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<document id="079F3F78D98A06F544F51EAE1E391744" ID-CLB-Dataset="73097" ID-DOI="10.11646/zootaxa.5154.2.2" ID-GBIF-Dataset="8cb6ae7b-3b75-4480-81d6-fed58cf5e5a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="6641280" ID-ZooBank="89445A18-515F-4A80-8515-7CB39E4DC73F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1655194333503" checkinUser="plazi" docAuthor="Webster, Grant N. &amp; Bool, Ian" docDate="2022" docId="03B98792FFDDFF941ACF429BCD149FC1" docLanguage="en" docName="zootaxa.5154.2.2.pdf" docOrigin="Zootaxa 5154 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anstisia Webster &amp; Bool 2022, new genus" docType="treatment" docUuid="BC602384-BB5E-4A84-A6E5-A6B0A71C80C9" docUuidSource="ZooBank" docVersion="7" lastPageNumber="145" masterDocId="FF80FFEAFFCCFF861A58420CC97E9D30" masterDocTitle="A new genus for four myobatrachid frogs from the South Western Australian Ecoregion" masterLastPageNumber="151" masterPageNumber="127" pageNumber="144" updateTime="1699339323944" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CD44E6B430E1336DEF2B3AFCAA1815C8">
<mods:title id="A93D205D5C7178FF5A275EED34196233">A new genus for four myobatrachid frogs from the South Western Australian Ecoregion</mods:title>
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<mods:namePart id="BC2038DFB769A6D26F9B1CBA044EFF14">Webster, Grant N.</mods:namePart>
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<mods:affiliation id="AE20F4738174D9BF812025F8DE07B990">School of Environmental and Rural Science, University of New England, Armidale, New South Wales, 2350, Australia. grantwebster. aecs @ gmail. com; https: // orcid. org / 0000 - 0002 - 7728 - 4107</mods:affiliation>
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<mods:namePart id="1894A479422DBBED77321F8BF534560D">Bool, Ian</mods:namePart>
<mods:affiliation id="8ABD2AB8E6074C800F1862209E4861EE">School of Environmental, Atmospheric and Life Sciences, Centre for Sustainable Ecosystem Solutions, University of Wollongong, Northfields Avenue, Wollongong, New South Wales, 2522, Australia.</mods:affiliation>
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<mods:title id="FCACCD556B6CECDA523A162A900E9DC7">Zootaxa</mods:title>
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<mods:date id="C09CEBA3E541A8AFC5F73FA3415251E0">2022</mods:date>
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<mods:number id="759288A31D374AD87A36A8E7398335F9">2022-06-14</mods:number>
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<treatment id="03B98792FFDDFF941ACF429BCD149FC1" ID-DOI="http://doi.org/10.5281/zenodo.6646909" ID-GBIF-Taxon="195917558" ID-Zenodo-Dep="6646909" LSID="urn:lsid:zoobank.org:act:BC602384-BB5E-4A84-A6E5-A6B0A71C80C9" httpUri="http://treatment.plazi.org/id/03B98792FFDDFF941ACF429BCD149FC1" lastPageId="18" lastPageNumber="145" pageId="17" pageNumber="144">
<subSubSection id="C30A650FFFDDFF971ACF429BC8FC9D81" box="[151,386,151,177]" pageId="17" pageNumber="144" type="nomenclature">
<paragraph id="8BAF3684FFDDFF971ACF429BC8FC9D81" blockId="17.[151,386,151,177]" box="[151,386,151,177]" pageId="17" pageNumber="144">
<heading id="D0E781E8FFDDFF971ACF429BC8FC9D81" bold="true" box="[151,386,151,177]" fontSize="11" level="1" pageId="17" pageNumber="144" reason="1">
<emphasis id="B964EA96FFDDFF971ACF429BC8FC9D81" bold="true" box="[151,386,151,177]" pageId="17" pageNumber="144">
<taxonomicName id="4C104D07FFDDFF971ACF429BC9879D81" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[151,249,151,177]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDDFF971ACF429BC9879D81" bold="true" box="[151,249,151,177]" italics="true" pageId="17" pageNumber="144">Anstisia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757EDFFDDFF971B58429BC8FC9D81" box="[256,386,151,177]" pageId="17" pageNumber="144" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30A650FFFDDFF971ACF42D0CA369CB4" pageId="17" pageNumber="144" type="reference_group">
<paragraph id="8BAF3684FFDDFF971ACF42D0CBCE9DC4" blockId="17.[151,688,220,244]" box="[151,688,220,244]" pageId="17" pageNumber="144">
<taxonomicName id="4C104D07FFDDFF971ACF42D0C9979DC3" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[151,233,220,243]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDDFF971ACF42D0C9979DC3" box="[151,233,220,243]" italics="true" pageId="17" pageNumber="144">Anstisia</emphasis>
</taxonomicName>
.
