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<document id="9834DC3C5EBC905ED3DB52BDBD89A36F" ID-DOI="10.11646/zootaxa.3710.2.1" ID-GBIF-Dataset="867ca84b-ed17-40e4-a603-b2efd0e6fef3" ID-ISSN="1175-5326" ID-Zenodo-Dep="248125" ID-ZooBank="6E42B926-DAA9-4BAE-B995-8BDB19B93268" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460675777785" checkinUser="plazi" docAuthor="Winston, Judith E. &amp; Vieira, Leandro M." docDate="2013" docId="03B10F76FF93DB73FF7BBCC67942227C" docLanguage="en" docName="zt03710p146.pdf" docOrigin="Zootaxa 3710 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psammocleidochasma Winston &amp; Vieira, 2013, gen. nov." docType="treatment" docVersion="9" lastPageNumber="133" masterDocId="FF88770EFF8CDB52FFECBA0E7B0B2361" masterDocTitle="Systematics of interstitial encrusting bryozoans from southeastern Brazil" masterLastPageNumber="146" masterPageNumber="101" pageNumber="132" updateTime="1698349855894" updateUser="plazi">
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<mods:title id="59EF9B32D3A7A89ED57006BCAD6C9972">Systematics of interstitial encrusting bryozoans from southeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="50EA4DD4764F3A540B5AFC73EFCD3DA2" type="personal">
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<mods:namePart id="813A3DC7DDF059381B28516AD9977DE4">Winston, Judith E.</mods:namePart>
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<mods:namePart id="698B4C94B5F5B2ABA18E252AB8D5BAE8">Vieira, Leandro M.</mods:namePart>
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<mods:title id="38B6ED315747E1B41C18C9CC9F01E636">Zootaxa</mods:title>
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<mods:part id="D82553E05D8E53D8AF0A1CD89BCF5B69">
<mods:date id="9A8E2DF8E0908A495622B93C3C6796A7">2013</mods:date>
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<mods:number id="BCC8B3F49597744399C0E8673274D7F1">3710</mods:number>
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<mods:identifier id="700DA1D331B98B23BEC705CBF19E35C7" type="DOI">10.11646/zootaxa.3710.2.1</mods:identifier>
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<treatment id="03B10F76FF93DB73FF7BBCC67942227C" ID-DOI="http://doi.org/10.5281/zenodo.6163660" ID-GBIF-Taxon="119581245" ID-Zenodo-Dep="6163660" LSID="urn:lsid:plazi:treatment:03B10F76FF93DB73FF7BBCC67942227C" httpUri="http://treatment.plazi.org/id/03B10F76FF93DB73FF7BBCC67942227C" lastPageId="33" lastPageNumber="133" pageId="31" pageNumber="132">
<subSubSection id="C302EDEBFF93DB4DFF7BBCC679642583" box="[151,623,1736,1762]" pageId="31" pageNumber="132" type="nomenclature">
<paragraph id="8BA7BE60FF93DB4DFF7BBCC679642583" blockId="31.[151,623,1736,1762]" box="[151,623,1736,1762]" pageId="31" pageNumber="132">
<heading id="D0EF090CFF93DB4DFF7BBCC679642583" bold="true" box="[151,623,1736,1762]" fontSize="11" level="1" pageId="31" pageNumber="132" reason="1">
<emphasis id="B96C6272FF93DB4DFF7BBCC679642583" bold="true" box="[151,623,1736,1762]" pageId="31" pageNumber="132">
Genus
<taxonomicName id="4C18C5E3FF93DB4DFF02BCC67AF02583" ID-CoL="84SV9" box="[238,507,1736,1762]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="31" pageNumber="132" phylum="Bryozoa" rank="genus" status="gen. nov.">
<emphasis id="B96C6272FF93DB4DFF02BCC67AF02583" bold="true" box="[238,507,1736,1762]" italics="true" pageId="31" pageNumber="132">Psammocleidochasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25FDF09FF93DB4DFDEEBCC679642583" box="[514,623,1736,1762]" pageId="31" pageNumber="132" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302EDEBFF93DB4DFF7BBD0078152446" box="[151,798,1806,1831]" pageId="31" pageNumber="132" type="materials_examined">
<paragraph id="8BA7BE60FF93DB4DFF7BBD0078152446" blockId="31.[151,1436,1806,2011]" box="[151,798,1806,1831]" pageId="31" pageNumber="132">
<emphasis id="B96C6272FF93DB4DFF7BBD007A392446" bold="true" box="[151,306,1806,1831]" pageId="31" pageNumber="132">
<typeStatus id="54A300C2FF93DB4DFF7BBD007BD92446" box="[151,210,1806,1831]" pageId="31" pageNumber="132">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C18C5E3FF93DB4DFED5BD0079B62446" box="[313,701,1806,1831]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="31" pageNumber="132" phylum="Bryozoa" rank="species" species="tridentatum" status="sp. nov.">
<emphasis id="B96C6272FF93DB4DFED5BD0079B62446" box="[313,701,1806,1831]" italics="true" pageId="31" pageNumber="132">Psammocleidochasma tridentatum</emphasis>
</taxonomicName>
<emphasis id="B96C6272FF93DB4DFD28BD0078152446" bold="true" box="[708,798,1806,1831]" pageId="31" pageNumber="132">
<taxonomicNameLabel id="A25FDF09FF93DB4DFD28BD0078152446" box="[708,798,1806,1831]" pageId="31" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C302EDEBFF93DB4DFF2BBD3C79CE242A" box="[199,709,1842,1867]" pageId="31" pageNumber="132" type="etymology">
<paragraph id="8BA7BE60FF93DB4DFF2BBD3C79CE242A" blockId="31.[151,1436,1806,2011]" box="[199,709,1842,1867]" pageId="31" pageNumber="132">
<emphasis id="B96C6272FF93DB4DFF2BBD3C7A45242A" bold="true" box="[199,334,1842,1867]" pageId="31" pageNumber="132">Etymology.</emphasis>
From the Greek,
<emphasis id="B96C6272FF93DB4DFDF8BD3A7977242A" box="[532,636,1844,1867]" italics="true" pageId="31" pageNumber="132">psammos</emphasis>
, sand.
</paragraph>
</subSubSection>
<subSubSection id="C302EDEBFF93DB4DFF2BBD5879EB24BB" pageId="31" pageNumber="132" type="diagnosis">
<paragraph id="8BA7BE60FF93DB4DFF2BBD5879EB24BB" blockId="31.[151,1436,1806,2011]" pageId="31" pageNumber="132">
<emphasis id="B96C6272FF93DB4DFF2BBD587A4A240E" bold="true" box="[199,321,1878,1903]" pageId="31" pageNumber="132">Diagnosis.</emphasis>
<taxonomicName id="4C18C5E3FF93DB4DFEBBBD587AFC240E" box="[343,503,1878,1903]" class="Gymnolaemata" family="Cleidochasmatidae" genus="Cleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="31" pageNumber="132" phylum="Bryozoa" rank="genus">
<emphasis id="B96C6272FF93DB4DFEBBBD587AFC240E" box="[343,503,1878,1903]" italics="true" pageId="31" pageNumber="132">Cleidochasma</emphasis>
</taxonomicName>
-like, but sand-emcrusting and having a relatively smaller size of colony and autozooids, a tatiform ancestrula, no avicularia, uniserial to pluriserial growth, and thickened calcification of ovicells, as well as a raised peristome with 34 projecting solid tubercles arranged distally and laterally on its rim. Ooecium imperforate with semicircular frontal area.
</paragraph>
</subSubSection>
<caption id="DF67EEE8FFACDB72FF7BBCCD7A8C2454" httpUri="https://zenodo.org/record/248146/files/figure.png" pageId="32" pageNumber="133" targetBox="[151,1436,195,1707]" targetPageId="32">
<paragraph id="8BA7BE60FFACDB72FF7BBCCD7A8C2454" blockId="32.[151,1437,1731,1845]" pageId="32" pageNumber="133">
<emphasis id="B96C6272FFACDB72FF7BBCCD7A2925B9" bold="true" box="[151,290,1731,1753]" pageId="32" pageNumber="133">FIGURE 21.</emphasis>
<taxonomicName id="4C18C5E3FFACDB72FEC6BCCD798025B8" box="[298,651,1731,1753]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="32" pageNumber="133" phylum="Bryozoa" rank="species" species="tridentatum" status="sp. nov.">
<emphasis id="B96C6272FFACDB72FEC6BCCD798025B8" box="[298,651,1731,1753]" italics="true" pageId="32" pageNumber="133">Psammocleidochasma tridentatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25FDF09FFACDB72FD7FBCCA79ED25B8" box="[659,742,1732,1753]" pageId="32" pageNumber="133" rank="species">
<emphasis id="B96C6272FFACDB72FD7FBCCA79ED25B8" bold="true" box="[659,742,1732,1753]" pageId="32" pageNumber="133">sp. nov.</emphasis>
</taxonomicNameLabel>
: A, a colony attached to one side of a grain. B, a uniserial colony encrusting the edge of a grain. C, three zooids of the colony in B. D, profile of zooids showing projecting peristomial tubercles. E, ovicell and orifice with three tubercles surrounding it. F, zooidal orifice. Scale bars: A, B, 0.5 mm; C, 0.3 mm; D, 0.15 mm; E, 0.1 mm; F, 0.05 mm.
</paragraph>
</caption>
<subSubSection id="C302EDEBFFACDB73FF2BBD6C7942227C" lastPageId="33" lastPageNumber="134" pageId="32" pageNumber="133" type="discussion">
<paragraph id="8BA7BE60FFACDB73FF2BBD6C7942227C" blockId="32.[151,1436,1890,2023]" lastBlockId="33.[151,1436,151,285]" lastPageId="33" lastPageNumber="134" pageId="32" pageNumber="133">
<emphasis id="B96C6272FFACDB72FF2BBD6C7A30241A" bold="true" box="[199,315,1890,1915]" pageId="32" pageNumber="133">Remarks.</emphasis>
Recent revisions of species originally placed in the the genus
<emphasis id="B96C6272FFACDB72FBC8BD6C7FEB241A" box="[1060,1248,1890,1915]" italics="true" pageId="32" pageNumber="133">Cleidodochasma</emphasis>
have led to the discovery that at least two groups of species, belonging to different families, are involved (Soule
<emphasis id="B96C6272FFACDB72FB35BD897E1A24FE" box="[1241,1297,1926,1951]" italics="true" pageId="32" pageNumber="133">et al.</emphasis>
1991; Cook &amp; Bock 1996). The two sand-dwelling species attributed to this genus,
<taxonomicName id="4C18C5E3FFACDB72FC5BBDA47E3024A2" box="[951,1339,1962,1987]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="32" pageNumber="133" phylum="Bryozoa" rank="species" species="tridentatum" status="sp. nov.">
<emphasis id="B96C6272FFACDB72FC5BBDA47E3024A2" box="[951,1339,1962,1987]" italics="true" pageId="32" pageNumber="133">Psammocleidochasma tridentatum</emphasis>
</taxonomicName>
<emphasis id="B96C6272FFACDB72FAAEBDA47E9724A2" bold="true" box="[1346,1436,1962,1987]" pageId="32" pageNumber="133">
<taxonomicNameLabel id="A25FDF09FFACDB72FAAEBDA47E9724A2" box="[1346,1436,1962,1987]" pageId="32" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<collectingCountry id="F30FFEF0FFACDB72FF35BDC07A162486" box="[217,285,1998,2023]" name="Brazil" pageId="32" pageNumber="133">Brazil</collectingCountry>
and
<taxonomicName id="4C18C5E3FFACDB72FEB1BDC07F342486" authority="Winston &amp; Hakansson, 1986" authorityName="Winston &amp; Hakansson" authorityYear="1986" box="[349,1087,1998,2023]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="32" pageNumber="133" phylum="Bryozoa" rank="species" species="angustum" status="comb. nov.">
<emphasis id="B96C6272FFACDB72FEB1BDC079DB2487" box="[349,720,1998,2023]" italics="true" pageId="32" pageNumber="133">Psammocleidochasma angustum</emphasis>
(Winston &amp; Håkansson, 1986)
</taxonomicName>
<emphasis id="B96C6272FFACDB72FBA6BDC07FC42486" bold="true" box="[1098,1231,1998,2023]" pageId="32" pageNumber="133">
<taxonomicNameLabel id="A25FDF09FFACDB72FBA6BDC07FC42486" box="[1098,1231,1998,2023]" pageId="32" pageNumber="133">comb. nov.</taxonomicNameLabel>
</emphasis>
from Florida are distinctive in habitat and morphology and appear to belong with encrusting genera such as
<taxonomicName id="4C18C5E3FFADDB73FB50BA997AF723B5" authority="Soule, Soule &amp; Channey, 1991" authorityName="Soule, Soule &amp; Channey" authorityYear="1991" class="Gymnolaemata" family="Phidoloporidae" genus="Plesiocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="134" phylum="Bryozoa" rank="genus">
<emphasis id="B96C6272FFADDB73FB50BA997E9723D1" box="[1212,1436,151,176]" italics="true" pageId="33" pageNumber="134">Plesiocleidochasma</emphasis>
Soule, Soule &amp; Channey, 1991
</taxonomicName>
(=
<taxonomicName id="4C18C5E3FFADDB73FDC6BAB27F8C23B5" authority="Soule, Soule &amp; Channey, 1991" authorityName="Soule, Soule &amp; Channey" authorityYear="1991" box="[554,1159,188,213]" class="Gymnolaemata" family="Phidoloporidae" genus="Schedocleidochasma" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="134" phylum="Bryozoa" rank="genus">
<emphasis id="B96C6272FFADDB73FDC6BAB2781E23B4" box="[554,789,188,213]" italics="true" pageId="33" pageNumber="134">Schedocleidochasma</emphasis>
Soule, Soule &amp; Channey, 1991
</taxonomicName>
; see Berning 2012) and relatives, now placed in the Phidolophoridae. Owing to their distinctive suite of characters we have placed the two sand-dwelling forms in the new genus.
</paragraph>
</subSubSection>
</treatment>
</document>