EABDFFE4FFDDFE39FBD830D8F305" box="[409,429,1124,1149]" italics="true" pageId="4" pageNumber="155">le</emphasis>
|
||
) inserted slightly proximal to mid-length of prodorsum, erect but moderately curved; relatively thick, attenuate, ca. 420 Μm long, with strong barbs in three longitudinal rows for much of length; pair widely spaced (ca. 255 Μm mutual distance). Remaining four pairs of prodorsal setae aligned at edge of occipital wall; all nearly erect, with slight posterior direction. Interlamellar (
|
||
<emphasis id="B964EABDFFE4FFDDFC27FB6D32E8F390" box="[903,925,1233,1256]" italics="true" pageId="4" pageNumber="155">in</emphasis>
|
||
) and upper bothridial setae (
|
||
<emphasis id="B964EABDFFE4FFDDFB40FB6D3582F390" box="[1248,1271,1233,1256]" italics="true" pageId="4" pageNumber="155">xs</emphasis>
|
||
) larger than
|
||
<emphasis id="B964EABDFFE4FFDDFA28FB7334E9F390" box="[1416,1436,1231,1256]" italics="true" pageId="4" pageNumber="155">le</emphasis>
|
||
and bearing up to five rows of barbs in proximal region:
|
||
<emphasis id="B964EABDFFE4FFDDFCB5FB49325EF274" box="[789,811,1269,1292]" italics="true" pageId="4" pageNumber="155">in</emphasis>
|
||
about 570 Μm long, moderately attenuate;
|
||
<emphasis id="B964EABDFFE4FFDDFAAEFB493450F274" box="[1294,1317,1269,1292]" italics="true" pageId="4" pageNumber="155">xs</emphasis>
|
||
about 480 Μm, acicular, with strong barbs along whole length. Lower exobothridial seta (
|
||
<emphasis id="B964EABDFFE4FFDDFBBCFAA53545F248" box="[1052,1072,1305,1328]" italics="true" pageId="4" pageNumber="155">xi</emphasis>
|
||
) small (about 50 Μm); acicular, with minute barbs visible only at high magnification. Bothridial seta (
|
||
<emphasis id="B964EABDFFE4FFDDFC07FA8032B6F22D" box="[935,963,1340,1365]" italics="true" pageId="4" pageNumber="155">bo</emphasis>
|
||
, = ‘sensillus’) thinner than setae on either side (
|
||
<emphasis id="B964EABDFFE4FFDDFF7AFADD3185F200" box="[218,240,1377,1400]" italics="true" pageId="4" pageNumber="155">in</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE4FFDDFEA4FADD306EF200" box="[260,283,1377,1400]" italics="true" pageId="4" pageNumber="155">xs</emphasis>
|
||
); about 500 Μm long, tapering to thin flexible tip, with 2–4 rows of barbs along whole length. Bothridium (
|
||
<figureCitation id="132B2A2AFFE4FFDDFE87FA3830F1F2E5" box="[295,388,1412,1437]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">Figs 6D</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFE31FA3930BAF2E4" box="[401,463,1412,1436]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="4" pageNumber="155">7E, F</figureCitation>
|
||
) with simple conical form, no proximal curvature; outer half lined with approximately 20 concentric closely-spaced, fine circular ridges (some merging), which separate even finer honeycomb pattern; inner half abruptly narrower, with coarser ridges directed internally. Bothridial rim projecting slightly from surface; posterolaterally with tongue-like flange (
|
||
<emphasis id="B964EABDFFE4FFDDFDC2FA533304F170" box="[610,625,1519,1544]" italics="true" pageId="4" pageNumber="155">fl</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFE4FFDDFD20FA5333ACF170" box="[640,729,1519,1544]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">Fig. 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE4FFDDFF67F9A934E9F0C0" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="155">
|
||
<emphasis id="B964EABDFFE4FFDDFF67F9A9303DF154" box="[199,328,1557,1580]" italics="true" pageId="4" pageNumber="155">Notogaster.</emphasis>
|
||
Length about three times that of prodorsal aspis (excluding nuchal constriction); anterior width similar to that of prodorsum, about half that of maximum. Strongly but unevenly convex in both dorsal and lateral aspects (
|
||
<figureCitation id="132B2A2AFFE4FFDDFF59F9E03034F10C" box="[249,321,1628,1653]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="4" pageNumber="155">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFEF1F9E03001F10C" box="[337,372,1628,1652]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="4" pageNumber="155">2B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFE22F9E030D2F10C" box="[386,423,1628,1652]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="4" pageNumber="155">3A</figureCitation>
|
||
); maximum width and height at about level of setal row
|
||
<emphasis id="B964EABDFFE4FFDDFB96F9E0354BF10C" box="[1078,1086,1628,1652]" italics="true" pageId="4" pageNumber="155">f</emphasis>
|
||
; curvature steeper in posterior third (more marked in fully distended individuals). Notogaster without marginal tectum; weakly folded laterally in anterior third (humeral crease;
|
||
<emphasis id="B964EABDFFE4FFDDFE57F9183356F1C5" box="[503,547,1700,1725]" italics="true" pageId="4" pageNumber="155">h.cr</emphasis>
|
||
), forming pleural concavity (more defined in contracted individuals;
|
||
<figureCitation id="132B2A2AFFE4FFDDFA9AF91834E3F1C4" box="[1338,1430,1700,1725]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="4" pageNumber="155">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE4FFDDFF37F97431C8F198" box="[151,189,1736,1760]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="4" pageNumber="155">4D</figureCitation>
|
||
); concavity effaces posterior to setal row
|
||
<emphasis id="B964EABDFFE4FFDDFD2BF97533E2F198" box="[651,663,1737,1760]" italics="true" pageId="4" pageNumber="155">e</emphasis>
|
||
, pleural region becoming slightly convex posteriorly. General cuticle rather uniform and featureless, except as noted below; sclerotized cuticle with pores finer and less conspicuous than those of prodorsum. Well-spaced, minute surface granules in soft bordering cuticle, continuing onto anterior (
|
||
<figureCitation id="132B2A2AFFE4FFDDFF3FF888318EF035" box="[159,251,1844,1869]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">Fig. 6A</figureCitation>
|
||
, arrow) and lateral notogastral surface, becoming less dense in middle region of notogaster and absent posteriorly. Narrow transverse band of fine reticulation present in region of setal pairs
|
||
<emphasis id="B964EABDFFE4FFDDFBCAF8E435F0F008" box="[1130,1157,1880,1904]" italics="true" pageId="4" pageNumber="155">c1</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE4FFDDFB1CF8E435A2F008" box="[1212,1239,1880,1904]" italics="true" pageId="4" pageNumber="155">c2</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFE4FFDDFB47F8EB344BF008" box="[1255,1342,1879,1904]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="4" pageNumber="155">Fig. 6E</figureCitation>
|
||
, arrow). Notogaster and adanal plates fused posterolaterally, commencing at about level of seta
|
||
<emphasis id="B964EABDFFE4FFDDFBD4F8C035EAF0ED" box="[1140,1183,1916,1941]" italics="true" pageId="4" pageNumber="155">ad2</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFE4FFDDFB0FF8C0347FF0EC" box="[1199,1290,1916,1941]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="4" pageNumber="155">Fig. 3A</figureCitation>
|
||
; see Remark 9); narrow separating cuticle anterior to
|
||
<emphasis id="B964EABDFFE4FFDDFDF7F81C33F4F0C1" box="[599,641,1952,1977]" italics="true" pageId="4" pageNumber="155">ad2</emphasis>
|
||
pale, finely porose, hidden in fold in contracted individuals (
|
||
<figureCitation id="132B2A2AFFE4FFDDFA89F81C34FAF0C0" box="[1321,1423,1952,1977]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="4" pageNumber="155">Fig. 10B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE5FFDCFF67FF2B307FF59F" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="156">
|
||
Dorsal curvature interrupted by five transverse formations (
|
||
<figureCitation id="132B2A2AFFE5FFDCFCD0FF2B32ADF7C8" box="[880,984,151,176]" captionStart-0="FIGURE 2. A – E" captionStart-1="FIGURE 3. A – D" captionStart-2="FIGURE 4. A – G" captionStart-3="FIGURE 5. A – G" captionStartId-0="7.[151,250,1851,1873]" captionStartId-1="8.[151,250,1858,1880]" captionStartId-2="9.[151,250,1782,1804]" captionStartId-3="10.[151,250,1619,1641]" captionTargetBox-0="[161,1425,198,1839]" captionTargetBox-1="[193,1407,191,1840]" captionTargetBox-2="[151,1436,193,1759]" captionTargetBox-3="[151,1436,193,1596]" captionTargetId-0="figure@7.[151,1436,193,1844]" captionTargetId-1="figure@8.[189,1413,180,1840]" captionTargetId-2="figure@9.[151,1436,193,1760]" captionTargetId-3="figure@10.[151,1436,193,1598]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." captionText-1="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." captionText-2="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." captionText-3="FIGURE 5. A – G Nanohystrix hammerae n. sp., transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (pr. mu; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (po. mu) and deeper plane (right) with cuticular disk cd 5 (with underlying fibers, fib) and seta h 1 (with depressor muscle, h 1. mu); F, section of middle scissure (more lateral than indicated in A) showing base of seta f 1 and associated depressor muscle (f 1. mu); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations: bng (anterior border of notogaster), cr. mu (cheliceral retractor muscles); cw (caecum wall), ep 1 – 3 (plates of epimeres 1 – 3), nc (nuchal constriction), t. gr (transverse groove), sej (sejugal articulation), t. sul (transverse sulcus). All composites of 5 – 20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C – G)." httpUri-0="https://zenodo.org/record/238766/files/figure.png" httpUri-1="https://zenodo.org/record/238767/files/figure.png" httpUri-2="https://zenodo.org/record/238768/files/figure.png" httpUri-3="https://zenodo.org/record/238769/files/figure.png" pageId="5" pageNumber="156">Figs 2–5</figureCitation>
|
||
). (a) Shallow, simple transverse groove (
|
||
<emphasis id="B964EABDFFE5FFDCFF00FF0631B9F7A9" box="[160,204,186,209]" italics="true" pageId="5" pageNumber="156">t.gr.</emphasis>
|
||
) immediately posterior to setal pairs
|
||
<emphasis id="B964EABDFFE5FFDCFD24FF05338DF7A9" box="[644,760,185,210]" italics="true" pageId="5" pageNumber="156">c1 and c2</emphasis>
|
||
. (b) Deeper, broader transverse sulcus (
|
||
<emphasis id="B964EABDFFE5FFDCFB6CFF05358EF7AA" box="[1228,1275,185,210]" italics="true" pageId="5" pageNumber="156">t.sul</emphasis>
|
||
) immediately posterior to setal row
|
||
<emphasis id="B964EABDFFE5FFDCFE2BFF6630ECF78B" box="[395,409,218,243]" italics="true" pageId="5" pageNumber="156">d</emphasis>
|
||
. (c) Transverse type-S scissure bearing setal pairs
|
||
<emphasis id="B964EABDFFE5FFDCFC71FF673299F78B" box="[977,1004,219,243]" italics="true" pageId="5" pageNumber="156">e1</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFB82FF673549F78B" box="[1058,1084,219,243]" italics="true" pageId="5" pageNumber="156">e2</emphasis>
|
||
. (d) Transverse type-S scissure bearing setal pairs
|
||
<emphasis id="B964EABDFFE5FFDCFED1FF4030F3F66C" box="[369,390,252,276]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFE37FF4030D8F66C" box="[407,429,252,276]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFE49FF403370F66D" box="[489,517,252,277]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
. (e) Transverse type-S scissure bearing setal pair
|
||
<emphasis id="B964EABDFFE5FFDCFBE5FF403514F66D" box="[1093,1121,252,277]" italics="true" pageId="5" pageNumber="156">h1</emphasis>
|
||
. Setae of row
|
||
<emphasis id="B964EABDFFE5FFDCFAAEFF403469F66D" box="[1294,1308,252,277]" italics="true" pageId="5" pageNumber="156">d</emphasis>
|
||
on modest tubercles within narrowly elliptical, irregularly-bordered region of lighter-colored cuticle (
|
||
<figureCitation id="132B2A2AFFE5FFDCFB05FEA13470F64E" box="[1189,1285,285,310]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="5" pageNumber="156">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFAB4FEA2344DF64E" box="[1300,1336,286,310]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="5" pageNumber="156">2B</figureCitation>
|
||
); central region of ellipse (around pair
|
||
<emphasis id="B964EABDFFE5FFDCFE44FE823374F62F" box="[484,513,318,343]" italics="true" pageId="5" pageNumber="156">d1</emphasis>
|
||
) pale, lateral regions around pair
|
||
<emphasis id="B964EABDFFE5FFDCFCDBFE8232E2F62F" box="[891,919,318,343]" italics="true" pageId="5" pageNumber="156">d2</emphasis>
|
||
intermediate in color. Edges of pale cuticle of type-S scissures generally regular anteriorly but irregular posteriorly. Erectile setae of type-S scissures each on large tubercle with associated muscles (see Discussion). Scissures differ in: setal spacing, presence and form of side extensions from posterior slope of tubercle, presence of incursions (from sclerite behind scissure) or excursions (of pale cuticle into posterior sclerite). Anterior scissure (
|
||
<figureCitation id="132B2A2AFFE5FFDCFC0CFE783571F6A4" box="[940,1028,452,477]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="5" pageNumber="156">Fig. 2C</figureCitation>
|
||
) with setae of row
|
||
<emphasis id="B964EABDFFE5FFDCFB48FE793581F6A4" box="[1256,1268,453,476]" italics="true" pageId="5" pageNumber="156">e</emphasis>
|
||
almost evenly spaced; tubercles of pair
|
||
<emphasis id="B964EABDFFE5FFDCFE6AFE5A3091F686" box="[458,484,486,510]" italics="true" pageId="5" pageNumber="156">e1</emphasis>
|
||
with medial extensions fusing to form narrow bridge; with small incursion of sclerotization between
|
||
<emphasis id="B964EABDFFE5FFDCFE3AFDBB30C1F567" box="[410,436,519,543]" italics="true" pageId="5" pageNumber="156">e1</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFE4CFDBB3372F567" box="[492,519,519,543]" italics="true" pageId="5" pageNumber="156">e2</emphasis>
|
||
(*), sometimes appearing like separate, intercalary sclerite. Middle scissure (
|
||
<figureCitation id="132B2A2AFFE5FFDCFACEFDBA31CFF538" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="5" pageNumber="156">Fig. 2D</figureCitation>
|
||
) with mixed setation, including two pairs of row
|
||
<emphasis id="B964EABDFFE5FFDCFD51FD94338CF538" box="[753,761,552,576]" italics="true" pageId="5" pageNumber="156">f</emphasis>
|
||
setae and seta
|
||
<emphasis id="B964EABDFFE5FFDCFC06FD9B32B4F538" box="[934,961,551,576]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
; pair
|
||
<emphasis id="B964EABDFFE5FFDCFBA6FD94356EF538" box="[1030,1051,552,576]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
widely separated; central third of scissure narrow, without setae or noticeable incursions or intercalary sclerites; tubercles of
|
||
<emphasis id="B964EABDFFE5FFDCFB2AFDF535EAF519" box="[1162,1183,585,609]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFB74FDF5359FF519" box="[1236,1258,585,609]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
each with basal extensions, separated by narrow articulation; tubercle of
|
||
<emphasis id="B964EABDFFE5FFDCFC83FDD7324DF5FB" box="[803,824,619,643]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
also extended laterally, reaching near tubercle of
|
||
<emphasis id="B964EABDFFE5FFDCFAD8FDD634E1F5FB" box="[1400,1428,618,643]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
; tubercle of
|
||
<emphasis id="B964EABDFFE5FFDCFEB5FD303047F5DD" box="[277,306,652,677]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
without extension; scissure with irregular triangular region of pale cuticle extending posteriorly from near setae
|
||
<emphasis id="B964EABDFFE5FFDCFEB2FD123052F5BE" box="[274,295,686,710]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFEC3FD12300DF5BE" box="[355,376,686,710]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
. Posterior type-S scissure (
|
||
<figureCitation id="132B2A2AFFE5FFDCFD15FD11327EF5BE" box="[693,779,685,710]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="5" pageNumber="156">Fig. 2E</figureCitation>
|
||
) with only setal pair
|
||
<emphasis id="B964EABDFFE5FFDCFBA5FD113554F5BE" box="[1029,1057,685,710]" italics="true" pageId="5" pageNumber="156">h1</emphasis>
|
||
; each tubercle with short medial extension.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE5FFDCFF67FD533428F3A3" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="156">
|
||
With 19 pairs of glassy notogastral setae (including row
|
||
<emphasis id="B964EABDFFE5FFDCFC99FD533220F470" box="[825,853,751,776]" italics="true" pageId="5" pageNumber="156">ad</emphasis>
|
||
); some (
|
||
<emphasis id="B964EABDFFE5FFDCFC17FD4C32A7F470" box="[951,978,752,776]" italics="true" pageId="5" pageNumber="156">e1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFC7FFD4C328CF470" box="[991,1017,752,776]" italics="true" pageId="5" pageNumber="156">e2</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFBA7FD533557F470" box="[1031,1058,751,776]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
) extremely long, some (
|
||
<emphasis id="B964EABDFFE5FFDCFA93FD4C343DF470" box="[1331,1352,752,776]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFAF6FD4C341EF470" box="[1366,1387,752,776]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFAD9FD5334E1F470" box="[1401,1428,751,776]" italics="true" pageId="5" pageNumber="156">h1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFF37FCAE31C6F452" box="[151,179,786,810]" italics="true" pageId="5" pageNumber="156">p1</emphasis>
|
||
) merging posteriorly in brush-like effect when held horizontally (
|
||
<figureCitation id="132B2A2AFFE5FFDCFC08FCAD3572F452" box="[936,1031,785,810]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="5" pageNumber="156">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFBB5FCAE354EF452" box="[1045,1083,786,810]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="5" pageNumber="156">4A</figureCitation>
|
||
). Long setae of various forms: acicular setae and proximal regions of large attenuate setae with four (sometimes five) straight rows of dense barbs, equally spaced around seta (
|
||
<figureCitation id="132B2A2AFFE5FFDCFE7FFCE8334CF414" box="[479,569,852,877]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="5" pageNumber="156">Fig. 4E</figureCitation>
|
||
), forming imbricate ridges where barbs long and closely spaced (
|
||
<figureCitation id="132B2A2AFFE5FFDCFA91FCE834FAF414" box="[1329,1423,852,877]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="5" pageNumber="156">Fig. 4G</figureCitation>
|
||
); attenuate setae usually with two or three rows of more widely spaced barbs, at least in distal half (
|
||
<figureCitation id="132B2A2AFFE5FFDCFB58FCC93425F4F6" box="[1272,1360,885,910]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="5" pageNumber="156">Fig. 4F</figureCitation>
|
||
). Setal shapes and representative lengths (to show relativity in average female; all significantly variable) as follows:
|
||
<emphasis id="B964EABDFFE5FFDCFA21FC2B34EEF4D7" box="[1409,1435,919,943]" italics="true" pageId="5" pageNumber="156">c1</emphasis>
|
||
(170 Μm) finely attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFE65FC043095F4A8" box="[453,480,952,976]" italics="true" pageId="5" pageNumber="156">c2</emphasis>
|
||
(230 Μm) narrowly acicular;
|
||
<emphasis id="B964EABDFFE5FFDCFC8DFC043232F4A8" box="[813,839,952,976]" italics="true" pageId="5" pageNumber="156">c3</emphasis>
|
||
(750 Μm) attenuate);
|
||
<emphasis id="B964EABDFFE5FFDCFB9DFC05352DF4A8" box="[1085,1112,953,976]" italics="true" pageId="5" pageNumber="156">cp</emphasis>
|
||
(950 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFAE5FC0B3414F4A8" box="[1349,1377,951,976]" italics="true" pageId="5" pageNumber="156">d1</emphasis>
|
||
(970 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFEE6FC653017F48A" box="[326,354,985,1010]" italics="true" pageId="5" pageNumber="156">d2</emphasis>
|
||
(320 Μm) narrowly acicular, but curved;
|
||
<emphasis id="B964EABDFFE5FFDCFCE7FC663214F48A" box="[839,865,986,1010]" italics="true" pageId="5" pageNumber="156">e1</emphasis>
|
||
(1900 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFBC6FC6635F4F48A" box="[1126,1153,986,1010]" italics="true" pageId="5" pageNumber="156">e2</emphasis>
|
||
(1750 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFA26FC6634EEF48A" box="[1414,1435,986,1010]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
(1200 Μm) acicular;
|
||
<emphasis id="B964EABDFFE5FFDCFE20FC4730E3F36B" box="[384,406,1019,1043]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
(1000 Μm) acicular;
|
||
<emphasis id="B964EABDFFE5FFDCFD27FC4633D7F36B" box="[647,674,1018,1043]" italics="true" pageId="5" pageNumber="156">h1</emphasis>
|
||
(880 Μm) acicular;
|
||
<emphasis id="B964EABDFFE5FFDCFC26FC4632D6F36B" box="[902,931,1018,1043]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
(1350 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFB00FC4635C9F36B" box="[1184,1212,1018,1043]" italics="true" pageId="5" pageNumber="156">h3</emphasis>
|
||
(430 Μm) narrowly acicular;
|
||
<emphasis id="B964EABDFFE5FFDCFF5FFBA0306EF34C" box="[255,283,1052,1076]" italics="true" pageId="5" pageNumber="156">p1</emphasis>
|
||
(790 Μm) acicular;
|
||
<emphasis id="B964EABDFFE5FFDCFE5CFBA0336CF34C" box="[508,537,1052,1076]" italics="true" pageId="5" pageNumber="156">p2</emphasis>
|
||
(1300 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFCB4FBA03245F34C" box="[788,816,1052,1076]" italics="true" pageId="5" pageNumber="156">p3</emphasis>
|
||
(330 Μm) narrowly acicular;
|
||
<emphasis id="B964EABDFFE5FFDCFBDEFBA035DDF34D" box="[1150,1192,1052,1077]" italics="true" pageId="5" pageNumber="156">ad1</emphasis>
|
||
(1250 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFF37FB8131B4F32E" box="[151,193,1085,1110]" italics="true" pageId="5" pageNumber="156">ad2</emphasis>
|
||
(370 Μm) attenuate;
|
||
<emphasis id="B964EABDFFE5FFDCFE12FB8130A9F32E" box="[434,476,1085,1110]" italics="true" pageId="5" pageNumber="156">ad3</emphasis>
|
||
(200 Μm) attenuate. Six large pairs, including
|
||
<emphasis id="B964EABDFFE5FFDCFC54FB82357BF32E" box="[1012,1038,1086,1110]" italics="true" pageId="5" pageNumber="156">e1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFBBEFB82354DF32E" box="[1054,1080,1086,1110]" italics="true" pageId="5" pageNumber="156">e2</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFBE8FB82352BF32E" box="[1096,1118,1086,1110]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFBCEFB8235F6F32E" box="[1134,1155,1086,1110]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFB33FB8135DAF32E" box="[1171,1199,1085,1110]" italics="true" pageId="5" pageNumber="156">h1</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFB49FB813473F32E" box="[1257,1286,1085,1110]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
, inserting on intercalary sclerites within scissures (described above), apparently erectile. Several setal pairs with unusual locations (
|
||
<figureCitation id="132B2A2AFFE5FFDCFEABFBC33012F3E0" box="[267,359,1151,1176]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="5" pageNumber="156">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFED2FB3C30E3F3E0" box="[370,406,1152,1176]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="5" pageNumber="156">3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFE02FB3C30B2F3E0" box="[418,455,1152,1176]" captionStart="FIGURE 9. A – B" captionStartId="14.[151,250,1638,1660]" captionTargetBox="[155,1431,192,1610]" captionTargetId="figure@14.[151,1436,181,1617]" captionTargetPageId="14" captionText="FIGURE 9. A – B Nanohystrix hammerae n. sp., adult female. A, several of the large setae that are abbreviated in B and in Fig. 2 B; B, ventral aspect, legs and some setae only partially shown. Abbreviations: ips (lyrifissure); other labels indicate setae (see text). Scale bar: 100 Μm." httpUri="https://zenodo.org/record/238773/files/figure.png" pageId="5" pageNumber="156">9B</figureCitation>
|
||
):
|
||
<emphasis id="B964EABDFFE5FFDCFE7BFB3C3080F3E0" box="[475,501,1152,1176]" italics="true" pageId="5" pageNumber="156">c3</emphasis>
|
||
high on pleural surface, aligned with setal row
|
||
<emphasis id="B964EABDFFE5FFDCFBABFBC3356CF3E0" box="[1035,1049,1151,1176]" italics="true" pageId="5" pageNumber="156">d</emphasis>
|
||
and immediately anterior to
|
||
<emphasis id="B964EABDFFE5FFDCFAFDFB3D340DF3E0" box="[1373,1400,1153,1176]" italics="true" pageId="5" pageNumber="156">cp</emphasis>
|
||
;
|
||
<emphasis id="B964EABDFFE5FFDCFA26FB3C34EEF3E0" box="[1414,1435,1152,1176]" italics="true" pageId="5" pageNumber="156">f1</emphasis>
|
||
somewhat displaced laterally;
|
||
<emphasis id="B964EABDFFE5FFDCFE4BFB1D3372F3C2" box="[491,519,1185,1210]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
uniquely incorporated as third erectile setal pair in middle transverse scissure;
|
||
<emphasis id="B964EABDFFE5FFDCFA20FB1D34E9F3C2" box="[1408,1436,1185,1210]" italics="true" pageId="5" pageNumber="156">h3</emphasis>
|
||
more anterior and dorsal than usual in oribatid mites;
|
||
<emphasis id="B964EABDFFE5FFDCFD50FB7F327EF3A3" box="[752,779,1219,1243]" italics="true" pageId="5" pageNumber="156">p2</emphasis>
|
||
dorsoventrally aligned with
|
||
<emphasis id="B964EABDFFE5FFDCFBEEFB7F351FF3A3" box="[1102,1130,1219,1243]" italics="true" pageId="5" pageNumber="156">p3</emphasis>