<typeStatus id="54AB8826FFDDFF971AAD42D0C8599DC4" box="[245,295,220,244]" pageId="17" pageNumber="144">Type</typeStatus>
species:
<taxonomicName id="4C104D07FFDDFF971BDC42D0CBD29DC4" ID-CoL="ZGD9" authority=", Harrison, 1927" authorityName=", Harrison" authorityYear="1927" box="[388,684,220,244]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="rosea">
<emphasis id="B964EA96FFDDFF971BDC42D0CB7A9DC3" box="[388,516,220,243]" italics="true" pageId="17" pageNumber="144">Crinia rosea</emphasis>
,
<bibRefCitation id="EF814B75FFDDFF97184942D0CBD29DC4" author="Harrison, L." box="[529,684,220,244]" pageId="17" pageNumber="144" pagination="277 - 287" refId="ref16603" refString="Harrison, L. (1927) Notes on some Western Australian frogs, with descriptions of new species. Records of the Australian Museum, 15 (4), 277 - 287. https: // doi. org / 10.3853 / j. 0067 - 1975.15.1927.815" type="journal article" year="1927">Harrison, 1927</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAF3684FFDDFF971ACF432DCA369CB4" blockId="17.[151,1437,289,676]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971ACF432DC8689C0B" bold="true" box="[151,278,289,315]" pageId="17" pageNumber="144">Definition.</emphasis>
Nidicolous life history, terrestrial endotrophic larvae with vestigial non-feeding mouthparts, larvae &lt;
<quantity id="4CE89B61FFDDFF971FD8432EC9BD9C50" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="144" unit="mm" value="20.0">20 mm</quantity>
in maximum length, adults &lt;
<quantity id="4CE89B61FFDDFF971849434ACB1F9C50" box="[529,609,326,353]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="144" unit="mm" value="25.0">25 mm</quantity>
in maximum length, fourth toe length &lt;
<quantity id="4CE89B61FFDDFF971E72434ACD139C50" box="[1066,1133,326,352]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="144" unit="mm" value="7.0">7 mm</quantity>
, testes mass &lt;
<quantity id="4CE89B61FFDDFF971F57434ACC199C50" box="[1295,1383,326,352]" metricMagnitude="-6" metricUnit="kg" metricValue="1.0" pageId="17" pageNumber="144" unit="g" value="0.001">0.001 g</quantity>
, egg compliment &lt;50, monophasic call, primarily spring breeding.
</paragraph>
</subSubSection>
<subSubSection id="C30A650FFFDDFF971A9F4381CB629CFD" pageId="17" pageNumber="144" type="description">
<paragraph id="8BAF3684FFDDFF971A9F4381CB629CFD" blockId="17.[151,1437,289,676]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971A9F4381C8519C97" bold="true" box="[199,303,397,423]" pageId="17" pageNumber="144">Content.</emphasis>
Four species:
<taxonomicName id="4C104D07FFDDFF971B8C4382CABE9C99" authority="(Wardell-Johnson &amp; Roberts)" baseAuthorityName="Wardell-Johnson &amp; Roberts" box="[468,960,398,425]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="alba">
<emphasis id="B964EA96FFDDFF971B8C4382CB179C98" box="[468,617,398,424]" italics="true" pageId="17" pageNumber="144">Anstisia alba</emphasis>
(Wardell-Johnson &amp; Roberts)
</taxonomicName>
,
<taxonomicName id="4C104D07FFDDFF9719944382CD239C99" authority="(Main)" baseAuthorityName="Main" baseAuthorityYear="1963" box="[972,1117,398,425]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="lutea">
<emphasis id="B964EA96FFDDFF9719944382CD7A9C98" box="[972,1028,398,424]" italics="true" pageId="17" pageNumber="144">lutea</emphasis>
(Main)
</taxonomicName>
,
<taxonomicName id="4C104D07FFDDFF971E314383CC589C99" authority="(Harrison)" baseAuthorityName="Harrison" box="[1129,1318,398,425]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="rosea">
<emphasis id="B964EA96FFDDFF971E314383CDD89C98" box="[1129,1190,399,424]" italics="true" pageId="17" pageNumber="144">rosea</emphasis>
(Harrison)
</taxonomicName>
, and
<taxonomicName id="4C104D07FFDDFF971F3B4382CB669CFD" authority="(Wardell-Johnson &amp; Roberts)" baseAuthorityName="Wardell-Johnson &amp; Roberts" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="vitellina">
<emphasis id="B964EA96FFDDFF971F3B4382C9BD9CFC" italics="true" pageId="17" pageNumber="144">vitellina</emphasis>
(Wardell-Johnson &amp; Roberts)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A650FFFDDFF971A9F43D9CA1F9CC0" box="[199,865,469,496]" pageId="17" pageNumber="144" type="distribution">
<paragraph id="8BAF3684FFDDFF971A9F43D9CA1F9CC0" blockId="17.[151,1437,289,676]" box="[199,865,469,496]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971A9F43D9C81F9CDF" bold="true" box="[199,353,469,495]" pageId="17" pageNumber="144">Distribution.</emphasis>
Endemic to south-western
<collectingRegion id="49D4F866FFDDFF9718CA43DACA239CC0" box="[658,861,470,496]" country="Australia" name="Western Australia" pageId="17" pageNumber="144">Western Australia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A650FFFDDFF971A9F43F6CBAE9F94" pageId="17" pageNumber="144" type="etymology">
<paragraph id="8BAF3684FFDDFF971A9F43F6CBAE9F94" blockId="17.[151,1437,289,676]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971A9F43F6C8309F24" bold="true" box="[199,334,506,532]" pageId="17" pageNumber="144">Etymology.</emphasis>
The generic name
<taxonomicName id="4C104D07FFDDFF97187643F6CBF69F24" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[558,648,506,532]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDDFF97187643F6CBF69F24" box="[558,648,506,532]" italics="true" pageId="17" pageNumber="144">Anstisia</emphasis>
</taxonomicName>
honours the extensive life work of Dr Marion Anstis, work that has concisely outlined the developmental differences of the three current and four former
<taxonomicName id="4C104D07FFDDFF971E394012CC519F08" box="[1121,1327,542,568]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971E394012CDAD9F08" box="[1121,1235,542,568]" italics="true" pageId="17" pageNumber="144">Geocrinia</emphasis>
species
</taxonomicName>
, allowing for delineation between the two genera. Relevantly, given that her work has largely focused on the life histories of Australian anurans, in our opinion it is fitting that her name be applied to a genus that is distinguished largely on its unique life history strategy and larval morphology.
</paragraph>
</subSubSection>
<caption id="DF6F660CFFDDFF971ACF47E7CB629B7C" ID-DOI="http://doi.org/10.5281/zenodo.6641298" ID-Zenodo-Dep="6641298" httpUri="https://zenodo.org/record/6641298/files/figure.png" pageId="17" pageNumber="144" startId="17.[151,250,1515,1539]" targetBox="[302,1285,715,1489]" targetPageId="17">
<paragraph id="8BAF3684FFDDFF971ACF47E7CB629B7C" blockId="17.[151,1437,1515,1612]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971ACF47E7C86A9B34" bold="true" box="[151,276,1515,1540]" pageId="17" pageNumber="144">FIGURE 7.</emphasis>
The species of
<taxonomicName id="4C104D07FFDDFF971BED47E0CB799B33" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[437,519,1516,1539]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDDFF971BED47E0CB799B33" box="[437,519,1516,1539]" italics="true" pageId="17" pageNumber="144">Anstisia</emphasis>
</taxonomicName>
<emphasis id="B964EA96FFDDFF97185647E0CB139B34" bold="true" box="[526,621,1516,1540]" pageId="17" pageNumber="144">
<taxonomicNameLabel id="A25757EDFFDDFF97185647E0CB139B34" box="[526,621,1516,1540]" pageId="17" pageNumber="144" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
in life. A)
<taxonomicName id="4C104D07FFDDFF97188747E0CA4D9B33" box="[735,819,1516,1539]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="rosea">
<emphasis id="B964EA96FFDDFF97188747E0CA4D9B33" box="[735,819,1516,1539]" italics="true" pageId="17" pageNumber="144">A. rosea</emphasis>
</taxonomicName>
, from Pemberton, Western Australia; B)
<taxonomicName id="4C104D07FFDDFF971E8F47E0CC599B34" baseAuthorityName="Main" baseAuthorityYear="1963" box="[1239,1319,1516,1540]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="lutea">
<emphasis id="B964EA96FFDDFF971E8F47E0CC599B34" box="[1239,1319,1516,1540]" italics="true" pageId="17" pageNumber="144">A. lutea</emphasis>
</taxonomicName>
, from Walpole, Western Australia; C)
<taxonomicName id="4C104D07FFDDFF971BEB441CC8839B18" box="[435,509,1552,1576]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="alba">
<emphasis id="B964EA96FFDDFF971BEB441CC8839B18" box="[435,509,1552,1576]" italics="true" pageId="17" pageNumber="144">A. alba</emphasis>
</taxonomicName>
, from Witchcliffe, Western Australia; and D)
<taxonomicName id="4C104D07FFDDFF97198A441CCD3C9B18" box="[978,1090,1552,1576]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="vitellina">
<emphasis id="B964EA96FFDDFF97198A441CCD3C9B18" box="[978,1090,1552,1576]" italics="true" pageId="17" pageNumber="144">A. vitellina</emphasis>
</taxonomicName>
, from Spearwood Creek, Western Australia. Photographs by G. Webster.
</paragraph>
</caption>
<subSubSection id="C30A650FFFDDFF941A9F4475CD149FC1" lastPageId="18" lastPageNumber="145" pageId="17" pageNumber="144" type="diagnosis">
<paragraph id="8BAF3684FFDDFF941A9F4475CA019DC9" blockId="17.[151,1437,1657,2008]" lastBlockId="18.[151,1437,151,753]" lastPageId="18" lastPageNumber="145" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971A9F4475C83C9BA3" bold="true" box="[199,322,1657,1683]" pageId="17" pageNumber="144">Diagnosis.</emphasis>
The appearance in life of frog species within
<taxonomicName id="4C104D07FFDDFF9719664476CAE69BA4" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[830,920,1658,1684]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDDFF9719664476CAE69BA4" box="[830,920,1658,1684]" italics="true" pageId="17" pageNumber="144">Anstisia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D07FFDDFF9719964476CD3E9BA4" authorityName="Blake" authorityYear="1973" box="[974,1088,1658,1684]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="genus">
<emphasis id="B964EA96FFDDFF9719964476CD3E9BA4" box="[974,1088,1658,1684]" italics="true" pageId="17" pageNumber="144">Geocrinia</emphasis>
</taxonomicName>
is similar (see
<figureCitation id="132B2A01FFDDFF971EBF4476CC2E9BA4" box="[1255,1360,1658,1684]" captionStart="FIGURE 6" captionStartId="16.[151,250,1972,1996]" captionTargetBox="[330,1237,1222,1954]" captionTargetId="figure-476@16.[330,1237,1222,1954]" captionTargetPageId="16" captionText="FIGURE 6. The species of Geocrinia Blake 1973 in life.A) G. laevis, from Cradle Mountain, Tasmania; B) G. victoriana, from Eden, New South Wales; and C) G. leai, from Pemberton, Western Australia. Photographs by G. Webster." figureDoi="http://doi.org/10.5281/zenodo.6641296" httpUri="https://zenodo.org/record/6641296/files/figure.png" pageId="17" pageNumber="144">Figures 6</figureCitation>
and
<figureCitation id="132B2A01FFDDFF971FDD4476CCEA9BA4" box="[1413,1428,1658,1684]" captionStart="FIGURE 7" captionStartId="17.[151,250,1515,1539]" captionTargetBox="[302,1285,715,1489]" captionTargetId="figure-420@17.[302,1285,715,1489]" captionTargetPageId="17" captionText="FIGURE 7. The species of Anstisia gen. nov. in life. A) A. rosea, from Pemberton, Western Australia; B) A. lutea, from Walpole, Western Australia; C) A. alba, from Witchcliffe, Western Australia; and D) A. vitellina, from Spearwood Creek, Western Australia. Photographs by G. Webster." figureDoi="http://doi.org/10.5281/zenodo.6641298" httpUri="https://zenodo.org/record/6641298/files/figure.png" pageId="17" pageNumber="144">7</figureCitation>
) although key differences exist. The three species of
<taxonomicName id="4C104D07FFDDFF9718B94492CA2D9B88" authorityName="Blake" authorityYear="1973" box="[737,851,1694,1720]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="genus">
<emphasis id="B964EA96FFDDFF9718B94492CA2D9B88" box="[737,851,1694,1720]" italics="true" pageId="17" pageNumber="144">Geocrinia</emphasis>
</taxonomicName>
can be reliably distinguished from the four
<taxonomicName id="4C104D07FFDDFF971F1A4492C9969BEC" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971F1A4492CCE29B88" box="[1346,1436,1694,1720]" italics="true" pageId="17" pageNumber="144">Anstisia</emphasis>
species
</taxonomicName>
by ventral surface patterning in most cases. Ventral surfaces of
<taxonomicName id="4C104D07FFDDFF9719FC44CECD789BEC" box="[932,1030,1730,1756]" class="Amphibia" family="Myobatrachidae" genus="Geocinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B964EA96FFDDFF9719FC44CECD789BEC" box="[932,1030,1730,1756]" italics="true" pageId="17" pageNumber="144">G. laevis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D07FFDDFF971E6244CECDB09BEC" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[1082,1230,1730,1756]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="victoriana">
<emphasis id="B964EA96FFDDFF971E6244CECDB09BEC" box="[1082,1230,1730,1756]" italics="true" pageId="17" pageNumber="144">G. victoriana</emphasis>
</taxonomicName>
always have some degree of marbling in the form of light grey or brown blotches. This patterning can at times be present in
<taxonomicName id="4C104D07FFDDFF971F7844EACC159A30" box="[1312,1387,1766,1792]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="leai">
<emphasis id="B964EA96FFDDFF971F7844EACC159A30" box="[1312,1387,1766,1792]" italics="true" pageId="17" pageNumber="144">G. leai</emphasis>
</taxonomicName>
, but this species can also present a uniform ventral surface with pale yellow hues. Males of all three species can feature yellow throats. All
<taxonomicName id="4C104D07FFDDFF971B374522CB5C9A78" box="[367,546,1838,1864]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971B374522C8B79A78" box="[367,457,1838,1864]" italics="true" pageId="17" pageNumber="144">Anstisia</emphasis>
species
</taxonomicName>
however have distinctly individual ventral colouring. The ventral surface of
<taxonomicName id="4C104D07FFDDFF971FDC4522C9B79A5C" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="alba">
<emphasis id="B964EA96FFDDFF971FDC4522C9B79A5C" italics="true" pageId="17" pageNumber="144">A. alba</emphasis>
</taxonomicName>
is entirely uniform white; almost entirely coloured anteriorly with yolk orange in
<taxonomicName id="4C104D07FFDDFF971E01455ECDAD9A5C" box="[1113,1235,1874,1900]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="vitellina">
<emphasis id="B964EA96FFDDFF971E01455ECDAD9A5C" box="[1113,1235,1874,1900]" italics="true" pageId="17" pageNumber="144">A. vitellina</emphasis>
</taxonomicName>
; rose with pink to orange hues in
<taxonomicName id="4C104D07FFDDFF971B67457AC8E49AA0" box="[319,410,1910,1936]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="rosea">
<emphasis id="B964EA96FFDDFF971B67457AC8E49AA0" box="[319,410,1910,1936]" italics="true" pageId="17" pageNumber="144">A. rosea</emphasis>
</taxonomicName>
; and off-white or cream usually with a dark yellow or lemon-yellow wash in
<taxonomicName id="4C104D07FFDDFF971EAC457ACC359AA0" baseAuthorityName="Main" baseAuthorityYear="1963" box="[1268,1355,1910,1936]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="17" pageNumber="144" phylum="Chordata" rank="species" species="lutea">
<emphasis id="B964EA96FFDDFF971EAC457ACC359AA0" box="[1268,1355,1910,1936]" italics="true" pageId="17" pageNumber="144">A. lutea</emphasis>
</taxonomicName>
. Males of the latter two species are known to have dark grey or black throats, with females having pink and yellow throats respectively. Dorsally,
<taxonomicName id="4C104D07FFDDFF971BC045B2CB1F9AE8" box="[408,609,1982,2008]" pageId="17" pageNumber="144">
<emphasis id="B964EA96FFDDFF971BC045B2CB749AE8" box="[408,522,1982,2008]" italics="true" pageId="17" pageNumber="144">Geocrinia</emphasis>
species
</taxonomicName>
are highly variable but frequently feature a broad darker central marking. The dorsal pattern of
<taxonomicName id="4C104D07FFDEFF941BD4429BC8979D80" box="[396,489,151,177]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="rosea">
<emphasis id="B964EA96FFDEFF941BD4429BC8979D80" box="[396,489,151,177]" italics="true" pageId="18" pageNumber="145">A. rosea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D07FFDEFF941879429BCB049D81" baseAuthorityName="Main" baseAuthorityYear="1963" box="[545,634,151,177]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="lutea">
<emphasis id="B964EA96FFDEFF941879429BCB049D81" box="[545,634,151,177]" italics="true" pageId="18" pageNumber="145">A. lutea</emphasis>
</taxonomicName>
can resemble that of
<taxonomicName id="4C104D07FFDEFF941936429BCA9E9D81" authorityName="Blake" authorityYear="1973" box="[878,992,151,177]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="genus">
<emphasis id="B964EA96FFDEFF941936429BCA9E9D81" box="[878,992,151,177]" italics="true" pageId="18" pageNumber="145">Geocrinia</emphasis>
</taxonomicName>
but these species are reliably less variable, with the dorsal marking forming an inverted “V” on the lower back. Both
<taxonomicName id="4C104D07FFDEFF941E4142B7CD159DE5" box="[1049,1131,187,213]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="alba">
<emphasis id="B964EA96FFDEFF941E4142B7CD159DE5" box="[1049,1131,187,213]" italics="true" pageId="18" pageNumber="145">A. alba</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D07FFDEFF941EFA42B7CC609DE5" box="[1186,1310,187,213]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="vitellina">
<emphasis id="B964EA96FFDEFF941EFA42B7CC609DE5" box="[1186,1310,187,213]" italics="true" pageId="18" pageNumber="145">A. vitellina</emphasis>
</taxonomicName>
are similar in dorsal appearance with broken streaks of scattered darker spots.
</paragraph>
<paragraph id="8BAF3684FFDEFF941A9F430FCDD69F0D" blockId="18.[151,1437,151,753]" pageId="18" pageNumber="145">
In life, the appearance of tadpoles differs noticeably and cannot be confused, as
<taxonomicName id="4C104D07FFDEFF941E1D430FCDC99C2D" authorityName="Blake" authorityYear="1973" box="[1093,1207,259,285]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="genus">
<emphasis id="B964EA96FFDEFF941E1D430FCDC99C2D" box="[1093,1207,259,285]" italics="true" pageId="18" pageNumber="145">Geocrinia</emphasis>
</taxonomicName>
tadpoles have a kertinised oral disc with feeding mouthparts, while
<taxonomicName id="4C104D07FFDEFF9418F3432BCA7B9C71" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[683,773,295,321]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDEFF9418F3432BCA7B9C71" box="[683,773,295,321]" italics="true" pageId="18" pageNumber="145">Anstisia</emphasis>
</taxonomicName>
tadpoles do not. Tadpoles of the three
<taxonomicName id="4C104D07FFDEFF941EF4432BCC0B9C71" box="[1196,1397,295,321]" pageId="18" pageNumber="145">
<emphasis id="B964EA96FFDEFF941EF4432BCC609C71" box="[1196,1310,295,321]" italics="true" pageId="18" pageNumber="145">Geocrinia</emphasis>
species
</taxonomicName>
are uniformly brown on the dorsum, with patchy lighter colouration on the sides of the body and venter. In
<taxonomicName id="4C104D07FFDEFF941F534347CC109C55" box="[1291,1390,331,357]" class="Amphibia" family="Myobatrachidae" genus="Geocinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B964EA96FFDEFF941F534347CC109C55" box="[1291,1390,331,357]" italics="true" pageId="18" pageNumber="145">G. laevis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D07FFDEFF941ACF4363C8529CB8" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[151,300,367,393]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="victoriana">
<emphasis id="B964EA96FFDEFF941ACF4363C8529CB8" box="[151,300,367,393]" italics="true" pageId="18" pageNumber="145">G. victoriana</emphasis>
</taxonomicName>
the tail muscle is very light brown and fins mostly clear, while the tail muscle in
<taxonomicName id="4C104D07FFDEFF941EE24363CC789CB9" box="[1210,1286,367,393]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="leai">
<emphasis id="B964EA96FFDEFF941EE24363CC789CB9" box="[1210,1286,367,393]" italics="true" pageId="18" pageNumber="145">G. leai</emphasis>
</taxonomicName>
is pigmented with dark and light blotches with some irregular markings across the fins. Tadpoles of all
<taxonomicName id="4C104D07FFDEFF941ED6439FCC3F9C9D" box="[1166,1345,403,429]" pageId="18" pageNumber="145">
<emphasis id="B964EA96FFDEFF941ED6439FCD969C9D" box="[1166,1256,403,429]" italics="true" pageId="18" pageNumber="145">Anstisia</emphasis>
species
</taxonomicName>
possess vibrant light blue flecks across the body and tail, which are most apparent when viewed laterally, against a pigmentation of deep red-brown (
<taxonomicName id="4C104D07FFDEFF941BE343D7CB759CC5" box="[443,523,475,501]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="alba">
<emphasis id="B964EA96FFDEFF941BE343D7CB759CC5" box="[443,523,475,501]" italics="true" pageId="18" pageNumber="145">A. alba</emphasis>
</taxonomicName>
/
<taxonomicName id="4C104D07FFDEFF94184B43D7CBF39CC5" box="[531,653,475,501]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="vitellina">
<emphasis id="B964EA96FFDEFF94184B43D7CBF39CC5" box="[531,653,475,501]" italics="true" pageId="18" pageNumber="145">A. vitellina</emphasis>
</taxonomicName>
) to dark yellow-brown (
<taxonomicName id="4C104D07FFDEFF9419C643D7CA8A9CC5" baseAuthorityName="Main" baseAuthorityYear="1963" box="[926,1012,475,501]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="lutea">
<emphasis id="B964EA96FFDEFF9419C643D7CA8A9CC5" box="[926,1012,475,501]" italics="true" pageId="18" pageNumber="145">A. lutea</emphasis>
</taxonomicName>
/
<taxonomicName id="4C104D07FFDEFF9419A443D7CD269CC4" box="[1020,1112,475,501]" class="Amphibia" family="Myobatrachidae" genus="Crinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="rosea">
<emphasis id="B964EA96FFDEFF9419A443D7CD269CC4" box="[1020,1112,475,501]" italics="true" pageId="18" pageNumber="145">A. rosea</emphasis>
</taxonomicName>
). In the
<taxonomicName id="4C104D07FFDEFF941EED43D7CC149CC5" box="[1205,1386,475,501]" pageId="18" pageNumber="145">
<emphasis id="B964EA96FFDEFF941EED43D7CC719CC5" box="[1205,1295,475,501]" italics="true" pageId="18" pageNumber="145">Anstisia</emphasis>
species
</taxonomicName>
, this pigmentation is lighter on the tail muscle than body, and fins are opaque. Metamorph frogs of both genera resemble the adults, although
<taxonomicName id="4C104D07FFDEFF941B22402FCB529F0D" box="[378,556,547,573]" pageId="18" pageNumber="145">
<emphasis id="B964EA96FFDEFF941B22402FC8AA9F0D" box="[378,468,547,573]" italics="true" pageId="18" pageNumber="145">Anstisia</emphasis>
species
</taxonomicName>
possess the vibrant blue flecking typical of the tadpoles.
</paragraph>
<paragraph id="8BAF3684FFDEFF941A9F404BCD149FC1" blockId="18.[151,1437,151,753]" pageId="18" pageNumber="145">
In terms of distinguishing the two genera by advertisement call,
<taxonomicName id="4C104D07FFDEFF9419FB404BCD109F51" box="[931,1134,583,609]" pageId="18" pageNumber="145">
<emphasis id="B964EA96FFDEFF9419FB404BCD6B9F51" box="[931,1045,583,609]" italics="true" pageId="18" pageNumber="145">Geocrinia</emphasis>
species
</taxonomicName>
can be identified by a call consisting of two distinct note
<typeStatus id="54AB8826FFDEFF941BA44067CB499FB5" box="[508,567,619,645]" pageId="18" pageNumber="145">types</typeStatus>
(biphasic), compared to a singular note
<typeStatus id="54AB8826FFDEFF941E524067CD449FB5" box="[1034,1082,619,645]" pageId="18" pageNumber="145">type</typeStatus>
(monophasic) in
<taxonomicName id="4C104D07FFDEFF941F5F4067CC1F9FB5" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[1287,1377,619,645]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDEFF941F5F4067CC1F9FB5" box="[1287,1377,619,645]" italics="true" pageId="18" pageNumber="145">Anstisia</emphasis>
</taxonomicName>
. The introductory note alone, present in
<taxonomicName id="4C104D07FFDEFF94187D4083CBE99F99" authorityName="Blake" authorityYear="1973" box="[549,663,655,681]" class="Amphibia" family="Myobatrachidae" genus="Geocrinia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="genus">
<emphasis id="B964EA96FFDEFF94187D4083CBE99F99" box="[549,663,655,681]" italics="true" pageId="18" pageNumber="145">Geocrinia</emphasis>
</taxonomicName>
, is sufficient to distinguish this genus from all
<taxonomicName id="4C104D07FFDEFF941EF74083CC149F99" box="[1199,1386,655,681]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B964EA96FFDEFF941EF74083CC779F99" box="[1199,1289,655,681]" italics="true" pageId="18" pageNumber="145">Anstisia</emphasis>
species.
</taxonomicName>
The closely related genus,
<taxonomicName id="4C104D07FFDEFF941BC940BFC8B89FFD" authorityName="Tyler" authorityYear="1972" box="[401,454,691,717]" class="Amphibia" family="Myobatrachidae" genus="Assa" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="genus">
<emphasis id="B964EA96FFDEFF941BC940BFC8B89FFD" box="[401,454,691,717]" italics="true" pageId="18" pageNumber="145">Assa</emphasis>
</taxonomicName>
, shares a similar monophasic ticking call to
<taxonomicName id="4C104D07FFDEFF9419E240BFCD6A9FFD" authority="Webster &amp; Bool, 2022" authorityName="Webster &amp; Bool" authorityYear="2022" box="[954,1044,691,717]" class="Amphibia" family="Myobatrachidae" genus="Anstisia" kingdom="Animalia" order="Anura" pageId="18" pageNumber="144" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B964EA96FFDEFF9419E240BFCD6A9FFD" box="[954,1044,691,717]" italics="true" pageId="18" pageNumber="145">Anstisia</emphasis>
</taxonomicName>
; however these genera are entirely allopatric, with
<taxonomicName id="4C104D07FFDEFF941B1040DBC8039FC1" authorityName="Tyler" authorityYear="1972" box="[328,381,727,753]" class="Amphibia" family="Myobatrachidae" genus="Assa" kingdom="Animalia" order="Anura" pageId="18" pageNumber="145" phylum="Chordata" rank="genus">
<emphasis id="B964EA96FFDEFF941B1040DBC8039FC1" box="[328,381,727,753]" italics="true" pageId="18" pageNumber="145">Assa</emphasis>
</taxonomicName>
occurring in northern
<collectingRegion id="49D4F866FFDEFF94182140DBCA3E9FC1" box="[633,832,727,753]" country="Australia" name="New South Wales" pageId="18" pageNumber="145">New South Wales</collectingRegion>
and southern
<collectingRegion id="49D4F866FFDEFF94198640DBCD189FC1" box="[990,1126,727,753]" country="Australia" name="Queensland" pageId="18" pageNumber="145">Queensland</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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