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<mods:title id="78255AC92462ABFE727D6F7A284C9EE5">Feeding convergence among ray-finned fishes: Teeth of the herbivorous actinopterygians from the latest Permian of East European Platform, Russia</mods:title>
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<paragraph id="8BA7C37B8119FFE4FC934DD1FB5FFE63" blockId="3.[808,1233,415,443]" box="[808,1233,415,443]" pageId="3" pageNumber="74">
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<taxonomicName id="4C18B8F88119FFE4FC934DD1FB5FFE63" authority="Minikh, 1986" authorityName="Minikh" authorityYear="1986" box="[808,1233,415,443]" class="Insecta" family="Tunguskapteridae" genus="Isadia" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="suchonensis">
<emphasis id="B96C1F698119FFE4FC934DD1FB95FE63" box="[808,1051,415,443]" italics="true" pageId="3" pageNumber="74">Isadia suchonensis</emphasis>
Minikh, 1986
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<subSubSection id="C30290F08119FFE4FC934D82FC41F918" pageId="3" pageNumber="74" type="description">
<paragraph id="8BA7C37B8119FFE4FC934D82FC17FE39" blockId="3.[808,921,460,481]" box="[808,921,460,481]" pageId="3" pageNumber="74">
<figureCitation id="1323DFFE8119FFE4FC934D82FC1AFE39" box="[808,916,460,481]" captionStart="Fig" captionStartId="4.[151,184,1797,1817]" captionTargetBox="[174,1512,159,1772]" captionTargetId="figure-9@4.[174,1512,159,1772]" captionTargetPageId="4" captionText="Fig. 2. The isolated teeth of actinopterygian fish Isadia from the Sokovka outcrop, Vyazniki, Russia, late Permian (Upper Vyatkian). AD. Isadia aristoviensis Minikh, 1990, mandibulary teeth. A. ZPAL V.51/1, lingual view. B. ZPAL V.51/2, labial view. C. ZPAL V.51/3, lingual view. D. ZPAL V.51/4, labial view. EI. Isadia aristoviensis Minikh, 1990, maxillary teeth. E. ZPAL V.51/6, lingual view. F. ZPAL V.51/7, labial view. G. ZPAL V.51/5, lingual view. H. ZPAL V.51/8, lingual view. I. ZPAL V.51/9, labial view. J. Isadia arefievi Minikh, 2015, ZPAL V.51/10, mandibular tooth,?lingual view. K, L. Isadia suchonensis Minikh, 1986, mandibular teeth. K. ZPAL V.51/11, lingual (K1) and lateral (K2) views. L. ZPAL V.51/12, labial view. M. Isadia suchonensis Minikh, 1986, ZPAL V.51/13, maxillary teeth,?labial view. Scale bars 1 mm (AI), 0.5 mm (J, K, M), 0.2 mm (L)." figureDoi="http://doi.org/10.5281/zenodo.10980958" httpUri="https://zenodo.org/record/10980958/files/figure.png" pageId="3" pageNumber="74">Fig. 2KM</figureCitation>
.
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<paragraph id="8BA7C37B8119FFE4FC934DBAFC78FDB7" blockId="3.[808,1481,500,623]" pageId="3" pageNumber="74">
<emphasis id="B96C1F698119FFE4FC934DBAFC04FDD5" box="[808,906,500,525]" italics="true" pageId="3" pageNumber="74">Material</emphasis>
.—Isolated teeth (
<collectionCode id="ED095BBE8119FFE4FBF44DBAFB18FDD5" box="[1103,1174,500,525]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="3" pageNumber="74">ZPAL</collectionCode>
<collectionCode id="ED095BBE8119FFE4FB214DBAFB20FDD5" box="[1178,1198,500,525]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="74" type="Museum">V</collectionCode>
.51/1113) and 25 unnumbered teeth: 14 maxillary and 11 mandibular from Sokovka outcrop, Vyazniki in
<collectingRegion id="49DC0D998119FFE4FBAD4E7BFB57FD96" box="[1046,1241,565,590]" country="Russia" name="Vladimir" pageId="3" pageNumber="74">Vladimir Region</collectingRegion>
,
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, late Permian (Upper Vyatkian).
</paragraph>
<paragraph id="8BA7C37B8119FFE4FC934ECFFB1AFB7B" blockId="3.[808,1481,641,1187]" pageId="3" pageNumber="74">
<emphasis id="B96C1F698119FFE4FC934ECFFC20FD42" box="[808,942,641,666]" italics="true" pageId="3" pageNumber="74">Description</emphasis>
.—The teeth are curved in anterior view, with tips directed slightly lingually. A shallow row in the middle of the crown is visible on the labial side in some specimens. It is oriented parallel to the tooth axis. Teeth are small and have interior passage larger than
<taxonomicName id="4C18B8F88119FFE4FB084F4DFA1BFCC4" authorityName="Minikh" authorityYear="1990" box="[1203,1429,771,796]" class="Insecta" family="Tunguskapteridae" genus="Isadia" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="aristoviensis">
<emphasis id="B96C1F698119FFE4FB084F4DFA1BFCC4" box="[1203,1429,771,796]" italics="true" pageId="3" pageNumber="74">Isadia aristoviensis</emphasis>
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and it ends close to the tips of the crown. The crown of the collected teeth has 47 cusps (in some specimens of this species collected from other outcrops show 39 cusps on the crown). The tips of the cusps are not covered by acrodine (in our investigations, the acrodine tips in discussed species are developed and clearly distinct from tooth bases in absence of tubercle microstructure of the external surface). The length of the teeth is 1.161.47 mm. Two morphotypes of these teeth were found. One with 45 cusps, and second with 67 smaller cusps. The mandibular teeth have straight base of the crown with less cusplets than crowns of the maxillary teeth (
<bibRefCitation id="EF89BE8A8119FFE4FC0948C4FB06FB7B" author="Minikh, A. V. &amp; Arefiev, M. P. &amp; Golubev, V. K." box="[946,1160,1162,1187]" pageId="3" pageNumber="74" pagination="615 - 626" refId="ref6931" refString="Minikh, A. V., Arefiev, M. P., and Golubev, V. K. 2015. A new fish species of the genus Isadia (Actinopterygii, Eurynototoidiformes) from the new locality on the Malaya Northern Dvina River (terminal Permian, Vologda Region). Paleontological Journal 49 (6): 615 - 626." type="journal article" year="2015">Minikh et al. 2015</bibRefCitation>
).
</paragraph>
<paragraph id="8BA7C37B8119FFE4FC9348FBFB6CF9AD" blockId="3.[808,1481,1205,1653]" pageId="3" pageNumber="74">
<emphasis id="B96C1F698119FFE4FC9348FBFC02FB16" box="[808,908,1205,1230]" italics="true" pageId="3" pageNumber="74">Remarks</emphasis>
.—
<taxonomicName id="4C18B8F88119FFE4FC0B48FBFAC7FB16" authority="Minikh, 1986" authorityName="Minikh" authorityYear="1986" box="[944,1353,1205,1230]" class="Insecta" family="Tunguskapteridae" genus="Isadia" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="suchonensis">
<emphasis id="B96C1F698119FFE4FC0B48FBFB1DFB16" box="[944,1171,1205,1230]" italics="true" pageId="3" pageNumber="74">Isadia suchonensis</emphasis>
Minikh, 1986
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(
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SGU 104 Б/P-2, Mutovino locality) is the type species of this genus. It is distinguished by elongated and narrow teeth on a short maxilla (Minikh and Minikh 1986). All maxillary teeth are morphologically similar to each other. The mandibular teeth differ by width of cusplets. All have wide crown with many small cusps. The teeth from Sokovka outcrop are not as large as in previously known specimens of
<taxonomicName id="4C18B8F88119FFE4FC9349F7FC49FA0A" authorityName="Minikh" authorityYear="1986" box="[808,967,1465,1490]" class="Insecta" family="Tunguskapteridae" genus="Isadia" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="suchonensis">
<emphasis id="B96C1F698119FFE4FC9349F7FC49FA0A" box="[808,967,1465,1490]" italics="true" pageId="3" pageNumber="74">I. suchonensis</emphasis>
</taxonomicName>
. In the
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of
<taxonomicName id="4C18B8F88119FFE4FB1D49F7FAC8FA0A" authorityName="Minikh" authorityYear="1986" box="[1190,1350,1465,1490]" class="Insecta" family="Tunguskapteridae" genus="Isadia" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="suchonensis">
<emphasis id="B96C1F698119FFE4FB1D49F7FAC8FA0A" box="[1190,1350,1465,1490]" italics="true" pageId="3" pageNumber="74">I. suchonensis</emphasis>
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acrodine is very small but still noticeable. The small size of specimens from Sokovka suggests that it might be a subspecies of
<taxonomicName id="4C18B8F88119FFE4FC934A55FC44F9EC" authorityName="Minikh" authorityYear="1986" box="[808,970,1563,1588]" class="Insecta" family="Tunguskapteridae" genus="Isadia" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="suchonensis">
<emphasis id="B96C1F698119FFE4FC934A55FC44F9EC" box="[808,970,1563,1588]" italics="true" pageId="3" pageNumber="74">I. suchonensis</emphasis>
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that developed lack of acrodine or this is an effect of ontogeny. Likewise, this could be a preservational artifact. This requires further research.
</paragraph>
<paragraph id="8BA7C37B8119FFE4FC934AC9FC41F918" blockId="3.[808,1481,1671,1728]" pageId="3" pageNumber="74">
<emphasis id="B96C1F698119FFE4FC934AC9FB4EF978" box="[808,1216,1671,1696]" italics="true" pageId="3" pageNumber="74">Stratigraphic and geographic range</emphasis>
.—Permian, East European Platform.
</paragraph>
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