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<document id="D9CAD5CB93A7F0F224A80158642D566D" ID-DOI="10.26107/RBZ-2021-0010" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351956" ID-ZooBank="DD656B82-2FF6-444F-B259-827EDC7F3715" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630207335308" checkinUser="diego" docAuthor="Páll-Gergely, Barna &amp; Hunyadi, András" docDate="2021" docId="03B2EC0BFFC6FFAAFEA5FB93FA19BE4F" docLanguage="en" docName="RBZ-2021-0010.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Laotia luongi Páll-Gergely &amp; Hunyadi 2021, new species" docType="treatment" docVersion="3" lastPageNumber="121" masterDocId="FF8B9473FFC5FFA9FF94FFA7FFDBB823" masterDocTitle="A new species and a new subspecies of Laotia Saurin, 1953 (Gastropoda: Caenogastropoda: Alycaeidae) from Sơn La Province, Vietnam" masterLastPageNumber="123" masterPageNumber="118" pageNumber="121" updateTime="1699185198036" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="00FE3C3A49522B884B7641AF826D3AC9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="80D6CDA55062458A9C8EECCA2268107F">
<mods:title id="534E5A9D5A6257D60D141A1741FF3CBC">A new species and a new subspecies of Laotia Saurin, 1953 (Gastropoda: Caenogastropoda: Alycaeidae) from Sơn La Province, Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="CB2FC301D10DA78E97F50AF2D96183FF" type="personal">
<mods:role id="AD5C48D1A49B8D7FB5D70541D732A449">
<mods:roleTerm id="45A9CD9FC1B2C4F05B062A862065CF76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9E7CCA5189DC45F19C46242E912427EF">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="EFAA552278718D619C8E06347F76C3FE" type="personal">
<mods:role id="AE55561D95537B27CF1E196BEB902689">
<mods:roleTerm id="64B51E7975FCBD72EA7756F95CF68DED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9E1AF38A54AD15F122173EE466D5F446">Hunyadi, András</mods:namePart>
</mods:name>
<mods:typeOfResource id="C08BC24934C3A44E5E175BB1F1491E60">text</mods:typeOfResource>
<mods:relatedItem id="A2D31C872281C76930F93577B06DA10D" type="host">
<mods:titleInfo id="0ADFD55E5BD3151C3D50AEEFA56B56B4">
<mods:title id="05BC0E7596E35E839609C75320E55941">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="379D4C1FAA6BB50865838420817A36D8">
<mods:date id="D7B332799339EAC6D0C25612D438C1E9">2021</mods:date>
<mods:detail id="08C5E202314416E89D5BA4E290338F8C" type="pubDate">
<mods:number id="A4C486581E3B3D2999F24DA0E1AAF140">2021-05-07</mods:number>
</mods:detail>
<mods:detail id="5EDE8E0EAE8C55902A69C7B25C5FF97D" type="volume">
<mods:number id="54BD2ED285299814A21654DD7813E4A3">69</mods:number>
</mods:detail>
<mods:extent id="C4F64D52219C1DC0479418CDED421249" unit="page">
<mods:start id="A5DE9C8777359F80F71B350DE39F94E8">118</mods:start>
<mods:end id="453CCC70EB5458BA59600E62E6E84068">123</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EE9BEC581022DA458E6EB100C2000A16">journal article</mods:classification>
<mods:identifier id="D32CB923103AE87FFD04E0FFB37E7EF4" type="DOI">10.26107/RBZ-2021-0010</mods:identifier>
<mods:identifier id="13746C5507165580B058C50BEA160F82" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="D84CC4C6714226AB4F584147DA8AF25A" type="Zenodo-Dep">5351956</mods:identifier>
<mods:identifier id="DDD96E2876A13B2741BD08BBCC096951" type="ZooBank">DD656B82-2FF6-444F-B259-827EDC7F3715</mods:identifier>
</mods:mods>
<treatment id="03B2EC0BFFC6FFAAFEA5FB93FA19BE4F" LSID="urn:lsid:plazi:treatment:03B2EC0BFFC6FFAAFEA5FB93FA19BE4F" httpUri="http://treatment.plazi.org/id/03B2EC0BFFC6FFAAFEA5FB93FA19BE4F" lastPageNumber="121" pageId="3" pageNumber="121">
<subSubSection id="C3010E96FFC6FFAAFEA5FB93FD86BC6F" box="[305,605,1075,1100]" pageId="3" pageNumber="121" type="nomenclature">
<paragraph id="8BA45D1DFFC6FFAAFEA5FB93FD86BC6F" blockId="3.[305,605,1075,1132]" box="[305,605,1075,1100]" pageId="3" pageNumber="121">
<heading id="D0ECEA71FFC6FFAAFEA5FB93FD86BC6F" bold="true" box="[305,605,1075,1100]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="121" reason="2">
<emphasis id="B96F810FFFC6FFAAFEA5FB93FD86BC6F" bold="true" box="[305,605,1075,1100]" pageId="3" pageNumber="121">
<taxonomicName id="4C1B269EFFC6FFAAFEA5FB93FE13BC68" ID-CoL="8H9TW" authority="Páll-Gergely &amp; Hunyadi, 2021" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2021" box="[305,456,1075,1100]" class="Gastropoda" family="Alycaeidae" genus="Laotia" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="121" phylum="Mollusca" rank="species" species="luongi" status="sp. nov.">
<emphasis id="B96F810FFFC6FFAAFEA5FB93FE13BC68" bold="true" box="[305,456,1075,1100]" italics="true" pageId="3" pageNumber="121">Laotia luongi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C3C74FFC6FFAAFE4CFB93FD86BC6F" box="[472,605,1076,1100]" pageId="3" pageNumber="121" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3010E96FFC6FFAAFE10FBF3FDD1BC4F" box="[388,522,1108,1132]" pageId="3" pageNumber="121" type="description">
<paragraph id="8BA45D1DFFC6FFAAFE10FBF3FDD1BC4F" blockId="3.[305,605,1075,1132]" box="[388,522,1108,1132]" pageId="3" pageNumber="121">
(
<figureCitation id="13204198FFC6FFAAFE18FBF3FDD9BC4F" box="[396,514,1108,1132]" captionStart="Fig" captionStartId="4.[113,148,2035,2057]" captionTargetBox="[151,1413,189,2019]" captionTargetId="figure-13@4.[151,1413,189,2019]" captionTargetPageId="4" captionText="Fig. 3. Shells of Laotia Saurin, 1953 species. A, Laotia luongi, new species, holotype (VNMN-Z-Inv.000220); B, L. luongi, new species, paratype (coll. HA); C, syntype of Laotia pahiensis Saurin, 1953 (MNHN-IM-2000-28217. Photos: B. Páll-Gergely (A, B), M. Caballer (C, downloaded from the website of MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5351962" httpUri="https://zenodo.org/record/5351962/files/figure.png" pageId="3" pageNumber="121">Fig. 3A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3010E96FFC6FFAAFF1CFB33FE9BBE2F" pageId="3" pageNumber="121" type="materials_examined">
<paragraph id="8BA45D1DFFC6FFAAFF1CFB33FDF3BD6F" blockId="3.[135,775,1172,1356]" pageId="3" pageNumber="121">
<emphasis id="B96F810FFFC6FFAAFF1CFB33FEF4BC8F" bold="true" box="[136,303,1172,1196]" pageId="3" pageNumber="121">Type material.</emphasis>
<materialsCitation id="3B735740FFC6FFAAFEA0FB33FDF3BD6F" collectingDate="2020-02-09" collectionCode="V" collectorName="A. Hunyadi &amp; Hunyadi, H. &amp; Luong, J. U &amp; Pham" country="Vietnam" county="Yen Chau District" elevation="970" latitude="20.991383" location="On Commune" longLatPrecision="1" longitude="104.187836" municipality="Chi" pageId="3" pageNumber="121" specimenCount="1" stateProvince="Son La" typeStatus="holotype">
<typeStatus id="54A0E3BFFFC6FFAAFEA0FB33FE4CBC8F" box="[308,407,1172,1196]" pageId="3" pageNumber="121" type="holotype">Holotype</typeStatus>
(D:
<quantity id="4CE3F0F8FFC6FFAAFE52FB33FDC2BC88" box="[454,537,1172,1196]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="121" unit="mm" value="5.1">5.1 mm</quantity>
, H:
<quantity id="4CE3F0F8FFC6FFAAFDD0FB33FD4DBC8F" box="[580,662,1172,1196]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="121" unit="mm" value="2.9">2.9 mm</quantity>