|
||
or even anterior to it.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE5FFDCFF67FB583008F2BC" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="156">
|
||
Typical five pairs of notogastral proprioceptors present, but in unusual locations (
|
||
<figureCitation id="132B2A2AFFE5FFDCFBF5FB5835DAF384" box="[1109,1199,1252,1277]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="5" pageNumber="156">Fig. 3A</figureCitation>
|
||
); each minute, ca. 6– 7 µm, cupular in form, round or slightly elliptical when seen face-on (
|
||
<figureCitation id="132B2A2AFFE5FFDCFC3EFAB93570F266" box="[926,1029,1285,1310]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="5" pageNumber="156">Fig. 11G</figureCitation>
|
||
). Two pairs located within scissures:
|
||
<emphasis id="B964EABDFFE5FFDCFF37FA9B31C6F246" box="[151,179,1319,1342]" italics="true" pageId="5" pageNumber="156">im</emphasis>
|
||
in pale or intermediately pigmented cuticle at lateral extreme of anterior scissure, closely posterodorsal to seta
|
||
<emphasis id="B964EABDFFE5FFDCFF37FAF531C7F218" box="[151,178,1353,1376]" italics="true" pageId="5" pageNumber="156">cp</emphasis>
|
||
;
|
||
<emphasis id="B964EABDFFE5FFDCFF63FAF531ACF218" box="[195,217,1353,1376]" italics="true" pageId="5" pageNumber="156">ip</emphasis>
|
||
within middle scissure, between setae
|
||
<emphasis id="B964EABDFFE5FFDCFD3EFAF433C6F218" box="[670,691,1352,1376]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFD50FAFB3278F218" box="[752,781,1351,1376]" italics="true" pageId="5" pageNumber="156">h2</emphasis>
|
||
, anterior to lateral extension of
|
||
<emphasis id="B964EABDFFE5FFDCFB24FAF435ECF218" box="[1156,1177,1352,1376]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
. Three pairs in lateral region:
|
||
<emphasis id="B964EABDFFE5FFDCFF4DFAD63076F2F9" box="[237,259,1386,1409]" italics="true" pageId="5" pageNumber="156">ia</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFE9AFAD63023F2F9" box="[314,342,1386,1409]" italics="true" pageId="5" pageNumber="156">im</emphasis>
|
||
on sclerotized cuticle ca. 30–50 µm from ventral edge of notogaster,
|
||
<emphasis id="B964EABDFFE5FFDCFBC1FAD63502F2F9" box="[1121,1143,1386,1409]" italics="true" pageId="5" pageNumber="156">ia</emphasis>
|
||
about at level of
|
||
<emphasis id="B964EABDFFE5FFDCFA9BFAD63420F2F9" box="[1339,1365,1386,1409]" italics="true" pageId="5" pageNumber="156">cp</emphasis>
|
||
or
|
||
<emphasis id="B964EABDFFE5FFDCFADAFAD634E1F2FA" box="[1402,1428,1386,1410]" italics="true" pageId="5" pageNumber="156">c3</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE5FFDCFF37FA3631D8F2DB" box="[151,173,1418,1443]" italics="true" pageId="5" pageNumber="156">ih</emphasis>
|
||
at level of middle scissure;
|
||
<emphasis id="B964EABDFFE5FFDCFE43FA303371F2DB" box="[483,516,1420,1443]" italics="true" pageId="5" pageNumber="156">ips</emphasis>
|
||
in pale cuticle near anterior end of separation between main notogaster and adanal region (
|
||
<figureCitation id="132B2A2AFFE5FFDCFF4EFA103005F2BC" box="[238,368,1452,1477]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="5" pageNumber="156">Fig. 11F, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE5FFDCFF67FA713441F06A" blockId="5.[151,1437,151,1943]" pageId="5" pageNumber="156">
|
||
Five pairs of conspicuous cuticular disks present, two large, three small (
|
||
<figureCitation id="132B2A2AFFE5FFDCFB9CFA7135EBF29E" box="[1084,1182,1485,1510]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="5" pageNumber="156">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFB10FA7235A0F29E" box="[1200,1237,1486,1510]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="5" pageNumber="156">3A</figureCitation>
|
||
; see Remark 2), discernable in both transmitted and reflected light by lack of color (
|
||
<figureCitation id="132B2A2AFFE5FFDCFC3BFA52328CF17F" box="[923,1017,1518,1543]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="5" pageNumber="156">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFBA7FA533538F17F" box="[1031,1101,1519,1543]" captionStart="FIGURE 8. A – E" captionStartId="13.[151,250,1220,1242]" captionTargetBox="[151,1436,193,1198]" captionTargetId="figure@13.[151,1436,193,1198]" captionTargetPageId="13" captionText="FIGURE 8. A – E Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (A, C uncleared). A, Anterior portion of notogaster dissected from body (midline indicated by thin black line; prodorsal retractor muscles torn anteriorly) under strong DIC illumination to show muscles; B, left cuticular disk cd 4 and pale extension from middle scissure (posterodorsal view, anterior to right); C, region of cuticular disk cd 3 showing tendon insertion files and proximal parts of thin tendons (lateral aspect, anterior to right); D, portion of seta f 1, transmitted bright-field illumination; E, same, but with polarized light. A – C composites of 5 - 15 layers. Abbreviations: cd 1, cd 3 (cuticular disks), e 1. mu, f 1. mu (muscles acting on respective setae), pr. mu (dorsal components of prodorsal retractor muscles), ti (tendon insertions), t. sul (transverse sulcus, with concave wall appearing brown in transmitted light; other labels indicate setae (see text). Scale bars 20 Μm (A); 10 Μm (B – E)." httpUri="https://zenodo.org/record/238772/files/figure.png" pageId="5" pageNumber="156">8B, C</figureCitation>
|
||
); cuticle finely porose. Small disks all in anterodorsal region, usually slightly elliptical:
|
||
<emphasis id="B964EABDFFE5FFDCFCBDF9B33233F150" box="[797,838,1551,1576]" italics="true" pageId="5" pageNumber="156">cd1</emphasis>
|
||
smallest (ca. 15 µm length), about 30 µm posterior to seta
|
||
<emphasis id="B964EABDFFE5FFDCFF6AF98E3190F132" box="[202,229,1586,1610]" italics="true" pageId="5" pageNumber="156">c1</emphasis>
|
||
;
|
||
<emphasis id="B964EABDFFE5FFDCFF54F98D3069F132" box="[244,284,1585,1610]" italics="true" pageId="5" pageNumber="156">cd2</emphasis>
|
||
(ca. 30 µm) about 40–50 µm ventral to seta
|
||
<emphasis id="B964EABDFFE5FFDCFCB7F98E3247F132" box="[791,818,1586,1610]" italics="true" pageId="5" pageNumber="156">c2</emphasis>
|
||
;
|
||
<emphasis id="B964EABDFFE5FFDCFCE1F98D321CF132" box="[833,873,1585,1610]" italics="true" pageId="5" pageNumber="156">cd3</emphasis>
|
||
(ca. 25 µm) about 60–70 µm posterior to seta
|
||
<emphasis id="B964EABDFFE5FFDCFADAF98E34E0F132" box="[1402,1429,1586,1610]" italics="true" pageId="5" pageNumber="156">c2</emphasis>
|
||
. Larger two disks circular to slightly elliptical (ca. 55–60 µm);
|
||
<emphasis id="B964EABDFFE5FFDCFCF0F9EE320CF113" box="[848,889,1618,1643]" italics="true" pageId="5" pageNumber="156">cd4</emphasis>
|
||
located midway between erectile setae
|
||
<emphasis id="B964EABDFFE5FFDCFA94F9EF343CF113" box="[1332,1353,1619,1643]" italics="true" pageId="5" pageNumber="156">f2</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFE5FFDCFA20F9EE34EEF113" box="[1408,1435,1618,1643]" italics="true" pageId="5" pageNumber="156">h1</emphasis>
|
||
at end of pale triangular extension from middle scissure;
|
||
<emphasis id="B964EABDFFE5FFDCFCB3F9C83249F1F5" box="[787,828,1652,1677]" italics="true" pageId="5" pageNumber="156">cd5</emphasis>
|
||
immediately posterolateral to seta
|
||
<emphasis id="B964EABDFFE5FFDCFB63F9C835AAF1F5" box="[1219,1247,1652,1677]" italics="true" pageId="5" pageNumber="156">h1</emphasis>
|
||
. Two linear files of tendon insertions present anterolaterally (
|
||
<emphasis id="B964EABDFFE5FFDCFD28F92A33E2F1D5" box="[648,663,1686,1709]" italics="true" pageId="5" pageNumber="156">ti</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFE5FFDCFD07F9293277F1D6" box="[679,770,1685,1710]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="5" pageNumber="156">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFCADF92A3246F1D6" box="[781,819,1686,1710]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="5" pageNumber="156">3A</figureCitation>
|
||
). One file of 6–10 minute, uniform, darkly sclerotized, closely spaced internal projections directed posteriorly from near disk
|
||
<emphasis id="B964EABDFFE5FFDCFC0FF90A32A2F1B7" box="[943,983,1718,1743]" italics="true" pageId="5" pageNumber="156">cd3</emphasis>
|
||
, each projection serving as insertion for one thin tendon (
|
||
<figureCitation id="132B2A2AFFE5FFDCFEF7F96B30C5F188" box="[343,432,1751,1776]" captionStart="FIGURE 8. A – E" captionStartId="13.[151,250,1220,1242]" captionTargetBox="[151,1436,193,1198]" captionTargetId="figure@13.[151,1436,193,1198]" captionTargetPageId="13" captionText="FIGURE 8. A – E Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (A, C uncleared). A, Anterior portion of notogaster dissected from body (midline indicated by thin black line; prodorsal retractor muscles torn anteriorly) under strong DIC illumination to show muscles; B, left cuticular disk cd 4 and pale extension from middle scissure (posterodorsal view, anterior to right); C, region of cuticular disk cd 3 showing tendon insertion files and proximal parts of thin tendons (lateral aspect, anterior to right); D, portion of seta f 1, transmitted bright-field illumination; E, same, but with polarized light. A – C composites of 5 - 15 layers. Abbreviations: cd 1, cd 3 (cuticular disks), e 1. mu, f 1. mu (muscles acting on respective setae), pr. mu (dorsal components of prodorsal retractor muscles), ti (tendon insertions), t. sul (transverse sulcus, with concave wall appearing brown in transmitted light; other labels indicate setae (see text). Scale bars 20 Μm (A); 10 Μm (B – E)." httpUri="https://zenodo.org/record/238772/files/figure.png" pageId="5" pageNumber="156">Fig. 8C</figureCitation>
|
||
). Second, less distinct row of insertions, perpendicular to first, directed dorsomedially in transverse sulcus behind seta
|
||
<emphasis id="B964EABDFFE5FFDCFE42F945308BF06A" box="[482,510,1785,1810]" italics="true" pageId="5" pageNumber="156">d2</emphasis>
|
||
; only most lateral few insertions of second row with internal projections.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFE5FFD4FF67F8A03294F06E" blockId="5.[151,1437,151,1943]" lastBlockId="13.[151,1437,1501,1994]" lastPageId="13" lastPageNumber="164" pageId="5" pageNumber="156">
|
||
<emphasis id="B964EABDFFE5FFDCFF67F8A03065F04B" box="[199,272,1820,1843]" italics="true" pageId="5" pageNumber="156">Venter.</emphasis>
|
||
Coxisternum comprising about one-third of ventral surface (
|
||
<figureCitation id="132B2A2AFFE5FFDCFC60F8A63568F04B" box="[960,1053,1818,1843]" captionStart="FIGURE 9. A – B" captionStartId="14.[151,250,1638,1660]" captionTargetBox="[155,1431,192,1610]" captionTargetId="figure@14.[151,1436,181,1617]" captionTargetPageId="14" captionText="FIGURE 9. A – B Nanohystrix hammerae n. sp., adult female. A, several of the large setae that are abbreviated in B and in Fig. 2 B; B, ventral aspect, legs and some setae only partially shown. Abbreviations: ips (lyrifissure); other labels indicate setae (see text). Scale bar: 100 Μm." httpUri="https://zenodo.org/record/238773/files/figure.png" pageId="5" pageNumber="156">Figs 9B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFE5FFDCFB89F8A73509F04B" box="[1065,1148,1819,1843]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="5" pageNumber="156">10B, C</figureCitation>
|
||
); nearly quadrate overall, with trochanter of each leg articulating at about same distance from midline. Muscle sigilla weakly defined in transmitted light. Paired plates of epimeres I and II strongly convex and independent both from other member of pair and from adjacent epimeral plates; soft cuticle medially separating plates of epimere I very broad, that of epimere II very narrow (
|
||
<figureCitation id="132B2A2AFFEDFFD4FE10FA61336CF28E" box="[432,537,1501,1526]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="13" pageNumber="164">Fig. 11A</figureCitation>
|
||
). By contrast, epimeres III and IV without soft medial cuticle (but with line of cuticular weakness visible in transmitted light;
|
||
<figureCitation id="132B2A2AFFEDFFD4FD1FF9BD326CF162" box="[703,793,1537,1562]" captionStart="FIGURE 9. A – B" captionStartId="14.[151,250,1638,1660]" captionTargetBox="[155,1431,192,1610]" captionTargetId="figure@14.[151,1436,181,1617]" captionTargetPageId="14" captionText="FIGURE 9. A – B Nanohystrix hammerae n. sp., adult female. A, several of the large setae that are abbreviated in B and in Fig. 2 B; B, ventral aspect, legs and some setae only partially shown. Abbreviations: ips (lyrifissure); other labels indicate setae (see text). Scale bar: 100 Μm." httpUri="https://zenodo.org/record/238773/files/figure.png" pageId="13" pageNumber="164">Fig. 9B</figureCitation>
|
||
); epimeres fused to each other in medial half, forming single unit. Central fused region of unit relatively flat, but lateral half of each epimere discrete, cylindrical. Epimere III articulating with epimere II through narrow ventrosejugal articulation having weak transverse striation; posterior edge of epimere IV strongly curved, sharply defined, slightly overhanging genital plates and hiding broad articulation. Without coxisternal apodemes; epimere IV laterally with thickened internal ridge supporting articulation with leg trochanter. Coxisternal setation 3-1-3-4; no variation noted. Setae
|
||
<emphasis id="B964EABDFFEDFFD4FBD1F90A35FBF1B6" box="[1137,1166,1718,1742]" italics="true" pageId="13" pageNumber="164">1a</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFEDFFD4FB3CF90A35CDF1B6" box="[1180,1208,1718,1742]" italics="true" pageId="13" pageNumber="164">2a</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFEDFFD4FB68F90A3591F1B6" box="[1224,1252,1718,1742]" italics="true" pageId="13" pageNumber="164">3a</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFEDFFD4FABCF90A344DF1B6" box="[1308,1336,1718,1742]" italics="true" pageId="13" pageNumber="164">4a</emphasis>
|
||
straight, tapered only distally, conspicuously barbed (
|
||
<figureCitation id="132B2A2AFFEDFFD4FD39F9653277F18A" box="[665,770,1753,1778]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="13" pageNumber="164">Fig. 10E</figureCitation>
|
||
); others attenuate to various degrees, with fewer barbs or nearly glabrous. Setae range in size from about 85 Μm (
|
||
<emphasis id="B964EABDFFEDFFD4FCA5F9423254F06E" box="[773,801,1790,1814]" italics="true" pageId="13" pageNumber="164">3a</emphasis>
|
||
) to 150 Μm (
|
||
<emphasis id="B964EABDFFEDFFD4FC14F94132A4F06E" box="[948,977,1789,1814]" italics="true" pageId="13" pageNumber="164">1b</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6F6627FFE6FFDFFF37F8823571F095" httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="6" pageNumber="157" targetBox="[164,1407,195,1812]" targetPageId="6">
|
||
<paragraph id="8BAF36AFFFE6FFDFFF37F8823571F095" blockId="6.[151,1436,1854,2029]" pageId="6" pageNumber="157">
|
||
<emphasis id="B964EABDFFE6FFDFFF37F8823022F02B" bold="true" box="[151,343,1854,1876]" pageId="6" pageNumber="157">FIGURE 1. A–H</emphasis>
|
||
<taxonomicName id="4C104D2CFFE6FFDFFEC4F882332DF02C" box="[356,600,1854,1876]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFE6FFDFFEC4F882332DF02C" box="[356,600,1854,1876]" italics="true" pageId="6" pageNumber="157">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFE6FFDFFDC4F88333D6F02C" box="[612,675,1855,1876]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="B964EABDFFE6FFDFFDC4F88333D6F02C" bold="true" box="[612,675,1855,1876]" pageId="6" pageNumber="157">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations:
|
||
<emphasis id="B964EABDFFE6FFDFFB20F80535DDF0B7" box="[1152,1192,1977,1999]" italics="true" pageId="6" pageNumber="157">h.cr</emphasis>
|
||
(humeral crease); other labels indicate setae (see text). All composites of 10–20 layers. Scale bars: all 100 Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFE7FFDEFF37F887356AF092" httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="7" pageNumber="158" targetBox="[161,1425,198,1839]" targetPageId="7">
|
||
<paragraph id="8BAF36AFFFE7FFDEFF37F887356AF092" blockId="7.[151,1437,1851,2026]" pageId="7" pageNumber="158">
|
||
<emphasis id="B964EABDFFE7FFDEFF37F887303EF028" bold="true" box="[151,331,1851,1873]" pageId="7" pageNumber="158">FIGURE 2. A–E</emphasis>
|
||
<taxonomicName id="4C104D2CFFE7FFDEFEF2F8873335F029" box="[338,576,1851,1873]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFE7FFDEFEF2F8873335F029" box="[338,576,1851,1873]" italics="true" pageId="7" pageNumber="158">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFE7FFDEFDE7F88033F4F029" box="[583,641,1852,1873]" pageId="7" pageNumber="158" rank="species">
|
||
<emphasis id="B964EABDFFE7FFDEFDE7F88033F4F029" bold="true" box="[583,641,1852,1873]" pageId="7" pageNumber="158">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk
|
||
<emphasis id="B964EABDFFE7FFDEFD28F82B33D8F0D5" box="[648,685,1943,1965]" italics="true" pageId="7" pageNumber="158">cd4</emphasis>
|
||
; E, entire posterior scissure and cuticular disks
|
||
<emphasis id="B964EABDFFE7FFDEFB2DF82B35C7F0D5" box="[1165,1202,1943,1965]" italics="true" pageId="7" pageNumber="158">cd5</emphasis>
|
||
(posterodorsal aspect). Abbreviations:
|
||
<emphasis id="B964EABDFFE7FFDEFE94F80A302FF0B4" box="[308,346,1974,1996]" italics="true" pageId="7" pageNumber="158">bng</emphasis>
|
||
(border of notogaster),
|
||
<emphasis id="B964EABDFFE7FFDEFDEFF80A33F8F0B4" box="[591,653,1974,1996]" italics="true" pageId="7" pageNumber="158">cd1–5</emphasis>
|
||
(cuticular disks),
|
||
<emphasis id="B964EABDFFE7FFDEFCE7F80B322EF0B4" box="[839,859,1975,1996]" italics="true" pageId="7" pageNumber="158">ia</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE7FFDEFCC8F80B32F4F0B4" box="[872,897,1975,1996]" italics="true" pageId="7" pageNumber="158">im</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE7FFDEFC2FF80B32D6F0B4" box="[911,931,1975,1996]" italics="true" pageId="7" pageNumber="158">ip</emphasis>
|
||
(lyrifissures),
|
||
<emphasis id="B964EABDFFE7FFDEFB99F80B3525F0B4" box="[1081,1104,1975,1996]" italics="true" pageId="7" pageNumber="158">nc</emphasis>
|
||
(nuchal constriction),
|
||
<emphasis id="B964EABDFFE7FFDEFA99F80B3432F0B4" box="[1337,1351,1975,1996]" italics="true" pageId="7" pageNumber="158">ti</emphasis>
|
||
(tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C–E to same scale).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFE8FFD1FF37F8FE3302F089" httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="8" pageNumber="159" targetBox="[193,1407,191,1840]" targetPageId="8">
|
||
<paragraph id="8BAF36AFFFE8FFD1FF37F8FE3302F089" blockId="8.[151,1436,1858,2033]" pageId="8" pageNumber="159">
|
||
<emphasis id="B964EABDFFE8FFD1FF37F8FE3022F02F" bold="true" box="[151,343,1858,1880]" pageId="8" pageNumber="159">FIGURE 3. A–D</emphasis>
|
||
<taxonomicName id="4C104D2CFFE8FFD1FEC3F8FE3322F020" box="[355,599,1858,1880]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFE8FFD1FEC3F8FE3322F020" box="[355,599,1858,1880]" italics="true" pageId="8" pageNumber="159">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFE8FFD1FDC3F8FE33D7F02F" box="[611,674,1858,1879]" pageId="8" pageNumber="159" rank="species">
|
||
<emphasis id="B964EABDFFE8FFD1FDC3F8FE33D7F02F" bold="true" box="[611,674,1858,1879]" pageId="8" pageNumber="159">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations:
|
||
<emphasis id="B964EABDFFE8FFD1FCB2F8223243F0CC" box="[786,822,1950,1972]" italics="true" pageId="8" pageNumber="159">ca1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE8FFD1FCEDF8223207F0CC" box="[845,882,1950,1972]" italics="true" pageId="8" pageNumber="159">ca2</emphasis>
|
||
(prodorsal carinae),
|
||
<emphasis id="B964EABDFFE8FFD1FBC4F82235D7F0CC" box="[1124,1186,1950,1972]" italics="true" pageId="8" pageNumber="159">cd1–5</emphasis>
|
||
(cuticular disks),
|
||
<emphasis id="B964EABDFFE8FFD1FAD9F82334E9F0CC" box="[1401,1436,1951,1972]" italics="true" pageId="8" pageNumber="159">cpc</emphasis>
|
||
(podocephalic canal),
|
||
<emphasis id="B964EABDFFE8FFD1FEDCF80230E5F0AB" box="[380,400,1982,2003]" italics="true" pageId="8" pageNumber="159">ia</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE8FFD1FE3EF80230C2F0AB" box="[414,439,1982,2003]" italics="true" pageId="8" pageNumber="159">im</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFE8FFD1FE67F80230AEF0AB" box="[455,475,1982,2003]" italics="true" pageId="8" pageNumber="159">ip</emphasis>
|
||
(lyrifissures),
|
||
<emphasis id="B964EABDFFE8FFD1FDD3F80233FEF0AB" box="[627,651,1982,2003]" italics="true" pageId="8" pageNumber="159">nc</emphasis>
|
||
(nuchal constriction),
|
||
<emphasis id="B964EABDFFE8FFD1FCD7F80232F0F0AB" box="[887,901,1982,2003]" italics="true" pageId="8" pageNumber="159">ti</emphasis>
|
||
(tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFE9FFD0FF37F94A30EEF0BC" httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="9" pageNumber="160" targetBox="[151,1436,193,1759]" targetPageId="9">
|
||
<paragraph id="8BAF36AFFFE9FFD0FF37F94A30EEF0BC" blockId="9.[151,1437,1782,1988]" pageId="9" pageNumber="160">
|
||
<emphasis id="B964EABDFFE9FFD0FF37F94A303EF074" bold="true" box="[151,331,1782,1804]" pageId="9" pageNumber="160">FIGURE 4. A–G</emphasis>
|
||
<taxonomicName id="4C104D2CFFE9FFD0FEF1F94A334BF074" box="[337,574,1782,1804]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFE9FFD0FEF1F94A334BF074" box="[337,574,1782,1804]" italics="true" pageId="9" pageNumber="160">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFE9FFD0FDE4F94A3309F073" box="[580,636,1782,1803]" pageId="9" pageNumber="160" rank="species">
|
||
<emphasis id="B964EABDFFE9FFD0FDE4F94A3309F073" bold="true" box="[580,636,1782,1803]" pageId="9" pageNumber="160">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk
|
||
<emphasis id="B964EABDFFE9FFD0FBB2F88F3542F031" box="[1042,1079,1843,1865]" italics="true" pageId="9" pageNumber="160">cd1</emphasis>
|
||
, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks
|
||
<emphasis id="B964EABDFFE9FFD0FE3EF8CC30B6F0FE" box="[414,451,1904,1926]" italics="true" pageId="9" pageNumber="160">cd5</emphasis>
|
||
(insert is enlargement of right disk
|
||
<emphasis id="B964EABDFFE9FFD0FC92F8CC3222F0FE" box="[818,855,1904,1926]" italics="true" pageId="9" pageNumber="160">cd5</emphasis>
|
||
showing exudate); E, base of seta
|
||
<emphasis id="B964EABDFFE9FFD0FB1CF8CC35A1F0FE" box="[1212,1236,1904,1926]" italics="true" pageId="9" pageNumber="160">e2</emphasis>
|
||
; F, basal portion of seta
|
||
<emphasis id="B964EABDFFE9FFD0FF65F83331ABF0DD" box="[197,222,1935,1957]" italics="true" pageId="9" pageNumber="160">d1</emphasis>
|
||
; G, basal portion of seta
|
||
<emphasis id="B964EABDFFE9FFD0FE42F8333083F0DD" box="[482,502,1935,1957]" italics="true" pageId="9" pageNumber="160">f1</emphasis>
|
||
. Abbreviations;
|
||
<emphasis id="B964EABDFFE9FFD0FD01F83333BCF0DD" box="[673,713,1935,1957]" italics="true" pageId="9" pageNumber="160">h.cr</emphasis>
|
||
(humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A–D); 10 Μm (E–G).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFEAFFD3FF37F9EF3537F0C2" httpUri="https://zenodo.org/record/238769/files/figure.png" pageId="10" pageNumber="161" targetBox="[151,1436,193,1596]" targetPageId="10">
|
||
<paragraph id="8BAF36AFFFEAFFD3FF37F9EF3537F0C2" blockId="10.[151,1436,1619,1978]" pageId="10" pageNumber="161">
|
||
<emphasis id="B964EABDFFEAFFD3FF37F9EF303AF111" bold="true" box="[151,335,1619,1641]" pageId="10" pageNumber="161">FIGURE 5. A–G</emphasis>
|
||
<taxonomicName id="4C104D2CFFEAFFD3FEF6F9EF3330F111" box="[342,581,1619,1641]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFEAFFD3FEF6F9EF3330F111" box="[342,581,1619,1641]" italics="true" pageId="10" pageNumber="161">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFEAFFD3FDEDF9E833FDF111" box="[589,648,1620,1641]" pageId="10" pageNumber="161" rank="species">
|
||
<emphasis id="B964EABDFFEAFFD3FDEDF9E833FDF111" bold="true" box="[589,648,1620,1641]" pageId="10" pageNumber="161">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (
|
||
<emphasis id="B964EABDFFEAFFD3FBD9F97335C7F19C" box="[1145,1202,1743,1764]" italics="true" pageId="10" pageNumber="161">pr.mu</emphasis>
|
||
; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (
|
||
<emphasis id="B964EABDFFEAFFD3FE18F8B03083F059" box="[440,502,1804,1825]" italics="true" pageId="10" pageNumber="161">po.mu</emphasis>
|
||
) and deeper plane (right) with cuticular disk
|
||
<emphasis id="B964EABDFFEAFFD3FC71F8B73283F059" box="[977,1014,1803,1825]" italics="true" pageId="10" pageNumber="161">cd5</emphasis>
|
||
(with underlying fibers,
|
||
<emphasis id="B964EABDFFEAFFD3FB5AF8B73460F059" box="[1274,1301,1803,1825]" italics="true" pageId="10" pageNumber="161">fib</emphasis>
|
||
) and seta
|
||
<emphasis id="B964EABDFFEAFFD3FA23F8B734E9F059" box="[1411,1436,1803,1825]" italics="true" pageId="10" pageNumber="161">h1</emphasis>
|
||
(with depressor muscle,
|
||
<emphasis id="B964EABDFFEAFFD3FE34F89630A7F038" box="[404,466,1834,1856]" italics="true" pageId="10" pageNumber="161">h1.mu</emphasis>
|
||
