279 lines
36 KiB
XML
279 lines
36 KiB
XML
<document id="B7A2EC2CABE5903AB02D26DA9188918C" ID-DOI="10.11646/zootaxa.3633.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261931" ID-ZooBank="60F0956C-A1F5-44CA-9D14-8C04343AD01F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629849986103" checkinUser="felipe" docAuthor="Tedesco, Anazélia M. & Aguiar, Alexandre P." docDate="2013" docId="03B987AEFF80FFE10AE1F965FE43FC9F" docLanguage="en" docName="zt03633p138.pdf" docOrigin="Zootaxa 3633 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3633.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Toechorychus darienus Tedesco & Aguiar 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="FF80FFD6FFA0FFC30A76FFC1FF88FFE0" masterDocTitle="<strong> Phylogeny and Revision of <em> Toechorychus </ em> Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species </ strong>" masterLastPageNumber="138" masterPageNumber="1" pageNumber="33" updateTime="1699120729359" updateUser="plazi">
|
||
<mods:mods id="F94E6768E91AED7ABC7DFA9AEAB758CB" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4739F95583AEED4BCF75D8B71050B362">
|
||
<mods:title id="CFCAFEDE8108E9A3F70E04E03A042A6A"><strong> Phylogeny and Revision of <em> Toechorychus </ em> Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species </ strong></mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="EC523828D5DDE34C23D42334907D88A6" type="personal">
|
||
<mods:role id="626D85E6E07470E0B6BDF25DE0297BF4">
|
||
<mods:roleTerm id="F8F2E30DE64F4BD8D6F75BA9AB7B4E7F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="12062EA4CABCAA3A8FCBA6FE51E8FD77">Tedesco, Anazélia M.</mods:namePart>
|
||
<mods:affiliation id="E30B4FC88D3813B7D209BAD0AE123739">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Zoologia, Av. Fernando Ferrari 514 - Goiabeiras,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="728DBC0D5FAEA947B79B7279A2DD571E" type="personal">
|
||
<mods:role id="BB626ECF4E03E6A4D3383F908C66495C">
|
||
<mods:roleTerm id="338EEE30816DBD23E94F7D2B6F205DB1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7CBB10C4238687CB2AA916270C8173C8">Aguiar, Alexandre P.</mods:namePart>
|
||
<mods:affiliation id="3343D529C62CC0B7EC5761D6BA05988B">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Zoologia, Av. Fernando Ferrari 514 - Goiabeiras,</mods:affiliation>
|
||
<mods:nameIdentifier id="A828C319FE859011E3E664BEF47D1FD2" type="email">aguiar.2@osu.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D167FBAD92C4454221FBF5432EE58223">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E39654D333908B1F60BF9E5D4938CA6E" type="host">
|
||
<mods:titleInfo id="D6B69A433B16B027BED17CC971012C0B">
|
||
<mods:title id="8F8E546E5533AC2FAA98BD43EAA06D4E">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="862944204543A4F9952BD4BB677B2F7E">
|
||
<mods:date id="892217EF377493A0FF3405A41A97C114">2013</mods:date>
|
||
<mods:detail id="ACE4BE9F2CB954D4957947B4B259136E" type="pubDate">
|
||
<mods:number id="8D17201119AEDE4C5DF131CABB5EA17E">2013-03-22</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C5CFF46B42168BF4699D17CAB5AA910F" type="volume">
|
||
<mods:number id="F13DE79833BB77A51A75289B52A71C9E">3633</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="85F2353E84DB9690D035FAA7E7B35FF6" type="issue">
|
||
<mods:number id="A9006FCD2370E78EB9D29F801E2BF60B">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6679334F93F555FE9CDA09854F3872AC" unit="page">
|
||
<mods:start id="CA281B14F77515019F1E48B4C6D8A722">1</mods:start>
|
||
<mods:end id="01736F6012191F16824EE6075ED3CCCC">138</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="06D1A6B4D46736794C7D6EF57F9C2543">
|
||
<mods:url id="563F046D1AF73AD70BA515059A635BA8">http://dx.doi.org/10.11646/zootaxa.3633.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="D19454B37155E9153DA41F959CB6F716">journal article</mods:classification>
|
||
<mods:identifier id="4AB30406A63F22596959D02AB38E7A6B" type="DOI">10.11646/zootaxa.3633.1.1</mods:identifier>
|
||
<mods:identifier id="20E696E4E88FA78DA0C643FAF0A461D2" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="FDF3D448F575CEFDF75FCF870A590AC6" type="Zenodo-Dep">5261931</mods:identifier>
|
||
<mods:identifier id="06A9CC0F99C819E073004B777BA5B786" type="ZooBank">60F0956C-A1F5-44CA-9D14-8C04343AD01F</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B987AEFF80FFE10AE1F965FE43FC9F" ID-DOI="http://doi.org/10.5281/zenodo.5261799" ID-Zenodo-Dep="5261799" LSID="urn:lsid:plazi:treatment:03B987AEFF80FFE10AE1F965FE43FC9F" httpUri="http://treatment.plazi.org/id/03B987AEFF80FFE10AE1F965FE43FC9F" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
|
||
<subSubSection id="C30A6533FF80FFE30AE1F965FD0EF95D" box="[151,646,1699,1726]" pageId="32" pageNumber="33" type="nomenclature">
|
||
<paragraph id="8BAF36B8FF80FFE30AE1F965FD0EF95D" blockId="32.[151,646,1699,1758]" box="[151,646,1699,1726]" pageId="32" pageNumber="33">
|
||
<heading id="D0E781D4FF80FFE30AE1F965FD0EF95D" bold="true" box="[151,646,1699,1726]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
|
||
<emphasis id="B964EAAAFF80FFE30AE1F965FD0EF95D" bold="true" box="[151,646,1699,1726]" pageId="32" pageNumber="33">
|
||
<taxonomicName id="4C104D3BFF80FFE30AE1F965FD93F95D" authority="Tedesco" authorityName="Tedesco & Aguiar" authorityYear="2013" box="[151,539,1699,1726]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="darienus" status="sp. nov.">
|
||
<emphasis id="B964EAAAFF80FFE30AE1F965FE38F95E" bold="true" box="[151,432,1700,1726]" italics="true" pageId="32" pageNumber="33">Toechorychus darienus</emphasis>
|
||
Tedesco
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25757D1FF80FFE3085EF962FD0EF95D" box="[552,646,1699,1725]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A6533FF80FFE10AE1F907FA8AFEFC" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BAF36B8FF80FFE30AE1F907FE48F93E" blockId="32.[151,646,1699,1758]" box="[151,448,1734,1758]" pageId="32" pageNumber="33">
|
||
(
|
||
<figureCitation id="132B2A3DFF80FFE30AE9F907FF7DF93E" box="[159,245,1734,1758]" captionStart="FIGURES 28–31" captionStartId="109.[151,264,1888,1911]" captionTargetBox="[151,1435,193,1867]" captionTargetId="figure-62@109.[151,1435,193,1867]" captionTargetPageId="109" captionText="FIGURES 28–31. Lateral pictures of Toechorychus. 28, T. ondensis sp. nov., holotype. 29, T. albimaculatus Taschenberg. 30, T. darienus sp. nov., holotype. 31, T. zulius sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261961" httpUri="https://zenodo.org/record/5261961/files/figure.png" pageId="32" pageNumber="33">Figs 30</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A3DFF80FFE30B77F907FE97F93E" box="[257,287,1734,1758]" captionStart="FIGURES 70–84" captionStartId="115.[151,264,1840,1863]" captionTargetBox="[151,1435,193,1819]" captionTargetId="figure-173@115.[151,1435,193,1819]" captionTargetPageId="115" captionText="FIGURES 70–84. Head, frontal view. 70, T. martinus sp. nov., holotype. 71, T. morelus sp. nov., holotype. 72, T. zulius sp. nov., holotype. 73, T. amazonensis sp. nov., holotype. 74, T. paramaribus sp. nov., holotype. 75, T. heredius sp. nov., holotype. 76, T. linaresius sp. nov., holotype. 77, T. marowijnus sp. nov., holotype. 78, T. darienus sp. nov., holotype. 79, T. kawus sp. nov., holotype. 80, T. jatainus sp. nov., holotype. 81, T. taperinus sp. nov., holotype. 82, T. teutonius sp. nov., holotype. 83, T. calius sp. nov., holotype. 84, T. surinamus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261981" httpUri="https://zenodo.org/record/5261981/files/figure.png" pageId="32" pageNumber="33">78</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A3DFF80FFE30B5DF907FEC2F93E" box="[299,330,1734,1758]" captionStart="FIGURES 91–96" captionStartId="117.[151,264,1674,1697]" captionTargetBox="[151,1435,193,1653]" captionTargetId="figure-84@117.[151,1435,193,1653]" captionTargetPageId="117" captionText="FIGURES 91–96. Mesoscutum, dorsal view. 91, T. marcapatus sp. nov., holotype. 92, T. darienus sp. nov., holotype. 93, T. coaracius sp. nov., holotype. 94, T. itapuensis sp. nov., holotype. 95, T. martinus sp. nov., holotype. 96, T. zulius sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261985" httpUri="https://zenodo.org/record/5261985/files/figure.png" pageId="32" pageNumber="33">92</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A3DFF80FFE30B23F907FE09F93E" box="[341,385,1734,1758]" captionStart="FIGURES 157–162" captionStartId="126.[151,264,1672,1695]" captionTargetBox="[151,1435,193,1651]" captionTargetId="figure-85@126.[151,1435,193,1651]" captionTargetPageId="126" captionText="FIGURES 157–162. Propodeum, dorsal view. 157, T. kawus sp. nov., holotype. 158, T. zulius sp. nov., holotype. 159, T. taperinus sp. nov., holotype. 160, T. heredius sp. nov., holotype. 161, T. bombuscarus sp. nov., holotype. 162, T. darienus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5262009" httpUri="https://zenodo.org/record/5262009/files/figure.png" pageId="32" pageNumber="33">162</figureCitation>
|
||
,
|
||
<figureCitation id="132B2A3DFF80FFE30BFBF907FE30F93E" box="[397,440,1734,1758]" captionStart="FIGURES 163–174" captionStartId="127.[151,264,1631,1654]" captionTargetBox="[151,1435,193,1610]" captionTargetId="figure-141@127.[151,1435,193,1610]" captionTargetPageId="127" captionText="FIGURES 163–174. Hind coxa and metasoma color patterns, dorsal view. 163, T. napus sp. nov., holotype. 164, T. teutonius sp. nov., holotype. 165, T. kawus sp. nov., holotype. 166, T. taperinus sp. nov., holotype. 167, T. darienus sp. nov., holotype. 168, T. amapaeus sp. nov., holotype. 169, T. ondensis sp. nov., holotype. 170, T. jatainus sp. nov., holotype. 171, T. albimaculatus Taschenberg. 172, T. vilhenus sp. nov., holotype. 173, T. benius sp. nov., holotype. 174, T. cassunungae Brauns." figureDoi="http://doi.org/10.5281/zenodo.5262013" httpUri="https://zenodo.org/record/5262013/files/figure.png" pageId="32" pageNumber="33">167</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF80FFE30AE1F8CFFCCEF8C6" blockId="32.[151,1436,1806,2010]" box="[151,838,1806,1830]" pageId="32" pageNumber="33">
|
||
Description.
|
||
<typeStatus id="54AB881AFF80FFE30B51F8CFFE07F8C6" box="[295,399,1806,1830]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
FEMALE. Fore wing length
|
||
<quantity id="4CE89B5DFF80FFE308ADF8CFFCCAF8C6" box="[731,834,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="6.06" pageId="32" pageNumber="33" unit="mm" value="6.06">6.06 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF80FFE20AB1F8F2FA86FE67" blockId="32.[151,1436,1806,2010]" lastBlockId="33.[151,1437,151,2012]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
<emphasis id="B964EAAAFF80FFE30AB1F8F2FE8CF8AB" box="[199,260,1843,1867]" italics="true" pageId="32" pageNumber="33">Head</emphasis>
|
||
. Mandible 1.38 as long as basal width, densely pilose; ventral margin projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 1.51 as wide as high, subquadrate, entirely finely strigulate; apex 1.45 as long as base, truncate; apical margin sharp, medially straight. Supraclypeal area entirely strigulate, also densely punctate, moderately pilose, medially slightly prominent; between antennal foramens with V-shaped carina, medially interrupted; radicle foveolate. Antenna with 25 flagellomeres; white band starting at flagellomere 4, reaching flagellomere 11; flagellum slender; subapical flagellomeres slightly flattened. Supra-antennal area with very faint but distinct median longitudinal carina, medially without longitudinal elevation, near antennal sockets striate; dorsal half medially rugulose. Paraocular area smooth. Vertex rugulose between ocelli, at anterior third with short, longitudinal sulcus, posterior two-thirds smooth; gena and vertex behind ocelli minutely strigulate, sparsely pilose; gena in lateral view at level of dorsal portion of occipital carina narrow, ventrally wide; occipital carina faint, dorsally absent, ventrally slightly projected as crest, reaching hypostomal carina at mandible base or nearly so; hypostomal carina slightly projected as crest; malar space 1.06 as long as basal width of mandible.
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF81FFE20AB1FE54FB9DFB07" blockId="33.[151,1437,151,2012]" pageId="33" pageNumber="34">
|
||
<taxonomicName id="4C104D3BFF81FFE20AB1FE54FEB7FE4C" box="[199,319,405,428]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964EAAAFF81FFE20AB1FE54FEB7FE4C" box="[199,319,405,428]" italics="true" pageId="33" pageNumber="34">Mesosoma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C104D3BFF81FFE20B3BFE55FE4FFE4B" box="[333,455,404,427]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="33" pageNumber="75" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
|
||
in dorsal view 1.80 as long as wide; in lateral view, mesosoma middle width
|
||
<quantity id="4CE89B5DFF81FFE20F47FE55FA10FE4B" box="[1329,1432,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="mm" value="0.2">0.20 mm</quantity>
|
||
. Pronotum centrally smooth, latero-ventrally smooth, margin near mesopleuron ventrally corrugated; pronotal swelling finely rugulose; collar dorso-laterally rounded, distinctly swollen, anteriorly smooth; epomia absent. Mesoscutum sparsely pilose, lobes dorsally punctulate, laterally markedly corrugated, central lobe with dense coarse punctures, central lobe without longitudinal sulcus or carina; notauli deeply impressed, posteriorly abruptly curved, U-shaped. Scutoscutellar groove smooth, polished; scutellum 0.98 as long as wide, scarcely punctate; scutellar carina incomplete, restricted approximately to anterior 0.6; postscutellum 0.30 as long as wide; hind margin of metanotum with two lateral teeth. Subalar ridge somewhat elongate; mesopleuron entirely markedly strigate, ventrally strigulate; mesopleural groove corrugated; epicnemial carina stout, dorsally curving forwards; dorsal end of epicnemial carina reaching 0.2 of posterior margin of pronotum, behind epicnemial carina with rounded confused-rugulose area; sternaulus complete, reaching base of mid coxa, markedly sinuate, deeply impressed; markedly corrugated; juxtacoxal carina of mesopleuron stout and long; scrobe deeply impressed, forming pit. Mesothoracic venter strigulate; median portion of posterior transverse carina of mesothoracic venter V-shaped, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.09 as long as anterior portion of propodeum, moderately deep and narrow, medially wide, very faintly corrugated, almost smooth; propodeum 1.16 as long as wide medially; anterior margin of propodeum with two faint lateral teeth; anterior portion of propodeum punctulate or with sparse coarse punctures, medially with two posteriorly convergent faint longitudinal carinae; lateral longitudinal carina of propodeum represented by short curved carina; posterior portion of propodeum entirely markedly strigate, medially without distinctly stout longitudinal carina; behind posterior transverse carina with two divergent longitudinal carinae; anterior transverse carina of propodeum complete, stout, medially slightly arched forwards; posterior transverse carina absent, only with lateral slight swellings; propodeal spiracle 1.45 as long as wide; pleural carina of propodeum irregular and faint, obsolescent posteriorly, fused with sculpture of lower division of metapleuron and propodeum; lower division of metapleuron strigulate or sparsely punctate, sparsely pilose, juxtacoxal carina long and stout.
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF81FFE20AB1FB34FD36FA7C" blockId="33.[151,1437,151,2012]" pageId="33" pageNumber="34">
|
||
<emphasis id="B964EAAAFF81FFE20AB1FB34FE83FAEC" box="[199,267,1269,1292]" italics="true" pageId="33" pageNumber="34">Wings</emphasis>
|
||
. Fore wing: vein 2+3Rs slightly concave; ramellus absent; vein 1M+Rs posteriorly faintly sinuous; vein 1cu-a straight, arising far from vein 1M+Rs base; vein 2Cu 0.46 as long as vein 2cu-a; vein 4Rs 0.93 as long as vein 4M, uniformly slightly convex; cell 1+2Rs 0.72 as high as pterostigma, rectangular; vein 2M approximately as long as vein 3M; vein 3M distinct, 0.64 as long as vein
|
||
<specimenCount id="9D16FD31FF81FFE20975FA9EFCB9FA97" box="[771,817,1375,1399]" pageId="33" pageNumber="34" type="generic">2M.</specimenCount>
|
||
Hind wing: vein 1Cu 1.33 as long as vein cu-a; vein 2- 1A reaching 0.83 of distance to posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF81FFE20AB1FA68FCCAFA5F" blockId="33.[151,1437,151,2012]" box="[199,834,1447,1472]" pageId="33" pageNumber="34">
|
||
<emphasis id="B964EAAAFF81FFE20AB1FA68FF74FA20" box="[199,252,1449,1472]" italics="true" pageId="33" pageNumber="34">Legs</emphasis>
|
||
. Tibia with sparse short bristles; hind coxa globose.
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF81FFE20AB1FA0CFCFBF9AF" blockId="33.[151,1437,151,2012]" pageId="33" pageNumber="34">
|
||
<emphasis id="B964EAAAFF81FFE20AB1FA0CFEB4FA04" box="[199,316,1485,1508]" italics="true" pageId="33" pageNumber="34">Metasoma</emphasis>
|
||
. T1 0.52 as long as hind femur, with basolateral tooth; anteriorly smooth, posteriorly coriarious; spiracle at anterior 0.54, slightly prominent; postpetiole dorso-anteriorly convex; T2–8 minutely coriarious; almost glabrous; T2 1.09 as long as wide at apex; apex of T2 1.93 as wide as base. Dorsal valve of ovipositor with nodus indistinct; apex of lower valve with series of teeth; notch present.
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF81FFE10AB1F99DFC83FF34" blockId="33.[151,1437,151,2012]" lastBlockId="34.[151,1436,151,895]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
<emphasis id="B964EAAAFF81FFE20AB1F99DFE8FF994" box="[199,263,1628,1652]" italics="true" pageId="33" pageNumber="34">Color</emphasis>
|
||
. Head and mesosoma yellow and black, metasoma mostly orange. Head: pale yellow (208,133,077); mandible, ventral half of malar space, apex of clypeus, M-shaped mark at ventral half of supraclypeal area, supraantennal area medially, vertex, and occiput, black; scape ventrally pale yellow; pedicel, and flagellum brown; flagellum dorsally entirely white at f5–10 entirely, partially at f4 and f11.
|
||
<taxonomicName id="4C104D3BFF81FFE20E6EF909FB1EF93F" box="[1048,1174,1736,1759]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="33" pageNumber="75" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
|
||
: black, except for the following pale yellow marks: propleuron centrally, collar laterally, pronotal swelling, longitudinal narrow mark at posterior 0.9 of central lobe of mesoscutum, tegula, axillary carina, scutellum, postscutellum, large mark extending from subalar ridge to posterior ventral margin of mesopleuron and over epicnecium, dorsal 0.8 of hypoepimeron, mesothoracic venter, upper division of metapleuron, lower division of metapleuron except ventral rounded spot, hind margin of metanotum, and two lateral longitudinal marks at posterior portion of propodeum. Legs: mostly orange (212,147,043); fore coxa pale yellow, posteriorly fulvous; mid coxa pale yellow, laterally with black strip reaching first trochanter; hind coxa pale yellow with lateral black stripes at entire length. Metasoma: mostly orange; T1 dorsally black, laterally and at posterior 0.2, pale yellow, brownish around spiracle; T2–8 yellowish towards apex; S2–3 laterally dark brown (108,062,031).
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF82FFE10AB1FF1EFA8AFEFC" blockId="34.[151,1436,151,895]" pageId="34" pageNumber="35">
|
||
<emphasis id="B964EAAAFF82FFE10AB1FF1EFEBFFF18" bold="true" box="[199,311,223,248]" pageId="34" pageNumber="35">Variation</emphasis>
|
||
. Mandible basally white. Occipital carina dorsally widely interrupted, completely absent. Mid coxa dorsally black only at apical 0.4. Hind coxa laterally with small black spot. T1 black around spiracle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A6533FF82FFE10AB1FEE6FE00FDDC" pageId="34" pageNumber="35" type="discussion">
|
||
<paragraph id="8BAF36B8FF82FFE10AB1FEE6FE54FDF7" blockId="34.[151,1436,151,895]" pageId="34" pageNumber="35">
|
||
<emphasis id="B964EAAAFF82FFE10AB1FEE6FEC0FEA0" bold="true" box="[199,328,295,320]" pageId="34" pageNumber="35">Comments</emphasis>
|
||
. Very similar to
|
||
<taxonomicName id="4C104D3BFF82FFE1087FFEE8FD1EFEA0" authority="Tedesco & Aguiar, 2013" authorityName="Tedesco & Aguiar" authorityYear="2013" box="[521,662,296,320]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="amapaeus" status="sp. nov.">
|
||
<emphasis id="B964EAAAFF82FFE1087FFEE8FD9EFEA0" box="[521,534,297,320]" italics="true" pageId="34" pageNumber="35">T</emphasis>
|
||
.
|
||
<emphasis id="B964EAAAFF82FFE10853FEE8FD1EFEA0" box="[549,662,297,320]" italics="true" pageId="34" pageNumber="35">amapaeus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757D1FF82FFE108E9FEE9FD71FEDF" box="[671,761,296,320]" pageId="34" pageNumber="35" rank="species">
|
||
<emphasis id="B964EAAAFF82FFE108E9FEE9FD31FEA0" bold="true" box="[671,697,296,320]" pageId="34" pageNumber="35">sp</emphasis>
|
||
.
|
||
<emphasis id="B964EAAAFF82FFE108BFFEE9FD7BFEA0" bold="true" box="[713,755,296,320]" pageId="34" pageNumber="35">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, from which it can be isolated by having supraclypeal area ventrally with M-shaped mark (
|
||
<emphasis id="B964EAAAFF82FFE10873FE8CFD94FE84" box="[517,540,333,356]" italics="true" pageId="34" pageNumber="35">vs</emphasis>
|
||
. with two lateral longitudinal black stripes in
|
||
<taxonomicName id="4C104D3BFF82FFE10E45FE8CFB4AFE84" authority="Tedesco & Aguiar, 2013" authorityName="Tedesco & Aguiar" authorityYear="2013" box="[1075,1218,332,356]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="amapaeus" status="sp. nov.">
|
||
<emphasis id="B964EAAAFF82FFE10E45FE8CFBC8FE84" box="[1075,1088,333,356]" italics="true" pageId="34" pageNumber="35">T</emphasis>
|
||
.
|
||
<emphasis id="B964EAAAFF82FFE10E27FE8CFB4AFE84" box="[1105,1218,333,356]" italics="true" pageId="34" pageNumber="35">amapaeus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25757D1FF82FFE10EBAFE8DFAA0FE83" box="[1228,1320,332,356]" pageId="34" pageNumber="35" rank="species">
|
||
<emphasis id="B964EAAAFF82FFE10EBAFE8DFB6EFE84" bold="true" box="[1228,1254,332,356]" pageId="34" pageNumber="35">sp</emphasis>
|
||
.
|
||
<emphasis id="B964EAAAFF82FFE10E81FE8DFAA9FE84" bold="true" box="[1271,1313,332,356]" pageId="34" pageNumber="35">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
;
|
||
<figureCitation id="132B2A3DFF82FFE10F4FFE8DFA07FE84" box="[1337,1423,332,356]" captionStart="FIGURES 55–69" captionStartId="114.[151,264,1840,1863]" captionTargetBox="[151,1435,193,1819]" captionTargetId="figure-163@114.[151,1435,193,1819]" captionTargetPageId="114" captionText="FIGURES 55–69. Head, frontal view. 55, T. surinamus sp. nov., holotype. 56, T. callangus sp. nov., holotype. 57, T. coaracius sp. nov., holotype. 58, T. guarapuavus sp. nov., holotype. 59, T. itapuensis sp. nov., holotype. 60, T. benius sp. nov., holotype. 61, T. melgassus sp. nov., holotype. 62, T. stramineus Taschenberg. 63, T. tumazulus sp. nov., holotype. 64, T. ondensis sp. nov., holotype. 65, T. fluminensis sp. nov., holotype. 66, T. amapaeus sp. nov., holotype. 67, T. barticus sp. nov., holotype. 68, T. albimaculatus Taschenberg. 69, T. cassunungae Brauns." figureDoi="http://doi.org/10.5281/zenodo.5261977" httpUri="https://zenodo.org/record/5261977/files/figure.png" pageId="34" pageNumber="35">Fig. 66</figureCitation>
|
||
); lateral black stripes at supraclypeal area reaching antennal foramens, even if medially interrupted (
|
||
<emphasis id="B964EAAAFF82FFE10E9DFEB0FA8AFE68" box="[1259,1282,369,392]" italics="true" pageId="34" pageNumber="35">vs</emphasis>
|
||
. not reaching antennal foramens); malar space ventrally black (
|
||
<emphasis id="B964EAAAFF82FFE108B2FE54FD53FE4C" box="[708,731,405,428]" italics="true" pageId="34" pageNumber="35">vs</emphasis>
|
||
. completely yellow); occipital carina near mandible black (
|
||
<emphasis id="B964EAAAFF82FFE10F0BFE54FA1CFE4C" box="[1405,1428,405,428]" italics="true" pageId="34" pageNumber="35">vs</emphasis>
|
||
. bright yellow); juxtacoxal carina of mesopleuron stout and long (
|
||
<emphasis id="B964EAAAFF82FFE109F9FE78FC2EFE30" box="[911,934,441,464]" italics="true" pageId="34" pageNumber="35">vs</emphasis>
|
||
. faint and short); scrobe deeply impressed, forming a pit (
|
||
<emphasis id="B964EAAAFF82FFE10B4AFE1CFEDBFE14" box="[316,339,477,500]" italics="true" pageId="34" pageNumber="35">vs</emphasis>
|
||
. very shallow, forming a sulcus); lower division of metapleuron with ventral rounded black spot (
|
||
<emphasis id="B964EAAAFF82FFE10AD6FDC0FF3FFDF8" box="[160,183,513,536]" italics="true" pageId="34" pageNumber="35">vs</emphasis>
|
||
. entirely yellow;
|
||
<figureCitation id="132B2A3DFF82FFE10B0BFE3EFE46FDF7" box="[381,462,511,535]" captionStart="FIGURES 12–15" captionStartId="105.[151,264,1920,1943]" captionTargetBox="[151,1435,186,1891]" captionTargetId="figure-65@105.[151,1435,186,1891]" captionTargetPageId="105" captionText="FIGURES 12–15. Lateral pictures of Toechorychus. 12, T. vilhenus sp. nov., holotype. 13, T. amapaeus sp. nov., holotype. 14, T. martinus sp. nov., holotype. 15, T. catarinus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261949" httpUri="https://zenodo.org/record/5261949/files/figure.png" pageId="34" pageNumber="35">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF36B8FF82FFE10AB1FDE5FE00FDDC" blockId="34.[151,1436,151,895]" box="[199,392,548,572]" pageId="34" pageNumber="35">
|
||
<emphasis id="B964EAAAFF82FFE10AB1FDE5FE8CFDDC" bold="true" box="[199,260,548,572]" pageId="34" pageNumber="35">Male</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A6533FF82FFE10AB1FD86FA95FD64" pageId="34" pageNumber="35" type="distribution">
|
||
<paragraph id="8BAF36B8FF82FFE10AB1FD86FA95FD64" blockId="34.[151,1436,151,895]" pageId="34" pageNumber="35">
|
||
<emphasis id="B964EAAAFF82FFE10AB1FD86FED1FD80" bold="true" box="[199,345,583,608]" pageId="34" pageNumber="35">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3077628FF82FFE10B1EFD86FE6AFDBF" box="[360,482,583,607]" name="Costa Rica" pageId="34" pageNumber="35">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3077628FF82FFE1086CFD89FDFEFDBF" box="[538,630,584,607]" name="Panama" pageId="34" pageNumber="35">Panama</collectingCountry>
|
||
. Record from
|
||
<emphasis id="B964EAAAFF82FFE10961FD89FCFAFD80" box="[791,882,584,608]" italics="true" pageId="34" pageNumber="35">Heredia</emphasis>
|
||
and
|
||
<emphasis id="B964EAAAFF82FFE109DFFD89FC70FD80" box="[937,1016,584,608]" italics="true" pageId="34" pageNumber="35">
|
||
<collectingRegion id="49D4F85AFF82FFE109DFFD89FC70FD80" box="[937,1016,584,608]" country="Panama" name="Darien" pageId="34" pageNumber="35">Darién</collectingRegion>
|
||
</emphasis>
|
||
. The
|
||
<typeStatus id="54AB881AFF82FFE10E4CFD89FBE2FD80" box="[1082,1130,584,608]" pageId="34" pageNumber="35">type</typeStatus>
|
||
locality is
|
||
<collectingRegion id="49D4F85AFF82FFE10E9EFD86FABCFDBF" box="[1256,1332,583,607]" country="Panama" name="Darien" pageId="34" pageNumber="35">Darién</collectingRegion>
|
||
National Park,
|
||
<emphasis id="B964EAAAFF82FFE10AAEFDADFEAFFD64" box="[216,295,620,644]" italics="true" pageId="34" pageNumber="35">
|
||
<collectingRegion id="49D4F85AFF82FFE10AAEFDADFEAFFD64" box="[216,295,620,644]" country="Panama" name="Darien" pageId="34" pageNumber="35">Darién</collectingRegion>
|
||
</emphasis>
|
||
(
|
||
<geoCoordinate id="EE24507FFF82FFE10B41FDADFE3BFD64" box="[311,435,620,644]" degrees="7" direction="north" minutes="44" orientation="latitude" pageId="34" pageNumber="35" precision="15" seconds="10" value="7.736111">7°44’10”N</geoCoordinate>
|
||
<geoCoordinate id="EE24507FFF82FFE10BCCFDADFDC2FD64" box="[442,586,620,644]" degrees="77" direction="west" minutes="32" orientation="longitude" pageId="34" pageNumber="35" precision="15" seconds="50" value="-77.54722">77°32’50”W</geoCoordinate>
|
||
) (
|
||
<figureCitation id="132B2A3DFF82FFE10814FDADFD4CFD64" box="[610,708,620,644]" captionStart="FIGURES 232–239" captionStartId="134.[151,264,1946,1969]" captionTargetBox="[172,1416,183,1934]" captionTargetId="figure-103@134.[172,1416,179,1934]" captionTargetPageId="134" captionText="FIGURES 232–239. Distribution records. 232, T. linaresius sp. nov., holotype. 233, T. vilhenus sp. nov., holotype. 234, T. marowijnus sp. nov., holotype. 235, T. darienus sp. nov., holotype. 236, T. kawus sp. nov., holotype. 237, T. jatainus sp. nov., holotype. 238, T. pirrus sp. nov., holotype. 239, T. benius sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5262033" httpUri="https://zenodo.org/record/5262033/files/figure.png" pageId="34" pageNumber="35">Fig. 235</figureCitation>
|
||
). These records comprise a range of 3°42’ in latitude.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A6533FF82FFE10AB1FD4EFEFDFD2C" pageId="34" pageNumber="35" type="biology_ecology">
|
||
<paragraph id="8BAF36B8FF82FFE10AB1FD4EFEFDFD2C" blockId="34.[151,1436,151,895]" pageId="34" pageNumber="35">
|
||
<emphasis id="B964EAAAFF82FFE10AB1FD4EFE97FD48" bold="true" box="[199,287,655,680]" pageId="34" pageNumber="35">Biology</emphasis>
|
||
. Collected near
|
||
<taxonomicName id="4C104D3BFF82FFE10BA2FD51FCE1FD47" authority="Richards" authorityName="Richards" box="[468,873,655,680]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="collarellus">
|
||
<emphasis id="B964EAAAFF82FFE10BA2FD51FD76FD48" box="[468,766,656,680]" italics="true" pageId="34" pageNumber="35">Mischocyttarus collarellus</emphasis>
|
||
Richards
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C104D3BFF82FFE10900FD4EFC68FD47" box="[886,992,655,679]" class="Insecta" family="Vespidae" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Vespidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C104D3BFF82FFE1099CFD4EFBD2FD47" authorityName="Lepeletier de Saint Fargeau" authorityYear="1836" box="[1002,1114,655,679]" class="Insecta" family="Vespidae" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Polistinae">Polistinae</taxonomicName>
|
||
) nest, in La Selva Biological Station,
|
||
<collectingCountry id="F3077628FF82FFE10A85FD75FEF9FD2C" box="[243,369,692,716]" name="Costa Rica" pageId="34" pageNumber="35">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A6533FF82FFE10AB1FD16FE43FC9F" pageId="34" pageNumber="35" type="materials_examined">
|
||
<paragraph id="8BAF36B8FF82FFE10AB1FD16FE43FC9F" blockId="34.[151,1436,151,895]" pageId="34" pageNumber="35">
|
||
<emphasis id="B964EAAAFF82FFE10AB1FD16FE3AFD10" bold="true" box="[199,434,727,752]" pageId="34" pageNumber="35">Material examined</emphasis>
|
||
.
|
||
<typeStatus id="54AB881AFF82FFE10BBEFD16FDB8FD0F" box="[456,560,727,751]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D16FD31FF82FFE10836FD19FDDDFD0F" box="[576,597,728,751]" pageId="34" pageNumber="35" type="female">♀</specimenCount>
|
||
,
|
||
<emphasis id="B964EAAAFF82FFE1081AFD16FD6EFD10" bold="true" box="[620,742,727,752]" pageId="34" pageNumber="35">
|
||
<collectingCountry id="F3077628FF82FFE1081AFD16FD6EFD10" box="[620,742,727,752]" name="Panama" pageId="34" pageNumber="35">PANAMA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B964EAAAFF82FFE1088BFD19FCC4FD10" box="[765,844,728,752]" italics="true" pageId="34" pageNumber="35">
|
||
<collectingRegion id="49D4F85AFF82FFE1088BFD19FCC4FD10" box="[765,844,728,752]" country="Panama" name="Darien" pageId="34" pageNumber="35">Darién</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49D4F85AFF82FFE10912FD16FC38FD0F" box="[868,944,727,751]" country="Panama" name="Darien" pageId="34" pageNumber="35">Darién</collectingRegion>
|
||
National Park, Pirre, Est. Rancho Frio,
|
||
<date id="FFAE1078FF82FFE10AE1FD3DFE8CFCF4" box="[151,260,764,788]" pageId="34" pageNumber="35" value="2000-01-17">17.I.2000</date>
|
||
, 80 m, R.Cambra & A.Santos
|
||
<emphasis id="B964EAAAFF82FFE1082AFD3DFDF6FCF4" box="[604,638,764,788]" italics="true" pageId="34" pageNumber="35">leg</emphasis>
|
||
.,
|
||
<taxonomicName id="4C104D3BFF82FFE108E5FD3DFC1BFCF3" authority=", Townes" authorityName="Townes" box="[659,915,764,788]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964EAAAFF82FFE108E5FD3DFCBCFCF4" box="[659,820,764,788]" italics="true" pageId="34" pageNumber="35">Toechorychus,</emphasis>
|
||
Townes
|
||
</taxonomicName>
|
||
det. (
|
||
<collectionCode id="ED01AE7DFF82FFE109A5FD3DFB92FCF4" box="[979,1050,764,788]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1008" lsid="urn:lsid:biocol.org:col:1008" name="American Entomological Institute" pageId="34" pageNumber="35">AEIC</collectionCode>
|
||
). Complete, in good shape, except for mesoscutum somewhat damaged by pin.
|
||
<typeStatus id="54AB881AFF82FFE108DDFCE1FC85FCD8" box="[683,781,800,824]" pageId="34" pageNumber="35" type="paratype">Paratype</typeStatus>
|
||
<specimenCount id="9D16FD31FF82FFE1096CFCE1FCA7FCD7" box="[794,815,800,823]" pageId="34" pageNumber="35" type="female">♀</specimenCount>
|
||
from
|
||
<emphasis id="B964EAAAFF82FFE10908FCDEFBB8FCD8" bold="true" box="[894,1072,799,824]" pageId="34" pageNumber="35">
|
||
<collectingCountry id="F3077628FF82FFE10908FCDEFBB8FCD8" box="[894,1072,799,824]" name="Costa Rica" pageId="34" pageNumber="35">COSTA RICA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B964EAAAFF82FFE10E32FCE1FB17FCD8" box="[1092,1183,800,824]" italics="true" pageId="34" pageNumber="35">
|
||
<collectingRegion id="49D4F85AFF82FFE10E32FCE1FB17FCD8" box="[1092,1183,800,824]" country="Costa Rica" name="Heredia" pageId="34" pageNumber="35">Heredia</collectingRegion>
|
||
</emphasis>
|
||
, La Selva Biological Station,
|
||
<date id="FFAE1078FF82FFE10A8EFC85FEE4FCBC" box="[248,364,836,860]" pageId="34" pageNumber="35" value="1999-06-08">8.VI.1999</date>
|
||
, E.F.Smith
|
||
<taxonomicName id="4C104D3BFF82FFE10B80FC85FD7BFCBC" authorityName=": Townes" authorityYear="1946" box="[502,755,836,860]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="abactus">
|
||
<emphasis id="B964EAAAFF82FFE10B80FC85FD7BFCBC" box="[502,755,836,860]" italics="true" pageId="34" pageNumber="35">Toechorychus abactus</emphasis>
|
||
</taxonomicName>
|
||
, det. I.Gauld 2001, flying near
|
||
<taxonomicName id="4C104D3BFF82FFE10E1BFC85FA14FCBC" box="[1133,1436,836,860]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="collarellus">
|
||
<emphasis id="B964EAAAFF82FFE10E1BFC85FA14FCBC" box="[1133,1436,836,860]" italics="true" pageId="34" pageNumber="35">Mischocyttarus collarellus</emphasis>
|
||
</taxonomicName>
|
||
nest, Nest P. 03B (
|
||
<collectionCode id="ED01AE7DFF82FFE10B12FCA6FE36FC9F" box="[356,446,871,895]" pageId="34" pageNumber="35">BMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |