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<document ID-DOI="http://doi.org/10.5281/zenodo.5999639" ID-GBIF-Dataset="a1d56c0b-d41f-42c1-91d2-417ab340969f" ID-GBIF-Taxon="134839584" ID-ISSN="0007-196X" ID-Zenodo-Dep="5999639" checkinTime="1506069121042" checkinUser="plazi" docAuthor="Gabriel Emiliano Ferreira, Andréa Onofre De Araújo, Michael John Gilbert Hopkins &amp; Alain Chautems" docDate="2017" docId="03B98796A031FFEFE6C3FC0E75AFFD13" docLanguage="en" docName="Ferreira etal 2017_Besleria.pdf" docOrigin="Brittonia 69 (2)" docStyle="DocumentStyle{}" docTitle="Besleria iara Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems 2017, sp. nov." docType="treatment" docVersion="5" lastPageId="4" lastPageNumber="245" masterDocId="FF80FFEEA031FFEBE468FFE4745DFFFD" masterDocTitle="A new species of Besleria (Gesneriaceae) from the western Amazon rainforest" masterLastPageNumber="245" masterPageNumber="241" pageId="0" pageNumber="241" updateTime="1644265614912" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Besleria (Gesneriaceae) from the western Amazon rainforest</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gabriel Emiliano Ferreira</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Andréa Onofre De Araújo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Michael John Gilbert Hopkins</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alain Chautems</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Brittonia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-02-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>241</mods:start>
<mods:end>245</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1007/s12228-017-9464-6</mods:identifier>
<mods:identifier type="GBIF-Dataset">a1d56c0b-d41f-42c1-91d2-417ab340969f</mods:identifier>
<mods:identifier type="ISSN">0007-196X</mods:identifier>
<mods:identifier type="Zenodo-Dep">924479</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5999639" ID-GBIF-Taxon="134839584" ID-Zenodo-Dep="5999639" LSID="urn:lsid:plazi:treatment:03B98796A031FFEFE6C3FC0E75AFFD13" httpUri="http://treatment.plazi.org/id/03B98796A031FFEFE6C3FC0E75AFFD13" lastPageId="4" lastPageNumber="245" pageId="0" pageNumber="241">
<subSubSection pageId="0" pageNumber="241" type="nomenclature">
<paragraph blockId="0.[683,1196,1002,1142]" pageId="0" pageNumber="241">
<taxonomicName authority="G. E. Ferreira &amp; M. J. G. Hopkins" authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[683,1192,1002,1025]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="241" phylum="Tracheophyta" rank="species" species="iara" status="sp. nov.">
<emphasis bold="true" box="[683,824,1002,1025]" pageId="0" pageNumber="241">Besleria iara</emphasis>
G. E. Ferreira &amp; M. J. G. Hopkins
</taxonomicName>
,
<taxonomicNameLabel box="[711,800,1032,1055]" pageId="0" pageNumber="241" rank="species">
<emphasis bold="true" box="[711,800,1032,1055]" pageId="0" pageNumber="241">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="241" type="materials_examined">
<paragraph blockId="0.[683,1196,1002,1142]" pageId="0" pageNumber="241">
<materialsCitation ID-GBIF-Occurrence="1655986898" collectingDate="2016-02-08" collectionCode="INPA" collectorName="G. E. Ferreira" country="Brazil" location="Benjamin Constant" municipality="Type" pageId="0" pageNumber="241" specimenCount="2" stateProvince="Amazonas" typeStatus="holotype">
<collectingMunicipality box="[811,868,1032,1055]" pageId="0" pageNumber="241">
<typeStatus box="[811,868,1032,1055]" pageId="0" pageNumber="241">Type</typeStatus>
</collectingMunicipality>
:
<collectingCountry box="[882,949,1032,1055]" name="Brazil" pageId="0" pageNumber="241">Brazil</collectingCountry>
.
<collectingRegion box="[964,1079,1032,1055]" country="Peru" name="Amazonas" pageId="0" pageNumber="241">Amazonas</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:03B98796A031FFEFE6C3FC0E75AFFD13:8ECF605BA031FFEBE02DFBEC777BFBC1" country="Brazil" municipality="Type" name="Benjamin Constant" pageId="0" pageNumber="241" stateProvince="Amazonas">Benjamin Constant</location>
, sítio Prof. Agno Nonato, trilha prox. ao açude,
<collectingDate box="[815,990,1090,1113]" pageId="0" pageNumber="241" value="2016-02-08">
<date box="[815,990,1090,1113]" pageId="0" pageNumber="241" value="2016-02-08">8 February 2016</date>
</collectingDate>
,
<emphasis box="[1000,1195,1090,1113]" italics="true" pageId="0" pageNumber="241">
<collectorName box="[1000,1150,1090,1113]" pageId="0" pageNumber="241">G. E. Ferreira</collectorName>
355
</emphasis>
(
<typeStatus box="[718,814,1119,1142]" pageId="0" pageNumber="241">holotype</typeStatus>
:
<collectionCode box="[823,885,1119,1142]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="0" pageNumber="241">INPA</collectionCode>
; isotype: G). (
<figureCitation box="[1033,1105,1119,1142]" captionStart="FIG" captionStartId="1.[151,164,1518,1537]" captionTargetBox="[128,1173,169,1502]" captionTargetId="figure@1.[128,1173,168,1502]" captionTargetPageId="1" captionText="FIG. 1. Besleria iara. A. Habit. B. Open Corolla showing stamens and staminode. C. Gynoecium and Nectary. D. Close up of the inflorescence and stem. E. Calyx and Corolla in side view. (A E from G. E. Ferreira 355)." httpUri="https://zenodo.org/record/924481/files/figure.png" pageId="0" pageNumber="241">Figs. 1</figureCitation>
,
<figureCitation box="[1118,1131,1119,1142]" captionStart="FIG" captionStartId="2.[170,183,946,965]" captionTargetBox="[343,997,168,930]" captionTargetId="figure@2.[343,997,168,930]" captionTargetPageId="2" captionText="FIG. 2. Morphology and habitat of Besleria iara. A. Flowers in frontal and side view and a portion of the stem. B. Flower in side view. C. Habit. D. Inflorescence. E. Detail of the leaf abaxial surface (A E from G. E. Ferreira 355)." httpUri="https://zenodo.org/record/924485/files/figure.png" pageId="0" pageNumber="241">2</figureCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="241" type="diagnosis">
<paragraph blockId="0.[685,1196,1181,1331]" pageId="0" pageNumber="241">
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[712,819,1181,1200]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="241" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[712,819,1181,1200]" italics="true" pageId="0" pageNumber="241">Besleria iara</emphasis>
</taxonomicName>
is similar to
<taxonomicName authority="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems, 2017" authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" baseAuthorityName="Mart." box="[937,1044,1181,1200]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="241" phylum="Tracheophyta" rank="species" sensu="lato" species="aggregata" status="s.l.">
<emphasis box="[937,1044,1181,1200]" italics="true" pageId="0" pageNumber="241">B. aggregata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1053,1077,1181,1200]" pageId="0" pageNumber="241" sensu="lato">s.l.</taxonomicNameLabel>
in its axillary inflorescences, and flowers with orange imbricate
<taxonomicName box="[1083,1125,1207,1226]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="0" pageNumber="242" phylum="Porifera" rank="genus">calyx</taxonomicName>
lobes. It differs by having a long tubular-ventricose yellow corolla (vs. ventricose-urceolate orange to scarlet), and larger leaf blades, 18 30 ×
<quantity box="[728,826,1285,1305]" metricMagnitude="-2" metricUnit="m" metricValue="9.35" metricValueMax="11.4" metricValueMin="7.3" pageId="0" pageNumber="241" unit="cm" value="9.35" valueMax="11.4" valueMin="7.3">7.311.4 cm</quantity>
, with serrate margins (vs. 1519 ×
<quantity box="[1103,1191,1285,1305]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.4" metricValueMin="5.5" pageId="0" pageNumber="241" unit="cm" value="5.95" valueMax="6.4" valueMin="5.5">5.56.4 cm</quantity>
, with entire margins).
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="243" pageId="0" pageNumber="241" type="description">
<paragraph blockId="0.[685,1196,1363,1650]" lastBlockId="2.[143,654,1056,1430]" lastPageId="2" lastPageNumber="243" pageId="0" pageNumber="241">
<emphasis box="[712,891,1363,1386]" italics="true" pageId="0" pageNumber="241">Terrestrial shrub,</emphasis>
<quantity box="[896,954,1363,1386]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="0" pageNumber="241" unit="m" value="1.5">1.5 m</quantity>
tall.
<emphasis box="[1003,1063,1363,1386]" italics="true" pageId="0" pageNumber="241">Stems</emphasis>
unbranched, subterete, up to
<quantity box="[866,942,1392,1415]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="241" unit="cm" value="0.5">0.5 cm</quantity>
diam., woody at base, densely hirsute, internodes 3.011.0 cm.
<emphasis box="[1124,1196,1422,1445]" italics="true" pageId="0" pageNumber="241">Leaves</emphasis>
opposite, pairs subequal; petioles
<quantity box="[1050,1132,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="0" pageNumber="241" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">14 cm</quantity>
long, green, densely hirsute; leaf blades oblanceolate, 1830 ×
<quantity box="[783,917,1509,1533]" metricMagnitude="-2" metricUnit="m" metricValue="9.35" metricValueMax="11.4" metricValueMin="7.3" pageId="0" pageNumber="241" unit="cm" value="9.35" valueMax="11.4" valueMin="7.3">7.311.4 cm</quantity>
, chartaceous, apex attenuate, base long-attenuate, margin sparsely serrate; adaxial surface dark green, hirsute; abaxial surface pale green, puberulous; secondary veins in 71 1 pairs.
<emphasis box="[830,982,1626,1649]" italics="true" pageId="0" pageNumber="241">Inflorescence</emphasis>
axillary, cymose, epedunculate, with 37 flowers per node; pedicels terete,
<quantity box="[196,315,1640,1664]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="1" pageNumber="242" unit="cm" value="1.75" valueMax="2.3" valueMin="1.2">1.22.3 cm</quantity>
long, orangish, hirsute.
<emphasis box="[574,634,1641,1664]" italics="true" pageId="1" pageNumber="242">Calyx</emphasis>
lobes 5, fused at base and imbricate for 3/4 of length, adherent to corolla base, essentially equal, ovate, 1214 ×
<quantity box="[843,937,1611,1635]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="242" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
, orange, hirsute, apex obtuse, margin entire-ciliate.
<emphasis box="[980,1060,1641,1664]" italics="true" pageId="1" pageNumber="242">Corolla</emphasis>
erect with respect to
<taxonomicName box="[771,830,1670,1693]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="1" pageNumber="242" phylum="Porifera" rank="genus">calyx</taxonomicName>
,
<quantity box="[839,951,1670,1694]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.7" metricValueMin="1.8" pageId="1" pageNumber="242" unit="mm" value="22.5" valueMax="27.0" valueMin="18.0">1827 mm</quantity>
long, tubular, slightly ventricose, yellow, outside puberulous, inside glabrous, tube contracted at base,
<quantity box="[505,568,1056,1079]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="243" unit="mm" value="5.0">5 mm</quantity>
wide at base,
<quantity box="[204,266,1086,1109]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="243" unit="mm" value="7.0">7 mm</quantity>
wide at middle,
<quantity box="[444,506,1086,1109]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="243" unit="mm" value="5.0">5 mm</quantity>
wide at apex; limb regular, lobes 5, patent, yellow, unequal, the 2 larger ones 4 ×
<quantity box="[327,391,1144,1167]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="243" unit="mm" value="4.0">4 mm</quantity>
, the 3 smaller ones 2 ×
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="243" unit="mm" value="3.0">3 mm</quantity>
, obtuse, margin entire, ciliate, pilose.
<emphasis box="[572,654,1173,1196]" italics="true" pageId="2" pageNumber="243">Nectary</emphasis>
annular,
<quantity box="[232,293,1202,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="243" unit="mm" value="1.0">1 mm</quantity>
wide, glabrous.
<emphasis box="[462,549,1202,1225]" italics="true" pageId="2" pageNumber="243">Stamens</emphasis>
4, includ- ed; filaments
<quantity box="[288,390,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="243" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
long, epipetal, adnate to base of corolla for
<quantity box="[347,412,1261,1284]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="243" unit="mm" value="4.0">4 mm</quantity>
, glabrous but pilose at insertion; anthers 2 ×
<quantity box="[367,430,1290,1313]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="243" unit="mm" value="2.0">2 mm</quantity>
, longitudinally dehiscent; staminode
<quantity box="[315,377,1319,1342]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="243" unit="mm" value="2.0">2 mm</quantity>
long.
<emphasis box="[445,510,1319,1342]" italics="true" pageId="2" pageNumber="243">Ovary</emphasis>
ovoid, 24 ×
<quantity box="[143,236,1348,1372]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="2" pageNumber="243" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
, pilose; style included within corolla throat,
<quantity box="[219,282,1378,1401]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="243" unit="mm" value="9.0">9 mm</quantity>
long, pilose; stigma bifid, puberulent.
<emphasis box="[195,248,1407,1430]" italics="true" pageId="2" pageNumber="243">Fruit</emphasis>
not seen.
</paragraph>
</subSubSection>
<footnote pageId="0" pageNumber="241">
<paragraph blockId="0.[143,794,1762,1823]" box="[143,665,1762,1780]" pageId="0" pageNumber="241">
<taxonomicName authorityName="Houghton &amp; C.A. Armstrong" authorityYear="1934" box="[143,212,1763,1779]" class="Magnoliopsida" family="Cactaceae" genus="Brittonia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="0" pageNumber="241" phylum="Tracheophyta" rank="genus">
<docJournal box="[143,212,1763,1779]" pageId="0" pageNumber="241">
<emphasis box="[143,212,1763,1779]" italics="true" pageId="0" pageNumber="241">Brittonia</emphasis>
</docJournal>
</taxonomicName>
<docVolume box="[218,240,1762,1780]" pageId="0" pageNumber="241">69</docVolume>
(2): 241245 (
<docYear box="[349,391,1762,1780]" pageId="0" pageNumber="241">2017</docYear>
), DOI 10.1007/s12228-017-9464-6
</paragraph>
<paragraph blockId="0.[143,794,1762,1823]" box="[143,592,1784,1802]" pageId="0" pageNumber="241">ISSN: 0007-196X (print) ISSN: 1938-436X (electronic)</paragraph>
<paragraph blockId="0.[143,794,1762,1823]" box="[143,794,1805,1823]" pageId="0" pageNumber="241">© 2017, by The New York Botanical Garden Press, Bronx, NY 10458-5126 U.S.A.</paragraph>
<paragraph blockId="0.[143,426,1845,1863]" box="[143,426,1845,1863]" pageId="0" pageNumber="241">Published Online: 15 February 2017</paragraph>
</footnote>
<caption httpUri="https://zenodo.org/record/924481/files/figure.png" pageId="1" pageNumber="242" targetBox="[128,1173,169,1502]" targetPageId="1">
<paragraph blockId="1.[124,1176,1518,1561]" pageId="1" pageNumber="242">
<emphasis bold="true" box="[151,164,1518,1537]" pageId="1" pageNumber="242">
<emphasis bold="true" box="[164,181,1522,1535]" pageId="1" pageNumber="242">FIG</emphasis>
</emphasis>
<emphasis bold="true" box="[180,205,1518,1537]" pageId="1" pageNumber="242">. 1.</emphasis>
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[222,329,1518,1537]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="242" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[222,329,1518,1537]" italics="true" pageId="1" pageNumber="242">
<taxonomicName box="[222,290,1518,1537]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="242" phylum="Tracheophyta" rank="genus">Besleria</taxonomicName>
<taxonomicName box="[295,329,1518,1537]" pageId="1" pageNumber="242" rank="species" species="iara">iara</taxonomicName>
</emphasis>
</taxonomicName>
.
<emphasis bold="true" box="[338,358,1518,1537]" pageId="1" pageNumber="242">A.</emphasis>
Habit.
<emphasis bold="true" box="[417,436,1518,1537]" pageId="1" pageNumber="242">B.</emphasis>
Open Corolla showing stamens and staminode.
<emphasis bold="true" box="[827,847,1518,1537]" pageId="1" pageNumber="242">C.</emphasis>
Gynoecium and Nectary.
<emphasis bold="true" box="[1059,1079,1518,1537]" pageId="1" pageNumber="242">D.</emphasis>
Close up of the inflorescence and stem.
<emphasis bold="true" box="[353,373,1542,1561]" pageId="1" pageNumber="242">E.</emphasis>
Calyx and Corolla in side view. (AE from
<emphasis box="[740,889,1542,1561]" italics="true" pageId="1" pageNumber="242">G.E. Ferreira 355</emphasis>
).
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/924485/files/figure.png" pageId="2" pageNumber="243" targetBox="[343,997,168,930]" targetPageId="2">
<paragraph blockId="2.[143,1196,946,989]" pageId="2" pageNumber="243">
<emphasis bold="true" box="[170,183,946,965]" pageId="2" pageNumber="243">
<emphasis bold="true" box="[183,200,950,963]" pageId="2" pageNumber="243">FIG</emphasis>
</emphasis>
<emphasis bold="true" box="[199,225,946,965]" pageId="2" pageNumber="243">. 2.</emphasis>
Morphology and habitat of
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[467,574,946,965]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="243" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[467,574,946,965]" italics="true" pageId="2" pageNumber="243">Besleria iara</emphasis>
</taxonomicName>
.
<emphasis bold="true" box="[585,605,946,965]" pageId="2" pageNumber="243">A.</emphasis>
Flowers in frontal and side view and a portion of the stem.
<emphasis bold="true" box="[1092,1111,946,965]" pageId="2" pageNumber="243">B.</emphasis>
Flower in side view.
<emphasis bold="true" box="[231,251,970,989]" pageId="2" pageNumber="243">C.</emphasis>
Habit.
<emphasis bold="true" box="[313,333,970,989]" pageId="2" pageNumber="243">D.</emphasis>
Inflorescence.
<emphasis bold="true" box="[459,478,970,989]" pageId="2" pageNumber="243">E.</emphasis>
Detail of the leaf abaxial surface (AE from
<emphasis box="[851,1001,970,989]" italics="true" pageId="2" pageNumber="243">G.E. Ferreira 355</emphasis>
).
</paragraph>
</caption>
<subSubSection pageId="2" pageNumber="243" type="distribution">
<paragraph blockId="2.[143,653,1465,1722]" lastBlockId="2.[685,1196,1058,1557]" pageId="2" pageNumber="243">
<emphasis box="[170,586,1465,1489]" italics="true" pageId="2" pageNumber="243">
Habitat and distribution.—
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[451,586,1465,1488]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="243" phylum="Tracheophyta" rank="species" species="iara">Besleria iara</taxonomicName>
</emphasis>
is currently known only from the Javari River, in the municipalities of Atalaia do Norte and Benjamin Constant,
<collectingRegion box="[247,355,1553,1576]" country="Peru" name="Amazonas" pageId="2" pageNumber="243">Amazonas</collectingRegion>
,
<collectingCountry box="[367,429,1553,1576]" name="Brazil" pageId="2" pageNumber="243">Brazil</collectingCountry>
, and from
<collectingRegion box="[546,612,1553,1576]" country="Peru" name="Loreto" pageId="2" pageNumber="243">Loreto</collectingRegion>
department,
<collectingCountry box="[241,287,1582,1605]" name="Peru" pageId="2" pageNumber="243">Peru</collectingCountry>
(Fig. 3). It occurs in the understory of lowland Amazonian rainforests on terra firme, along the margins of small streams, at about
<quantity box="[533,586,1641,1664]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="243" unit="m" value="50.0">50 m</quantity>
elevation. We observed few individuals at the
<typeStatus box="[549,592,1670,1693]" pageId="2" pageNumber="243">type</typeStatus>
locality. The larger area of distribution is poorly known from a floristic context. It is expected that additional fieldwork in the region will result in the discovery of additional populations.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="243" type="biology_ecology">
<paragraph blockId="2.[685,1196,1058,1557]" pageId="2" pageNumber="243">
<emphasis box="[712,852,1148,1171]" italics="true" pageId="2" pageNumber="243">Phenology.—</emphasis>
Flowers recorded from October to February.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="243" type="etymology">
<paragraph blockId="2.[685,1196,1058,1557]" pageId="2" pageNumber="243">
<emphasis box="[712,851,1206,1229]" italics="true" pageId="2" pageNumber="243">Etymology.—</emphasis>
The specific epithet B
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[1088,1131,1206,1229]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="243" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[1088,1131,1206,1229]" italics="true" pageId="2" pageNumber="243">iara</emphasis>
</taxonomicName>
^ is a reference to the Brazilian indigenous legend of the water spirit
<emphasis box="[869,926,1265,1288]" italics="true" pageId="2" pageNumber="243">yîara</emphasis>
, meaning BLady of the waters^ in the Tupí-Guaraní language.
<emphasis box="[1110,1168,1294,1317]" italics="true" pageId="2" pageNumber="243">Yîara</emphasis>
is generally envisaged as a beautiful young lady, similar to a mermaid (
<bibRefCitation author="Chiaradia, C." box="[915,1082,1352,1375]" journalOrPublisher="Limiar" pageId="2" pageNumber="243" refString="Chiaradia, C. 2008. Dicionario de palavras brasileiras de origem indigena. Limiar." title="Dicionario de palavras brasileiras de origem indigena" type="book" year="2008">Chiaradia, 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="243" type="conservation">
<paragraph blockId="2.[685,1196,1058,1557]" pageId="2" pageNumber="243">
<emphasis box="[712,947,1382,1405]" italics="true" pageId="2" pageNumber="243">Conservation status.—</emphasis>
Endangered (EN) B1ab, according to the
<collectionCode box="[863,927,1412,1435]" pageId="2" pageNumber="243">IUCN</collectionCode>
criteria, based on the extent of occurrence estimated to be less than
<quantity metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="243" unit="km" value="5000.0">5,000 km</quantity>
2 from only 45 locations and due to continued habitat loss (
<bibRefCitation author="IUCN" box="[822,955,1504,1527]" journalOrPublisher="IUCN, Gland, Switzerland and Cambridge, UK." pageId="2" pageNumber="243" publicationUrl="www.iucnredlist.org/technicaldocuments/categories-and-criteria" refString="IUCN. 2012. IUCN Red List categories and criteria. Version 3.1. Second edition. IUCN, Gland, Switzerland and Cambridge, UK. Available at: www. iucnredlist. org / technicaldocuments / categories-and-criteria." title="IUCN Red List categories and criteria. Version 3.1. Second edition" type="book" year="2012">
<collectionCode box="[822,890,1504,1527]" pageId="2" pageNumber="243">IUCN</collectionCode>
, 2012
</bibRefCitation>
;
<bibRefCitation author="IUCN Standards and Petitions Subcommittee" journalOrPublisher="Version 12. Available at" pageId="2" pageNumber="243" publicationUrl="http://www.iucnredlist. org/documents/RedListGuidelines.pdf" refString="IUCN Standards and Petitions Subcommittee. 2016. Guidelines for using the IUCN Red List categories and criteria. Version 12. Available at: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf." title="Guidelines for using the IUCN Red List categories and criteria" type="book" year="2016">
<collectionCode box="[973,1037,1504,1527]" pageId="2" pageNumber="243">IUCN</collectionCode>
Standards and Petitions Subcommittee, 2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="244" pageId="2" pageNumber="243" type="materials_examined">
<paragraph blockId="2.[685,1197,1598,1721]" pageId="2" pageNumber="243">
<materialsCitation ID-GBIF-Occurrence="1655986896" collectingDate="1989-01-18" collectionCode="CEN, INPA" collectorName="de Sao Joaquim &amp; C. A. C. Ferreira" country="Brazil" county="Municipio Atalaia do Norte" location="Brasil" municipality="Rio Javari" pageId="2" pageNumber="243" specimenCount="1" stateProvince="Amazonas">
<emphasis bold="true" box="[712,1196,1598,1617]" pageId="2" pageNumber="243">
Additional specimens examined.
<collectingCountry box="[1005,1086,1598,1617]" name="Brazil" pageId="2" pageNumber="243">BRAZIL</collectingCountry>
.
<collectingRegion box="[1098,1193,1598,1617]" country="Peru" name="Amazonas" pageId="2" pageNumber="243">Amazonas</collectingRegion>
:
</emphasis>
<collectingCounty box="[685,919,1624,1643]" pageId="2" pageNumber="243">Município Atalaia do Norte</collectingCounty>
, localidade
<collectorName box="[1020,1155,1624,1643]" pageId="2" pageNumber="243">de São Joaquim</collectorName>
,
<collectingMunicipality pageId="2" pageNumber="243">Rio Javari</collectingMunicipality>
, fronteira entre
<location LSID="urn:lsid:plazi:treatment:03B98796A031FFEFE6C3FC0E75AFFD13:8ECF605BA033FFE9E707F99677FDF978" box="[879,928,1650,1669]" country="Brazil" county="Municipio Atalaia do Norte" municipality="Rio Javari" name="Brasil" pageId="2" pageNumber="243" stateProvince="Amazonas">Brasil</location>
e
<location LSID="urn:lsid:plazi:treatment:03B98796A031FFEFE6C3FC0E75AFFD13:8ECF605BA033FFE9E7D3F99677BEF978" box="[955,995,1650,1669]" country="Brazil" county="Municipio Atalaia do Norte" municipality="Rio Javari" name="Peru" pageId="2" pageNumber="243" stateProvince="Amazonas">Peru</location>
,
<collectingDate box="[1007,1118,1650,1669]" pageId="2" pageNumber="243" value="1989-01-18">
<date box="[1007,1118,1650,1669]" pageId="2" pageNumber="243" value="1989-01-18">18 Jan 1989</date>
</collectingDate>
,
<emphasis italics="true" pageId="2" pageNumber="243">
<collectorName pageId="2" pageNumber="243">C. A.C. Ferreira</collectorName>
et al. 993 3
</emphasis>
(
<collectionCode box="[858,902,1676,1695]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="2" pageNumber="243">CEN</collectionCode>
,
<collectionCode box="[910,959,1676,1695]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="243">INPA</collectionCode>
, NY, US)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1655986897" collectingDate="1976-10-20" collectorName="IUCN Standards and Petitions Subcommittee" country="Ecuador" location="Rio Javari" municipality="Municipio Atalaia do Norte" pageId="2" pageNumber="243" specimenCount="1">
<collectingMunicipality pageId="2" pageNumber="243">Município Atalaia do Norte</collectingMunicipality>
, Estirão do
<collectingCountry box="[857,928,1702,1721]" name="Ecuador" pageId="2" pageNumber="243">Equador</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:03B98796A031FFEFE6C3FC0E75AFFD13:8ECF605BA033FFE9E7C0F94277A6F944" box="[936,1019,1702,1721]" country="Ecuador" municipality="Municipio Atalaia do Norte" name="Rio Javari" pageId="2" pageNumber="243">Rio Javari</location>
,
<collectingDate box="[1030,1134,1702,1721]" pageId="2" pageNumber="243" value="1976-10-20">
<date box="[1030,1134,1702,1721]" pageId="2" pageNumber="243" value="1976-10-20">20 Oct 1976</date>
</collectingDate>
,
<emphasis box="[1142,1194,1702,1721]" italics="true" pageId="2" pageNumber="243">
P.
<collectorName box="[1163,1194,1702,1721]" name="IUCN Standards and Petitions Subcommittee" pageId="2" pageNumber="243">I. S.</collectorName>
</emphasis>
</materialsCitation>
</paragraph>
<caption box="[151,463,844,863]" httpUri="https://zenodo.org/record/924489/files/figure.png" pageId="3" pageNumber="244" targetBox="[196,1104,169,828]" targetPageId="3">
<paragraph blockId="3.[151,463,844,863]" box="[151,463,844,863]" pageId="3" pageNumber="244">
<emphasis bold="true" box="[151,164,844,863]" pageId="3" pageNumber="244">
<emphasis bold="true" box="[164,181,848,861]" pageId="3" pageNumber="244">FIG</emphasis>
</emphasis>
<emphasis bold="true" box="[180,206,844,863]" pageId="3" pageNumber="244">. 3.</emphasis>
Distribution of
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[350,458,844,863]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[350,458,844,863]" italics="true" pageId="3" pageNumber="244">Besleria iara</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="3.[124,636,940,1101]" pageId="3" pageNumber="244">
<emphasis box="[124,390,940,959]" italics="true" pageId="3" pageNumber="244">
<taxonomicName authority="J. R. Nascimento" authorityName="Schioedte &amp; Meinert" authorityYear="1881" box="[124,176,940,959]" class="Malacostraca" family="Cymothoidae" genus="Braga" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="244" phylum="Arthropoda" rank="genus">Braga</taxonomicName>
&amp; J. R. Nascimento 3273
</emphasis>
(
<collectionCode box="[402,453,940,959]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="244">INPA</collectionCode>
).
<emphasis bold="true" box="[467,598,940,959]" pageId="3" pageNumber="244">
<collectionCode box="[467,523,940,959]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14398" name="Universit<69> di Perugia" pageId="3" pageNumber="244">
<collectingCountry box="[467,523,940,959]" name="Peru" pageId="3" pageNumber="244">PERU</collectingCountry>
</collectionCode>
.
<collectingRegion box="[534,593,940,959]" country="Peru" name="Loreto" pageId="3" pageNumber="244">Loreto</collectingRegion>
:
</emphasis>
Rio Javari, 2 hours above Rio Javarizinho,
<date box="[467,579,966,985]" pageId="3" pageNumber="244" value="1976-10-24">24 Oct 1976</date>
,
<emphasis italics="true" pageId="3" pageNumber="244">G. T. Prance et al. 24081</emphasis>
(
<collectionCode box="[298,347,992,1011]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="244">INPA</collectionCode>
, MG).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="245" pageId="3" pageNumber="244" type="discussion">
<paragraph blockId="3.[124,636,940,1101]" lastBlockId="4.[143,654,179,750]" lastPageId="4" lastPageNumber="245" pageId="3" pageNumber="244">
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[151,289,1020,1043]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[151,289,1020,1043]" italics="true" pageId="3" pageNumber="244">Besleria iara</emphasis>
</taxonomicName>
and
<taxonomicName authority="(Mart.) Hanst." authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" baseAuthorityName="Mart." box="[342,635,1020,1043]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="aggregata">
<emphasis box="[342,478,1020,1043]" italics="true" pageId="3" pageNumber="244">B. aggregata</emphasis>
(Mart.) Hanst.
</taxonomicName>
are sympatric and share the same habitat and vegetative characters. Both species occur in the understory of humid forest, close to streams, and are shrubs completely covered by indumentum. However,
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[773,845,995,1018]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[773,845,995,1018]" italics="true" pageId="3" pageNumber="244">B. iara</emphasis>
</taxonomicName>
has a yellow, tubular-ventricose corolla, ovate
<taxonomicName box="[816,872,1023,1046]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="242" phylum="Porifera" rank="genus">calyx</taxonomicName>
lobes with the obtuse apices adherent to the corolla base, wider leaf blades (
<quantity box="[674,803,1078,1102]" metricMagnitude="-2" metricUnit="m" metricValue="9.35" metricValueMax="11.4" metricValueMin="7.3" pageId="3" pageNumber="244" unit="cm" value="9.35" valueMax="11.4" valueMin="7.3">7.311.4 cm</quantity>
wide) with sparsely serrate margins, and densely hirsute indumentum on the stems, whereas
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" baseAuthorityName="Mart." box="[239,378,207,230]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="245" phylum="Tracheophyta" rank="species" species="aggregata">
<emphasis box="[239,378,207,230]" italics="true" pageId="4" pageNumber="245">B. aggregata</emphasis>
</taxonomicName>
has an orange to scarlet, ventricose-urceolate corolla, lanceolate
<taxonomicName box="[595,654,235,258]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="242" phylum="Porifera" rank="genus">calyx</taxonomicName>
lobes with the attenuate apices spreading from the corolla, narrower leaf blades (
<quantity box="[528,653,291,315]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.4" metricValueMin="5.5" pageId="4" pageNumber="245" unit="cm" value="5.95" valueMax="6.4" valueMin="5.5">5.56.4 cm</quantity>
wide) with entire margins, and sericeous indumentum on the stems (Table 1).
</paragraph>
<caption box="[136,759,1201,1220]" pageId="3" pageNumber="244">
<paragraph blockId="3.[136,759,1201,1220]" box="[136,759,1201,1220]" pageId="3" pageNumber="244">
<emphasis bold="true" box="[136,149,1201,1220]" pageId="3" pageNumber="244">
<emphasis bold="true" box="[149,192,1204,1218]" pageId="3" pageNumber="244">TABLE</emphasis>
</emphasis>
<emphasis bold="true" box="[197,208,1201,1220]" pageId="3" pageNumber="244">1</emphasis>
Morphological comparisons of
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[484,592,1201,1220]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[484,592,1201,1220]" italics="true" pageId="3" pageNumber="244">
<taxonomicName box="[484,552,1201,1220]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="genus">Besleria</taxonomicName>
iara
</emphasis>
</taxonomicName>
and related species.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="244">
<table box="[124,1176,1263,1707]" gridcols="4" gridrows="18" pageId="3" pageNumber="244">
<tr box="[124,1176,1263,1282]" gridrow="0" pageId="3" pageNumber="244">
<th box="[124,450,1263,1282]" gridcol="0" gridrow="0" pageId="3" pageNumber="244">Traits/Species</th>
<emphasis box="[524,1030,1263,1282]" italics="true" pageId="3" pageNumber="244">
<th box="[522,659,1263,1282]" gridcol="1" gridrow="0" pageId="3" pageNumber="244">
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" baseAuthorityName="Mart." box="[524,631,1263,1282]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="aggregata">B. aggregata</taxonomicName>
</th>
<th box="[732,878,1263,1282]" gridcol="2" gridrow="0" pageId="3" pageNumber="244">
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[735,793,1263,1282]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="iara">B. iara</taxonomicName>
</th>
<th box="[950,1176,1263,1282]" gridcol="3" gridrow="0" pageId="3" pageNumber="244">
<taxonomicName box="[951,1030,1263,1282]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="244" phylum="Tracheophyta" rank="species" species="obtusa">
B.
<taxonomicName box="[975,1030,1263,1282]" pageId="3" pageNumber="244" rank="species" species="obtusa">obtusa</taxonomicName>
</taxonomicName>
</th>
</emphasis>
</tr>
<tr box="[124,1176,1305,1324]" gridrow="1" pageId="3" pageNumber="244">
<th box="[124,450,1305,1324]" gridcol="0" gridrow="1" pageId="3" pageNumber="244">Petiole length [cm]</th>
<td box="[522,659,1305,1324]" gridcol="1" gridrow="1" pageId="3" pageNumber="244">1.63.9</td>
<td box="[732,878,1305,1324]" gridcol="2" gridrow="1" pageId="3" pageNumber="244">1.04.0</td>
<td box="[950,1176,1305,1324]" gridcol="3" gridrow="1" pageId="3" pageNumber="244">&lt;1.0</td>
</tr>
<tr box="[124,1176,1329,1348]" gridrow="2" pageId="3" pageNumber="244">
<th box="[124,450,1329,1348]" gridcol="0" gridrow="2" pageId="3" pageNumber="244">Leaf blade size [cm]</th>
<td box="[522,659,1329,1348]" gridcol="1" gridrow="2" pageId="3" pageNumber="244">1519 × 5.56.4</td>
<td box="[732,878,1329,1348]" gridcol="2" gridrow="2" pageId="3" pageNumber="244">1830 × 7.311.4</td>
<td box="[950,1176,1329,1348]" gridcol="3" gridrow="2" pageId="3" pageNumber="244">1823 × 5.79.5</td>
</tr>
<tr box="[124,1176,1353,1372]" gridrow="3" pageId="3" pageNumber="244">
<th box="[124,450,1353,1372]" gridcol="0" gridrow="3" pageId="3" pageNumber="244">Leaf blade indumentum abaxial surface</th>
<td box="[522,659,1353,1372]" gridcol="1" gridrow="3" pageId="3" pageNumber="244">sericeous</td>
<td box="[732,878,1353,1372]" gridcol="2" gridrow="3" pageId="3" pageNumber="244">puberulous</td>
<td box="[950,1176,1353,1372]" gridcol="3" gridrow="3" pageId="3" pageNumber="244">tomentose-hirsute at midrib</td>
</tr>
<tr box="[124,1176,1377,1396]" gridrow="4" pageId="3" pageNumber="244">
<th box="[124,450,1377,1396]" gridcol="0" gridrow="4" pageId="3" pageNumber="244">Leaf blade margin</th>
<td box="[522,659,1377,1396]" gridcol="1" gridrow="4" pageId="3" pageNumber="244">entire</td>
<td box="[732,878,1377,1396]" gridcol="2" gridrow="4" pageId="3" pageNumber="244">sparsely serrate</td>
<td box="[950,1176,1377,1396]" gridcol="3" gridrow="4" pageId="3" pageNumber="244">entire-ciliate</td>
</tr>
<tr box="[124,1176,1401,1420]" gridrow="5" pageId="3" pageNumber="244">
<th box="[124,450,1401,1420]" gridcol="0" gridrow="5" pageId="3" pageNumber="244">Number of flowers per axil</th>
<td box="[522,659,1401,1420]" gridcol="1" gridrow="5" pageId="3" pageNumber="244">15</td>
<td box="[732,878,1401,1420]" gridcol="2" gridrow="5" pageId="3" pageNumber="244">37</td>
<td box="[950,1176,1401,1420]" gridcol="3" gridrow="5" pageId="3" pageNumber="244">310</td>
</tr>
<tr box="[124,1176,1425,1444]" gridrow="6" pageId="3" pageNumber="244">
<th box="[124,450,1425,1444]" gridcol="0" gridrow="6" pageId="3" pageNumber="244">Pedicel indumentum</th>
<td box="[522,659,1425,1444]" gridcol="1" gridrow="6" pageId="3" pageNumber="244">pilose</td>
<td box="[732,878,1425,1444]" gridcol="2" gridrow="6" pageId="3" pageNumber="244">hirsute</td>
<td box="[950,1176,1425,1444]" gridcol="3" gridrow="6" pageId="3" pageNumber="244">pilose</td>
</tr>
<tr box="[124,1176,1448,1468]" gridrow="7" pageId="3" pageNumber="244">
<th box="[124,450,1448,1468]" gridcol="0" gridrow="7" pageId="3" pageNumber="244">Pedicel length [cm]</th>
<td box="[522,659,1448,1468]" gridcol="1" gridrow="7" pageId="3" pageNumber="244">0.50.8</td>
<td box="[732,878,1448,1468]" gridcol="2" gridrow="7" pageId="3" pageNumber="244">1.22.3</td>
<td box="[950,1176,1448,1468]" gridcol="3" gridrow="7" pageId="3" pageNumber="244">0.5</td>
</tr>
<tr box="[124,1176,1473,1492]" gridrow="8" pageId="3" pageNumber="244">
<th box="[124,450,1473,1492]" gridcol="0" gridrow="8" pageId="3" pageNumber="244">Calyx color</th>
<td box="[522,659,1473,1492]" gridcol="1" gridrow="8" pageId="3" pageNumber="244">orange</td>
<td box="[732,878,1473,1492]" gridcol="2" gridrow="8" pageId="3" pageNumber="244">orange</td>
<td box="[950,1176,1473,1492]" gridcol="3" gridrow="8" pageId="3" pageNumber="244">-</td>
</tr>
<tr box="[124,1176,1496,1516]" gridrow="9" pageId="3" pageNumber="244">
<th box="[124,450,1496,1516]" gridcol="0" gridrow="9" pageId="3" pageNumber="244">
<taxonomicName box="[125,174,1497,1516]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="242" phylum="Porifera" rank="genus">Calyx</taxonomicName>
lobe length [mm]
</th>
<td box="[522,659,1496,1516]" gridcol="1" gridrow="9" pageId="3" pageNumber="244">1417</td>
<td box="[732,878,1496,1516]" gridcol="2" gridrow="9" pageId="3" pageNumber="244">1214</td>
<td box="[950,1176,1496,1516]" gridcol="3" gridrow="9" pageId="3" pageNumber="244">815</td>
</tr>
<tr box="[124,1176,1520,1540]" gridrow="10" pageId="3" pageNumber="244">
<th box="[124,450,1520,1540]" gridcol="0" gridrow="10" pageId="3" pageNumber="244">Calyx lobe width [mm]</th>
<td box="[522,659,1520,1540]" gridcol="1" gridrow="10" pageId="3" pageNumber="244">34</td>
<td box="[732,878,1520,1540]" gridcol="2" gridrow="10" pageId="3" pageNumber="244">35</td>
<td box="[950,1176,1520,1540]" gridcol="3" gridrow="10" pageId="3" pageNumber="244">12.5</td>
</tr>
<tr box="[124,1176,1545,1564]" gridrow="11" pageId="3" pageNumber="244">
<th box="[124,450,1545,1564]" gridcol="0" gridrow="11" pageId="3" pageNumber="244">
<taxonomicName box="[125,174,1545,1564]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="242" phylum="Porifera" rank="genus">Calyx</taxonomicName>
lobe shape
</th>
<td box="[522,659,1545,1564]" gridcol="1" gridrow="11" pageId="3" pageNumber="244">lanceolate</td>
<td box="[732,878,1545,1564]" gridcol="2" gridrow="11" pageId="3" pageNumber="244">ovate</td>
<td box="[950,1176,1545,1564]" gridcol="3" gridrow="11" pageId="3" pageNumber="244">linear-lanceolate</td>
</tr>
<tr box="[124,1176,1569,1588]" gridrow="12" pageId="3" pageNumber="244">
<th box="[124,450,1569,1588]" gridcol="0" gridrow="12" pageId="3" pageNumber="244">Calyx lobe apex shape</th>
<td box="[522,659,1569,1588]" gridcol="1" gridrow="12" pageId="3" pageNumber="244">attenuate</td>
<td box="[732,878,1569,1588]" gridcol="2" gridrow="12" pageId="3" pageNumber="244">obtuse</td>
<td box="[950,1176,1569,1588]" gridcol="3" gridrow="12" pageId="3" pageNumber="244">long-acuminate</td>
</tr>
<tr box="[124,1176,1592,1611]" gridrow="13" pageId="3" pageNumber="244">
<th box="[124,450,1592,1611]" gridcol="0" gridrow="13" pageId="3" pageNumber="244">Calyx lobes orientation</th>
<td box="[522,659,1592,1611]" gridcol="1" gridrow="13" pageId="3" pageNumber="244">spreading</td>
<td box="[732,878,1592,1611]" gridcol="2" gridrow="13" pageId="3" pageNumber="244">adherent</td>
<td box="[950,1176,1592,1611]" gridcol="3" gridrow="13" pageId="3" pageNumber="244">spreading</td>
</tr>
<tr box="[124,1176,1616,1635]" gridrow="14" pageId="3" pageNumber="244">
<th box="[124,450,1616,1635]" gridcol="0" gridrow="14" pageId="3" pageNumber="244">Calyx indumentum</th>
<td box="[522,659,1616,1635]" gridcol="1" gridrow="14" pageId="3" pageNumber="244">pilose</td>
<td box="[732,878,1616,1635]" gridcol="2" gridrow="14" pageId="3" pageNumber="244">hirsute</td>
<td box="[950,1176,1616,1635]" gridcol="3" gridrow="14" pageId="3" pageNumber="244">pilose</td>
</tr>
<tr box="[124,1176,1640,1659]" gridrow="15" pageId="3" pageNumber="244">
<th box="[124,450,1640,1659]" gridcol="0" gridrow="15" pageId="3" pageNumber="244">Corolla color</th>
<td box="[522,659,1640,1659]" gridcol="1" gridrow="15" pageId="3" pageNumber="244">orange to scarlet</td>
<td box="[732,878,1640,1659]" gridcol="2" gridrow="15" pageId="3" pageNumber="244">yellow</td>
<td box="[950,1176,1640,1659]" gridcol="3" gridrow="15" pageId="3" pageNumber="244">yellow</td>
</tr>
<tr box="[124,1176,1664,1683]" gridrow="16" pageId="3" pageNumber="244">
<th box="[124,450,1664,1683]" gridcol="0" gridrow="16" pageId="3" pageNumber="244">Corolla length [mm]</th>
<td box="[522,659,1664,1683]" gridcol="1" gridrow="16" pageId="3" pageNumber="244">1620</td>
<td box="[732,878,1664,1683]" gridcol="2" gridrow="16" pageId="3" pageNumber="244">1827</td>
<td box="[950,1176,1664,1683]" gridcol="3" gridrow="16" pageId="3" pageNumber="244">1825</td>
</tr>
<tr box="[124,1176,1688,1707]" gridrow="17" pageId="3" pageNumber="244">
<th box="[124,450,1688,1707]" gridcol="0" gridrow="17" pageId="3" pageNumber="244">Corolla lobe length [mm]</th>
<td box="[522,659,1688,1707]" gridcol="1" gridrow="17" pageId="3" pageNumber="244">23</td>
<td box="[732,878,1688,1707]" gridcol="2" gridrow="17" pageId="3" pageNumber="244">24</td>
<td box="[950,1176,1688,1707]" gridcol="3" gridrow="17" pageId="3" pageNumber="244">56</td>
</tr>
</table>
</paragraph>
<paragraph blockId="4.[143,654,179,750]" pageId="4" pageNumber="245">
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[170,304,377,400]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="245" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[170,304,377,400]" italics="true" pageId="4" pageNumber="245">Besleria iara</emphasis>
</taxonomicName>
is also morphologically similar to
<taxonomicName authority="C. V. Morton" authorityName="C. V. Morton" box="[143,382,406,429]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="245" phylum="Tracheophyta" rank="species" species="obtusa">
<emphasis box="[143,241,406,429]" italics="true" pageId="4" pageNumber="245">B. obtusa</emphasis>
C. V. Morton
</taxonomicName>
, known only from the type collection (
<emphasis box="[258,468,435,458]" italics="true" pageId="4" pageNumber="245">J. Cuatrecasas 4636</emphasis>
, Colombia, Meta, 1939). Both species have oblanceolate leaves and a yellow corolla, but
<taxonomicName authorityName="Gabriel Emiliano Ferreira &amp; Andréa Onofre De Araújo &amp; Michael John Gilbert Hopkins &amp; Alain Chautems" authorityYear="2017" box="[363,435,494,517]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="245" phylum="Tracheophyta" rank="species" species="iara">
<emphasis box="[363,435,494,517]" italics="true" pageId="4" pageNumber="245">B. iara</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName box="[202,303,523,546]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="245" phylum="Tracheophyta" rank="species" species="obtusa">
<emphasis box="[202,303,523,546]" italics="true" pageId="4" pageNumber="245">B. obtusa</emphasis>
</taxonomicName>
by having a longer petiole (1.0 4.0 cm long vs. &lt;
<quantity box="[336,389,552,575]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="245" unit="cm" value="1.0">1 cm</quantity>
long), hirsute leaf blades (vs. glabrous with tomentose-hirsute midrib), sparsely serrate margins (vs. entire-ciliate), longer pedicels (
<quantity box="[242,359,639,663]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="4" pageNumber="245" unit="cm" value="1.75" valueMax="2.3" valueMin="1.2">1.22.3 cm</quantity>
long vs.
<quantity box="[453,524,640,663]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="245" unit="cm" value="0.5">0.5 cm</quantity>
long), ovate
<taxonomicName box="[143,199,669,692]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="242" phylum="Porifera" rank="genus">calyx</taxonomicName>
lobes with the obtuse apices adherent to the corolla (vs.
<taxonomicName box="[267,323,698,721]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="242" phylum="Porifera" rank="genus">calyx</taxonomicName>
lobes linear-lanceolate with the long-acuminate apices spreading).
</paragraph>
</subSubSection>
</treatment>
</document>