) (VNMN- Z-Inv.000220),
<collectingCountry id="F30C1D8DFFC6FFAAFEBFFB13FE5CBCEF" box="[299,391,1204,1228]" name="Vietnam" pageId="3" pageNumber="121">Vietnam</collectingCountry>
,
<collectingRegion id="49DF93FFFFC6FFAAFE04FB13FDE5BCEF" box="[400,574,1204,1230]" country="Vietnam" name="Son La" pageId="3" pageNumber="121">SƠn La Province</collectingRegion>
,
<collectingCounty id="62C52591FFC6FFAAFDD2FB13FCDFBCEF" box="[582,772,1204,1228]" pageId="3" pageNumber="121">Yên Châu District</collectingCounty>
,
<collectingMunicipality id="6BC0C767FFC6FFAAFF13FB73FF76BCCD" box="[135,173,1236,1262]" pageId="3" pageNumber="121">Chi</collectingMunicipality>
ềng
<location id="8EC40BC6FFC6FFAAFF4EFB73FEA9BCCF" LSID="urn:lsid:plazi:treatment:03B2EC0BFFC6FFAAFEA5FB93FA19BE4F:8EC40BC6FFC6FFAAFF4EFB73FEA9BCCF" box="[218,370,1236,1260]" country="Vietnam" county="Yen Chau District" latitude="20.991383" longLatPrecision="1" longitude="104.187836" municipality="Chi" name="On Commune" pageId="3" pageNumber="121" stateProvince="Son La">On Commune</location>
,
<location id="8EC40BC6FFC6FFAAFEE8FB73FE4EBCC8" LSID="urn:lsid:plazi:treatment:03B2EC0BFFC6FFAAFEA5FB93FA19BE4F:8EC40BC6FFC6FFAAFEE8FB73FE4EBCC8" box="[380,405,1236,1259]" country="Vietnam" county="Yen Chau District" latitude="20.991383" longLatPrecision="1" longitude="104.187836" municipality="Chi" name="Tr" pageId="3" pageNumber="121" stateProvince="Son La">Tr</location>
ạm Hốc
<location id="8EC40BC6FFC6FFAAFE79FB73FDE5BCCF" LSID="urn:lsid:plazi:treatment:03B2EC0BFFC6FFAAFEA5FB93FA19BE4F:8EC40BC6FFC6FFAAFE79FB73FDE5BCCF" box="[493,574,1236,1260]" country="Vietnam" county="Yen Chau District" latitude="20.991383" longLatPrecision="1" longitude="104.187836" municipality="Chi" name="Village" pageId="3" pageNumber="121" stateProvince="Son La">Village</location>
,
<location id="8EC40BC6FFC6FFAAFDDCFB73FDB2BCCF" LSID="urn:lsid:plazi:treatment:03B2EC0BFFC6FFAAFEA5FB93FA19BE4F:8EC40BC6FFC6FFAAFDDCFB73FDB2BCCF" box="[584,617,1236,1260]" country="Vietnam" county="Yen Chau District" latitude="20.991383" longLatPrecision="1" longitude="104.187836" municipality="Chi" name="Nh" pageId="3" pageNumber="121" stateProvince="Son La">Nh</location>
ả
<location id="8EC40BC6FFC6FFAAFDEEFB73FCD9BCCF" LSID="urn:lsid:plazi:treatment:03B2EC0BFFC6FFAAFEA5FB93FA19BE4F:8EC40BC6FFC6FFAAFDEEFB73FCD9BCCF" box="[634,770,1236,1260]" country="Vietnam" county="Yen Chau District" latitude="20.991383" longLatPrecision="1" longitude="104.187836" municipality="Chi" name="Nhung Cave" pageId="3" pageNumber="121" stateProvince="Son La">Nhung Cave</location>
, around the cave,
<elevation id="0036BA2EFFC6FFAAFEDCFB53FE17BD2F" box="[328,460,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="121" unit="m" value="970.0">
<quantity id="4CE3F0F8FFC6FFAAFEDCFB53FE54BD28" box="[328,399,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="121" unit="m" value="970.0">970 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE2F3BDAFFC6FFAAFE48FB55FDBDBD28" box="[476,614,1266,1291]" degrees="20" direction="north" minutes="59.483" orientation="latitude" pageId="3" pageNumber="121" precision="1" value="20.991383">20°59.483′N</geoCoordinate>
,
<geoCoordinate id="EE2F3BDAFFC6FFAAFDE0FB55FCDCBD28" box="[628,775,1266,1291]" degrees="104" direction="east" minutes="11.270" orientation="longitude" pageId="3" pageNumber="121" precision="1" value="104.187836">104°11.270′E</geoCoordinate>
(locality code: 2020/20), coll.
<collectorName id="26EE38CBFFC6FFAAFE53FAB3FD9DBD0F" box="[455,582,1300,1324]" pageId="3" pageNumber="121">A. Hunyadi</collectorName>
, H.
<collectionCode id="ED0AC5D8FFC6FFAAFDFEFAB3FD5BBD08" box="[618,640,1300,1323]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="121" type="Museum">V</collectionCode>
.
<collectorName id="26EE38CBFFC6FFAAFD1DFAB3FCD9BD08" box="[649,770,1300,1324]" pageId="3" pageNumber="121">Luong, J.U</collectorName>
. Otani &amp; S.
<collectionCode id="ED0AC5D8FFC6FFAAFF68FA93FECFBD6F" box="[252,276,1332,1356]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="121" type="Museum">V</collectionCode>
.
<collectorName id="26EE38CBFFC6FFAAFEB4FA93FEBBBD6F" box="[288,352,1332,1356]" pageId="3" pageNumber="121">Pham</collectorName>
,
<date id="FFA57BDDFFC6FFAAFEF8FA93FDFFBD6F" box="[364,548,1332,1356]" pageId="3" pageNumber="121" value="2020-02-09">
<collectingDate id="EFE18235FFC6FFAAFEF8FA93FDFFBD6F" box="[364,548,1332,1356]" pageId="3" pageNumber="121" value="2020-02-09">9 February 2020</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA45D1DFFC6FFAAFF1CFAD3FD37BD8F" blockId="3.[136,775,1396,1452]" pageId="3" pageNumber="121">
<materialsCitation id="3B735740FFC6FFAAFF1CFAD3FCD8BDAF" box="[136,771,1396,1420]" collectingDate="2020-02-09" collectionCode="V" collectorName="A. Hunyadi &amp; Hunyadi, H. &amp; Luong, J. U &amp; Pham" country="Vietnam" county="Yen Chau District" elevation="970" latitude="20.991383" location="On Commune" longLatPrecision="1" longitude="104.187836" municipality="Chi" pageId="3" pageNumber="121" specimenCount="5" specimenCount-adult="5" stateProvince="Son La" typeStatus="paratype">
<typeStatus id="54A0E3BFFFC6FFAAFF1CFAD3FF28BDAF" box="[136,243,1396,1420]" pageId="3" pageNumber="121" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D1D9694FFC6FFAAFF6AFAD3FE93BDAF" box="[254,328,1396,1420]" pageId="3" pageNumber="121" type="adult">5 adult</specimenCount>
shells (coll. HA), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B735740FFC6FFAAFF1DFA33FD33BD8F" box="[137,744,1428,1452]" collectingDate="2020-02-09" collectionCode="V" collectorName="A. Hunyadi &amp; Hunyadi, H. &amp; Luong, J. U &amp; Pham" country="Vietnam" county="Yen Chau District" elevation="970" latitude="20.991383" location="On Commune" longLatPrecision="1" longitude="104.187836" municipality="Chi" pageId="3" pageNumber="121" specimenCount="8" specimenCount-adult="8" stateProvince="Son La" typeStatus="paratype">
<specimenCount id="9D1D9694FFC6FFAAFF1DFA33FF0FBD8F" box="[137,212,1428,1452]" pageId="3" pageNumber="121" type="adult">8 adult</specimenCount>
broken shells (coll. HA), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BA45D1DFFC6FFAAFF1CFA73FE9BBE2F" blockId="3.[136,775,1492,1548]" pageId="3" pageNumber="121">
<materialsCitation id="3B735740FFC6FFAAFF1CFA73FEE6BE2F" collectingDate="2020-02-09" collectionCode="V" collectorName="A. Hunyadi &amp; Hunyadi, H. &amp; Luong, J. U &amp; Pham" country="Vietnam" county="Yen Chau District" elevation="970" latitude="20.991383" location="On Commune" longLatPrecision="1" longitude="104.187836" municipality="Chi" pageId="3" pageNumber="121" specimenCount="5" specimenCount-juvenile="5" stateProvince="Son La">
<emphasis id="B96F810FFFC6FFAAFF1CFA73FEA2BDCF" bold="true" box="[136,377,1492,1516]" pageId="3" pageNumber="121">Additional material.</emphasis>
<specimenCount id="9D1D9694FFC6FFAAFE10FA73FE2EBDCF" box="[388,501,1492,1516]" pageId="3" pageNumber="121" type="juvenile">5 juvenile</specimenCount>
shells (coll. HA), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3010E96FFC6FFAAFF1CF993FBA5BC4F" pageId="3" pageNumber="121" type="description">
<paragraph id="8BA45D1DFFC6FFAAFF1CF993FD7DBFCF" blockId="3.[134,776,1588,2028]" pageId="3" pageNumber="121">
<emphasis id="B96F810FFFC6FFAAFF1CF993FED4BE6F" bold="true" box="[136,271,1588,1612]" pageId="3" pageNumber="121">Description.</emphasis>
Shell dextral, depressed, 4.25–4.75 whorls; shell shape and size matches
<taxonomicName id="4C1B269EFFC6FFAAFEE7F9F2FD26BE4F" authority="christahemmenae" authorityName="christahemmenae Pall-Gergely" authorityYear="2014" box="[371,765,1620,1644]" class="Gastropoda" family="Alycaeidae" genus="Laotia" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="121" phylum="Mollusca" rank="subSpecies" species="christahemmenae" subSpecies="christahemmenae">
<emphasis id="B96F810FFFC6FFAAFEE7F9F2FD26BE4F" box="[371,765,1620,1644]" italics="true" pageId="3" pageNumber="121">L. christahemmenae christahemmenae</emphasis>
</taxonomicName>
: approximately quarter of last whorl turns obliquely upwards; aperture directed towards apex; protoconch elevated (more elevated than it would be expected from the shell shape), 1.25–1.5 whorls, finely granular, without any notable sculptural elements; teleoconch with widely-spaced, straight, lamella-like, fragile ribs; last whorls with 36–62 ribs (n = 4); area between ribs glossy without spiral lines; rib density constant from initial whorls until the aperture; umbilicus relatively wide, ca. one third of minor shell diameter; aperture rounded; inner peristome slightly protruding, expanded, smeared onto the penultimate whorl; outer peristome strongly expanded, slightly reflected; operculum unknown.
</paragraph>
<paragraph id="8BA45D1DFFC6FFAAFCD7FB93FBA5BC4F" blockId="3.[835,1474,1076,1132]" pageId="3" pageNumber="121">
<emphasis id="B96F810FFFC6FFAAFCD7FB93FB83BC6F" bold="true" box="[835,1112,1076,1100]" pageId="3" pageNumber="121">Measurements (in mm).</emphasis>
D: 4.3–5.1, H: 2.3–2.9 (smallest and largest shells measured).
</paragraph>
</subSubSection>
<subSubSection id="C3010E96FFC6FFAAFCD7FB33FB01BDAF" pageId="3" pageNumber="121" type="diagnosis">
<paragraph id="8BA45D1DFFC6FFAAFCD7FB33FB01BDAF" blockId="3.[834,1475,1172,1420]" pageId="3" pageNumber="121">
<emphasis id="B96F810FFFC6FFAAFCD7FB33FB9BBC8F" bold="true" box="[835,1088,1172,1196]" pageId="3" pageNumber="121">Differential diagnosis.</emphasis>
In shell size and shape this species is similar to
<taxonomicName id="4C1B269EFFC6FFAAFC56FB12FB42BCEF" authorityName="Pall-Gergely" authorityYear="2014" box="[962,1177,1204,1228]" class="Gastropoda" family="Alycaeidae" genus="Laotia" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="121" phylum="Mollusca" rank="species" species="christahemmenae">
<emphasis id="B96F810FFFC6FFAAFC56FB12FB42BCEF" box="[962,1177,1204,1228]" italics="true" pageId="3" pageNumber="121">L. christahemmenae</emphasis>
</taxonomicName>
. However, the latter species has much denser, bent ribs (straight in the new species) and the area behind the peristome is without ribs (ribbed in the new species).
<taxonomicName id="4C1B269EFFC6FFAAFC4EFAB2FB57BD0F" authorityName="Saurin" authorityYear="1953" box="[986,1164,1300,1324]" class="Gastropoda" family="Alycaeidae" genus="Laotia" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="121" phylum="Mollusca" rank="species" species="pahiensis">
<emphasis id="B96F810FFFC6FFAAFC4EFAB2FB57BD0F" box="[986,1164,1300,1324]" italics="true" pageId="3" pageNumber="121">Laotia pahiensis</emphasis>
</taxonomicName>
(
<figureCitation id="13204198FFC6FFAAFB0EFAB3FB37BD0F" box="[1178,1260,1300,1324]" captionStart="Fig" captionStartId="4.[113,148,2035,2057]" captionTargetBox="[151,1413,189,2019]" captionTargetId="figure-13@4.[151,1413,189,2019]" captionTargetPageId="4" captionText="Fig. 3. Shells of Laotia Saurin, 1953 species. A, Laotia luongi, new species, holotype (VNMN-Z-Inv.000220); B, L. luongi, new species, paratype (coll. HA); C, syntype of Laotia pahiensis Saurin, 1953 (MNHN-IM-2000-28217. Photos: B. Páll-Gergely (A, B), M. Caballer (C, downloaded from the website of MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5351962" httpUri="https://zenodo.org/record/5351962/files/figure.png" pageId="3" pageNumber="121">Fig. 3C</figureCitation>
) also possesses ribs behind the peristome and has widely-spaced, straight ribs. Nevertheless, it is much smaller than the new species, and its aperture is less distorted upwards.
</paragraph>
</subSubSection>
<subSubSection id="C3010E96FFC6FFAAFCD7FA13FBE1BE2F" pageId="3" pageNumber="121" type="etymology">
<paragraph id="8BA45D1DFFC6FFAAFCD7FA13FBE1BE2F" blockId="3.[835,1474,1460,1548]" pageId="3" pageNumber="121">
<emphasis id="B96F810FFFC6FFAAFCD7FA13FC1CBDEF" bold="true" box="[835,967,1460,1484]" pageId="3" pageNumber="121">Etymology.</emphasis>
This new species is dedicated to and named after our much-valued friend, Hào Văn LƯƠng, who helped during field collection.
</paragraph>
</subSubSection>
<subSubSection id="C3010E96FFC6FFAAFCD7F993FA19BE4F" pageId="3" pageNumber="121" type="distribution">
<paragraph id="8BA45D1DFFC6FFAAFCD7F993FA19BE4F" blockId="3.[835,1474,1588,1646]" pageId="3" pageNumber="121">
<emphasis id="B96F810FFFC6FFAAFCD7F993FC03BE6F" bold="true" box="[835,984,1588,1612]" pageId="3" pageNumber="121">Distribution.</emphasis>
<taxonomicName id="4C1B269EFFC6FFAAFC76F992FBA2BE6F" authority="Páll-Gergely &amp; Hunyadi, 2021" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2021" box="[994,1145,1588,1612]" class="Gastropoda" family="Alycaeidae" genus="Laotia" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="121" phylum="Mollusca" rank="species" species="luongi" status="sp. nov.">
<emphasis id="B96F810FFFC6FFAAFC76F992FBA2BE6F" box="[994,1145,1588,1612]" italics="true" pageId="3" pageNumber="121">Laotia luongi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C3C74FFC6FFAAFB1EF993FAC8BE6F" box="[1162,1299,1588,1612]" pageId="3" pageNumber="121" rank="species">new species</taxonomicNameLabel>
, is known only from the
<typeStatus id="54A0E3BFFFC6FFAAFC30F9F3FC09BE4F" box="[932,978,1620,1644]" pageId="3" pageNumber="121">type</typeStatus>
locality in
<collectingRegion id="49DF93FFFFC6FFAAFBDEF9F3FADBBE4F" box="[1098,1280,1620,1646]" country="Vietnam" name="Son La" pageId="3" pageNumber="121">SƠn La Province</collectingRegion>
,
<collectingCountry id="F30C1D8DFFC6FFAAFA9DF9F3FABEBE4F" box="[1289,1381,1620,1644]" name="Vietnam" pageId="3" pageNumber="121">Vietnam</collectingCountry>
(
<figureCitation id="13204198FFC6FFAAFAE0F9F3FA6EBE4F" box="[1396,1461,1620,1644]" captionStart="Fig" captionStartId="3.[136,171,970,992]" captionTargetBox="[136,1474,189,955]" captionTargetId="figure-8@3.[136,1474,189,955]" captionTargetPageId="3" captionText="Fig. 2. Distribution of Laotia Saurin, 1953 species in Vietnam. Triangle: Laotia christahemmenae phami, new subspecies; inverse triangle: Laotia luongi, new species; square:Laotia christahemmenae christahemmenae Páll-Gergely, 2014 (filled square represents the type locality)." figureDoi="http://doi.org/10.5281/zenodo.5351960" httpUri="https://zenodo.org/record/5351960/files/figure.png" pageId="3" pageNumber="121">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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