); F, section of middle scissure (more lateral than indicated in A) showing base of seta
|
||
<emphasis id="B964EABDFFEAFFD3FAFCF8963405F038" box="[1372,1392,1834,1856]" italics="true" pageId="10" pageNumber="161">f1</emphasis>
|
||
and associated depressor muscle (
|
||
<emphasis id="B964EABDFFEAFFD3FE62F8F4308FF026" box="[450,506,1864,1886]" italics="true" pageId="10" pageNumber="161">f1.mu</emphasis>
|
||
); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations:
|
||
<emphasis id="B964EABDFFEAFFD3FDE1F8DB3312F005" box="[577,615,1895,1917]" italics="true" pageId="10" pageNumber="161">bng</emphasis>
|
||
(anterior border of notogaster),
|
||
<emphasis id="B964EABDFFEAFFD3FC0FF8D43292F005" box="[943,999,1896,1917]" italics="true" pageId="10" pageNumber="161">cr.mu</emphasis>
|
||
(cheliceral retractor muscles);
|
||
<emphasis id="B964EABDFFEAFFD3FA83F8D4344AF005" box="[1315,1343,1896,1917]" italics="true" pageId="10" pageNumber="161">cw</emphasis>
|
||
(caecum wall),
|
||
<emphasis id="B964EABDFFEAFFD3FF7DF83A3069F0E4" box="[221,284,1926,1948]" italics="true" pageId="10" pageNumber="161">ep1–3</emphasis>
|
||
(plates of epimeres 1–3),
|
||
<emphasis id="B964EABDFFEAFFD3FDE2F83B332FF0E4" box="[578,602,1927,1948]" italics="true" pageId="10" pageNumber="161">nc</emphasis>
|
||
(nuchal constriction),
|
||
<emphasis id="B964EABDFFEAFFD3FCF1F83B3201F0E4" box="[849,884,1927,1948]" italics="true" pageId="10" pageNumber="161">t.gr</emphasis>
|
||
(transverse groove),
|
||
<emphasis id="B964EABDFFEAFFD3FBFEF83B350FF0E4" box="[1118,1146,1927,1948]" italics="true" pageId="10" pageNumber="161">sej</emphasis>
|
||
(sejugal articulation),
|
||
<emphasis id="B964EABDFFEAFFD3FAD1F83A34E9F0E4" box="[1393,1436,1926,1948]" italics="true" pageId="10" pageNumber="161">t.sul</emphasis>
|
||
(transverse sulcus). All composites of 5–20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C–G).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFEBFFD2FF37FA4D35EAF184" httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="11" pageNumber="162" targetBox="[151,1436,193,1498]" targetPageId="11">
|
||
<paragraph id="8BAF36AFFFEBFFD2FF37FA4D35EAF184" blockId="11.[151,1436,1521,1788]" pageId="11" pageNumber="162">
|
||
<emphasis id="B964EABDFFEBFFD2FF37FA4D3027F17F" bold="true" box="[151,338,1521,1543]" pageId="11" pageNumber="162">FIGURE 6. A–G</emphasis>
|
||
<taxonomicName id="4C104D2CFFEBFFD2FEFCFA4D3338F17F" box="[348,589,1521,1543]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFEBFFD2FEFCFA4D3338F17F" box="[348,589,1521,1543]" italics="true" pageId="11" pageNumber="162">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFEBFFD2FDF7FA4D33E6F17E" box="[599,659,1521,1542]" pageId="11" pageNumber="162" rank="species">
|
||
<emphasis id="B964EABDFFEBFFD2FDF7FA4D33E6F17E" bold="true" box="[599,659,1521,1542]" pageId="11" pageNumber="162">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left)—arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I–III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations:
|
||
<emphasis id="B964EABDFFEBFFD2FC09F93632BBF1D8" box="[937,974,1674,1696]" italics="true" pageId="11" pageNumber="162">ca1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFEBFFD2FC7EF9363577F1D8" box="[990,1026,1674,1696]" italics="true" pageId="11" pageNumber="162">ca2</emphasis>
|
||
(prodorsal carinae),
|
||
<emphasis id="B964EABDFFEBFFD2FB7FF9373477F1D8" box="[1247,1282,1675,1696]" italics="true" pageId="11" pageNumber="162">cpc</emphasis>
|
||
(podocephalic canal),
|
||
<emphasis id="B964EABDFFEBFFD2FF42F9153075F1C7" box="[226,256,1705,1727]" italics="true" pageId="11" pageNumber="162">Ch</emphasis>
|
||
(chelicera),
|
||
<emphasis id="B964EABDFFEBFFD2FE20F91530EDF1C7" box="[384,408,1705,1727]" italics="true" pageId="11" pageNumber="162">dc</emphasis>
|
||
(dorsal carina of chelicera),
|
||
<emphasis id="B964EABDFFEBFFD2FD1DF915338EF1C7" box="[701,763,1705,1727]" italics="true" pageId="11" pageNumber="162">ep1–3</emphasis>
|
||
(plates of epimeres 1–3),
|
||
<emphasis id="B964EABDFFEBFFD2FBA6F9153561F1C7" box="[1030,1044,1705,1727]" italics="true" pageId="11" pageNumber="162">fl</emphasis>
|
||
(flange of bothridial wall),
|
||
<emphasis id="B964EABDFFEBFFD2FA90F9163438F1C7" box="[1328,1357,1706,1727]" italics="true" pageId="11" pageNumber="162">Ge</emphasis>
|
||
(gena),
|
||
<emphasis id="B964EABDFFEBFFD2FF37F97531DAF1A6" box="[151,175,1737,1758]" italics="true" pageId="11" pageNumber="162">Ls</emphasis>
|
||
(labrum),
|
||
<emphasis id="B964EABDFFEBFFD2FEBFF9753043F1A6" box="[287,310,1737,1758]" italics="true" pageId="11" pageNumber="162">nc</emphasis>
|
||
(nuchal constriction),
|
||
<emphasis id="B964EABDFFEBFFD2FD84F97B333CF1A5" box="[548,585,1735,1757]" italics="true" pageId="11" pageNumber="162">oc.l</emphasis>
|
||
(lateral eye),
|
||
<emphasis id="B964EABDFFEBFFD2FD7DF97B3381F1A5" box="[733,756,1735,1757]" italics="true" pageId="11" pageNumber="162">rb</emphasis>
|
||
(rostral bulge),
|
||
<emphasis id="B964EABDFFEBFFD2FC3FF97532CEF1A6" box="[927,955,1737,1758]" italics="true" pageId="11" pageNumber="162">rm</emphasis>
|
||
(rostral margin),
|
||
<emphasis id="B964EABDFFEBFFD2FBD4F97B35D3F1A5" box="[1140,1190,1735,1757]" italics="true" pageId="11" pageNumber="162">sub.t</emphasis>
|
||
(subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFECFFD5FF00F9A433DFF019" httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="12" pageNumber="163" targetBox="[151,1436,193,1537]" targetPageId="12">
|
||
<paragraph id="8BAF36AFFFECFFD5FF00F9A433DFF019" blockId="12.[151,1436,1560,1889]" pageId="12" pageNumber="163">
|
||
<emphasis id="B964EABDFFECFFD5FF00F9A43020F155" bold="true" box="[160,341,1560,1582]" pageId="12" pageNumber="163">FIGURE 7. A–J</emphasis>
|
||
<taxonomicName id="4C104D2CFFECFFD5FEFEF9A4333AF156" box="[350,591,1560,1582]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFECFFD5FEFEF9A4333AF156" box="[350,591,1560,1582]" italics="true" pageId="12" pageNumber="163">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFECFFD5FDF8F9A533E0F156" box="[600,661,1561,1582]" pageId="12" pageNumber="163" rank="species">
|
||
<emphasis id="B964EABDFFECFFD5FDF8F9A533E0F156" bold="true" box="[600,661,1561,1582]" pageId="12" pageNumber="163">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5–20 layers. Abbreviations:
|
||
<emphasis id="B964EABDFFECFFD5FE21F95330EFF07D" box="[385,410,1775,1797]" italics="true" pageId="12" pageNumber="163">bo</emphasis>
|
||
(bothridium),
|
||
<emphasis id="B964EABDFFECFFD5FD92F9533333F07D" box="[562,582,1775,1797]" italics="true" pageId="12" pageNumber="163">bt</emphasis>
|
||
(base of rostral tectum),
|
||
<emphasis id="B964EABDFFECFFD5FCEEF9533206F07D" box="[846,883,1775,1797]" italics="true" pageId="12" pageNumber="163">ca1</emphasis>
|
||
(prodorsal carina),
|
||
<emphasis id="B964EABDFFECFFD5FBE0F953352BF07D" box="[1088,1118,1775,1797]" italics="true" pageId="12" pageNumber="163">Ch</emphasis>
|
||
(chelicera),
|
||
<emphasis id="B964EABDFFECFFD5FB41F94C3471F07D" box="[1249,1284,1776,1797]" italics="true" pageId="12" pageNumber="163">cpc</emphasis>
|
||
(podocephalic canal),
|
||
<emphasis id="B964EABDFFECFFD5FF42F8B13072F05B" box="[226,263,1805,1827]" italics="true" pageId="12" pageNumber="163">oc.l</emphasis>
|
||
(lateral eye),
|
||
<emphasis id="B964EABDFFECFFD5FE36F8B330B2F05C" box="[406,455,1807,1828]" italics="true" pageId="12" pageNumber="163">oc.m</emphasis>
|
||
(median eye),
|
||
<emphasis id="B964EABDFFECFFD5FDC1F8B133EDF05B" box="[609,664,1805,1827]" italics="true" pageId="12" pageNumber="163">mu.sl</emphasis>
|
||
(subcapitular levator muscle),
|
||
<emphasis id="B964EABDFFECFFD5FC75F8B13299F05B" box="[981,1004,1805,1827]" italics="true" pageId="12" pageNumber="163">rb</emphasis>
|
||
(rostral bulge),
|
||
<emphasis id="B964EABDFFECFFD5FB31F8B335DAF05C" box="[1169,1199,1807,1828]" italics="true" pageId="12" pageNumber="163">pK</emphasis>
|
||
(prodorsal condyle),
|
||
<emphasis id="B964EABDFFECFFD5FA2BF8B334E9F05C" box="[1419,1436,1807,1828]" italics="true" pageId="12" pageNumber="163">st</emphasis>
|
||
(constriction of tectum depth),
|
||
<emphasis id="B964EABDFFECFFD5FE77F8913087F03A" box="[471,498,1837,1858]" italics="true" pageId="12" pageNumber="163">str</emphasis>
|
||
(supporting strut),
|
||
<emphasis id="B964EABDFFECFFD5FD1AF890339EF03A" box="[698,747,1836,1858]" italics="true" pageId="12" pageNumber="163">sub.t</emphasis>
|
||
(subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G–J); 10 Μm (B); 5 Μm (C, E, F).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFEDFFD4FF37FB783440F2C9" httpUri="https://zenodo.org/record/238772/files/figure.png" pageId="13" pageNumber="164" targetBox="[151,1436,193,1198]" targetPageId="13">
|
||
<paragraph id="8BAF36AFFFEDFFD4FF37FB783440F2C9" blockId="13.[151,1436,1220,1457]" pageId="13" pageNumber="164">
|
||
<emphasis id="B964EABDFFEDFFD4FF37FB78303CF3A1" bold="true" box="[151,329,1220,1242]" pageId="13" pageNumber="164">FIGURE 8. A–E</emphasis>
|
||
<taxonomicName id="4C104D2CFFEDFFD4FEEFFB783349F3A2" box="[335,572,1220,1242]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFEDFFD4FEEFFB783349F3A2" box="[335,572,1220,1242]" italics="true" pageId="13" pageNumber="164">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFEDFFD4FDE2FB78330EF3A1" box="[578,635,1220,1241]" pageId="13" pageNumber="164" rank="species">
|
||
<emphasis id="B964EABDFFEDFFD4FDE2FB78330EF3A1" bold="true" box="[578,635,1220,1241]" pageId="13" pageNumber="164">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, transmitted light compound microscope images of adults (A, C uncleared). A, Anterior portion of notogaster dissected from body (midline indicated by thin black line; prodorsal retractor muscles torn anteriorly) under strong DIC illumination to show muscles; B, left cuticular disk
|
||
<emphasis id="B964EABDFFEDFFD4FC75FABD328FF26F" box="[981,1018,1281,1303]" italics="true" pageId="13" pageNumber="164">cd4</emphasis>
|
||
and pale extension from middle scissure (posterodorsal view, anterior to right); C, region of cuticular disk
|
||
<emphasis id="B964EABDFFEDFFD4FC90FA9C3220F24E" box="[816,853,1312,1334]" italics="true" pageId="13" pageNumber="164">cd3</emphasis>
|
||
showing tendon insertion files and proximal parts of thin tendons (lateral aspect, anterior to right); D, portion of seta
|
||
<emphasis id="B964EABDFFEDFFD4FD4EFA823277F22C" box="[750,770,1342,1364]" italics="true" pageId="13" pageNumber="164">f1</emphasis>
|
||
, transmitted bright-field illumination; E, same, but with polarized light. A–C composites of 5-15 layers. Abbreviations:
|
||
<emphasis id="B964EABDFFEDFFD4FD66FAE1326AF20B" box="[710,799,1373,1395]" italics="true" pageId="13" pageNumber="164">cd1, cd3</emphasis>
|
||
(cuticular disks),
|
||
<emphasis id="B964EABDFFEDFFD4FC7CFAE1356CF20B" box="[988,1049,1373,1395]" italics="true" pageId="13" pageNumber="164">e1.mu</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFEDFFD4FB88FAE13514F20B" box="[1064,1121,1373,1395]" italics="true" pageId="13" pageNumber="164">f1.mu</emphasis>
|
||
(muscles acting on respective setae),
|
||
<emphasis id="B964EABDFFEDFFD4FF40FAC1306CF2EA" box="[224,281,1405,1426]" italics="true" pageId="13" pageNumber="164">pr.mu</emphasis>
|
||
(dorsal components of prodorsal retractor muscles),
|
||
<emphasis id="B964EABDFFEDFFD4FC9AFAC1323DF2EA" box="[826,840,1405,1426]" italics="true" pageId="13" pageNumber="164">ti</emphasis>
|
||
(tendon insertions),
|
||
<emphasis id="B964EABDFFEDFFD4FBBDFAC0353DF2EA" box="[1053,1096,1404,1426]" italics="true" pageId="13" pageNumber="164">t.sul</emphasis>
|
||
(transverse sulcus, with concave wall appearing brown in transmitted light; other labels indicate setae (see text). Scale bars 20 Μm (A); 10 Μm (B–E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFEDFFD4FF67F89D32EEF0B2" blockId="13.[151,1437,1501,1994]" pageId="13" pageNumber="164">
|
||
Supracoxal seta
|
||
<emphasis id="B964EABDFFEDFFD4FE2CF89E30D7F041" box="[396,418,1826,1849]" italics="true" pageId="13" pageNumber="164">eI</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFEDFFD4FE18F89D3357F042" box="[440,546,1825,1850]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="13" pageNumber="164">Fig. 11B</figureCitation>
|
||
) minute, spiniform (ca. 13 Μm), apparently immobile (no clear alveolus); inconspicuously located on dorsal face of epimere I at edge of sclerotization. Anterodorsal corner of epimere II with sharp spine or cluster of up to four smaller spines (
|
||
<figureCitation id="132B2A2AFFEDFFD4FCA4F8D5321CF0FA" box="[772,873,1897,1922]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="13" pageNumber="164">Fig. 11C</figureCitation>
|
||
). Podocephalic canal (
|
||
<emphasis id="B964EABDFFEDFFD4FBC5F8D635F9F0F9" box="[1125,1164,1898,1921]" italics="true" pageId="13" pageNumber="164">cpc</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFEDFFD4FB3AF8D53580F0FA" box="[1178,1269,1897,1922]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="13" pageNumber="164">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFEDFFD4FAA1F8D63450F0FA" box="[1281,1317,1898,1922]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="13" pageNumber="164">7D</figureCitation>
|
||
) typical in form, running from supracoxal region of epimere I across soft pleural cuticle, then onto subcapitulum near articulation with prodorsum; often hidden in cuticular fold (
|
||
<figureCitation id="132B2A2AFFEDFFD4FC94F80D32FBF0B2" box="[820,910,1969,1994]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="13" pageNumber="164">Fig. 7G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6F6627FFEEFFD7FF37F9DA30E7F1C1" httpUri="https://zenodo.org/record/238773/files/figure.png" pageId="14" pageNumber="165" targetBox="[155,1431,192,1610]" targetPageId="14">
|
||
<paragraph id="8BAF36AFFFEEFFD7FF37F9DA30E7F1C1" blockId="14.[151,1436,1638,1721]" pageId="14" pageNumber="165">
|
||
<emphasis id="B964EABDFFEEFFD7FF37F9DA303DF103" bold="true" box="[151,328,1638,1660]" pageId="14" pageNumber="165">FIGURE 9. A–B</emphasis>
|
||
<taxonomicName id="4C104D2CFFEEFFD7FEEFF9DA3349F104" box="[335,572,1638,1660]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFEEFFD7FEEFF9DA3349F104" box="[335,572,1638,1660]" italics="true" pageId="14" pageNumber="165">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFEEFFD7FDE2F9DA330EF103" box="[578,635,1638,1659]" pageId="14" pageNumber="165" rank="species">
|
||
<emphasis id="B964EABDFFEEFFD7FDE2F9DA330EF103" bold="true" box="[578,635,1638,1659]" pageId="14" pageNumber="165">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female. A, several of the large setae that are abbreviated in B and in Fig. 2B; B, ventral aspect, legs and some setae only partially shown. Abbreviations:
|
||
<emphasis id="B964EABDFFEEFFD7FC1FF93A32A8F1E3" box="[959,989,1670,1691]" italics="true" pageId="14" pageNumber="165">ips</emphasis>
|
||
(lyrifissure); other labels indicate setae (see text). Scale bar: 100 Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFEEFFD7FF67F95C3347F08D" blockId="14.[151,1437,1760,2038]" pageId="14" pageNumber="165">
|
||
Genital plates about 1.2 times longer than broad (seen flat), with gently curving outer margin; pale inner margin often reflexed (
|
||
<figureCitation id="132B2A2AFFEEFFD7FE36F8B8308BF065" box="[406,510,1796,1821]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="14" pageNumber="165">Fig. 10A</figureCitation>
|
||
); cuticle connecting genital and anal plates mostly soft, sharply folded, reminiscent of shirt-collar, medially with small patch of sclerotized cuticle (not illustrated) observed only with dissection. With nine pairs of smooth, attenuate genital setae (ca. 70–85 Μm long) arranged in two longitudinal rows: six setae aligned with medial margin, anterior pair directed ventromedially and others sharply posteroventrally; three other pairs near lateral margin of plate. Aggenital region soft, without evidence of plate or aggenital setae. Cuticle of genital vestibule soft, granulate; genital papillae of simple form (ca. 50 Μm diameter), all three pairs similar (
|
||
<figureCitation id="132B2A2AFFEEFFD7FACAF80431CFF08D" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="14" pageNumber="165">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFEEFFD7FF65F861319DF08D" box="[197,232,2013,2037]" captionStart="FIGURE 9. A – B" captionStartId="14.[151,250,1638,1660]" captionTargetBox="[155,1431,192,1610]" captionTargetId="figure@14.[151,1436,181,1617]" captionTargetPageId="14" captionText="FIGURE 9. A – B Nanohystrix hammerae n. sp., adult female. A, several of the large setae that are abbreviated in B and in Fig. 2 B; B, ventral aspect, legs and some setae only partially shown. Abbreviations: ips (lyrifissure); other labels indicate setae (see text). Scale bar: 100 Μm." httpUri="https://zenodo.org/record/238773/files/figure.png" pageId="14" pageNumber="165">9B</figureCitation>
|
||
), with lightly pigmented cap.
|
||
</paragraph>
|
||
<caption id="DF6F6627FFEFFFD6FF37FA56350AF102" httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="15" pageNumber="166" targetBox="[151,1436,193,1492]" targetPageId="15">
|
||
<paragraph id="8BAF36AFFFEFFFD6FF37FA56350AF102" blockId="15.[151,1436,1514,1658]" pageId="15" pageNumber="166">
|
||
<emphasis id="B964EABDFFEFFFD6FF37FA56302EF287" bold="true" box="[151,347,1514,1536]" pageId="15" pageNumber="166">FIGURE 10. A–E</emphasis>
|
||
<taxonomicName id="4C104D2CFFEFFFD6FEC4FA563321F178" box="[356,596,1514,1536]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFEFFFD6FEC4FA563321F178" box="[356,596,1514,1536]" italics="true" pageId="15" pageNumber="166">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFEFFFD6FDFDFA5633ECF287" box="[605,665,1514,1535]" pageId="15" pageNumber="166" rank="species">
|
||
<emphasis id="B964EABDFFEFFFD6FDFDFA5633ECF287" bold="true" box="[605,665,1514,1535]" pageId="15" pageNumber="166">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta
|
||
<emphasis id="B964EABDFFEFFFD6FF5CF9D83060F102" box="[252,277,1636,1658]" italics="true" pageId="15" pageNumber="166">3a</emphasis>
|
||
. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFEFFFD6FF67F91D3280F0A2" blockId="15.[151,1437,1697,2010]" pageId="15" pageNumber="166">
|
||
Ovipositor with typical, double-walled tubular form and plicate cuticle (
|
||
<figureCitation id="132B2A2AFFEFFFD6FC5BF91D350FF1C2" box="[1019,1146,1697,1722]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="15" pageNumber="166">Figs 3B, C</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFEFFFD6FB28F91E35A9F1C2" box="[1160,1244,1698,1722]" captionStart="FIGURE 12. A – F" captionStartId="17.[151,250,775,797]" captionTargetBox="[151,1436,193,753]" captionTargetId="figure@17.[151,1436,193,754]" captionTargetPageId="17" captionText="FIGURE 12. A – F Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ 1 and coupled seta d; E, dorsolateral aspect of tibia I seta d (coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5 – 20 layers. Scale bars: 25 Μm (A – C, F); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238776/files/figure.png" pageId="15" pageNumber="166">12A, B</figureCitation>
|
||
); about 500–650 Μm when fully extended, half that when folded inside body. Distal lobes normal in form: paired anterior (dorsal) lobes and larger single posterior (ventral) lobe, the latter with convex, slightly sclerotized inner face that coapts with anterior lobes in retracted ovipositors (cf.
|
||
<figureCitation id="132B2A2AFFEFFFD6FD1BF8B13232F05E" box="[699,839,1805,1830]" captionStart="FIGURE 12. A – F" captionStartId="17.[151,250,775,797]" captionTargetBox="[151,1436,193,753]" captionTargetId="figure@17.[151,1436,193,754]" captionTargetPageId="17" captionText="FIGURE 12. A – F Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ 1 and coupled seta d; E, dorsolateral aspect of tibia I seta d (coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5 – 20 layers. Scale bars: 25 Μm (A – C, F); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238776/files/figure.png" pageId="15" pageNumber="166">Fig. 12A, B</figureCitation>
|
||
). With normal complement of nine pairs of simple, eupathidial setae, but with unusual distribution (see Remark 6). Three pairs of coronal setae (
|
||
<emphasis id="B964EABDFFEFFFD6FB67F88D35A6F032" box="[1223,1235,1841,1866]" italics="true" pageId="15" pageNumber="166">k</emphasis>
|
||
; 40–50 Μm); two pairs within primary fold constriction, one well below on anterior side. Each anterior lobe with three setae: τ1 (ca. 70 Μm) largest and most distal; τa (ca. 40 Μm) most proximal, at limit of plicate cuticle; τb (ca. 40 Μm) at intermediate level and more laterally positioned. Posterior lobe with three pairs of setae: ψ1 (ca. 75 Μm) most distal; ψa and ψb (each ca. 35 Μm) at same level, near limit of plicate cuticle.
|
||
</paragraph>
|
||
<caption id="DF6F6627FFF0FFC9FF37FA7D30D9F072" httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="16" pageNumber="167" targetBox="[151,1436,193,1451]" targetPageId="16">
|
||
<paragraph id="8BAF36AFFFF0FFC9FF37FA7D30D9F072" blockId="16.[151,1436,1473,1802]" pageId="16" pageNumber="167">
|
||
<emphasis id="B964EABDFFF0FFC9FF37FA7D3029F2AF" bold="true" box="[151,348,1473,1495]" pageId="16" pageNumber="167">FIGURE 11. A–G</emphasis>
|
||
<taxonomicName id="4C104D2CFFF0FFC9FEC5FA7D3320F2AF" box="[357,597,1473,1495]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF0FFC9FEC5FA7D3320F2AF" box="[357,597,1473,1495]" italics="true" pageId="16" pageNumber="167">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF0FFC9FDFEFA7E33EFF2AF" box="[606,666,1474,1495]" pageId="16" pageNumber="167" rank="species">
|
||
<emphasis id="B964EABDFFF0FFC9FDFEFA7E33EFF2AF" bold="true" box="[606,666,1474,1495]" pageId="16" pageNumber="167">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta
|
||
<emphasis id="B964EABDFFF0FFC9FBABFA5D356BF28E" box="[1035,1054,1505,1526]" italics="true" pageId="16" pageNumber="167">eI</emphasis>
|
||
, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9B), ventral aspect, anterior to top; G, close-up of cupule
|
||
<emphasis id="B964EABDFFF0FFC9FF45F9C63077F1F7" box="[229,258,1658,1679]" italics="true" pageId="16" pageNumber="167">ips</emphasis>
|
||
; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations:
|
||
<emphasis id="B964EABDFFF0FFC9FB7AF924358AF1D6" box="[1242,1279,1688,1710]" italics="true" pageId="16" pageNumber="167">ep1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFF0FFC9FAAEF9243447F1D6" box="[1294,1330,1688,1710]" italics="true" pageId="16" pageNumber="167">ep2</emphasis>
|
||
(epimeral plates),
|
||
<emphasis id="B964EABDFFF0FFC9FF4AF9043068F1B5" box="[234,285,1720,1741]" italics="true" pageId="16" pageNumber="167">g.mu</emphasis>
|
||
(genital plate muscles),
|
||
<emphasis id="B964EABDFFF0FFC9FD82F904334AF1B5" box="[546,575,1720,1741]" italics="true" pageId="16" pageNumber="167">ips</emphasis>
|
||
(lyrifissure),
|
||
<emphasis id="B964EABDFFF0FFC9FD71F904339EF1B5" box="[721,747,1720,1741]" italics="true" pageId="16" pageNumber="167">pa</emphasis>
|
||
(base of preanal seta),
|
||
<emphasis id="B964EABDFFF0FFC9FC46F9043578F1B5" box="[998,1037,1720,1741]" italics="true" pageId="16" pageNumber="167">pr.a</emphasis>
|
||
(preanal apodeme),
|
||
<emphasis id="B964EABDFFF0FFC9FB47F9043456F1B5" box="[1255,1315,1720,1741]" italics="true" pageId="16" pageNumber="167">pra.m</emphasis>
|
||
(transverse preanal muscles); other labels indicate setae (see text). All composites of 5–20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E–G).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFF0FFC9FF67F88A3074F0C3" blockId="16.[151,1436,1846,1979]" pageId="16" pageNumber="167">
|
||
Spermatopositor (
|
||
<figureCitation id="132B2A2AFFF0FFC9FE2FF88A3099F037" box="[399,492,1846,1871]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="16" pageNumber="167">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF0FFC9FE59F88B335CF037" box="[505,553,1847,1871]" captionStart="FIGURE 12. A – F" captionStartId="17.[151,250,775,797]" captionTargetBox="[151,1436,193,753]" captionTargetId="figure@17.[151,1436,193,754]" captionTargetPageId="17" captionText="FIGURE 12. A – F Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ 1 and coupled seta d; E, dorsolateral aspect of tibia I seta d (coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5 – 20 layers. Scale bars: 25 Μm (A – C, F); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238776/files/figure.png" pageId="16" pageNumber="167">12C</figureCitation>
|
||
) very short (ca. 50–60 Μm), conical, but with weakly striate lobes discernable; ovate in cross-section at base (ca. 50
|
||
<date id="FFAE106FFFF0FFC9FD9BF8E73319F00B" box="[571,620,1882,1907]" pageId="16" pageNumber="167" value="1960-10">x 60</date>
|
||
Μm). With eight pairs of small eupathidial setae (20–25 Μm), all tapered but blunt-ended; each anterior lobe with three at base (
|
||
<emphasis id="B964EABDFFF0FFC9FCB1F8C23252F0EF" box="[785,807,1918,1943]" italics="true" pageId="16" pageNumber="167">k)</emphasis>
|
||
and three distally (τ); posterior lobe with two pairs of setae (ψ).
|
||
</paragraph>
|
||
<caption id="DF6F6627FFF1FFC8FF37FCBB35FEF4E0" httpUri="https://zenodo.org/record/238776/files/figure.png" pageId="17" pageNumber="168" targetBox="[151,1436,193,753]" targetPageId="17">
|
||
<paragraph id="8BAF36AFFFF1FFC8FF37FCBB35FEF4E0" blockId="17.[151,1436,775,920]" pageId="17" pageNumber="168">
|
||
<emphasis id="B964EABDFFF1FFC8FF37FCBB3023F464" bold="true" box="[151,342,775,797]" pageId="17" pageNumber="168">FIGURE 12. A–F</emphasis>
|
||
<taxonomicName id="4C104D2CFFF1FFC8FEFEFCBB3338F465" box="[350,589,775,797]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF1FFC8FEFEFCBB3338F465" box="[350,589,775,797]" italics="true" pageId="17" pageNumber="168">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF1FFC8FDF4FCBB33FBF464" box="[596,654,775,796]" pageId="17" pageNumber="168" rank="species">
|
||
<emphasis id="B964EABDFFF1FFC8FDF4FCBB33FBF464" bold="true" box="[596,654,775,796]" pageId="17" pageNumber="168">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ1 and coupled seta
|
||
<emphasis id="B964EABDFFF1FFC8FB2FFCF835E9F422" box="[1167,1180,836,858]" italics="true" pageId="17" pageNumber="168">d</emphasis>
|
||
; E, dorsolateral aspect of tibia I seta
|
||
<emphasis id="B964EABDFFF1FFC8FEA5FCDF3067F401" box="[261,274,867,889]" italics="true" pageId="17" pageNumber="168">d</emphasis>
|
||
(coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5–20 layers. Scale bars: 25 Μm (A–C, F); 10 Μm (D, E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFF1FFC8FF67FC033334F264" blockId="17.[151,1437,959,2028]" pageId="17" pageNumber="168">
|
||
Plates of anal region collectively forming vague trapezoid in contracted individuals, widest posteriorly (
|
||
<figureCitation id="132B2A2AFFF1FFC8FACFFC03319BF484" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="17" pageNumber="168">Fig. 10A, B</figureCitation>
|
||
). Seen in distended individuals, anal plates widely separated from peranal plates and adanal region of notogaster (
|
||
<figureCitation id="132B2A2AFFF1FFC8FEBEFBBB300CF367" box="[286,377,1031,1056]" captionStart="FIGURE 9. A – B" captionStartId="14.[151,250,1638,1660]" captionTargetBox="[155,1431,192,1610]" captionTargetId="figure@14.[151,1436,181,1617]" captionTargetPageId="14" captionText="FIGURE 9. A – B Nanohystrix hammerae n. sp., adult female. A, several of the large setae that are abbreviated in B and in Fig. 2 B; B, ventral aspect, legs and some setae only partially shown. Abbreviations: ips (lyrifissure); other labels indicate setae (see text). Scale bar: 100 Μm." httpUri="https://zenodo.org/record/238773/files/figure.png" pageId="17" pageNumber="168">Fig. 9B</figureCitation>
|
||
), mostly by pale, porose cuticle with inconspicuously linear pattern. Each plate about four times longer than wide (ca. 280
|
||
<date id="FFAE106FFFF1FFC8FE5CFB903359F33C" box="[508,556,1068,1092]" pageId="17" pageNumber="168" value="1970-10">x 70</date>
|
||
Μm); tapered anteriorly to about one-third maximum width. Anal setae aligned along midline of plate; all attenuate, barbed, increasing in size posteriorly (
|
||
<emphasis id="B964EABDFFF1FFC8FC7CFBF33572F31F" box="[988,1031,1103,1127]" italics="true" pageId="17" pageNumber="168">an3</emphasis>
|
||
157 Μm,
|
||
<emphasis id="B964EABDFFF1FFC8FBD0FBF335EFF31F" box="[1136,1178,1103,1127]" italics="true" pageId="17" pageNumber="168">an2</emphasis>
|
||
270 Μm,
|
||
<emphasis id="B964EABDFFF1FFC8FAA4FBF3345AF31F" box="[1284,1327,1103,1127]" italics="true" pageId="17" pageNumber="168">an1</emphasis>
|
||
970 Μm);
|
||
<emphasis id="B964EABDFFF1FFC8FF37FBC831B4F3F4" box="[151,193,1140,1164]" italics="true" pageId="17" pageNumber="168">an2</emphasis>
|
||
closer to
|
||
<emphasis id="B964EABDFFF1FFC8FE97FBC83015F3F4" box="[311,352,1140,1164]" italics="true" pageId="17" pageNumber="168">an3</emphasis>
|
||
than to
|
||
<emphasis id="B964EABDFFF1FFC8FE64FBC8309BF3F4" box="[452,494,1140,1164]" italics="true" pageId="17" pageNumber="168">an1</emphasis>
|
||
. Peranal plates extremely narrow, strap-like (ca. 300
|
||
<date id="FFAE106FFFF1FFC8FBF9FBC835F8F3F4" box="[1113,1165,1139,1164]" pageId="17" pageNumber="168" value="1915-10">x 15</date>
|
||
Μm); medial edge soft, unpigmented, often flared outward. Plate pair fused anteriorly to form small ‘preanal’ apodeme for muscle attachment (
|
||
<figureCitation id="132B2A2AFFF1FFC8FE84FB073018F3AC" box="[292,365,1211,1236]" captionStart="FIGURE 11. A – G" captionStartId="16.[151,250,1473,1495]" captionTargetBox="[151,1436,193,1451]" captionTargetId="figure@16.[151,1436,193,1452]" captionTargetPageId="16" captionText="FIGURE 11. A – G Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images. A, mid-region of epimeres I and II, ventral aspect, showing granular soft cuticle; B, right supracoxal seta eI, anterolateral aspect; C, dorsal aspect (anterior to top) of left supracoxal region of epimeres I and II, showing spinous process at anterior border of epimere II (inserts show variation in process); D soft lateral cuticle above epimere I, showing cerotegument granules, some elongated, vermiform; E, same, enlargement of lower right corner, showing underlying fine striation and porosity; F, portion of incursion of pale cuticle lateral to adanal region of notogaster (cf. lower right side of Fig. 9 B), ventral aspect, anterior to top; G, close-up of cupule ips; H, shallow internal view of dissected (partial) anogenital region, showing preanal apodeme and genital plate muscles; I, same, but more internal focal plane, showing transverse compressor muscles. Abbreviations: ep 1, ep 2 (epimeral plates), g. mu (genital plate muscles), ips (lyrifissure), pa (base of preanal seta), pr. a (preanal apodeme), pra. m (transverse preanal muscles); other labels indicate setae (see text). All composites of 5 – 20 layers. Scale bars: 50 Μm (H, I); 20 Μm (A, D); 10 Μm (C); 5 Μm (B, E – G)." httpUri="https://zenodo.org/record/238775/files/figure.png" pageId="17" pageNumber="168">Fig.11</figureCitation>
|
||
H, I); similar small ‘postanal’ apodeme posteriorly for attachment of suspensor muscles (see Discussion). Single pair of peranal setae small (
|
||
<emphasis id="B964EABDFFF1FFC8FD1FFB5C33AEF38F" box="[703,731,1248,1271]" italics="true" pageId="17" pageNumber="168">pa</emphasis>
|
||
, ca. 60 Μm;
|
||
<figureCitation id="132B2A2AFFF1FFC8FCD1FB6332ACF38F" box="[881,985,1247,1272]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="17" pageNumber="168">Fig. 10D</figureCitation>
|
||
), attenuate, weakly barbed; inserted on lateral edge near anterior end of plate.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF1FFCBFF67FA9B33DBF6D4" blockId="17.[151,1437,959,2028]" lastBlockId="18.[151,1436,151,429]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
|
||
<emphasis id="B964EABDFFF1FFC8FF67FA9B3023F238" box="[199,342,1319,1344]" italics="true" pageId="17" pageNumber="168">Gnathosoma</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFF1FFC8FEC5FA9B3091F238" box="[357,484,1319,1344]" captionStart-0="FIGURE 13. A – E" captionStart-1="FIGURE 14. A – I" captionStart-2="FIGURE 15. A – I" captionStartId-0="18.[151,250,1862,1884]" captionStartId-1="19.[151,250,1685,1707]" captionStartId-2="20.[151,250,1459,1481]" captionTargetBox-0="[159,1425,491,1817]" captionTargetBox-1="[151,1436,193,1661]" captionTargetBox-2="[151,1436,193,1437]" captionTargetId-0="figure@18.[151,1436,481,1841]" captionTargetId-1="figure@19.[151,1436,193,1663]" captionTargetId-2="figure@20.[151,1436,193,1438]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." captionText-1="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." captionText-2="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri-0="https://zenodo.org/record/238777/files/figure.png" httpUri-1="https://zenodo.org/record/238779/files/figure.png" httpUri-2="https://zenodo.org/record/238780/files/figure.png" pageId="17" pageNumber="168">Figs 13–15</figureCitation>
|
||
). Subcapitulum and chelicerae usually oriented vertically or nearly so (
|
||
<figureCitation id="132B2A2AFFF1FFC8FAA1FA9B342BF247" box="[1281,1374,1319,1344]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="17" pageNumber="168">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF1FFC8FAC9FA9B34FBF238" box="[1385,1422,1319,1344]" captionStart="FIGURE 5. A – G" captionStartId="10.[151,250,1619,1641]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. A – G Nanohystrix hammerae n. sp., transmitted light compound microscope images of uncleared adult females, coarsely sectioned near sagittal plane (all lateral views, anterior to right). A, overview of fully distended specimen, with marked sections showing location of indicated close-ups; B, anterior third of fully contracted specimen, with close-up indicated; C, dorsal sejugal region, showing occipital wall and nuchal constriction of prodorsum, and extrinsic retractor muscles of chelicera; D, dorsal sejugal region and anterior region of notogaster, showing prodorsal retractor muscles (pr. mu; broken in C) inserted near border of nuchal constriction; E, posterior slope of hysterosoma in region of third scissure—at two focal planes, showing postanal suspensor muscle (po. mu) and deeper plane (right) with cuticular disk cd 5 (with underlying fibers, fib) and seta h 1 (with depressor muscle, h 1. mu); F, section of middle scissure (more lateral than indicated in A) showing base of seta f 1 and associated depressor muscle (f 1. mu); G, ventrosejugal region, showing narrow ventral part of sejugal articulation and broad soft cuticle of epimere I. Other abbreviations: bng (anterior border of notogaster), cr. mu (cheliceral retractor muscles); cw (caecum wall), ep 1 – 3 (plates of epimeres 1 – 3), nc (nuchal constriction), t. gr (transverse groove), sej (sejugal articulation), t. sul (transverse sulcus). All composites of 5 – 20 layers. Scale bars: 100 Μm (A, B); 25 Μm (C – G)." httpUri="https://zenodo.org/record/238769/files/figure.png" pageId="17" pageNumber="168">5A</figureCitation>
|
||
), even slightly posteroventrally in some individuals. Subcapitulum relatively small but unusually wide at base (ca. 1.2 times wider than long), abruptly narrowed by almost 50% at level of palp insertion (
|
||
<figureCitation id="132B2A2AFFF1FFC8FB39FAD33470F2F0" box="[1177,1285,1391,1416]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="17" pageNumber="168">Fig. 15A</figureCitation>
|
||
). Stenarthric, (
|
||
<figureCitation id="132B2A2AFFF1FFC8FF3FFA2F307CF2D4" box="[159,265,1427,1452]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="17" pageNumber="168">Fig. 13A</figureCitation>
|
||
): triangular mentum well delineated by pair of oblique labiogenal articulations (
|
||
<emphasis id="B964EABDFFF1FFC8FB33FA2F35DCF2D4" box="[1171,1193,1427,1452]" italics="true" pageId="17" pageNumber="168">lg</emphasis>
|
||
) closely approaching each other anteromedially, near base of lateral lips. Mentum (
|
||
<emphasis id="B964EABDFFF1FFC8FCCBFA04320AF2B7" box="[875,895,1464,1487]" italics="true" pageId="17" pageNumber="168">H</emphasis>
|
||
) paler than genae; with one pair of attenuate, sparsely barbed setae (
|
||
<emphasis id="B964EABDFFF1FFC8FE3AFA6730DDF28C" box="[410,424,1499,1524]" italics="true" pageId="17" pageNumber="168">h</emphasis>
|
||
, ca. 80 Μm). Gena (
|
||
<emphasis id="B964EABDFFF1FFC8FD30FA6033D1F28B" box="[656,676,1500,1523]" italics="true" pageId="17" pageNumber="168">G</emphasis>
|
||
) without dorsal rasp-like region of denticles; each gena with three weakly barbed ventral setae: posterior two (
|
||
<emphasis id="B964EABDFFF1FFC8FD2BF9BC33ECF16F" box="[651,665,1536,1559]" italics="true" pageId="17" pageNumber="168">n</emphasis>
|
||
, ca. 90 Μm;
|
||
<emphasis id="B964EABDFFF1FFC8FC8AF9BC324BF16F" box="[810,830,1536,1559]" italics="true" pageId="17" pageNumber="168">m</emphasis>
|
||
, 80 Μm) nearly aligned with
|
||
<emphasis id="B964EABDFFF1FFC8FB2CFA4335EFF160" box="[1164,1178,1535,1560]" italics="true" pageId="17" pageNumber="168">h</emphasis>
|
||
, distal seta (
|
||
<emphasis id="B964EABDFFF1FFC8FA8CF9BC344FF16F" box="[1324,1338,1536,1559]" italics="true" pageId="17" pageNumber="168">a</emphasis>
|
||
, 90 Μm) about at level with anterior corner of mentum. Lateral lips (
|
||
<emphasis id="B964EABDFFF1FFC8FCE9F998321DF143" box="[841,872,1572,1595]" italics="true" pageId="17" pageNumber="168">LL</emphasis>
|
||
) relatively large, fully exposed in ventral aspect, each clearly separated from gena by postadoral articulation (
|
||
<emphasis id="B964EABDFFF1FFC8FC9CF9FB323FF118" box="[828,842,1607,1632]" italics="true" pageId="17" pageNumber="168">d</emphasis>
|
||
); distal halves tapering to narrow adjacent tips. Each lip with triangular sclerite on ventral face and three pairs of adoral setae, each having few, minute barbs; distal seta
|
||
<emphasis id="B964EABDFFF1FFC8FF37F93331CBF1DF" box="[151,190,1679,1703]" italics="true" pageId="17" pageNumber="168">or1</emphasis>
|
||
(ca. 30 Μm) inserted in soft cuticle immediately distal to tip of sclerite, setal base distinctly flared, without normal alveolus; setae
|
||
<emphasis id="B964EABDFFF1FFC8FE3BF90830B4F1B4" box="[411,449,1716,1740]" italics="true" pageId="17" pageNumber="168">or2</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFF1FFC8FE59F9083354F1B4" box="[505,545,1716,1740]" italics="true" pageId="17" pageNumber="168">or3</emphasis>
|
||
(ca. 55 Μm) inserted on sclerite, at mid-length of lip, on small tubercles and in normal alveoli, in nearly transverse row. Dorsal surface with several longitudinal rows of extremely minute cilia or denticles, close to medial edge in distal half of lip. Rutellum (
|
||
<emphasis id="B964EABDFFF1FFC8FCD0F94032FAF06B" box="[880,911,1788,1811]" italics="true" pageId="17" pageNumber="168">Ru</emphasis>
|
||
) atelobasic; no rutellar brush; narrow, nearly uniform in width; distally rather chisel-like, with three aligned distal teeth. Rutellum basally incorporated smoothly with elongated manubrial zone (
|
||
<emphasis id="B964EABDFFF1FFC8FDB1F8F83343F023" box="[529,566,1860,1883]" italics="true" pageId="17" pageNumber="168">Mn</emphasis>
|
||
) of gena; manubrial-rutellar unit angled rather sharply anteromediad, with rutellar tips nearly meeting just anterior to lateral lips (
|
||
<figureCitation id="132B2A2AFFF1FFC8FCA1F8DB32DFF007" box="[769,938,1895,1920]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="17" pageNumber="168">Fig. 14B, D, E</figureCitation>
|
||
); distal limit of manubrium (
|
||
<emphasis id="B964EABDFFF1FFC8FB51F8D43588F007" box="[1265,1277,1896,1919]" italics="true" pageId="17" pageNumber="168">c</emphasis>
|
||
) well-marked in transmitted polarized light due to strong birefringence of rutellum (
|
||
<figureCitation id="132B2A2AFFF1FFC8FC0CF8373566F0DC" box="[940,1043,1931,1956]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="17" pageNumber="168">Fig. 15B</figureCitation>
|
||
); abaxial fissure (
|
||
<emphasis id="B964EABDFFF1FFC8FB7DF8303586F0DC" box="[1245,1267,1932,1956]" italics="true" pageId="17" pageNumber="168">αf</emphasis>
|
||
) marking base of manubrium visible in both SEM (
|
||
<figureCitation id="132B2A2AFFF1FFC8FDEAF81333C6F0BF" box="[586,691,1967,1992]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="17" pageNumber="168">Fig. 14B</figureCitation>
|
||
) and transmitted light (
|
||
<figureCitation id="132B2A2AFFF1FFC8FC6DF813354DF0B0" box="[973,1080,1967,1992]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="17" pageNumber="168">Fig. 15B</figureCitation>
|
||
). Dorsal face of subcapitulum without capitular apodeme; posterior margin supported by pair of narrow transverse bands of sclerotization interrupted by U-shaped medial notch (
|
||
<emphasis id="B964EABDFFF2FFCBFDF4FF2533F9F7C8" box="[596,652,153,176]" italics="true" pageId="18" pageNumber="169">m.no</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFF2FFCBFD3CFF2B3276F7C8" box="[668,771,151,176]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="18" pageNumber="169">Fig. 15A</figureCitation>
|
||
, insert). With flattened, sometimes pointed tubercle (
|
||
<emphasis id="B964EABDFFF2FFCBFAFDFF2B34E7F7C8" box="[1373,1426,151,176]" italics="true" pageId="18" pageNumber="169">sub.t</emphasis>
|
||
) on each dorsolateral corner of subcapitulum, at articulation with prodorsal condyle
|
||
<emphasis id="B964EABDFFF2FFCBFBF0FF013504F7AC" box="[1104,1137,189,212]" italics="true" pageId="18" pageNumber="169">pK</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFF2FFCBFB21FF0035A8F7AC" box="[1153,1245,188,213]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="18" pageNumber="169">Fig. 7G</figureCitation>
|
||
). Postpalpal seta (
|
||
<emphasis id="B964EABDFFF2FFCBFF00FF5D31B1F780" box="[160,196,225,248]" italics="true" pageId="18" pageNumber="169">ep;</emphasis>
|
||
ca. 8–10 Μm) spiniform, erect, inserted without alveolus (apparently immobile) just anterior to tubercle (
|
||
<figureCitation id="132B2A2AFFF2FFCBFACFFF6331BFF665" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="18" pageNumber="169">Fig. 15D</figureCitation>
|
||
). Labrum (
|
||
<emphasis id="B964EABDFFF2FFCBFEE7FEB93014F664" box="[327,353,261,284]" italics="true" pageId="18" pageNumber="169">Ls</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFF2FFCBFECFFEB83099F664" box="[367,492,260,285]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="18" pageNumber="169">Fig 15B, C</figureCitation>
|
||
) broad, with slightly raised medial ridge; distally rounded, extending almost to level of rutellar tips (if not elevated). Distal quarter of labrum with complex but slightly variable structure: with three transverse rows of dense spicules spanning plane of symmetry on dorsal face and fourth, most ventral row usually interrupted medially by larger distal spines, usually one pair (
|
||
<figureCitation id="132B2A2AFFF2FFCBFCF0FED332C3F6F0" box="[848,950,367,392]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="18" pageNumber="169">Figs 14F</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF2FFCBFC63FED33280F6F0" box="[963,1013,367,392]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="18" pageNumber="169">15C</figureCitation>
|
||
); ventral surface usually with several nipple-like lobes and proximal folds (
|
||
<figureCitation id="132B2A2AFFF2FFCBFD9CFE2833D7F6D4" box="[572,674,404,429]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="18" pageNumber="169">Fig. 14E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6F6627FFF2FFCBFF37F8FA35FFF08E" httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="18" pageNumber="169" targetBox="[159,1425,491,1817]" targetPageId="18">
|
||
<paragraph id="8BAF36AFFFF2FFCBFF37F8FA35FFF08E" blockId="18.[151,1436,1862,2038]" pageId="18" pageNumber="169">
|
||
<emphasis id="B964EABDFFF2FFCBFF37F8FA302CF023" bold="true" box="[151,345,1862,1884]" pageId="18" pageNumber="169">FIGURE 13. A–E</emphasis>
|
||
<taxonomicName id="4C104D2CFFF2FFCBFEC1F8FA3325F024" box="[353,592,1862,1884]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF2FFCBFEC1F8FA3325F024" box="[353,592,1862,1884]" italics="true" pageId="18" pageNumber="169">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF2FFCBFDF8F8FB33E6F024" box="[600,659,1863,1884]" pageId="18" pageNumber="169" rank="species">
|
||
<emphasis id="B964EABDFFF2FFCBFDF8F8FB33E6F024" bold="true" box="[600,659,1863,1884]" pageId="18" pageNumber="169">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations:
|
||
<emphasis id="B964EABDFFF2FFCBFAD7F83834E9F0E2" box="[1399,1436,1924,1946]" italics="true" pageId="18" pageNumber="169">c.fd</emphasis>
|
||
(carina of fixed digit),
|
||
<emphasis id="B964EABDFFF2FFCBFE2AF81E30CEF0C0" box="[394,443,1954,1976]" italics="true" pageId="18" pageNumber="169">c.md</emphasis>
|
||
(carina of movable digit),
|
||
<emphasis id="B964EABDFFF2FFCBFD7BF81E339DF0C0" box="[731,744,1954,1976]" italics="true" pageId="18" pageNumber="169">d</emphasis>
|
||
(postadoral articulation),
|
||
<emphasis id="B964EABDFFF2FFCBFC58F81E3579F0C0" box="[1016,1036,1954,1976]" italics="true" pageId="18" pageNumber="169">lg</emphasis>
|
||
(labiogenal articulation), G (gena),
|
||
<emphasis id="B964EABDFFF2FFCBFA2AF81F34E9F0C0" box="[1418,1436,1955,1976]" italics="true" pageId="18" pageNumber="169">H</emphasis>
|
||
(mentum),
|
||
<emphasis id="B964EABDFFF2FFCBFEA9F87E3050F0AF" box="[265,293,1986,2007]" italics="true" pageId="18" pageNumber="169">LL</emphasis>
|
||
(lateral lip),
|
||
<emphasis id="B964EABDFFF2FFCBFE0EF87E30BEF0AF" box="[430,459,1986,2007]" italics="true" pageId="18" pageNumber="169">Ru</emphasis>
|
||
(rutellum),
|
||
<emphasis id="B964EABDFFF2FFCBFDE8F87E3328F0AF" box="[584,605,1986,2007]" italics="true" pageId="18" pageNumber="169">sc</emphasis>
|
||
(scale-like flap),
|
||
<emphasis id="B964EABDFFF2FFCBFCB4F87E3245F0AF" box="[788,816,1986,2007]" italics="true" pageId="18" pageNumber="169">tr.t</emphasis>
|
||
(trochanteral tectum),
|
||
<emphasis id="B964EABDFFF2FFCBFBBDF87E3547F0AF" box="[1053,1074,1986,2007]" italics="true" pageId="18" pageNumber="169">Tr</emphasis>
|
||
(trochanter),
|
||
<emphasis id="B964EABDFFF2FFCBFB62F87D35AFF0AF" box="[1218,1242,1985,2007]" italics="true" pageId="18" pageNumber="169">vb</emphasis>
|
||
(vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFF3FFCAFF37F92935EEF01A" httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="19" pageNumber="170" targetBox="[151,1436,193,1661]" targetPageId="19">
|
||
<paragraph id="8BAF36AFFFF3FFCAFF37F92935EEF01A" blockId="19.[151,1436,1684,1890]" pageId="19" pageNumber="170">
|
||
<emphasis id="B964EABDFFF3FFCAFF37F9293027F1D2" bold="true" box="[151,338,1685,1707]" pageId="19" pageNumber="170">FIGURE 14. A–I</emphasis>
|
||
<taxonomicName id="4C104D2CFFF3FFCAFEFBF928333FF1D2" box="[347,586,1684,1706]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF3FFCAFEFBF928333FF1D2" box="[347,586,1684,1706]" italics="true" pageId="19" pageNumber="170">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF3FFCAFDF3F92933FBF1D2" box="[595,654,1685,1706]" pageId="19" pageNumber="170" rank="species">
|
||
<emphasis id="B964EABDFFF3FFCAFDF3F92933FBF1D2" bold="true" box="[595,654,1685,1706]" pageId="19" pageNumber="170">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations:
|
||
<emphasis id="B964EABDFFF3FFCAFA27F8B334EEF05D" box="[1415,1435,1807,1829]" italics="true" pageId="19" pageNumber="170">αf</emphasis>
|
||
(abaxial fissure),
|
||
<emphasis id="B964EABDFFF3FFCAFEE7F8923019F03C" box="[327,364,1838,1860]" italics="true" pageId="19" pageNumber="170">c.fd</emphasis>
|
||
(carina of fixed digit),
|
||
<emphasis id="B964EABDFFF3FFCAFDF7F8923300F03C" box="[599,629,1838,1860]" italics="true" pageId="19" pageNumber="170">Ch</emphasis>
|
||
(chelicera);
|
||
<emphasis id="B964EABDFFF3FFCAFD54F892327DF03C" box="[756,776,1838,1860]" italics="true" pageId="19" pageNumber="170">lg</emphasis>
|
||
(labiogenal articulation),
|
||
<emphasis id="B964EABDFFF3FFCAFBADF8933550F03C" box="[1037,1061,1839,1860]" italics="true" pageId="19" pageNumber="170">Ls</emphasis>
|
||
(labrum);
|
||
<emphasis id="B964EABDFFF3FFCAFB31F89335D8F03C" box="[1169,1197,1839,1860]" italics="true" pageId="19" pageNumber="170">Ru</emphasis>
|
||
(rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20Μm (B, C); 10 Μm (G, H); 5 Μm (D–F, I).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFF4FFCDFF37FA0F332EF062" httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="20" pageNumber="171" targetBox="[151,1436,193,1437]" targetPageId="20">
|
||
<paragraph id="8BAF36AFFFF4FFCDFF37FA0F332EF062" blockId="20.[151,1436,1459,1818]" pageId="20" pageNumber="171">
|
||
<emphasis id="B964EABDFFF4FFCDFF37FA0F3024F2B0" bold="true" box="[151,337,1459,1481]" pageId="20" pageNumber="171">FIGURE 15. A–I</emphasis>
|
||
<taxonomicName id="4C104D2CFFF4FFCDFEF9FA0F333DF2B1" box="[345,584,1459,1481]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF4FFCDFEF9FA0F333DF2B1" box="[345,584,1459,1481]" italics="true" pageId="20" pageNumber="171">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF4FFCDFDF0FA0833FFF2B1" box="[592,650,1460,1481]" pageId="20" pageNumber="171" rank="species">
|
||
<emphasis id="B964EABDFFF4FFCDFDF0FA0833FFF2B1" bold="true" box="[592,650,1460,1481]" pageId="20" pageNumber="171">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (
|
||
<emphasis id="B964EABDFFF4FFCDFAFBFA6F34F9F290" box="[1371,1420,1491,1512]" italics="true" pageId="20" pageNumber="171">m.no</emphasis>
|
||
); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5–20 layers. Abbreviations:
|
||
<emphasis id="B964EABDFFF4FFCDFA27F93634EEF1D8" box="[1415,1435,1674,1696]" italics="true" pageId="20" pageNumber="171">αf</emphasis>
|
||
(abaxial fissure),
|
||
<emphasis id="B964EABDFFF4FFCDFEF3F916302BF1C7" box="[339,350,1706,1727]" italics="true" pageId="20" pageNumber="171">c</emphasis>
|
||
(junction of manubrium with birefringent rutellum),
|
||
<emphasis id="B964EABDFFF4FFCDFC00F91632CDF1C7" box="[928,952,1706,1727]" italics="true" pageId="20" pageNumber="171">ep</emphasis>
|
||
(postpalpal seta),
|
||
<emphasis id="B964EABDFFF4FFCDFB22F91435D3F1C6" box="[1154,1190,1704,1726]" italics="true" pageId="20" pageNumber="171">l.or</emphasis>
|
||
(lamellated organ),
|
||
<emphasis id="B964EABDFFF4FFCDFA24F91634E9F1C7" box="[1412,1436,1706,1727]" italics="true" pageId="20" pageNumber="171">Ls</emphasis>
|
||
(labrum),
|
||
<emphasis id="B964EABDFFF4FFCDFF5CF97B305CF1A5" box="[252,297,1735,1757]" italics="true" pageId="20" pageNumber="171">l.mu</emphasis>
|
||
(levator muscle),
|
||
<emphasis id="B964EABDFFF4FFCDFE42F97B3377F1A5" box="[482,514,1735,1757]" italics="true" pageId="20" pageNumber="171">l.te</emphasis>
|
||
(levator tendon of movable digit),
|
||
<emphasis id="B964EABDFFF4FFCDFCC8F97432FCF1A5" box="[872,905,1736,1757]" italics="true" pageId="20" pageNumber="171">Mn</emphasis>
|
||
(manubrium),
|
||
<emphasis id="B964EABDFFF4FFCDFB8AF9743529F1A5" box="[1066,1116,1736,1757]" italics="true" pageId="20" pageNumber="171">m.no</emphasis>
|
||
(medial notch),
|
||
<emphasis id="B964EABDFFF4FFCDFAA4F9743446F1A5" box="[1284,1331,1736,1757]" italics="true" pageId="20" pageNumber="171">m.sp</emphasis>
|
||
(marginal spines),
|
||
<emphasis id="B964EABDFFF4FFCDFF4BF95B307DF184" box="[235,264,1767,1788]" italics="true" pageId="20" pageNumber="171">Ru</emphasis>
|
||
(rutellum),
|
||
<emphasis id="B964EABDFFF4FFCDFEDFF95A30C5F184" box="[383,432,1766,1788]" italics="true" pageId="20" pageNumber="171">sub.t</emphasis>
|
||
(subcapitular tubercle),
|
||
<emphasis id="B964EABDFFF4FFCDFD06F95B33CEF184" box="[678,699,1767,1788]" italics="true" pageId="20" pageNumber="171">Tr</emphasis>
|
||
(trochanter)
|
||
<emphasis id="B964EABDFFF4FFCDFCE0F95B322EF184" box="[832,859,1767,1788]" italics="true" pageId="20" pageNumber="171">tr.t</emphasis>
|
||
(trochanteral tectum),
|
||
<emphasis id="B964EABDFFF4FFCDFBE2F95A352FF184" box="[1090,1114,1766,1788]" italics="true" pageId="20" pageNumber="171">vb</emphasis>
|
||
(vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B–D, G, H).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFF5FFCCFF37F977336AF044" httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="21" pageNumber="172" targetBox="[155,1431,195,1713]" targetPageId="21">
|
||
<paragraph id="8BAF36AFFFF5FFCCFF37F977336AF044" blockId="21.[151,1436,1739,1852]" pageId="21" pageNumber="172">
|
||
<emphasis id="B964EABDFFF5FFCCFF37F9773028F199" bold="true" box="[151,349,1739,1761]" pageId="21" pageNumber="172">FIGURE 16. A–G</emphasis>
|
||
<taxonomicName id="4C104D2CFFF5FFCCFEC6F9773323F199" box="[358,598,1739,1761]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF5FFCCFEC6F9773323F199" box="[358,598,1739,1761]" italics="true" pageId="21" pageNumber="172">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF5FFCCFDFFF97733EEF198" box="[607,667,1739,1760]" pageId="21" pageNumber="172" rank="species">
|
||
<emphasis id="B964EABDFFF5FFCCFDFFF97733EEF198" bold="true" box="[607,667,1739,1760]" pageId="21" pageNumber="172">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (
|
||
<emphasis id="B964EABDFFF5FFCCFDD1F9563309F187" box="[625,636,1770,1791]" italics="true" pageId="21" pageNumber="172">e</emphasis>
|
||
); D, tibia I solenidion
|
||
<emphasis id="B964EABDFFF5FFCCFCD1F95532FEF187" box="[881,907,1769,1791]" italics="true" pageId="21" pageNumber="172">φ1</emphasis>
|
||
with minute coupled seta
|
||
<emphasis id="B964EABDFFF5FFCCFB04F95535C5F187" box="[1188,1200,1769,1791]" italics="true" pageId="21" pageNumber="172">d</emphasis>
|
||
; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (
|
||
<emphasis id="B964EABDFFF5FFCCFCCDF8B5320DF066" box="[877,888,1801,1822]" italics="true" pageId="21" pageNumber="172">e</emphasis>
|
||
). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D–F); 5 Μm (C, G).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFF5FFCFFF67F8D535A7F6D4" blockId="21.[151,1436,1897,2030]" lastBlockId="22.[151,1437,151,2013]" lastPageId="22" lastPageNumber="173" pageId="21" pageNumber="172">
|
||
Palp relatively large and conspicuous (
|
||
<figureCitation id="132B2A2AFFF5FFCCFDDDF8D533AFF0FA" box="[637,730,1897,1922]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="21" pageNumber="172">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF5FFCCFD47F8D6326FF0FA" box="[743,794,1898,1922]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="21" pageNumber="172">14A</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF5FFCCFC86F8D5322CF0FA" box="[806,857,1897,1922]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="21" pageNumber="172">15A</figureCitation>
|
||
. Simple in form (
|
||
<figureCitation id="132B2A2AFFF5FFCCFB82F8D535FCF0FA" box="[1058,1161,1897,1922]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="21" pageNumber="172">Fig. 13B</figureCitation>
|
||
); five freely articulating segments with typical proportions: trochanter, genu and tibia short; femur about twice length of genu; tarsus elongated, about equal to femur and genu combined. Tarsus finely striated distally; with dorsal lyrifissure proximally. Setal formula (trochanter to tarsus) 0-
|
||
<date id="FFAE106FFFF5FFCCFD75F86A3390F096" box="[725,741,2006,2030]" pageId="21" pageNumber="172" value="2013-01-02" valueMax="2013-03-02" valueMin="2013-01-02">2</date>
|
||
-1-3-13 (solenidion not included; tarsus count includes fused distal eupathidia as two); probable homologies indicated on
|
||
<figureCitation id="132B2A2AFFF6FFCFFCEAFF2B32A1F7C8" box="[842,980,151,176]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">Fig. 13B, C</figureCitation>
|
||
. All setae of femur, genu and tibia, and several proximal setae of tarsus, attenuate and noticeably barbed; more distal setae with fewer barbs, or none. Tarsal solenidion ω finely tapered, flexible; inserted unusually far distally, in terminal quarter of segment, extending slightly beyond most distal tarsal setae when straight. Homology of tarsal setation incompletely known (see Remark 10); most tarsal setae with soft cuticle of alveolus teardrop-shaped, pointed distally. Six distal tarsal setae eupathidial, including probable
|
||
<emphasis id="B964EABDFFF6FFCFFD9FFEF13318F61C" box="[575,621,333,356]" italics="true" pageId="22" pageNumber="173">acm</emphasis>
|
||
and ultimal (
|
||
<emphasis id="B964EABDFFF6FFCFFCA7FEF03269F61D" box="[775,796,332,357]" italics="true" pageId="22" pageNumber="173">ul</emphasis>
|
||
) pair (
|
||
<figureCitation id="132B2A2AFFF6FFCFFCC9FEF032A1F61C" box="[873,980,332,357]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">Figs 13C</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFC41FEF03560F61C" box="[993,1045,332,356]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="22" pageNumber="173">14G</figureCitation>
|
||
);
|
||
<emphasis id="B964EABDFFF6FFCFFB8BFEF03532F61D" box="[1067,1095,332,357]" italics="true" pageId="22" pageNumber="173">ul'</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFF6FFCFFBDFFEF135EDF61C" box="[1151,1176,333,356]" italics="true" pageId="22" pageNumber="173">su</emphasis>
|
||
fused in basal 2/3, but individual setae distinguishable, spread noticeably in distal third. Variations: one specimen with femoral seta
|
||
<emphasis id="B964EABDFFF6FFCFFAD5FECD34E9F6F0" box="[1397,1436,369,392]" italics="true" pageId="22" pageNumber="173">sup</emphasis>
|
||
absent; another with seta
|
||
<emphasis id="B964EABDFFF6FFCFFE13FE2830A5F6D4" box="[435,464,404,428]" italics="true" pageId="22" pageNumber="173">inf</emphasis>
|
||
doubled (represented by two closely-spaced, unusually small setae).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF6FFCFFF67FE0B3461F248" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
Chelicera usually oriented vertically; chelate-dentate in form, with typical outline in lateral view (
|
||
<figureCitation id="132B2A2AFFF6FFCFFAA9FE0B34FAF6A8" box="[1289,1423,439,464]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">Fig. 13D, E</figureCitation>
|
||
). Cheliceral frame not encroaching onto principal segment; cuticle of latter distinctly porose proximal to chela. Trochanter (
|
||
<emphasis id="B964EABDFFF6FFCFFE89FDBD3034F560" box="[297,321,513,536]" italics="true" pageId="22" pageNumber="173">Tr</emphasis>
|
||
) distinct, relatively large, well sclerotized ventrally; soft dorsally, merging with and losing distinctness from soft granular cuticle of cheliceral frame. Trochanter with tongue-like, well sclerotized tectum (
|
||
<emphasis id="B964EABDFFF6FFCFFF00FDF531CBF518" box="[160,190,585,608]" italics="true" pageId="22" pageNumber="173">tr.t</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFF6FFCFFF74FDFB302AF518" box="[212,351,583,608]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="22" pageNumber="173">Fig. 15E–G</figureCitation>
|
||
) ventrally, overhanging articulation with principal segment. Tectum with hyaline cuticular coating, extending distally as thin, colorless fringe with minute cilia; several large spines more basally on abaxial edge of tectum, distal to articulation with principal segment. Principal segment strongly asymmetrical, giving broad, flat-faced appearance to anterior of mite (
|
||
<figureCitation id="132B2A2AFFF6FFCFFD18FD083220F5B4" box="[696,853,692,717]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="22" pageNumber="173">Fig. 6B, C, G</figureCitation>
|
||
). Adaxial face almost flat, forming coaptive surface when chelicerae close together (
|
||
<figureCitation id="132B2A2AFFF6FFCFFE5FFD6B3323F588" box="[511,598,727,752]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="22" pageNumber="173">Fig. 6B</figureCitation>
|
||
), accentuated dorsally by carina (
|
||
<emphasis id="B964EABDFFF6FFCFFC70FD6B329EF588" box="[976,1003,727,752]" italics="true" pageId="22" pageNumber="173">dc</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFF6FFCFFC59FD6B353AF588" box="[1017,1103,727,752]" captionStart="FIGURE 6. A – G" captionStartId="11.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1498]" captionTargetId="figure@11.[151,1436,193,1499]" captionTargetPageId="11" captionText="FIGURE 6. A – G Nanohystrix hammerae n. sp., scanning electron micrographs of adults (most with small surface debris artifacts). A, sejugal region of partly distended specimen, dorsolateral aspect (anterior to left) — arrow indicates minute surface granules (probably cerotegument); B, anterior third, ventral aspect, showing rostrum, gnathosoma and epimeres I – III; C, same, ventrolateral aspect, with chelicerae spread; D, bothridium and base of bothridial seta; E, portion of sejugal region of contracted specimen, anterodorsal view (cf. left side of Fig. 4 B), showing reticulation (arrow) on notogaster; F, enlargement of C, right center; G, prodorsum, lateral aspect (slightly anterodorsal). Abbreviations: ca 1, ca 2 (prodorsal carinae), cpc (podocephalic canal), Ch (chelicera), dc (dorsal carina of chelicera), ep 1 – 3 (plates of epimeres 1 – 3), fl (flange of bothridial wall), Ge (gena), Ls (labrum), nc (nuchal constriction), oc. l (lateral eye), rb (rostral bulge), rm (rostral margin), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 50 Μm (B, C, G); 25 Μm (A, F); 10 Μm (E); 5 Μm (D)." httpUri="https://zenodo.org/record/238770/files/figure.png" pageId="22" pageNumber="173">Fig. 6C</figureCitation>
|
||
) bearing several short spines; abaxial face strongly convex, increasingly so as chelicera broadens proximally to maximum width, close to proximal end (
|
||
<figureCitation id="132B2A2AFFF6FFCFFEE0FCA330DDF440" box="[320,424,799,824]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="22" pageNumber="173">Fig. 15E</figureCitation>
|
||
). Abaxial face of principal segment with few features: proximal third with about dozen minute, sharp spicules, many at end of fine, sharply defined carinae angling posterodorsad (
|
||
<figureCitation id="132B2A2AFFF6FFCFFB19FCF8347AF424" box="[1209,1295,836,861]" captionStart="FIGURE 7. A – J" captionStartId="12.[160,259,1560,1582]" captionTargetBox="[151,1436,193,1537]" captionTargetId="figure@12.[151,1436,193,1539]" captionTargetPageId="12" captionText="FIGURE 7. A – J Nanohystrix hammerae n. sp., transmitted light compound microscope images of adults (cleared unless noted). A, right posterior region of prodorsum, dorsal aspect, showing muscle sigilla; B, close-up of general cuticular porosity; C, lateral eye, dorsal view; D, partial lateral aspect of prodorsum and supracoxal region, anterior to right—uncleared specimen showing pigment in lateral eye (insert shows eye from posterodorsal aspect); E, bothridium and base of bothridial seta, lateral aspect with focus near edge of cup; F, same, with focus at center of cup; G, similar to D, but after clearing; H, optical section of hyaline rostral bulge and median eye, lateral aspect, anterior to right (cf. Grandjean 1958 b, his Fig. 2); I, anterior view of central rostrum and cheliceral bases, focused on rostral bulge; J, same, focused deeper, on median eye. All composites of 5 – 20 layers. Abbreviations: bo (bothridium), bt (base of rostral tectum), ca 1 (prodorsal carina), Ch (chelicera), cpc (podocephalic canal), oc. l (lateral eye), oc. m (median eye), mu. sl (subcapitular levator muscle), rb (rostral bulge), pK (prodorsal condyle), st (constriction of tectum depth), str (supporting strut), sub. t (subcapitular tubercle); other labels indicate setae (see text). Scale bars: 20 Μm (A, D, G – J); 10 Μm (B); 5 Μm (C, E, F)." httpUri="https://zenodo.org/record/238771/files/figure.png" pageId="22" pageNumber="173">Figs 7J</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFABDFCF83438F424" box="[1309,1357,836,860]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">13E</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFAFBFCF834FBF425" box="[1371,1422,836,861]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="22" pageNumber="173">15H</figureCitation>
|
||
); fixed digit with low carina (
|
||
<emphasis id="B964EABDFFF6FFCFFE73FCDB3371F4F8" box="[467,516,871,896]" italics="true" pageId="22" pageNumber="173">c.fd.</emphasis>
|
||
) near middle (
|
||
<figureCitation id="132B2A2AFFF6FFCFFD09FCDB3264F4F8" box="[681,785,871,896]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">Figs 13E</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFCBDFCD4323AF4F8" box="[797,847,872,896]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="22" pageNumber="173">14B</figureCitation>
|
||
); movable digit with short carina (
|
||
<emphasis id="B964EABDFFF6FFCFFB75FCDB347EF4F8" box="[1237,1291,871,896]" italics="true" pageId="22" pageNumber="173">c.md</emphasis>
|
||
) proximally, which coapts to edge of fixed digit if chela closed. Adaxial face of principal segment (
|
||
<figureCitation id="132B2A2AFFF6FFCFFBC4FC3035BFF4DC" box="[1124,1226,908,933]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">Fig. 13D</figureCitation>
|
||
) deeply excavated in proximal half, accommodating asymmetrical insertion of trochanter; with about half-dozen spicules in distal half. Without Trägårdh’s organ, but with attached, irregular, ‘puffy’ (perhaps inflatable;
|
||
<figureCitation id="132B2A2AFFF6FFCFFBCCFC6835B8F495" box="[1132,1229,980,1005]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="22" pageNumber="173">Fig. 15E</figureCitation>
|
||
) vesicular band of soft cuticle (
|
||
<emphasis id="B964EABDFFF6FFCFFE89FC4B3031F368" box="[297,324,1015,1040]" italics="true" pageId="22" pageNumber="173">vb</emphasis>
|
||
) running distally from trochanter to end at strong spine (ca. 15 Μm) on mid-face of fixed digit. Small, freely projecting scale-like flap (
|
||
<emphasis id="B964EABDFFF6FFCFFDD0FBA133F2F34C" box="[624,647,1053,1076]" italics="true" pageId="22" pageNumber="173">sc</emphasis>
|
||
) present, with ciliate margin, arising at about level of articulation between segments. Lamellated organ (
|
||
<emphasis id="B964EABDFFF6FFCFFDEFFB83330EF320" box="[591,635,1087,1112]" italics="true" pageId="22" pageNumber="173">l.or.</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFF6FFCFFD2CFB83339BF320" box="[652,750,1087,1112]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="22" pageNumber="173">Fig. 15F</figureCitation>
|
||
, insert) present close to flap (see Remark 5). Fixed digit of chela strongly birefringent distal to seta
|
||
<emphasis id="B964EABDFFF6FFCFFDCEFBD833E2F305" box="[622,663,1124,1149]" italics="true" pageId="22" pageNumber="173">chb</emphasis>
|
||
; with five teeth (including terminal tooth and small tooth high on adaxial face). Movable digit birefringent in distal half; with four teeth (including terminal and very weak subterminal tooth). Principal segment with two attenuate setae, both aligned near midline, just abaxial to dorsal carina (
|
||
<figureCitation id="132B2A2AFFF6FFCFFF4AFB733007F390" box="[234,370,1231,1256]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">Fig. 13D, E</figureCitation>
|
||
):
|
||
<emphasis id="B964EABDFFF6FFCFFE26FB7330DAF390" box="[390,431,1231,1256]" italics="true" pageId="22" pageNumber="173">cha</emphasis>
|
||
(ca. 65 Μm) clearly barbed, inserted at base of fixed digit;
|
||
<emphasis id="B964EABDFFF6FFCFFB9EFB733513F390" box="[1086,1126,1231,1256]" italics="true" pageId="22" pageNumber="173">chb</emphasis>
|
||
(ca. 40 Μm) nearly smooth, inserted at mid-level of fixed digit. Pronounced, often pigmented tendon (
|
||
<emphasis id="B964EABDFFF6FFCFFC7EFB493284F274" box="[990,1009,1269,1292]" italics="true" pageId="22" pageNumber="173">ts</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFF6FFCFFBA1FB48352EF275" box="[1025,1115,1268,1293]" captionStart="FIGURE 15. A – I" captionStartId="20.[151,250,1459,1481]" captionTargetBox="[151,1436,193,1437]" captionTargetId="figure@20.[151,1436,193,1438]" captionTargetPageId="20" captionText="FIGURE 15. A – I Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images of dissected parts of gnathosoma. A, dorsal aspect of subcapitulum and palps with enlarged insert showing medial notch in posterior margin (m. no); B, distal subcapitulum in strong DIC, dorsal aspect, showing birefringence of rutellum; C, same, but with focus on tip of labrum; D, right posterolateral corner of subcapitulum, lateral aspect (anterior to right), showing postpalpal seta and tubercle; E, chelicera, ventral aspect; F. chelicera, adaxial aspect, with insert showing lamellated organ; G, enlargement of ventrodistal region of cheliceral trochanter (cf. E), showing marginal spines and tectum with hyaline fringe (black arrow); H, proximodorsal region of chelicera showing porose cuticle with spines; I, chelicera in strong DIC illumination, focused internally to show levator tendon of movable digit and associated muscles. All composites of 5 – 20 layers. Abbreviations: αf (abaxial fissure), c (junction of manubrium with birefringent rutellum), ep (postpalpal seta), l. or (lamellated organ), Ls (labrum), l. mu (levator muscle), l. te (levator tendon of movable digit), Mn (manubrium), m. no (medial notch), m. sp (marginal spines), Ru (rutellum), sub. t (subcapitular tubercle), Tr (trochanter) tr. t (trochanteral tectum), vb (vesicular band). Scale bars: 50 Μm (E, F, I); 25 Μm (A); 10 Μm (B – D, G, H)." httpUri="https://zenodo.org/record/238780/files/figure.png" pageId="22" pageNumber="173">
|
||
Fig.
|
||
<date id="FFAE106FFFF6FFCFFB95FB48352EF275" box="[1077,1115,1268,1293]" pageId="22" pageNumber="173" value="1915-01">15I</date>
|
||
</figureCitation>
|
||
) present, attaching bands of massive levator muscle to movable digit; articulation of digit with distinct onchophysis
|
||
<emphasis id="B964EABDFFF6FFCFFBCEFAA435E5F257" box="[1134,1168,1304,1327]" italics="true" pageId="22" pageNumber="173">op'</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFF6FFCFFB3FFAAB3472F248" box="[1183,1287,1303,1328]" captionStart="FIGURE 13. A – E" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[159,1425,491,1817]" captionTargetId="figure@18.[151,1436,481,1841]" captionTargetPageId="18" captionText="FIGURE 13. A – E Nanohystrix hammerae n. sp., gnathosomal elements of adult. A, central region of subcapitulum, ventral aspect; B, right palp, abaxial aspect (integumental granules shown only on basal segment); C, distal portion of palp tarsus with enlarged section of fused eupathidia; D, left chelicera, adaxial aspect; E. Right chelicera, abaxial aspect. Abbreviations: c. fd (carina of fixed digit), c. md (carina of movable digit), d (postadoral articulation), lg (labiogenal articulation), G (gena), H (mentum), LL (lateral lip), Ru (rutellum), sc (scale-like flap), tr. t (trochanteral tectum), Tr (trochanter), vb (vesicular band of chelicera); other labels indicate setae (see text). Scale bars: 50 Μm (D, E); 25 Μm (A, B); 10 Μm (C)." httpUri="https://zenodo.org/record/238777/files/figure.png" pageId="22" pageNumber="173">Fig. 13D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF6FFCFFF67FA81334AF008" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="173">
|
||
<emphasis id="B964EABDFFF6FFCFFF67FA813076F22C" box="[199,259,1341,1364]" italics="true" pageId="22" pageNumber="173">Legs.</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFF6FFCFFEB7FA803006F22D" box="[279,371,1340,1365]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="22" pageNumber="173">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFE24FA8030D6F22C" box="[388,419,1340,1364]" captionStart="FIGURE 17. A – C" captionStartId="24.[151,250,1858,1880]" captionTargetBox="[157,1436,193,1823]" captionTargetId="figure@24.[151,1436,193,1837]" captionTargetPageId="24" captionText="FIGURE 17. A – C Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, left leg III, tarsus partially shown; B, left tarsus III; C, left leg IV (slightly twisted at base). Labels all indicate setiform organs (see text). Scale bars: 100 Μm." httpUri="https://zenodo.org/record/238782/files/figure.png" pageId="22" pageNumber="173">17</figureCitation>
|
||
). Relatively long: leg IV longest, about as long as body (varies with degree of body distension); approximate proportions of others (relative to IV): 0.8 (I), 0.7 (II), 0.8 (III). Legs III and IV with normal, posterolateral direction in all specimens. Legs I and II unusually mobile (inferred from highly variable positions in preserved specimens); with normal anterolateral orientation in some specimens (
|
||
<figureCitation id="132B2A2AFFF6FFCFFB6AFA1B3433F2B8" box="[1226,1350,1447,1472]" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="22" pageNumber="173">Fig. 1D, F</figureCitation>
|
||
) but in others directed laterally, posterolaterally, or even posterodorsally (at nearly same angle as legs III and IV;
|
||
<figureCitation id="132B2A2AFFF6FFCFFA9CFA7031A5F170" captionStart="FIGURE 1. A – H" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[164,1407,195,1812]" captionTargetId="figure@6.[164,1408,195,1812]" captionTargetPageId="6" captionText="FIGURE 1. A – H Nanohystrix hammerae n. sp., reflected light stereomicroscope images of adults (genders unknown; multiyear preservation except D). A, dorsal aspect of specimen with legs I, II angled posteriorly; B, ventral aspect of same; C, anterior third in posterodorsal aspect; D, dorsolateral aspect of recently collected specimen; E, dorsal aspect of notogaster; F, lateral aspect with legs I in walking position; G, lateral aspect (anterior half) of specimen with right leg I elevated; H, posterior aspect showing granules (arrows) of paired digestive caeca, seen by transparency. Abbreviations: h. cr (humeral crease); other labels indicate setae (see text). All composites of 10 – 20 layers. Scale bars: all 100 Μm." httpUri="https://zenodo.org/record/238765/files/figure.png" pageId="22" pageNumber="173">Figs 1A, B, G</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFF7FFA4C3064F170" box="[223,273,1520,1544]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="22" pageNumber="173">10B</figureCitation>
|
||
). Leg I, and to small extent leg II, rotatable crab-like; i.e. ventral surface twisted forward in some specimens (
|
||
<figureCitation id="132B2A2AFFF6FFCFFEBEF9A8300EF154" box="[286,379,1556,1581]" captionStart="FIGURE 4. A – G" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,193,1759]" captionTargetId="figure@9.[151,1436,193,1760]" captionTargetPageId="9" captionText="FIGURE 4. A – G Nanohystrix hammerae n. sp., scanning electron micrographs (genders unknown; most with debris artifacts). A, dorsal overview of contracted specimen with erectile setae depressed and legs I, II directed laterally; B, anterodorsal aspect of same (prodorsum fractured centrally), insert includes enlargement of cuticular disk cd 1, indicated by black arrow in main figure; C, dorsolateral aspect of hysterosoma of partially distended specimen with setae erected; D, same, enlarged, short arrows on cuticular disks cd 5 (insert is enlargement of right disk cd 5 showing exudate); E, base of seta e 2; F, basal portion of seta d 1; G, basal portion of seta f 1. Abbreviations; h. cr (humeral crease); other labels indicate setae (see text). Scale bars 100 Μm (A – D); 10 Μm (E – G)." httpUri="https://zenodo.org/record/238768/files/figure.png" pageId="22" pageNumber="173">Fig. 4B</figureCitation>
|
||
). Surface of most leg segments nearly smooth, but tarsi finely striated distally (
|
||
<figureCitation id="132B2A2AFFF6FFCFFA83F9A834FAF154" box="[1315,1423,1556,1581]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="22" pageNumber="173">Fig. 14H</figureCitation>
|
||
); cuticle finely porose but without regions of larger pores (porose areas) or other surface patterns. Each tarsus with small proximodorsal lyrifissure. Pretarsus of all legs similar, tridactylous with empodial claw slightly longer and about twice width of lateral claws (14H); claws without denticles; ambulacrum without separate condylophores. Except for trochanters I and II and all tarsi, leg segments roughly tubular and similar in size (
|
||
<figureCitation id="132B2A2AFFF6FFCFFB6BF9183443F1C4" box="[1227,1334,1700,1725]" captionStart="FIGURE 10. A – E" captionStartId="15.[151,250,1514,1536]" captionTargetBox="[151,1436,193,1492]" captionTargetId="figure@15.[151,1436,193,1492]" captionTargetPageId="15" captionText="FIGURE 10. A – E Nanohystrix hammerae n. sp., scanning electron micrographs of adults, ventral aspect (asterisks denote artifacts; most images with unmarked small surface debris). A, anogenital region, peranal valves slightly opened; B, habitus of fully contracted specimen with legs I, II laterally directed, insert showing posterior region of more distended specimen with weaker imbrication of adanal region (cf. arrows); C, coxisternal region; D, peranal seta and soft, finely striated cuticle lateral to it; E, seta 3 a. Labels indicate setae (see text). Scale bars: 100 Μm (B); 50 Μm (A, C); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238774/files/figure.png" pageId="22" pageNumber="173">Figs 10B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFAE4F9183417F1C5" box="[1348,1378,1700,1725]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="22" pageNumber="173">16</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFAD0F91834FAF1C4" box="[1392,1423,1700,1724]" captionStart="FIGURE 17. A – C" captionStartId="24.[151,250,1858,1880]" captionTargetBox="[157,1436,193,1823]" captionTargetId="figure@24.[151,1436,193,1837]" captionTargetPageId="24" captionText="FIGURE 17. A – C Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, left leg III, tarsus partially shown; B, left tarsus III; C, left leg IV (slightly twisted at base). Labels all indicate setiform organs (see text). Scale bars: 100 Μm." httpUri="https://zenodo.org/record/238782/files/figure.png" pageId="22" pageNumber="173">17</figureCitation>
|
||
). Trochanters I and II relatively short, less than half length of respective femur. Tarsi much elongated; tarsus I about 6–7 times longer than proximal width, tapering gradually from base to tip; tarsus IV about 11–12 times longer than proximal width, with most tapering in proximal quarter, distal to which segment almost isodiametric, very thin and somewhat flexible (based on variations in preservation;
|
||
<figureCitation id="132B2A2AFFF6FFCFFCE8F88832C1F034" box="[840,948,1844,1869]" captionStart="FIGURE 12. A – F" captionStartId="17.[151,250,775,797]" captionTargetBox="[151,1436,193,753]" captionTargetId="figure@17.[151,1436,193,754]" captionTargetPageId="17" captionText="FIGURE 12. A – F Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ 1 and coupled seta d; E, dorsolateral aspect of tibia I seta d (coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5 – 20 layers. Scale bars: 25 Μm (A – C, F); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238776/files/figure.png" pageId="22" pageNumber="173">Fig. 12F</figureCitation>
|
||
); form of tarsi II and III progressively intermediate between that of I and IV.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF6FFCEFF67F8C1358BF6A8" blockId="22.[151,1437,151,2013]" lastBlockId="23.[151,1437,151,464]" lastPageId="23" lastPageNumber="174" pageId="22" pageNumber="173">
|
||
Leg setation, expressed as formula (I-II-III-IV; famuli and minute setae
|
||
<emphasis id="B964EABDFFF6FFCFFB85F8C03546F0ED" box="[1061,1075,1916,1941]" italics="true" pageId="22" pageNumber="173">d</emphasis>
|
||
included, solenidia excluded): trochanters (1-1-2-2); femora (6-6-4-4); genua (5-5-4-4); tibiae (6-6-4-4); tarsi (20-18-17-15). Setal homologies given in
|
||
<figureCitation id="132B2A2AFFF6FFCFFF5AF8783025F0A5" box="[250,336,1988,2013]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="22" pageNumber="173">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF6FFCFFEFDF878300CF0A4" box="[349,377,1988,2012]" captionStart="FIGURE 17. A – C" captionStartId="24.[151,250,1858,1880]" captionTargetBox="[157,1436,193,1823]" captionTargetId="figure@24.[151,1436,193,1837]" captionTargetPageId="24" captionText="FIGURE 17. A – C Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, left leg III, tarsus partially shown; B, left tarsus III; C, left leg IV (slightly twisted at base). Labels all indicate setiform organs (see text). Scale bars: 100 Μm." httpUri="https://zenodo.org/record/238782/files/figure.png" pageId="22" pageNumber="173">17</figureCitation>
|
||
and Table 1. Noted variations include two tarsi I lacking seta
|
||
<emphasis id="B964EABDFFF6FFCFFB96F8793539F0A4" box="[1078,1100,1989,2012]" italics="true" pageId="22" pageNumber="173">it'</emphasis>
|
||
and one tarsus II lacking seta
|
||
<emphasis id="B964EABDFFF7FFCEFF37FF2531DEF7C8" box="[151,171,153,176]" italics="true" pageId="23" pageNumber="174">tc</emphasis>
|
||
' (20 of each leg examined). Most setae moderately to strongly barbed, exceptions being distal setae of tarsi, on which barbs usually small and sparse as illustrated, and smooth eupathidial setae and regressive seta
|
||
<emphasis id="B964EABDFFF7FFCEFAA2FF003465F7AD" box="[1282,1296,188,213]" italics="true" pageId="23" pageNumber="174">d</emphasis>
|
||
of genu and tibia I (see below). As in body setae, with birefringent core extending into all barbs. Alveoli of tarsal setae dropshaped, pointed distally. Six setae eupathidial on tarsus I, including
|
||
<emphasis id="B964EABDFFF7FFCEFC0AFEB932C0F664" box="[938,949,261,284]" italics="true" pageId="23" pageNumber="174">s</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFF7FFCEFC66FEB93290F664" box="[966,997,261,284]" italics="true" pageId="23" pageNumber="174">m"</emphasis>
|
||
and pairs (
|
||
<emphasis id="B964EABDFFF7FFCEFBCBFEB9350FF664" box="[1131,1146,261,284]" italics="true" pageId="23" pageNumber="174">it</emphasis>
|
||
) and (
|
||
<emphasis id="B964EABDFFF7FFCEFB69FEB935A2F664" box="[1225,1239,261,284]" italics="true" pageId="23" pageNumber="174">p</emphasis>
|
||
). Famulus (
|
||
<emphasis id="B964EABDFFF7FFCEFAC5FEB93404F664" box="[1381,1393,261,284]" italics="true" pageId="23" pageNumber="174">e</emphasis>
|
||
) of tarsus I small (ca. 20 Μm) but conspicuous, being most proximodorsal setiform organ of segment and erect on small tubercle (
|
||
<figureCitation id="132B2A2AFFF7FFCEFEA0FEF0302AF61C" box="[256,351,332,357]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="23" pageNumber="174">
|
||
Figs
|
||
<date id="FFAE106FFFF7FFCEFE98FEF0302AF61C" box="[312,351,332,356]" pageId="23" pageNumber="174" value="2014-01">14I</date>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF7FFCEFECBFEF030E8F61D" box="[363,413,332,357]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="23" pageNumber="174">16C</figureCitation>
|
||
); with attenuate main tip and usually three simple or forked barbs clustered near mid-length (rarely fourth barb present); tip and barbs all open distally (
|
||
<figureCitation id="132B2A2AFFF7FFCEFCF5FED332C1F6F0" box="[853,948,367,392]" captionStart="FIGURE 14. A – I" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1661]" captionTargetId="figure@19.[151,1436,193,1663]" captionTargetPageId="19" captionText="FIGURE 14. A – I Nanohystrix hammerae n. sp., scanning electron micrographs of adult gnathosomal and leg elements (all with small surface debris artifacts). A, overview of proterosoma, ventral aspect; B, subcapitulum and chelicerae, ventral aspect; C, lateral aspect of gnathosoma and rostrum; D, enlargement of top-central region of B; E, enlargement of labrum, ventral aspect; F, distal part of labrum, anterolateral aspect; G, distal region of palp, lateral aspect; H, leg apotele and distal part of tarsus, near lateral aspect; I, famulus of tarsus I, lateral aspect, with insert showing open tip of hollow barb. Abbreviations: αf (abaxial fissure), c. fd (carina of fixed digit), Ch (chelicera); lg (labiogenal articulation), Ls (labrum); Ru (rutellum); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A); 20 Μm (B, C); 10 Μm (G, H); 5 Μm (D – F, I)." httpUri="https://zenodo.org/record/238779/files/figure.png" pageId="23" pageNumber="174">
|
||
Fig.
|
||
<date id="FFAE106FFFF7FFCEFC2DFECC32C1F6F0" box="[909,948,368,392]" pageId="23" pageNumber="174" value="2014-01">14I</date>
|
||
</figureCitation>
|
||
, insert). Famulus of tarsus II more distal, usually about mid-length on tarsus, smaller (ca. 10–12 Μm), strongly angled toward surface; variable in formattenuate to spiniform, sometimes with one small distal barb giving forked appearance (
|
||
<figureCitation id="132B2A2AFFF7FFCEFBCEFE0B3584F6A8" box="[1134,1265,439,464]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="23" pageNumber="174">Fig. 16F, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF7FFCEFF37FE41339FF286" blockId="23.[151,1306,507,532]" lastBlockId="23.[151,1435,1413,1598]" pageId="23" pageNumber="174">
|
||
<emphasis id="B964EABDFFF7FFCEFF37FE41307FF56C" bold="true" box="[151,266,509,532]" pageId="23" pageNumber="174">TABLE 1.</emphasis>
|
||
Ontogeny of setae (Roman letters) and solenidia (Greek letters) in
|
||
<taxonomicName id="4C104D2CFFF7FFCEFC77FE4035A4F56C" box="[983,1233,508,532]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF7FFCEFC77FE4035A4F56C" box="[983,1233,508,532]" italics="true" pageId="23" pageNumber="174">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF7FFCEFB77FE413466F56C" box="[1239,1299,509,532]" pageId="23" pageNumber="174" rank="species">
|
||
<emphasis id="B964EABDFFF7FFCEFB77FE413466F56C" bold="true" box="[1239,1299,509,532]" pageId="23" pageNumber="174">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
1 1Setae and solenidia are first added at the indicated instar (La, Pn, Dn, Tn, Ad = larva, protonymph, deutonymph, tritonymph, adult, respectively) and remain present in subsequent instars; dash indicates no addition. Setae in parentheses represent pseudosymmetrical pairs, brackets indicate physical change; for seta
|
||
<emphasis id="B964EABDFFF7FFCEFBDCFA7A35FCF2A6" box="[1148,1161,1478,1502]" italics="true" pageId="23" pageNumber="174">d</emphasis>
|
||
, subscripts
|
||
<emphasis id="B964EABDFFF7FFCEFAAEFA7B3454F2A6" box="[1294,1313,1479,1502]" italics="true" pageId="23" pageNumber="174">m</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFF7FFCEFA91FA7B344BF2A6" box="[1329,1342,1479,1502]" italics="true" pageId="23" pageNumber="174">n</emphasis>
|
||
indicate that the seta is minute or normally formed, respectively.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF7FFCEFE9FFD8C32B5F21C" pageId="23" pageNumber="174">
|
||
<table id="F910C40FFFF70026FF3FFD8C3405F21C" box="[159,1392,560,1380]" gridcols="5" gridrows="20" pageId="23" pageNumber="174">
|
||
<tr id="352034EDFFF70026FF3FFD8C3405F53E" box="[159,1392,560,582]" gridrow="0" pageId="23" pageNumber="174" rowspan-0="1">
|
||
<th id="76F15D91FFF70026FE9FFD8C334BF53E" box="[319,574,560,582]" gridcol="1" gridrow="0" pageId="23" pageNumber="174">Trochanter Femur</th>
|
||
<th id="76F15D91FFF70026FDD0FD8C33AFF53E" box="[624,730,560,582]" gridcol="2" gridrow="0" pageId="23" pageNumber="174">Genu</th>
|
||
<th id="76F15D91FFF70026FCA9FD8C32E5F53E" box="[777,912,560,582]" gridcol="3" gridrow="0" pageId="23" pageNumber="174">Tibia</th>
|
||
<th id="76F15D91FFF70026FC18FD8C3405F53E" box="[952,1392,560,582]" gridcol="4" gridrow="0" pageId="23" pageNumber="174">Tarsus</th>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFDE63405F517" box="[159,1392,602,623]" gridrow="1" pageId="23" pageNumber="174" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<th id="76F15D91FFF70026FF3FFDE63180F517" box="[159,245,602,623]" gridcol="0" gridrow="1" pageId="23" pageNumber="174">Leg I</th>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFD3E3405F5E0" box="[159,1392,642,664]" gridrow="2" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFD3E3180F5E0" box="[159,245,642,664]" gridcol="0" gridrow="2" pageId="23" pageNumber="174">La</th>
|
||
<td id="76F15D91FFF70026FE9FFD3E334BF5E0" box="[319,574,642,664]" gridcol="1" gridrow="2" pageId="23" pageNumber="174">- d, bv"</td>
|
||
<td id="76F15D91FFF70026FDD0FD3E33AFF5E0" box="[624,730,642,664]" gridcol="2" gridrow="2" pageId="23" pageNumber="174">(l), σ1, σ2</td>
|
||
<td id="76F15D91FFF70026FCA9FD3E32E5F5E0" box="[777,912,642,664]" gridcol="3" gridrow="2" pageId="23" pageNumber="174">d, c", l' (v), φ</td>
|
||
<td id="76F15D91FFF70026FC18FD3E3405F5E0" box="[952,1392,642,664]" gridcol="4" gridrow="2" pageId="23" pageNumber="174">(ft), (pl), (pv), (tc), (a), (u), (p), s, m", e, ω1</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFD103405F5BD" box="[159,1392,684,709]" gridrow="3" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFD103180F5BD" box="[159,245,684,709]" gridcol="0" gridrow="3" pageId="23" pageNumber="174">Pn</th>
|
||
<td id="76F15D91FFF70026FE9FFD10334BF5BD" box="[319,574,684,709]" gridcol="1" gridrow="3" pageId="23" pageNumber="174">v' -</td>
|
||
<td id="76F15D91FFF70026FDD0FD1033AFF5BD" box="[624,730,684,709]" gridcol="2" gridrow="3" pageId="23" pageNumber="174">d m, v"</td>
|
||
<td id="76F15D91FFF70026FCA9FD1032E5F5BD" box="[777,912,684,709]" gridcol="3" gridrow="3" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FD103405F5BD" box="[952,1392,684,709]" gridcol="4" gridrow="3" pageId="23" pageNumber="174">(it), m', ω2</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFD6B3405F595" box="[159,1392,727,749]" gridrow="4" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFD6B3180F595" box="[159,245,727,749]" gridcol="0" gridrow="4" pageId="23" pageNumber="174">Dn-Tn*</th>
|
||
<td id="76F15D91FFF70026FE9FFD6B334BF595" box="[319,574,727,749]" gridcol="1" gridrow="4" pageId="23" pageNumber="174">- (l) (v)</td>
|
||
<td id="76F15D91FFF70026FDD0FD6B33AFF595" box="[624,730,727,749]" gridcol="2" gridrow="4" pageId="23" pageNumber="174">v', σ3</td>
|
||
<td id="76F15D91FFF70026FCA9FD6B32E5F595" box="[777,912,727,749]" gridcol="3" gridrow="4" pageId="23" pageNumber="174">l"</td>
|
||
<td id="76F15D91FFF70026FC18FD6B3405F595" box="[952,1392,727,749]" gridcol="4" gridrow="4" pageId="23" pageNumber="174">ω3</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFCBD3405F46F" box="[159,1392,769,791]" gridrow="5" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFCBD3180F46F" box="[159,245,769,791]" gridcol="0" gridrow="5" pageId="23" pageNumber="174">Ad</th>
|
||
<td id="76F15D91FFF70026FE9FFCBD334BF46F" box="[319,574,769,791]" gridcol="1" gridrow="5" pageId="23" pageNumber="174">- -</td>
|
||
<td id="76F15D91FFF70026FDD0FCBD33AFF46F" box="[624,730,769,791]" gridcol="2" gridrow="5" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FCA9FCBD32E5F46F" box="[777,912,769,791]" gridcol="3" gridrow="5" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FCBD3405F46F" box="[952,1392,769,791]" gridcol="4" gridrow="5" pageId="23" pageNumber="174">-</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFC973405F438" box="[159,1392,811,832]" gridrow="6" pageId="23" pageNumber="174" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<th id="76F15D91FFF70026FF3FFC973180F438" box="[159,245,811,832]" gridcol="0" gridrow="6" pageId="23" pageNumber="174">Leg II</th>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFCE83405F415" box="[159,1392,852,877]" gridrow="7" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFCE83180F415" box="[159,245,852,877]" gridcol="0" gridrow="7" pageId="23" pageNumber="174">La</th>
|
||
<td id="76F15D91FFF70026FE9FFCE8334BF415" box="[319,574,852,877]" gridcol="1" gridrow="7" pageId="23" pageNumber="174">- d, bv", bx'</td>
|
||
<td id="76F15D91FFF70026FDD0FCE833AFF415" box="[624,730,852,877]" gridcol="2" gridrow="7" pageId="23" pageNumber="174">d m, (l), σ</td>
|
||
<td id="76F15D91FFF70026FCA9FCE832E5F415" box="[777,912,852,877]" gridcol="3" gridrow="7" pageId="23" pageNumber="174">d, c", l' (v), φ</td>
|
||
<td id="76F15D91FFF70026FC18FCE83405F415" box="[952,1392,852,877]" gridcol="4" gridrow="7" pageId="23" pageNumber="174">(ft), (pl), (pv), (tc), (a), (u), (p), s, e, ω1</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFCC33405F4E1" box="[159,1392,895,921]" gridrow="8" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFCC33180F4E1" box="[159,245,895,921]" gridcol="0" gridrow="8" pageId="23" pageNumber="174">Pn</th>
|
||
<td id="76F15D91FFF70026FE9FFCC3334BF4E1" box="[319,574,895,921]" gridcol="1" gridrow="8" pageId="23" pageNumber="174">v' -</td>
|
||
<td id="76F15D91FFF70026FDD0FCC333AFF4E1" box="[624,730,895,921]" gridcol="2" gridrow="8" pageId="23" pageNumber="174">v" [d n]</td>
|
||
<td id="76F15D91FFF70026FCA9FCC332E5F4E1" box="[777,912,895,921]" gridcol="3" gridrow="8" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FCC33405F4E1" box="[952,1392,895,921]" gridcol="4" gridrow="8" pageId="23" pageNumber="174">-</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFC173405F4B9" box="[159,1392,939,961]" gridrow="9" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFC173180F4B9" box="[159,245,939,961]" gridcol="0" gridrow="9" pageId="23" pageNumber="174">Dn-Tn*</th>
|
||
<td id="76F15D91FFF70026FE9FFC17334BF4B9" box="[319,574,939,961]" gridcol="1" gridrow="9" pageId="23" pageNumber="174">- (l), v"</td>
|
||
<td id="76F15D91FFF70026FDD0FC1733AFF4B9" box="[624,730,939,961]" gridcol="2" gridrow="9" pageId="23" pageNumber="174">v'</td>
|
||
<td id="76F15D91FFF70026FCA9FC1732E5F4B9" box="[777,912,939,961]" gridcol="3" gridrow="9" pageId="23" pageNumber="174">l"</td>
|
||
<td id="76F15D91FFF70026FC18FC173405F4B9" box="[952,1392,939,961]" gridcol="4" gridrow="9" pageId="23" pageNumber="174">(it), ω2</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFC683405F492" box="[159,1392,980,1002]" gridrow="10" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFC683180F492" box="[159,245,980,1002]" gridcol="0" gridrow="10" pageId="23" pageNumber="174">Ad</th>
|
||
<td id="76F15D91FFF70026FE9FFC68334BF492" box="[319,574,980,1002]" gridcol="1" gridrow="10" pageId="23" pageNumber="174">- -</td>
|
||
<td id="76F15D91FFF70026FDD0FC6833AFF492" box="[624,730,980,1002]" gridcol="2" gridrow="10" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FCA9FC6832E5F492" box="[777,912,980,1002]" gridcol="3" gridrow="10" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FC683405F492" box="[952,1392,980,1002]" gridcol="4" gridrow="10" pageId="23" pageNumber="174">-</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFC433405F36C" box="[159,1392,1023,1044]" gridrow="11" pageId="23" pageNumber="174" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<th id="76F15D91FFF70026FF3FFC433180F36C" box="[159,245,1023,1044]" gridcol="0" gridrow="11" pageId="23" pageNumber="174">Leg III</th>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFB9B3405F339" box="[159,1392,1063,1089]" gridrow="12" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFB9B3180F339" box="[159,245,1063,1089]" gridcol="0" gridrow="12" pageId="23" pageNumber="174">La</th>
|
||
<td id="76F15D91FFF70026FE9FFB9B334BF339" box="[319,574,1063,1089]" gridcol="1" gridrow="12" pageId="23" pageNumber="174">- d, ev'</td>
|
||
<td id="76F15D91FFF70026FDD0FB9B33AFF339" box="[624,730,1063,1089]" gridcol="2" gridrow="12" pageId="23" pageNumber="174">d m, l', σ</td>
|
||
<td id="76F15D91FFF70026FCA9FB9B32E5F339" box="[777,912,1063,1089]" gridcol="3" gridrow="12" pageId="23" pageNumber="174">d, l', (v), φ</td>
|
||
<td id="76F15D91FFF70026FC18FB9B3405F339" box="[952,1392,1063,1089]" gridcol="4" gridrow="12" pageId="23" pageNumber="174">(ft), pl', (pv), (tc), (a), (u), (p), s</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFBEF3405F315" box="[159,1392,1107,1133]" gridrow="13" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFBEF3180F315" box="[159,245,1107,1133]" gridcol="0" gridrow="13" pageId="23" pageNumber="174">Pn</th>
|
||
<td id="76F15D91FFF70026FE9FFBEF334BF315" box="[319,574,1107,1133]" gridcol="1" gridrow="13" pageId="23" pageNumber="174">v' -</td>
|
||
<td id="76F15D91FFF70026FDD0FBEF33AFF315" box="[624,730,1107,1133]" gridcol="2" gridrow="13" pageId="23" pageNumber="174">v" [d n]</td>
|
||
<td id="76F15D91FFF70026FCA9FBEF32E5F315" box="[777,912,1107,1133]" gridcol="3" gridrow="13" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FBEF3405F315" box="[952,1392,1107,1133]" gridcol="4" gridrow="13" pageId="23" pageNumber="174">-</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFBC33405F3ED" box="[159,1392,1151,1173]" gridrow="14" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFBC33180F3ED" box="[159,245,1151,1173]" gridcol="0" gridrow="14" pageId="23" pageNumber="174">Dn-Tn*</th>
|
||
<td id="76F15D91FFF70026FE9FFBC3334BF3ED" box="[319,574,1151,1173]" gridcol="1" gridrow="14" pageId="23" pageNumber="174">l' l', v"</td>
|
||
<td id="76F15D91FFF70026FDD0FBC333AFF3ED" box="[624,730,1151,1173]" gridcol="2" gridrow="14" pageId="23" pageNumber="174">v'</td>
|
||
<td id="76F15D91FFF70026FCA9FBC332E5F3ED" box="[777,912,1151,1173]" gridcol="3" gridrow="14" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FBC33405F3ED" box="[952,1392,1151,1173]" gridcol="4" gridrow="14" pageId="23" pageNumber="174">(it), pl"</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFB143405F3C6" box="[159,1392,1192,1214]" gridrow="15" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFB143180F3C6" box="[159,245,1192,1214]" gridcol="0" gridrow="15" pageId="23" pageNumber="174">Ad</th>
|
||
<td id="76F15D91FFF70026FE9FFB14334BF3C6" box="[319,574,1192,1214]" gridcol="1" gridrow="15" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FDD0FB1433AFF3C6" box="[624,730,1192,1214]" gridcol="2" gridrow="15" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FCA9FB1432E5F3C6" box="[777,912,1192,1214]" gridcol="3" gridrow="15" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FB143405F3C6" box="[952,1392,1192,1214]" gridcol="4" gridrow="15" pageId="23" pageNumber="174">-</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFB6F3405F390" box="[159,1392,1235,1256]" gridrow="16" pageId="23" pageNumber="174" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<th id="76F15D91FFF70026FF3FFB6F3180F390" box="[159,245,1235,1256]" gridcol="0" gridrow="16" pageId="23" pageNumber="174">Leg IV</th>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFB473405F269" box="[159,1392,1275,1297]" gridrow="17" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFB473180F269" box="[159,245,1275,1297]" gridcol="0" gridrow="17" pageId="23" pageNumber="174">Pn</th>
|
||
<td id="76F15D91FFF70026FE9FFB47334BF269" box="[319,574,1275,1297]" gridcol="1" gridrow="17" pageId="23" pageNumber="174">- -</td>
|
||
<td id="76F15D91FFF70026FDD0FB4733AFF269" box="[624,730,1275,1297]" gridcol="2" gridrow="17" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FCA9FB4732E5F269" box="[777,912,1275,1297]" gridcol="3" gridrow="17" pageId="23" pageNumber="174">v"</td>
|
||
<td id="76F15D91FFF70026FC18FB473405F269" box="[952,1392,1275,1297]" gridcol="4" gridrow="17" pageId="23" pageNumber="174">ft", (pv), (u), (p)</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFA983405F242" box="[159,1392,1316,1338]" gridrow="18" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFA983180F242" box="[159,245,1316,1338]" gridcol="0" gridrow="18" pageId="23" pageNumber="174">Dn-Tn*</th>
|
||
<td id="76F15D91FFF70026FE9FFA98334BF242" box="[319,574,1316,1338]" gridcol="1" gridrow="18" pageId="23" pageNumber="174">v', l' d, ev', l', v"</td>
|
||
<td id="76F15D91FFF70026FDD0FA9833AFF242" box="[624,730,1316,1338]" gridcol="2" gridrow="18" pageId="23" pageNumber="174">d, l', (v), σ</td>
|
||
<td id="76F15D91FFF70026FCA9FA9832E5F242" box="[777,912,1316,1338]" gridcol="3" gridrow="18" pageId="23" pageNumber="174">d, l', v', φ</td>
|
||
<td id="76F15D91FFF70026FC18FA983405F242" box="[952,1392,1316,1338]" gridcol="4" gridrow="18" pageId="23" pageNumber="174">ft', (tc), (pl), (a), s</td>
|
||
</tr>
|
||
<tr id="352034EDFFF70026FF3FFAF23405F21C" box="[159,1392,1358,1380]" gridrow="19" pageId="23" pageNumber="174">
|
||
<th id="76F15D91FFF70026FF3FFAF23180F21C" box="[159,245,1358,1380]" gridcol="0" gridrow="19" pageId="23" pageNumber="174">Ad</th>
|
||
<td id="76F15D91FFF70026FE9FFAF2334BF21C" box="[319,574,1358,1380]" gridcol="1" gridrow="19" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FDD0FAF233AFF21C" box="[624,730,1358,1380]" gridcol="2" gridrow="19" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FCA9FAF232E5F21C" box="[777,912,1358,1380]" gridcol="3" gridrow="19" pageId="23" pageNumber="174">-</td>
|
||
<td id="76F15D91FFF70026FC18FAF23405F21C" box="[952,1392,1358,1380]" gridcol="4" gridrow="19" pageId="23" pageNumber="174">-</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF7FFCEFF37F9BB3092F146" blockId="23.[151,1435,1413,1598]" pageId="23" pageNumber="174">* Setae in these rows are present in the tritonymph, but as no deutonymphs were studied they may have been added in either the deuto- or tritonymph.</paragraph>
|
||
<paragraph id="8BAF36AFFFF7FFCEFF67F9D735DFF06C" blockId="23.[151,1436,1643,2028]" pageId="23" pageNumber="174">
|
||
Disjunctions of tarsal setae as follows (' = anterior member more distal; "
|
||
<emphasis id="B964EABDFFF7FFCEFBA0F9D73565F1FC" bold="true" box="[1024,1040,1643,1668]" pageId="23" pageNumber="174">=</emphasis>
|
||
posterior member more distal; null = without noticeable disjunction): pair (
|
||
<emphasis id="B964EABDFFF7FFCEFDC1F9333305F1DF" box="[609,624,1679,1703]" italics="true" pageId="23" pageNumber="174">ft</emphasis>
|
||
) slightly ' on I, null on II, slightly " on III, IV; (
|
||
<emphasis id="B964EABDFFF7FFCEFB08F92C35C9F1DF" box="[1192,1212,1680,1703]" italics="true" pageId="23" pageNumber="174">tc</emphasis>
|
||
) strongly ' on I, II, weakly ' on III, weakly " on IV; (
|
||
<emphasis id="B964EABDFFF7FFCEFDB1F9083355F1B3" box="[529,544,1716,1739]" italics="true" pageId="23" pageNumber="174">it</emphasis>
|
||
) ' on I, II, almost null on III; (
|
||
<emphasis id="B964EABDFFF7FFCEFCDDF90832FEF1B3" box="[893,907,1716,1739]" italics="true" pageId="23" pageNumber="174">p</emphasis>
|
||
) null on all legs; (
|
||
<emphasis id="B964EABDFFF7FFCEFBFFF9083518F1B3" box="[1119,1133,1716,1739]" italics="true" pageId="23" pageNumber="174">u</emphasis>
|
||
) null on I–III, " on IV; (
|
||
<emphasis id="B964EABDFFF7FFCEFA24F90834E7F1B3" box="[1412,1426,1716,1739]" italics="true" pageId="23" pageNumber="174">a</emphasis>
|
||
) strongly ' on I, weakly ' or null on II, null to strongly " on III, IV; (
|
||
<emphasis id="B964EABDFFF7FFCEFCD4F96B32FFF188" box="[884,906,1751,1776]" italics="true" pageId="23" pageNumber="174">pl</emphasis>
|
||
) strongly ' on I–IV; (
|
||
<emphasis id="B964EABDFFF7FFCEFBD4F96435FBF197" box="[1140,1166,1752,1775]" italics="true" pageId="23" pageNumber="174">pv</emphasis>
|
||
) strongly " on I–IV; (
|
||
<emphasis id="B964EABDFFF7FFCEFADEF96434E7F197" box="[1406,1426,1752,1775]" italics="true" pageId="23" pageNumber="174">m</emphasis>
|
||
) strongly " on I. See Remark 16 for variation in disjunctions and Remark 17 for setal pair (
|
||
<emphasis id="B964EABDFFF7FFCEFB25F94035ECF06B" box="[1157,1177,1788,1811]" italics="true" pageId="23" pageNumber="174">m</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFF7FFC3FF67F8A330BEF0ED" blockId="23.[151,1436,1643,2028]" lastBlockId="26.[151,1436,1772,2013]" lastPageId="26" lastPageNumber="177" pageId="23" pageNumber="174">
|
||
Solenidial formulas: genua (3-1-1-1); tibiae (1-1-1-1); tarsi (3-2-0-0). Solenidia of tarsus I and all genua long, finely tapered (‘piliform’ in terminology of
|
||
<bibRefCitation id="EF814B5EFFF7FFCEFDDFF8FF3231F024" author="Grandjean" box="[639,836,1859,1884]" pageId="23" pageNumber="174" refString="Grandjean, F. (1935 b) Les poils et les organes sensitifs portes par les pattes et le palpe chez les Oribates. Bulletin de la Societe zoologique de France, 60, 6 - 39." type="journal article" year="1935" yearSuffix="b">Grandjean 1935b</bibRefCitation>
|
||
), similar in length to most leg setae. Solenidia of tibia and tarsus II distally tapered, but not finely so (‘ceratiform’). Tibia I solenidion φ longest (
|
||
<figureCitation id="132B2A2AFFF7FFCEFB13F8DB3469F0F8" box="[1203,1308,1895,1920]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="23" pageNumber="174">Fig. 16D</figureCitation>
|
||
), flagellate (‘tactile’), about 2/3 length of leg I. Seta
|
||
<emphasis id="B964EABDFFF7FFCEFDC8F8373303F0DC" box="[616,630,1931,1956]" italics="true" pageId="23" pageNumber="174">d</emphasis>
|
||
and respective solenidia with various relationships on genua and tibiae (
|
||
<figureCitation id="132B2A2AFFF7FFCEFF3FF8133182F0B0" box="[159,247,1967,1992]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="23" pageNumber="174">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFF7FFCEFEA5F8133051F0BF" box="[261,292,1967,1991]" captionStart="FIGURE 17. A – C" captionStartId="24.[151,250,1858,1880]" captionTargetBox="[157,1436,193,1823]" captionTargetId="figure@24.[151,1436,193,1837]" captionTargetPageId="24" captionText="FIGURE 17. A – C Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, left leg III, tarsus partially shown; B, left tarsus III; C, left leg IV (slightly twisted at base). Labels all indicate setiform organs (see text). Scale bars: 100 Μm." httpUri="https://zenodo.org/record/238782/files/figure.png" pageId="23" pageNumber="174">17</figureCitation>
|
||
): seta
|
||
<emphasis id="B964EABDFFF7FFCEFECFF8133008F0B0" box="[367,381,1967,1992]" italics="true" pageId="23" pageNumber="174">d</emphasis>
|
||
of genu I immediately adaxial to σ3, coupled in partially merged alveoli, minute (ca. 30–35 Μm), finely attenuate (
|
||
<figureCitation id="132B2A2AFFF7FFCEFE30F86F308DF094" box="[400,504,2003,2028]" captionStart="FIGURE 12. A – F" captionStartId="17.[151,250,775,797]" captionTargetBox="[151,1436,193,753]" captionTargetId="figure@17.[151,1436,193,754]" captionTargetPageId="17" captionText="FIGURE 12. A – F Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ 1 and coupled seta d; E, dorsolateral aspect of tibia I seta d (coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5 – 20 layers. Scale bars: 25 Μm (A – C, F); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238776/files/figure.png" pageId="23" pageNumber="174">Fig. 12D</figureCitation>
|
||
);
|
||
<emphasis id="B964EABDFFF7FFCEFDACF86F336FF094" box="[524,538,2003,2028]" italics="true" pageId="23" pageNumber="174">d</emphasis>
|
||
present but distantly proximal to solenidion on genua II–IV;
|
||
<emphasis id="B964EABDFFF7FFCEFB68F86F35A3F094" box="[1224,1238,2003,2028]" italics="true" pageId="23" pageNumber="174">d</emphasis>
|
||
of tibia I similar to that of genu, closely coupled adaxial to flagellate φ (
|
||
<figureCitation id="132B2A2AFFFAFFC3FCA5F950321BF07D" box="[773,878,1772,1797]" captionStart="FIGURE 12. A – F" captionStartId="17.[151,250,775,797]" captionTargetBox="[151,1436,193,753]" captionTargetId="figure@17.[151,1436,193,754]" captionTargetPageId="17" captionText="FIGURE 12. A – F Nanohystrix hammerae n. sp. adults, transmitted light compound microscope images except B (SEM). A, distal quarter of extended ovipositor with lobes spread, lateral aspect, anterior to right; B, similar, but distal lobes only slightly spread; C, spermatopositor, ventral aspect; D, lateral aspect of genu I solenidion σ 1 and coupled seta d; E, dorsolateral aspect of tibia I seta d (coupled solenidion removed); F, distal end of tarsus IV of male, showing potential for bending. All labels relate to setae (see text). Except B, all composites of 5 – 20 layers. Scale bars: 25 Μm (A – C, F); 10 Μm (D, E)." httpUri="https://zenodo.org/record/238776/files/figure.png" pageId="26" pageNumber="177">Figs 12E</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFFAFFC3FCDAF95032DBF07D" box="[890,942,1772,1797]" captionStart="FIGURE 16. A – G" captionStartId="21.[151,250,1739,1761]" captionTargetBox="[155,1431,195,1713]" captionTargetId="figure@21.[151,1436,193,1717]" captionTargetPageId="21" captionText="FIGURE 16. A – G Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, right leg I, tarsus partially shown; B, right tarsus I; C, enlargement of tarsus I famulus (e); D, tibia I solenidion φ 1 with minute coupled seta d; E, right leg II, tarsus partially shown; F, right tarsus; G, two variations of tarsus II famulus (e). Labels all indicate setiform organs (see text). Scale bars: 50 Μm (A, B, D – F); 5 Μm (C, G)." httpUri="https://zenodo.org/record/238781/files/figure.png" pageId="26" pageNumber="177">16D</figureCitation>
|
||
);
|
||
<emphasis id="B964EABDFFFAFFC3FC62F95032A5F07D" box="[962,976,1772,1797]" italics="true" pageId="26" pageNumber="177">d</emphasis>
|
||
of tibia II well posterior to φ;
|
||
<emphasis id="B964EABDFFFAFFC3FA89F9503442F07D" box="[1321,1335,1772,1797]" italics="true" pageId="26" pageNumber="177">d</emphasis>
|
||
of tibiae III and IV closely coupled to φ but proximal to solenidion and in separate alveolus (
|
||
<figureCitation id="132B2A2AFFFAFFC3FBE0F8AC35BDF051" box="[1088,1224,1808,1833]" captionStart="FIGURE 17. A – C" captionStartId="24.[151,250,1858,1880]" captionTargetBox="[157,1436,193,1823]" captionTargetId="figure@24.[151,1436,193,1837]" captionTargetPageId="24" captionText="FIGURE 17. A – C Nanohystrix hammerae n. sp., adult legs, abaxial aspect. A, left leg III, tarsus partially shown; B, left tarsus III; C, left leg IV (slightly twisted at base). Labels all indicate setiform organs (see text). Scale bars: 100 Μm." httpUri="https://zenodo.org/record/238782/files/figure.png" pageId="26" pageNumber="177">Fig. 17A, C</figureCitation>
|
||
). Solenidia ω1 and ω3 of tarsus I inserted near dorsal midline, ω1 loosely grouped with famulus and seta
|
||
<emphasis id="B964EABDFFFAFFC3FBCFF88935FFF035" box="[1135,1162,1845,1869]" italics="true" pageId="26" pageNumber="177">ft"</emphasis>
|
||
in basal quarter and ω3 located at about one-third segment length; ω2 with unusual distal position near level of iteral setae, almost ventral on abaxial face of segment.
|
||
</paragraph>
|
||
<caption id="DF6F6627FFF8FFC1FF37F8FE3593F00F" httpUri="https://zenodo.org/record/238782/files/figure.png" pageId="24" pageNumber="175" targetBox="[157,1436,193,1823]" targetPageId="24">
|
||
<paragraph id="8BAF36AFFFF8FFC1FF37F8FE3593F00F" blockId="24.[151,1436,1858,1911]" pageId="24" pageNumber="175">
|
||
<emphasis id="B964EABDFFF8FFC1FF37F8FE3023F020" bold="true" box="[151,342,1858,1880]" pageId="24" pageNumber="175">FIGURE 17. A–C</emphasis>
|
||
<taxonomicName id="4C104D2CFFF8FFC1FEFDF8FE333FF020" box="[349,586,1858,1880]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF8FFC1FEFDF8FE333FF020" box="[349,586,1858,1880]" italics="true" pageId="24" pageNumber="175">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF8FFC1FDF0F8FE33FCF02F" box="[592,649,1858,1879]" pageId="24" pageNumber="175" rank="species">
|
||
<emphasis id="B964EABDFFF8FFC1FDF0F8FE33FCF02F" bold="true" box="[592,649,1858,1879]" pageId="24" pageNumber="175">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult legs, abaxial aspect. A, left leg III, tarsus partially shown; B, left tarsus III; C, left leg IV (slightly twisted at base). Labels all indicate setiform organs (see text). Scale bars: 100 Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFF9FFC0FF37F9193329F00B" httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="25" pageNumber="176" targetBox="[157,1412,212,1665]" targetPageId="25">
|
||
<paragraph id="8BAF36AFFFF9FFC0FF37F9193329F00B" blockId="25.[151,1436,1701,1907]" pageId="25" pageNumber="176">
|
||
<emphasis id="B964EABDFFF9FFC0FF37F9193022F1C2" bold="true" box="[151,343,1701,1723]" pageId="25" pageNumber="176">FIGURE 18. A–K</emphasis>
|
||
<taxonomicName id="4C104D2CFFF9FFC0FEFEF919333EF1C3" box="[350,587,1701,1723]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="25" pageNumber="176" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFF9FFC0FEFEF919333EF1C3" box="[350,587,1701,1723]" italics="true" pageId="25" pageNumber="176">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFF9FFC0FDF0F91A33FCF1C3" box="[592,649,1702,1723]" pageId="25" pageNumber="176" rank="species">
|
||
<emphasis id="B964EABDFFF9FFC0FDF0F91A33FCF1C3" bold="true" box="[592,649,1702,1723]" pageId="25" pageNumber="176">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
juveniles: larva (A –I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta
|
||
<emphasis id="B964EABDFFF9FFC0FE5AF89C3372F04E" box="[506,519,1824,1846]" italics="true" pageId="25" pageNumber="176">d</emphasis>
|
||
broken in H, I. Abbreviations:
|
||
<emphasis id="B964EABDFFF9FFC0FCE5F89C321FF04E" box="[837,874,1824,1846]" italics="true" pageId="25" pageNumber="176">cd5</emphasis>
|
||
(cuticular disk),
|
||
<emphasis id="B964EABDFFF9FFC0FBB7F89C355EF04E" box="[1047,1067,1824,1846]" italics="true" pageId="25" pageNumber="176">ih</emphasis>
|
||
(lyrifissure),
|
||
<emphasis id="B964EABDFFF9FFC0FB15F89D35A1F04E" box="[1205,1236,1825,1846]" italics="true" pageId="25" pageNumber="176">t.e.</emphasis>
|
||
(vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I–K to same scale, except famulus enlargements in C, G).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6F6627FFFAFFC3FF37F9893459F1BD" httpUri="https://zenodo.org/record/238784/files/figure.png" pageId="26" pageNumber="177" targetBox="[151,1436,197,1564]" targetPageId="26">
|
||
<paragraph id="8BAF36AFFFFAFFC3FF37F9893459F1BD" blockId="26.[151,1436,1588,1733]" pageId="26" pageNumber="177">
|
||
<emphasis id="B964EABDFFFAFFC3FF37F989302FF132" bold="true" box="[151,346,1589,1611]" pageId="26" pageNumber="177">FIGURE 19. A–E</emphasis>
|
||
<taxonomicName id="4C104D2CFFFAFFC3FEC2F9883324F132" box="[354,593,1588,1610]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFFAFFC3FEC2F9883324F132" box="[354,593,1588,1610]" italics="true" pageId="26" pageNumber="177">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFFAFFC3FDFAF98933E0F132" box="[602,661,1589,1610]" pageId="26" pageNumber="177" rank="species">
|
||
<emphasis id="B964EABDFFFAFFC3FDFAF98933E0F132" bold="true" box="[602,661,1589,1610]" pageId="26" pageNumber="177">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
tritonymph (appendages and many setae incomplete). A, dorsal aspect; B, close-up of right humeral region of hysterosoma; C. lateral aspect; D, close-up of proterosomal coxisternum region (cf. C), some epimeral setae represented only by alveoli; E, ventral view of hysterosoma. Abbreviations:
|
||
<emphasis id="B964EABDFFFAFFC3FB20F9CE35CBF1F0" box="[1152,1214,1650,1672]" italics="true" pageId="26" pageNumber="177">cd1–5</emphasis>
|
||
(cuticular disks),
|
||
<emphasis id="B964EABDFFFAFFC3FAD9F9CF34E9F1F0" box="[1401,1436,1651,1672]" italics="true" pageId="26" pageNumber="177">cpc</emphasis>
|
||
(podocephalic canal),
|
||
<emphasis id="B964EABDFFFAFFC3FED8F92D30F9F1DE" box="[376,396,1681,1702]" italics="true" pageId="26" pageNumber="177">ia</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFAFFC3FE39F92D30C7F1DE" box="[409,434,1681,1702]" italics="true" pageId="26" pageNumber="177">im</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFAFFC3FE60F92D30A1F1DE" box="[448,468,1681,1702]" italics="true" pageId="26" pageNumber="177">ip</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFAFFC3FE41F92C336AF1DE" box="[481,543,1680,1702]" italics="true" pageId="26" pageNumber="177">ih, ips</emphasis>
|
||
(lyrifissures),
|
||
<emphasis id="B964EABDFFFAFFC3FD14F92C33ADF1DE" box="[692,728,1680,1702]" italics="true" pageId="26" pageNumber="177">sp2</emphasis>
|
||
(dorsal spine of epimere 2),
|
||
<emphasis id="B964EABDFFFAFFC3FC5FF92C3560F1DE" box="[1023,1045,1680,1702]" italics="true" pageId="26" pageNumber="177">Pf</emphasis>
|
||
(palp femur),
|
||
<emphasis id="B964EABDFFFAFFC3FB0AF92E35B2F1DF" box="[1194,1223,1682,1703]" italics="true" pageId="26" pageNumber="177">pK</emphasis>
|
||
(prodorsal condyle),
|
||
<emphasis id="B964EABDFFFAFFC3FF37F91331CCF1BD" box="[151,185,1711,1733]" italics="true" pageId="26" pageNumber="177">Tr1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFAFFC3FF67F90C319DF1BD" box="[199,232,1711,1733]" italics="true" pageId="26" pageNumber="177">Tr2</emphasis>
|
||
(trochanters of legs 1, 2); other labels indicate setae (see text). Scale bars: 100 Μm (A, C, E); 20 Μm (B, D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFFAFFC5FF67F81C32C4F665" blockId="26.[151,1436,1772,2013]" lastBlockId="28.[151,1437,151,2013]" lastPageId="28" lastPageNumber="179" pageId="26" pageNumber="177">
|
||
<emphasis id="B964EABDFFFAFFC3FF67F81C30A2F0C1" bold="true" box="[199,471,1952,1977]" pageId="26" pageNumber="177">
|
||
Ontogeny (
|
||
<figureCitation id="132B2A2AFFFAFFC3FEECF81C30BAF0C1" box="[332,463,1952,1977]" captionStart-0="FIGURE 18. A – K" captionStart-1="FIGURE 19. A – E" captionStart-2="FIGURE 20. A – I" captionStartId-0="25.[151,250,1701,1723]" captionStartId-1="26.[151,250,1589,1611]" captionStartId-2="27.[151,250,1753,1775]" captionTargetBox-0="[157,1412,212,1665]" captionTargetBox-1="[151,1436,197,1564]" captionTargetBox-2="[163,1406,193,1703]" captionTargetId-0="figure@25.[151,1436,193,1680]" captionTargetId-1="figure@26.[151,1436,193,1567]" captionTargetId-2="figure@27.[163,1407,193,1705]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." captionText-1="FIGURE 19. A – E Nanohystrix hammerae n. sp. tritonymph (appendages and many setae incomplete). A, dorsal aspect; B, close-up of right humeral region of hysterosoma; C. lateral aspect; D, close-up of proterosomal coxisternum region (cf. C), some epimeral setae represented only by alveoli; E, ventral view of hysterosoma. Abbreviations: cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip, ih, ips (lyrifissures), sp 2 (dorsal spine of epimere 2), Pf (palp femur), pK (prodorsal condyle), Tr 1, Tr 2 (trochanters of legs 1, 2); other labels indicate setae (see text). Scale bars: 100 Μm (A, C, E); 20 Μm (B, D)." captionText-2="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri-0="https://zenodo.org/record/238783/files/figure.png" httpUri-1="https://zenodo.org/record/238784/files/figure.png" httpUri-2="https://zenodo.org/record/238785/files/figure.png" pageId="26" pageNumber="177">Figs 18–20</figureCitation>
|
||
)
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B964EABDFFFAFFC3FE47F81E3307F0C1" box="[487,626,1954,1977]" italics="true" pageId="26" pageNumber="177">Facies, size.</emphasis>
|
||
All juvenile instars with general appearance of adult, including almost vertically oriented prodorsum, similar proportions of body regions, and presence of extremely long notogastral setae having proportions mostly similar to those of adult (
|
||
<figureCitation id="132B2A2AFFFCFFC5FCBAFF2B32D7F7C8" box="[794,930,151,176]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">Fig. 20A–E</figureCitation>
|
||
). Differ from adult in being mostly pale, with soft cuticle covered in fine granules; sclerotization and pigmentation more localized and more weakly developed than in adult, but still with fine pores. Body length (n = 1; not including setae or chelicerae) of larva 335 µm, protonymph 560 µm, tritonymph 900 µm (deutonymph not measured).
|
||
</paragraph>
|
||
<caption id="DF6F6627FFFBFFC2FF37F965355FF09B" httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="27" pageNumber="178" targetBox="[163,1406,193,1703]" targetPageId="27">
|
||
<paragraph id="8BAF36AFFFFBFFC2FF37F965355FF09B" blockId="27.[151,1437,1753,2020]" pageId="27" pageNumber="178">
|
||
<emphasis id="B964EABDFFFBFFC2FF37F9653027F196" bold="true" box="[151,338,1753,1775]" pageId="27" pageNumber="178">FIGURE 20. A–I</emphasis>
|
||
<taxonomicName id="4C104D2CFFFBFFC2FEFBF965333FF197" box="[347,586,1753,1775]" class="Arachnida" family="Trichthoniidae" genus="Nanohystrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="hammerae" status="sp. nov.">
|
||
<emphasis id="B964EABDFFFBFFC2FEFBF965333FF197" box="[347,586,1753,1775]" italics="true" pageId="27" pageNumber="178">Nanohystrix hammerae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757C6FFFBFFC2FDF3F96533FBF196" box="[595,654,1753,1774]" pageId="27" pageNumber="178" rank="species">
|
||
<emphasis id="B964EABDFFFBFFC2FDF3F96533FBF196" bold="true" box="[595,654,1753,1774]" pageId="27" pageNumber="178">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
juveniles, stereo- (A–C) and compound (D–I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22G),with insert showing enlargement of prodorsal strut; F, tritonymph, seta
|
||
<emphasis id="B964EABDFFFBFFC2FD6CF8EF3391F011" box="[716,740,1875,1897]" italics="true" pageId="27" pageNumber="178">e1</emphasis>
|
||
and cuticular disk
|
||
<emphasis id="B964EABDFFFBFFC2FC62F8EF3292F011" box="[962,999,1875,1897]" italics="true" pageId="27" pageNumber="178">cd4</emphasis>
|
||
, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta
|
||
<emphasis id="B964EABDFFFBFFC2FE3AF82D30C6F0DF" box="[410,435,1937,1959]" italics="true" pageId="27" pageNumber="178">ft"</emphasis>
|
||
torn away). Abbreviations:
|
||
<emphasis id="B964EABDFFFBFFC2FD43F82D327DF0DF" box="[739,776,1937,1959]" italics="true" pageId="27" pageNumber="178">cd4</emphasis>
|
||
(cuticular disk),
|
||
<emphasis id="B964EABDFFFBFFC2FC63F82D3574F0DF" box="[963,1025,1937,1959]" italics="true" pageId="27" pageNumber="178">e1.mu</emphasis>
|
||
(depressor muscles of seta
|
||
<emphasis id="B964EABDFFFBFFC2FA90F82D343DF0DF" box="[1328,1352,1937,1959]" italics="true" pageId="27" pageNumber="178">e1</emphasis>
|
||
);
|
||
<emphasis id="B964EABDFFFBFFC2FAC3F82E34E9F0DF" box="[1379,1436,1938,1959]" italics="true" pageId="27" pageNumber="178">pr.mu</emphasis>
|
||
(prodorsal retractor muscles),
|
||
<emphasis id="B964EABDFFFBFFC2FE6BF80D3090F0BE" box="[459,485,1969,1990]" italics="true" pageId="27" pageNumber="178">str</emphasis>
|
||
(prodorsal strut); other labels indicate setae (see text). D–I composites of 5–20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F–I). Scale not determined for C, probably similar to A.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF36AFFFFCFFC5FF67FE9B3042F560" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="179">
|
||
<emphasis id="B964EABDFFFCFFC5FF67FE9B3039F638" box="[199,332,295,320]" italics="true" pageId="28" pageNumber="179">Prodorsum.</emphasis>
|
||
Aspis pale, sclerotization hardly discernable in early instars, gradually becoming more noticeable and lightly pigmented in deuto- and tritonymph (cf.
|
||
<figureCitation id="132B2A2AFFFCFFC5FD7DFEF032F6F61C" box="[733,899,332,357]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">Fig. 20A, C, E</figureCitation>
|
||
). Structured as in adult, except vertical occipital region gradually decreasing in degree of sclerotization from top, and without nuchal constriction (
|
||
<figureCitation id="132B2A2AFFFCFFC5FAA2FED334FAF6F0" box="[1282,1423,367,392]" captionStart="FIGURE 19. A – E" captionStartId="26.[151,250,1589,1611]" captionTargetBox="[151,1436,197,1564]" captionTargetId="figure@26.[151,1436,193,1567]" captionTargetPageId="26" captionText="FIGURE 19. A – E Nanohystrix hammerae n. sp. tritonymph (appendages and many setae incomplete). A, dorsal aspect; B, close-up of right humeral region of hysterosoma; C. lateral aspect; D, close-up of proterosomal coxisternum region (cf. C), some epimeral setae represented only by alveoli; E, ventral view of hysterosoma. Abbreviations: cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip, ih, ips (lyrifissures), sp 2 (dorsal spine of epimere 2), Pf (palp femur), pK (prodorsal condyle), Tr 1, Tr 2 (trochanters of legs 1, 2); other labels indicate setae (see text). Scale bars: 100 Μm (A, C, E); 20 Μm (B, D)." httpUri="https://zenodo.org/record/238784/files/figure.png" pageId="28" pageNumber="179">Fig. 19A, C</figureCitation>
|
||
). Without lateral carina
|
||
<emphasis id="B964EABDFFFCFFC5FE3CFE2830B1F6D4" box="[412,452,404,428]" italics="true" pageId="28" pageNumber="179">ca2</emphasis>
|
||
; sclerotization in region below
|
||
<emphasis id="B964EABDFFFCFFC5FC9AFE283217F6D4" box="[826,866,404,428]" italics="true" pageId="28" pageNumber="179">ca1</emphasis>
|
||
hardly discernable, except for strut (
|
||
<emphasis id="B964EABDFFFCFFC5FAB5FE29345DF6D4" box="[1301,1320,405,428]" italics="true" pageId="28" pageNumber="179">st</emphasis>
|
||
= ‘lateral nervure’) ending in strong condyle
|
||
<emphasis id="B964EABDFFFCFFC5FD92FE053326F6A8" box="[562,595,441,464]" italics="true" pageId="28" pageNumber="179">pK</emphasis>
|
||
(
|
||
<figureCitation id="132B2A2AFFFCFFC5FDC5FE0B33B9F6A8" box="[613,716,439,464]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">Fig. 20E</figureCitation>
|
||
, insert). Eyes as in adult except pigmentation not observed in lateral eyes (possibly lost in specimen preparation). Prodorsal setae generally similar in form and distribution to those of adult.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFFCFFC5FF67FD9935A9F4F8" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="179">
|
||
<emphasis id="B964EABDFFFCFFC5FF67FD9930D5F544" box="[199,416,549,572]" italics="true" pageId="28" pageNumber="179">Gastronotic region</emphasis>
|
||
. Simple in structure, without general sclerotization or transverse scissures; noticeable sclerotization limited to tubercles at base of individual setae (
|
||
<figureCitation id="132B2A2AFFFCFFC5FCE1FDFB32BEF518" box="[833,971,583,608]" captionStart="FIGURE 19. A – E" captionStartId="26.[151,250,1589,1611]" captionTargetBox="[151,1436,197,1564]" captionTargetId="figure@26.[151,1436,193,1567]" captionTargetPageId="26" captionText="FIGURE 19. A – E Nanohystrix hammerae n. sp. tritonymph (appendages and many setae incomplete). A, dorsal aspect; B, close-up of right humeral region of hysterosoma; C. lateral aspect; D, close-up of proterosomal coxisternum region (cf. C), some epimeral setae represented only by alveoli; E, ventral view of hysterosoma. Abbreviations: cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip, ih, ips (lyrifissures), sp 2 (dorsal spine of epimere 2), Pf (palp femur), pK (prodorsal condyle), Tr 1, Tr 2 (trochanters of legs 1, 2); other labels indicate setae (see text). Scale bars: 100 Μm (A, C, E); 20 Μm (B, D)." httpUri="https://zenodo.org/record/238784/files/figure.png" pageId="28" pageNumber="179">Fig. 19A–C</figureCitation>
|
||
); in nymphs only setae
|
||
<emphasis id="B964EABDFFFCFFC5FB6FFDF4359CF518" box="[1231,1257,584,608]" italics="true" pageId="28" pageNumber="179">c3</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFFCFFC5FABFFDF5344FF518" box="[1311,1338,585,608]" italics="true" pageId="28" pageNumber="179">cp</emphasis>
|
||
on same sclerite (not examined in larva). Soft cuticle expansive and finely granulate. Cuticular disks well delineated: in tritonymph similar in size and distribution to those of adult; not studied in other instars, except posterior two large disks present in larva with same relative position as in adult. Distribution and form of setae generally similar to those of adult, except
|
||
<emphasis id="B964EABDFFFCFFC5FE3CFD6430C7F588" box="[412,434,728,752]" italics="true" pageId="28" pageNumber="179">f1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FE64FD6B3095F588" box="[452,480,727,752]" italics="true" pageId="28" pageNumber="179">h2</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FE52FD6B337BF588" box="[498,526,727,752]" italics="true" pageId="28" pageNumber="179">h3</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FD80FD64334EF588" box="[544,571,728,752]" italics="true" pageId="28" pageNumber="179">p2</emphasis>
|
||
with more normal positions relative to surrounding setae (not laterally or anteriorly displaced; cf.
|
||
<figureCitation id="132B2A2AFFFCFFC5FE0AFD403370F46C" box="[426,517,764,789]" captionStart="FIGURE 2. A – E" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[161,1425,198,1839]" captionTargetId="figure@7.[151,1436,193,1844]" captionTargetPageId="7" captionText="FIGURE 2. A – E Nanohystrix hammerae n. sp., adult female, dorsal aspect. A, habitus of fully distended specimen (legs and some setae incompletely drawn for overall clarity); B, enlargement of same (larger setae abbreviated, see Fig. 9 A); C, left side of anterior notogastral scissure (arrow at midline, setae indicated only by alveolus on basal tubercles); D, same view of middle scissure, showing extensive pale cuticle and disk cd 4; E, entire posterior scissure and cuticular disks cd 5 (posterodorsal aspect). Abbreviations: bng (border of notogaster), cd 1 – 5 (cuticular disks), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (C – E to same scale)." httpUri="https://zenodo.org/record/238766/files/figure.png" pageId="28" pageNumber="179">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFFCFFC5FDB1FD403342F46C" box="[529,567,764,788]" captionStart="FIGURE 3. A – D" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[193,1407,191,1840]" captionTargetId="figure@8.[189,1413,180,1840]" captionTargetPageId="8" captionText="FIGURE 3. A – D Nanohystrix hammerae n. sp. A, lateral aspect of fully distended female (legs omitted, some setae abbreviated or represented only by basal tubercles); B, sam e, genital region (with plate and genital papillae) and fully extended ovipositor, anterior to right; C, ovipositor, end-view (anterior to top); D, ventrolateral aspect of male genital region, showing spermatopositor in almost lateral aspect. Abbreviations: ca 1, ca 2 (prodorsal carinae), cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip (lyrifissures), nc (nuchal constriction), ti (tendon insertions); other labels indicate setae (see text). Scale bars: 100 Μm (A, B); 50 Μm (C, D)." httpUri="https://zenodo.org/record/238767/files/figure.png" pageId="28" pageNumber="179">3A</figureCitation>
|
||
); seta
|
||
<emphasis id="B964EABDFFFCFFC5FD20FD4033EFF46C" box="[640,666,764,788]" italics="true" pageId="28" pageNumber="179">e1</emphasis>
|
||
proportionally longer in larva (
|
||
<figureCitation id="132B2A2AFFFCFFC5FC5FFD403513F46C" box="[1023,1126,764,789]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">Fig. 20A</figureCitation>
|
||
) than adult (2 times vs. 1.6 times body length, respectively). Setae
|
||
<emphasis id="B964EABDFFFCFFC5FDF1FC9C331EF440" box="[593,619,800,824]" italics="true" pageId="28" pageNumber="179">e1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FDDAFC9C33E1F440" box="[634,660,800,824]" italics="true" pageId="28" pageNumber="179">e2</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FD02FC9C33CDF440" box="[674,696,800,824]" italics="true" pageId="28" pageNumber="179">f1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FD66FC9C33A9F440" box="[710,732,800,824]" italics="true" pageId="28" pageNumber="179">f2</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FD4AFCA33273F440" box="[746,774,799,824]" italics="true" pageId="28" pageNumber="179">h1</emphasis>
|
||
,
|
||
<emphasis id="B964EABDFFFCFFC5FCB4FCA33244F440" box="[788,817,799,824]" italics="true" pageId="28" pageNumber="179">h2</emphasis>
|
||
apparently erectile (held horizontally in larva but erect in studied nymphs; cf.
|
||
<figureCitation id="132B2A2AFFFCFFC5FE3DFCF83355F424" box="[413,544,836,861]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">Fig 20A, E</figureCitation>
|
||
); tritonymph with muscles attaching to tubercles of presumed erectile setae in same pattern as in adults (
|
||
<figureCitation id="132B2A2AFFFCFFC5FE19FCDB3354F4F8" box="[441,545,871,896]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">Fig. 20D</figureCitation>
|
||
; see Discussion). Files of sclerotized tendon insertions absent.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFFCFFC5FF67FC303365F3D8" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="179">
|
||
<emphasis id="B964EABDFFFCFFC5FF67FC3030D4F4DC" box="[199,417,908,933]" italics="true" pageId="28" pageNumber="179">Coxisternal region.</emphasis>
|
||
Incompletely studied. Larva (
|
||
<figureCitation id="132B2A2AFFFCFFC5FD57FC30322BF4DC" box="[759,862,908,933]" captionStart="FIGURE 18. A – K" captionStartId="25.[151,250,1701,1723]" captionTargetBox="[157,1412,212,1665]" captionTargetId="figure@25.[151,1436,193,1680]" captionTargetPageId="25" captionText="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="28" pageNumber="179">Fig. 18A</figureCitation>
|
||
) with weakly defined epimeral plates I–III similar in form to those of adult: epimere I with wide separation between paired plates, epimere II with narrow separation, epimere III without clear separation. Vestige of leg IV (
|
||
<emphasis id="B964EABDFFFCFFC5FCADFC69325AF494" box="[781,815,981,1004]" italics="true" pageId="28" pageNumber="179">t.e.</emphasis>
|
||
) present posterior to epimere III, in form of dome-like swelling (see Remark 12). Coxisternal setation (I to III) 3-1-2; seta
|
||
<emphasis id="B964EABDFFFCFFC5FC12FC4432B8F368" box="[946,973,1016,1040]" italics="true" pageId="28" pageNumber="179">1c</emphasis>
|
||
modified as scale covering Claparède’s organ;
|
||
<emphasis id="B964EABDFFFCFFC5FF45FBA03077F34C" box="[229,258,1052,1076]" italics="true" pageId="28" pageNumber="179">1a</emphasis>
|
||
and
|
||
<emphasis id="B964EABDFFFCFFC5FE99FBA03020F34C" box="[313,341,1052,1076]" italics="true" pageId="28" pageNumber="179">3a</emphasis>
|
||
shortest,
|
||
<emphasis id="B964EABDFFFCFFC5FE62FBA030ABF34D" box="[450,478,1052,1077]" italics="true" pageId="28" pageNumber="179">3b</emphasis>
|
||
longest; most with noticeable barbs, those of
|
||
<emphasis id="B964EABDFFFCFFC5FC43FBA0328AF34C" box="[995,1023,1052,1076]" italics="true" pageId="28" pageNumber="179">3a</emphasis>
|
||
minute. Tritonymph (
|
||
<figureCitation id="132B2A2AFFFCFFC5FB57FBA0342EF34C" box="[1271,1371,1052,1077]" captionStart="FIGURE 19. A – E" captionStartId="26.[151,250,1589,1611]" captionTargetBox="[151,1436,197,1564]" captionTargetId="figure@26.[151,1436,193,1567]" captionTargetPageId="26" captionText="FIGURE 19. A – E Nanohystrix hammerae n. sp. tritonymph (appendages and many setae incomplete). A, dorsal aspect; B, close-up of right humeral region of hysterosoma; C. lateral aspect; D, close-up of proterosomal coxisternum region (cf. C), some epimeral setae represented only by alveoli; E, ventral view of hysterosoma. Abbreviations: cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip, ih, ips (lyrifissures), sp 2 (dorsal spine of epimere 2), Pf (palp femur), pK (prodorsal condyle), Tr 1, Tr 2 (trochanters of legs 1, 2); other labels indicate setae (see text). Scale bars: 100 Μm (A, C, E); 20 Μm (B, D)." httpUri="https://zenodo.org/record/238784/files/figure.png" pageId="28" pageNumber="179">Fig. 19E</figureCitation>
|
||
) with epimeres structured as in adult but more weakly sclerotized; setation as in adult (3-1-3-4). Supracoxal region as in adult, except seta
|
||
<emphasis id="B964EABDFFFCFFC5FEC3FBD9300CF304" box="[355,377,1125,1148]" italics="true" pageId="28" pageNumber="179">eI</emphasis>
|
||
proportionally larger in larva (
|
||
<figureCitation id="132B2A2AFFFCFFC5FD7AFBD83234F304" box="[730,833,1124,1149]" captionStart="FIGURE 18. A – K" captionStartId="25.[151,250,1701,1723]" captionTargetBox="[157,1412,212,1665]" captionTargetId="figure@25.[151,1436,193,1680]" captionTargetPageId="25" captionText="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="28" pageNumber="179">Fig. 18B</figureCitation>
|
||
); spine of epimere 2 present in tritonymph (
|
||
<emphasis id="B964EABDFFFCFFC5FA99FBD83415F304" box="[1337,1376,1124,1148]" italics="true" pageId="28" pageNumber="179">sp2</emphasis>
|
||
;
|
||
<figureCitation id="132B2A2AFFFCFFC5FAD0FBD831BFF3D8" captionStart="FIGURE 19. A – E" captionStartId="26.[151,250,1589,1611]" captionTargetBox="[151,1436,197,1564]" captionTargetId="figure@26.[151,1436,193,1567]" captionTargetPageId="26" captionText="FIGURE 19. A – E Nanohystrix hammerae n. sp. tritonymph (appendages and many setae incomplete). A, dorsal aspect; B, close-up of right humeral region of hysterosoma; C. lateral aspect; D, close-up of proterosomal coxisternum region (cf. C), some epimeral setae represented only by alveoli; E, ventral view of hysterosoma. Abbreviations: cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip, ih, ips (lyrifissures), sp 2 (dorsal spine of epimere 2), Pf (palp femur), pK (prodorsal condyle), Tr 1, Tr 2 (trochanters of legs 1, 2); other labels indicate setae (see text). Scale bars: 100 Μm (A, C, E); 20 Μm (B, D)." httpUri="https://zenodo.org/record/238784/files/figure.png" pageId="28" pageNumber="179">Fig. 19D</figureCitation>
|
||
; not studied in other instars).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFFCFFC5FF67FB1030C7F29D" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="179">
|
||
<emphasis id="B964EABDFFFCFFC5FF67FB1030E7F3BC" box="[199,402,1196,1221]" italics="true" pageId="28" pageNumber="179">Anogenital region</emphasis>
|
||
. Ontogeny of genital papillae normal; larva without features in presumptive genital region; protonymph with single pair of genital setae on valves; tritonymph (
|
||
<figureCitation id="132B2A2AFFFCFFC5FC2FFB733284F390" box="[911,1009,1231,1256]" captionStart="FIGURE 19. A – E" captionStartId="26.[151,250,1589,1611]" captionTargetBox="[151,1436,197,1564]" captionTargetId="figure@26.[151,1436,193,1567]" captionTargetPageId="26" captionText="FIGURE 19. A – E Nanohystrix hammerae n. sp. tritonymph (appendages and many setae incomplete). A, dorsal aspect; B, close-up of right humeral region of hysterosoma; C. lateral aspect; D, close-up of proterosomal coxisternum region (cf. C), some epimeral setae represented only by alveoli; E, ventral view of hysterosoma. Abbreviations: cd 1 – 5 (cuticular disks), cpc (podocephalic canal), ia, im, ip, ih, ips (lyrifissures), sp 2 (dorsal spine of epimere 2), Pf (palp femur), pK (prodorsal condyle), Tr 1, Tr 2 (trochanters of legs 1, 2); other labels indicate setae (see text). Scale bars: 100 Μm (A, C, E); 20 Μm (B, D)." httpUri="https://zenodo.org/record/238784/files/figure.png" pageId="28" pageNumber="179">Fig. 19E</figureCitation>
|
||
) with seven pairs, three in lateral row, four in medial row (posterior two absent); not examined in deutonymph. Larva (
|
||
<figureCitation id="132B2A2AFFFCFFC5FBEEFB4835CDF274" box="[1102,1208,1268,1293]" captionStart="FIGURE 18. A – K" captionStartId="25.[151,250,1701,1723]" captionTargetBox="[157,1412,212,1665]" captionTargetId="figure@25.[151,1436,193,1680]" captionTargetPageId="25" captionText="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="28" pageNumber="179">Fig. 18A</figureCitation>
|
||
) with four pairs of pseudanal setae:
|
||
<emphasis id="B964EABDFFFCFFC5FEFCFAA430D7F248" box="[348,418,1304,1328]" italics="true" pageId="28" pageNumber="179">p2–p4</emphasis>
|
||
similar to epimeral setae in size but with fewer barbs,
|
||
<emphasis id="B964EABDFFFCFFC5FB91FAA43539F248" box="[1073,1100,1304,1328]" italics="true" pageId="28" pageNumber="179">p4</emphasis>
|
||
shortest;
|
||
<emphasis id="B964EABDFFFCFFC5FB61FAA435A8F248" box="[1217,1245,1304,1328]" italics="true" pageId="28" pageNumber="179">p1</emphasis>
|
||
large, diverging distally, longer than pseudanal valves; seta
|
||
<emphasis id="B964EABDFFFCFFC5FDDCFA8033EDF22C" box="[636,664,1340,1364]" italics="true" pageId="28" pageNumber="179">p4</emphasis>
|
||
(inguinal seta) lost in protonymph. Larva without inguinal seta of
|
||
<emphasis id="B964EABDFFFCFFC5FA24FA8034E7F22D" box="[1412,1426,1340,1365]" italics="true" pageId="28" pageNumber="179">h</emphasis>
|
||
- row;
|
||
<emphasis id="B964EABDFFFCFFC5FF71FAE33198F200" box="[209,237,1375,1400]" italics="true" pageId="28" pageNumber="179">h3</emphasis>
|
||
well formed. Cupule
|
||
<emphasis id="B964EABDFFFCFFC5FE41FAE33082F200" box="[481,503,1375,1400]" italics="true" pageId="28" pageNumber="179">ih</emphasis>
|
||
close to pseudanal valve in larva. Anal region of tritonymph similar to adult except for lack of noticeable sclerotization; peranal segment fully formed as well-delineated paraproctal valves; with small seta
|
||
<emphasis id="B964EABDFFFCFFC5FEB2FA15305BF2B8" box="[274,302,1449,1472]" italics="true" pageId="28" pageNumber="179">pa</emphasis>
|
||
and small preanal and postanal apodemes extending inward from ends of peranal valves and with similar attached muscles.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFFCFFC5FF67FA533534F10D" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="179">
|
||
<emphasis id="B964EABDFFFCFFC5FF67FA533028F170" box="[199,349,1519,1544]" italics="true" pageId="28" pageNumber="179">Gnathosoma.</emphasis>
|
||
Subcapitulum stenarthric in all juvenile instars; seta
|
||
<emphasis id="B964EABDFFFCFFC5FC63FA4D32A4F170" box="[963,977,1521,1544]" italics="true" pageId="28" pageNumber="179">n</emphasis>
|
||
of gena added in deuto- or tritonymph; adoral seta
|
||
<emphasis id="B964EABDFFFCFFC5FEBEF9A83031F154" box="[286,324,1556,1580]" italics="true" pageId="28" pageNumber="179">or3</emphasis>
|
||
added in protonymph. Palp seta
|
||
<emphasis id="B964EABDFFFCFFC5FD6EF9A83399F154" box="[718,748,1556,1580]" italics="true" pageId="28" pageNumber="179">inf</emphasis>
|
||
added in deuto- or tritonymph; setation of other segments similar in all post-larval instars, except in protonymph (and probably larva) terminal eupathidia
|
||
<emphasis id="B964EABDFFFCFFC5FB6CF98B347AF128" box="[1228,1295,1591,1616]" italics="true" pageId="28" pageNumber="179">ul', su</emphasis>
|
||
not fused as far distad as in adult. Chelicera as in adult except for generally lesser sclerotization.
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFFCFFC4FF67F93D35E4F780" blockId="28.[151,1437,151,2013]" lastBlockId="29.[151,1436,151,1040]" lastPageId="29" lastPageNumber="180" pageId="28" pageNumber="179">
|
||
<emphasis id="B964EABDFFFCFFC5FF67F93D3189F1E0" box="[199,252,1665,1688]" italics="true" pageId="28" pageNumber="179">Legs</emphasis>
|
||
. Pretarsi of all legs monodactylous in larva and protonymph (
|
||
<figureCitation id="132B2A2AFFFCFFC5FC16F9C3357BF1E0" box="[950,1038,1663,1688]" captionStart="FIGURE 18. A – K" captionStartId="25.[151,250,1701,1723]" captionTargetBox="[157,1412,212,1665]" captionTargetId="figure@25.[151,1436,193,1680]" captionTargetPageId="25" captionText="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="28" pageNumber="179">Figs 18</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A2AFFFCFFC5FBBAF93C3515F1E0" box="[1050,1120,1664,1688]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">20G, I</figureCitation>
|
||
), tridactylous in tritonymph (
|
||
<figureCitation id="132B2A2AFFFCFFC5FF3FF918307FF1C4" box="[159,266,1700,1725]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">Fig. 20H</figureCitation>
|
||
), not recorded for deutonymph (see Remark 13). Setal ontogeny (unknown for deutonymph) given in Table 1. Larva with femur II seta
|
||
<emphasis id="B964EABDFFFCFFC5FDB2F97B3346F198" box="[530,563,1735,1760]" italics="true" pageId="28" pageNumber="179">bx'</emphasis>
|
||
present (see Remark 21); famulus I with single barb. Genu ontogeny unusual (see Remarks 23–25): seta
|
||
<emphasis id="B964EABDFFFCFFC5FE6CF95030AFF07D" box="[460,474,1772,1797]" italics="true" pageId="28" pageNumber="179">d</emphasis>
|
||
absent from genu I of larva (
|
||
<figureCitation id="132B2A2AFFFCFFC5FC8BF95032E1F07C" box="[811,916,1772,1797]" captionStart="FIGURE 18. A – K" captionStartId="25.[151,250,1701,1723]" captionTargetBox="[157,1412,212,1665]" captionTargetId="figure@25.[151,1436,193,1680]" captionTargetPageId="25" captionText="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="28" pageNumber="179">Fig. 18D</figureCitation>
|
||
), forming as minute seta in protonymph (
|
||
<figureCitation id="132B2A2AFFFCFFC5FACFF95031B4F050" captionStart="FIGURE 18. A – K" captionStartId="25.[151,250,1701,1723]" captionTargetBox="[157,1412,212,1665]" captionTargetId="figure@25.[151,1436,193,1680]" captionTargetPageId="25" captionText="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="28" pageNumber="179">Fig. 18J</figureCitation>
|
||
), but distant from any solenidion until formation of σ
|
||
<quantity id="4CE89B4AFFFCFFC5FCBBF8AC3230F050" box="[795,837,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="28" pageNumber="179" unit="in" value="3.0">3 in</quantity>
|
||
deuto- or tritonymph;
|
||
<emphasis id="B964EABDFFFCFFC5FBE6F8B33521F050" box="[1094,1108,1807,1832]" italics="true" pageId="28" pageNumber="179">d</emphasis>
|
||
of genua II and III minute in larva (Fig, 18F), becoming normal, barbed in protonymph (
|
||
<figureCitation id="132B2A2AFFFCFFC5FCEFF88832CDF034" box="[847,952,1844,1869]" captionStart="FIGURE 18. A – K" captionStartId="25.[151,250,1701,1723]" captionTargetBox="[157,1412,212,1665]" captionTargetId="figure@25.[151,1436,193,1680]" captionTargetPageId="25" captionText="FIGURE 18. A – K Nanohystrix hammerae n. sp. juveniles: larva (A – I), protonymph (J, K). A, venter, legs partially shown; B, right leg I, abaxial aspect, tarsus simplified; C, right tarsus I, abaxial aspect, slightly rotated dorsally; D, right genu and tibia I, dorsal aspect; E, right leg II, abaxial aspect, tarsus simplified; F, right genu II, dorsal aspect; G, right tarsus II, abaxial aspect; H, left leg III, abaxial aspect, tarsus simplified; I, left tarsus III, abaxial aspect; J, right genu I, dorsal aspect (cf. D); K, right genu II, dorsal aspect (cf. F). Tibial seta d broken in H, I. Abbreviations: cd 5 (cuticular disk), ih (lyrifissure), t. e. (vestige of leg IV); other labels indicate setiform organs (see text). Scale bars: 50 Μm (A, B, E, H to same scale); 25 Μm (C, D, F, G, I – K to same scale, except famulus enlargements in C, G)." httpUri="https://zenodo.org/record/238783/files/figure.png" pageId="28" pageNumber="179">Fig. 18K</figureCitation>
|
||
) but proportionally smaller than in adult, reaching full adult proportion by tritonymph. Only proral setae (
|
||
<emphasis id="B964EABDFFFCFFC5FCCFF8E53208F008" box="[879,893,1881,1904]" italics="true" pageId="28" pageNumber="179">p</emphasis>
|
||
) eupathidial in larva;
|
||
<emphasis id="B964EABDFFFCFFC5FBD8F8E535F6F008" box="[1144,1155,1881,1904]" italics="true" pageId="28" pageNumber="179">s</emphasis>
|
||
becoming eupathidial in protonymph;
|
||
<emphasis id="B964EABDFFFCFFC5FE8EF8C13038F0EC" box="[302,333,1917,1940]" italics="true" pageId="28" pageNumber="179">m"</emphasis>
|
||
and iteral pair (
|
||
<emphasis id="B964EABDFFFCFFC5FE5EF8C1337BF0EC" box="[510,526,1917,1940]" italics="true" pageId="28" pageNumber="179">it</emphasis>
|
||
) normal in protonymph, becoming eupathidial in deuto- or tritonymph. Leg IV of protonymph (
|
||
<figureCitation id="132B2A2AFFFCFFC5FE93F81C30E4F0C0" box="[307,401,1952,1977]" captionStart="FIGURE 20. A – I" captionStartId="27.[151,250,1753,1775]" captionTargetBox="[163,1406,193,1703]" captionTargetId="figure@27.[163,1407,193,1705]" captionTargetPageId="27" captionText="FIGURE 20. A – I Nanohystrix hammerae n. sp. juveniles, stereo- (A – C) and compound (D – I) light microscope images. A, larva with all legs in walking position, lateral aspect; B, larva, anterior half in dorsal aspect, with right legs I, II; C, deutonymph with legs I and II laterally directed, dorsal aspect; D, tritonymph, left anterolateral region of hysterosoma showing muscles, cf. Fig. 8 A; E, tritonymph, lateral aspect (filled with food boluses and gregarine parasites, see Fig. 22 G), with insert showing enlargement of prodorsal strut; F, tritonymph, seta e 1 and cuticular disk cd 4, dorsolateral aspect, anterior to left; G, protonymph, distal part of tarsus III, lateral aspect; H, tritonymph, distal part of tarsus III, dorsal aspect; I, protonymph, left leg IV, abaxial aspect (seta ft " torn away). Abbreviations: cd 4 (cuticular disk), e 1. mu (depressor muscles of seta e 1); pr. mu (prodorsal retractor muscles), str (prodorsal strut); other labels indicate setae (see text). D – I composites of 5 – 20 layers. Scale bars: 100 Μm (A B, E); 25 Μm (D, F – I). Scale not determined for C, probably similar to A." httpUri="https://zenodo.org/record/238785/files/figure.png" pageId="28" pageNumber="179">
|
||
Fig.
|
||
<date id="FFAE106FFFFCFFC5FECAF81C30E4F0C0" box="[362,401,1952,1976]" pageId="28" pageNumber="179" value="1920-01">20I</date>
|
||
</figureCitation>
|
||
) with lyrifissure absent (otherwise present on all tarsi), tibia with
|
||
<emphasis id="B964EABDFFFCFFC5FB39F81D35C4F0C0" box="[1177,1201,1953,1976]" italics="true" pageId="28" pageNumber="179">v"</emphasis>
|
||
present, tarsus with typical set of seven setae: (
|
||
<emphasis id="B964EABDFFFCFFC5FE62F87930A5F0A4" box="[450,464,1989,2012]" italics="true" pageId="28" pageNumber="179">p</emphasis>
|
||
), (
|
||
<emphasis id="B964EABDFFFCFFC5FE50F879308BF0A4" box="[496,510,1989,2012]" italics="true" pageId="28" pageNumber="179">u</emphasis>
|
||
), (
|
||
<emphasis id="B964EABDFFFCFFC5FDBEF879334DF0A4" box="[542,568,1989,2012]" italics="true" pageId="28" pageNumber="179">pv</emphasis>
|
||
) and
|
||
<emphasis id="B964EABDFFFCFFC5FDD7F87833E7F0A4" box="[631,658,1988,2012]" italics="true" pageId="28" pageNumber="179">ft"</emphasis>
|
||
; other segments glabrous (see Remark 22). On leg I, solenidion ω2 of tarsus appearing in protonymph at same distal location as in adult; genu solenidion σ3 forming in deuto- or tritonymph, coupled to seta
|
||
<emphasis id="B964EABDFFFDFFC4FE72FF003095F7AD" box="[466,480,188,213]" italics="true" pageId="29" pageNumber="180">d</emphasis>
|
||
. Observed variations all on leg II of tritonymph (n =
|
||
<specimenCount id="9D16FD26FFFDFFC4FB9BFF0035B9F7AD" box="[1083,1228,188,213]" pageId="29" pageNumber="180" type="generic">3 specimens</specimenCount>
|
||
): (1) seta
|
||
<emphasis id="B964EABDFFFDFFC4FA99FF003438F7AD" box="[1337,1357,188,213]" italics="true" pageId="29" pageNumber="180">l"</emphasis>
|
||
absent unilaterally from tibia of one specimen; (2) famulus proximal to setae (
|
||
<emphasis id="B964EABDFFFDFFC4FC14FF5C32B6F780" box="[948,963,224,248]" italics="true" pageId="29" pageNumber="180">ft</emphasis>
|
||
) in one specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A6524FFFDFFC4FF67FEB8305FF425" pageId="29" pageNumber="180" type="materials_examined">
|
||
<paragraph id="8BAF36AFFFFDFFC4FF67FEB8357EF6D4" blockId="29.[151,1436,151,1040]" pageId="29" pageNumber="180">
|
||
<emphasis id="B964EABDFFFDFFC4FF67FEB830D9F665" bold="true" box="[199,428,260,285]" pageId="29" pageNumber="180">Material examined</emphasis>
|
||
.
|
||
<emphasis id="B964EABDFFFDFFC4FE1DFEB933C0F665" box="[445,693,260,285]" italics="true" pageId="29" pageNumber="180">
|
||
<typeStatus id="54AB880DFFFDFFC4FE1DFEB93084F664" box="[445,497,261,284]" pageId="29" pageNumber="180">Type</typeStatus>
|
||
series.
|
||
<typeStatus id="54AB880DFFFDFFC4FDEEFEB833C0F665" box="[590,693,260,285]" pageId="29" pageNumber="180" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
: adult female, deposited in
|
||
<collectionCode id="ED01AE6AFFFDFFC4FC5CFEB83539F664" LSID="urn:lsid:biocol.org:col:34070" box="[1020,1100,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="29" pageNumber="180">NZAC</collectionCode>
|
||
;
|
||
<collectingCountry id="F307763FFFFDFFC4FBFBFEB93582F665" box="[1115,1271,260,285]" name="New Zealand" pageId="29" pageNumber="180">New Zealand</collectingCountry>
|
||
, North Island, Waikato District, nr. Limestone Downs, ~
|
||
<geoCoordinate id="EE245068FFFDFFC4FDD3FE9433B8F638" box="[627,717,296,320]" direction="south" orientation="latitude" pageId="29" pageNumber="180" precision="925" value="-37.45">37°27'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE245068FFFDFFC4FD79FE9B3235F638" box="[729,832,295,320]" direction="east" orientation="longitude" pageId="29" pageNumber="180" precision="925" value="174.76666">174°46'E</geoCoordinate>
|
||
(Waikato Wind Farm, C-block); C.H. Watts, col.,
|
||
<date id="FFAE106FFFFDFFC4FAD7FE943191F61C" pageId="29" pageNumber="180" value="2010-01-28">28- I-2010</date>
|
||
, ex: pitfall trap on floor of native forest.
|
||
<typeStatus id="54AB880DFFFDFFC4FD14FEF13250F61C" box="[692,805,333,356]" pageId="29" pageNumber="180" type="paratype">
|
||
<emphasis id="B964EABDFFFDFFC4FD14FEF13250F61C" box="[692,805,333,356]" italics="true" pageId="29" pageNumber="180">Paratypes</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<specimenCount id="9D16FD26FFFDFFC4FC94FEF032EEF61D" box="[820,923,332,357]" pageId="29" pageNumber="180" type="adult">51 adults</specimenCount>
|
||
with same collection data as
|
||
<typeStatus id="54AB880DFFFDFFC4FB49FEF0343EF61D" box="[1257,1355,332,357]" pageId="29" pageNumber="180" type="holotype">holotype</typeStatus>
|
||
except dates
|
||
<date id="FFAE106FFFFDFFC4FF79FECC3013F6F0" box="[217,358,368,392]" pageId="29" pageNumber="180" value="2009-12-21">21-XII-2009</date>
|
||
through
|
||
<date id="FFAE106FFFFDFFC4FE6EFECC334BF6F0" box="[462,574,368,392]" pageId="29" pageNumber="180" value="2010-01-28">28-I-2010</date>
|
||
(13 deposited in
|
||
<collectionCode id="ED01AE6AFFFDFFC4FCA6FECC3220F6F0" LSID="urn:lsid:biocol.org:col:34070" box="[774,853,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="29" pageNumber="180">NZAC</collectionCode>
|
||
, four in
|
||
<collectionCode id="ED01AE6AFFFDFFC4FC16FECC3288F6F0" LSID="urn:lsid:biocol.org:col:32981" box="[950,1021,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="29" pageNumber="180">ANIC</collectionCode>
|
||
, four in
|
||
<collectionCode id="ED01AE6AFFFDFFC4FBFFFECC35EEF6F0" box="[1119,1179,368,392]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="29" pageNumber="180">CNC</collectionCode>
|
||
,
|
||
<quantity id="4CE89B4AFFFDFFC4FB08FED33597F6F0" box="[1192,1250,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="29" pageNumber="180" unit="in" value="15.0">15 in</quantity>
|
||
<collectionCode id="ED01AE6AFFFDFFC4FB4AFECC3420F6F0" box="[1258,1365,368,392]" pageId="29" pageNumber="180">CUMNH</collectionCode>
|
||
,
|
||
<quantity id="4CE89B4AFFFDFFC4FAC2FED334E9F6F0" box="[1378,1436,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="29" pageNumber="180" unit="in" value="15.0">15 in</quantity>
|
||
RAN). Preserved in alcohol, except 10
|
||
<typeStatus id="54AB880DFFFDFFC4FDEFFE2933CEF6D5" box="[591,699,405,429]" pageId="29" pageNumber="180" type="paratype">paratypes</typeStatus>
|
||
dissected on slides (in RAN).
|
||
</paragraph>
|
||
<paragraph id="8BAF36AFFFFDFFC4FF67FE0B305FF425" blockId="29.[151,1436,151,1040]" pageId="29" pageNumber="180">
|
||
<emphasis id="B964EABDFFFDFFC4FF67FE0B33DCF6A8" box="[199,681,439,464]" italics="true" pageId="29" pageNumber="180">Other studied material. Northland Region</emphasis>
|
||
. Four adults (in
|
||
<collectionCode id="ED01AE6AFFFDFFC4FCD0FE0432CBF6A8" LSID="urn:lsid:biocol.org:col:34070" box="[880,958,440,464]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="29" pageNumber="180">NZAC</collectionCode>
|
||
) from Te Paki Ecological District; Radar Bush, ~ 9.5km, SE of Cape Reinga, ca.
|
||
<geoCoordinate id="EE245068FFFDFFC4FDC2FE6033C9F68C" box="[610,700,476,500]" direction="south" orientation="latitude" pageId="29" pageNumber="180" precision="925" value="-34.466667">34°28'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE245068FFFDFFC4FD6AFE603240F68D" box="[714,821,476,501]" direction="east" orientation="longitude" pageId="29" pageNumber="180" precision="925" value="172.76666">172°46'E</geoCoordinate>
|
||
; O. Ball, col.,
|
||
<date id="FFAE106FFFFDFFC4FC7DFE603548F68D" box="[989,1085,476,501]" pageId="29" pageNumber="180" value="2006-09">IX-2006</date>
|
||
to
|
||
<date id="FFAE106FFFFDFFC4FBC3FE6035B6F68C" box="[1123,1219,476,500]" pageId="29" pageNumber="180" value="2007-03">III-2007</date>
|
||
, ex: pitfall traps in native shrubland. Three adults (in FM), one tritonymph (partly studied, lost), one deutonymph (lost), one protonymph and one larva (partly studied, lost) from Waipoua Forest (Toronui Track),
|
||
<quantity id="4CE89B4AFFFDFFC4FBC1FD9835DDF544" box="[1121,1192,548,573]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="180" unit="m" value="150.0">150 m</quantity>
|
||
a.s.l.; A. Newton and M. Thayer, col.,
|
||
<date id="FFAE106FFFFDFFC4FEFBFDF430AAF518" box="[347,479,584,608]" pageId="29" pageNumber="180" value="1980-04-13">13-IV-1980</date>
|
||
, ex: decaying nikau palm leaf bases and litter. One adult (in
|
||
<collectionCode id="ED01AE6AFFFDFFC4FB0BFDF43593F518" box="[1195,1254,584,608]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="29" pageNumber="180">CNC</collectionCode>
|
||
) from Waipoua Kauri Forest; L. Masner, col.,
|
||
<date id="FFAE106FFFFDFFC4FE48FDD03302F5FC" box="[488,631,620,644]" pageId="29" pageNumber="180" value="1983-12-12">12-XII-1983</date>
|
||
, ex: sweeping low vegetation in kauri forest.
|
||
<emphasis id="B964EABDFFFDFFC4FBD2FDD0344CF5FC" box="[1138,1337,620,645]" italics="true" pageId="29" pageNumber="180">Auckland Region.</emphasis>
|
||
Twentysix adults (
|
||
<quantity id="4CE89B4AFFFDFFC4FEB5FD2C3025F5D0" box="[277,336,655,680]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="29" pageNumber="180" unit="in" value="21.0">21 in</quantity>
|
||
<collectionCode id="ED01AE6AFFFDFFC4FEF9FD2C30DDF5D0" LSID="urn:lsid:biocol.org:col:34070" box="[345,424,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="29" pageNumber="180">NZAC</collectionCode>
|
||
, five in RAN) from vicinity of Laingholm, ~
|
||
<geoCoordinate id="EE245068FFFDFFC4FC16FD333565F5D0" box="[950,1040,655,680]" direction="south" orientation="latitude" pageId="29" pageNumber="180" precision="925" value="-36.95">36°57'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE245068FFFDFFC4FBBEFD2C35FDF5D0" box="[1054,1160,656,680]" direction="east" orientation="longitude" pageId="29" pageNumber="180" precision="925" value="174.63333">174°38'E</geoCoordinate>
|
||
; J.T. Pusateri col., dates ranging from
|
||
<date id="FFAE106FFFFDFFC4FE98FD0830C5F5B4" box="[312,432,692,716]" pageId="29" pageNumber="180" value="2007-05-19">19-V-2007</date>
|
||
to
|
||
<date id="FFAE106FFFFDFFC4FE7CFD08333BF5B4" box="[476,590,692,716]" pageId="29" pageNumber="180" value="2008-01-23">23-I-2008</date>
|
||
, ex: pitfall traps in semi-native forest. One adult from Huapai Scientific Reserve;
|
||
<date id="FFAE106FFFFDFFC4FF5FFD6B300CF588" box="[255,377,727,752]" pageId="29" pageNumber="180" value="2009-11-06">6-XI-2009</date>
|
||
, S.E. Thorpe, col.; ex: leaf litter in native forest.
|
||
<emphasis id="B964EABDFFFDFFC4FC3AFD6B3527F588" box="[922,1106,727,752]" italics="true" pageId="29" pageNumber="180">Waikato Region.</emphasis>
|
||
Three tritonymphs from
|
||
<typeStatus id="54AB880DFFFDFFC4FACCFD6434E9F588" box="[1388,1436,728,752]" pageId="29" pageNumber="180">type</typeStatus>
|
||
locality (one in
|
||
<collectionCode id="ED01AE6AFFFDFFC4FEEDFD4030CDF46C" box="[333,440,764,788]" pageId="29" pageNumber="180">CUMNH</collectionCode>
|
||
, two in RAN);
|
||
<date id="FFAE106FFFFDFFC4FDCAFD4033ABF46C" box="[618,734,764,788]" pageId="29" pageNumber="180" value="2010-01-28">28-I-2010</date>
|
||
. Fourteen adults from Te Akau, Waikato Wind Farm H-block (
|
||
<quantity id="4CE89B4AFFFDFFC4FF00FC9C31ACF440" box="[160,217,799,824]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="29" pageNumber="180" unit="in" value="11.0">11 in</quantity>
|
||
<collectionCode id="ED01AE6AFFFDFFC4FF41FC9C3045F440" LSID="urn:lsid:biocol.org:col:34070" box="[225,304,800,824]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="29" pageNumber="180">NZAC</collectionCode>
|
||
, three in RAN), ~
|
||
<geoCoordinate id="EE245068FFFDFFC4FDA6FC9C3315F440" box="[518,608,800,824]" direction="south" orientation="latitude" pageId="29" pageNumber="180" precision="925" value="-37.666668">37°40'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE245068FFFDFFC4FDCDFCA333ADF440" box="[621,728,799,824]" direction="east" orientation="longitude" pageId="29" pageNumber="180" precision="925" value="174.83333">174°50'E</geoCoordinate>
|
||
; C.H. Watts, col.,
|
||
<date id="FFAE106FFFFDFFC4FC09FC9C35B0F440" box="[937,1221,799,824]" pageId="29" pageNumber="180" value="2010-07-21" valueMax="2010-01-09" valueMin="2010-07-21">21-VII-09 and 13-I-2010</date>
|
||
, ex: pitfall traps in native forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A6524FFFDFFC4FF67FCDB3088F368" pageId="29" pageNumber="180" type="etymology">
|
||
<paragraph id="8BAF36AFFFFDFFC4FF67FCDB3088F368" blockId="29.[151,1436,151,1040]" pageId="29" pageNumber="180">
|
||
<emphasis id="B964EABDFFFDFFC4FF67FCDB3033F4F8" bold="true" box="[199,326,871,896]" pageId="29" pageNumber="180">Etymology</emphasis>
|
||
. The genitive species name honors the memory of Dutch soil biologist Marie Hammer (
|
||
<date id="FFAE106FFFFDFFC4FAF5FCD431A1F4DC" pageId="29" pageNumber="180" value="1907" valueMax="2002">1907– 2002</date>
|
||
). Her pioneering studies greatly expanded the geographic and taxonomic horizons of our knowledge of oribatid mites, especially those of the southern hemisphere (
|
||
<bibRefCitation id="EF814B5EFFFDFFC4FC23FC133557F4B0" author="Balogh" box="[899,1058,943,968]" pageId="29" pageNumber="180" refString="Balogh, J. (1985) Fifty years in the research of the world oribatid fauna. Ad honorem Dr. Marie Hammer. Opuscula Zoologica (Budapest), 21, 3 - 7." type="journal article" year="1985">Balogh 1985</bibRefCitation>
|
||
). In particular, her three large publications on
|
||
<collectingCountry id="F307763FFFFDFFC4FEEBFC693094F495" box="[331,481,980,1005]" name="New Zealand" pageId="29" pageNumber="180">New Zealand</collectingCountry>
|
||
oribatid mites (
|
||
<bibRefCitation id="EF814B5EFFFDFFC4FD32FC68324CF495" author="Hammer" box="[658,825,980,1005]" pageId="29" pageNumber="180" refString="Hammer, M. (1966) Investigations on the oribatid fauna of New Zealand. Part I. Biologiske Skrifter (det Kongelige Danske Videnskabernes Selskab), 15 (2), 1 - 108." type="journal article" year="1966">Hammer 1966</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF814B5EFFFDFFC4FCE4FC68320AF495" author="Hammer" box="[836,895,980,1005]" pageId="29" pageNumber="180" refString="Hammer, M. (1967) Investigations on the oribatid fauna of New Zealand. Part II. Biologiske Skrifter (det Kongelige Danske Videnskabernes Selskab), 15 (4), 1 - 60." type="journal article" year="1967">1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF814B5EFFFDFFC4FC2BFC6832B1F495" author="Hammer" box="[907,964,980,1005]" pageId="29" pageNumber="180" refString="Hammer, M. (1968) Investigations on the oribatid fauna of New Zealand. Part III. Biologiske Skrifter (det Kongelige Danske Videnskabernes Selskab), 16 (2), 1 - 96." type="journal article" year="1968">1968</bibRefCitation>
|
||
) provide a firm foundation for continuing studies of this interesting fauna.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |