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<document id="FD0660C32077ED9DC409157A6BE65A71" ID-CLB-Dataset="22649" ID-DOI="10.11646/zootaxa.4470.1.1" ID-GBIF-Dataset="615d5c4a-3b8d-4ba6-a05d-f37e432a862b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3770039" ID-ZooBank="60766083-99E8-48E9-86CE-161A9EEC066B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588000738706" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2018" docId="03AE6457792C9F7F4A9AFCCCFD5B1A96" docLanguage="en" docName="zootaxa.4470.1.1.pdf" docOrigin="Zootaxa 4470 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argia cuspidata Garrison &amp; von Ellenrieder 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="13" masterDocId="FF971C2F79279F734A0DFFD3FF981C75" masterDocTitle="Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species" masterLastPageNumber="69" masterPageNumber="1" pageNumber="12" updateTime="1698837067205" updateUser="ExternalLinkService">
<mods:mods id="BE5EDA8B7AFAA24C089F07951CD35AB3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8F9DB5E57F71FF48016A589ADB1BC5C5">
<mods:title id="D84E97DD2AB16C17B409F7EA31B9ABDE">Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species</mods:title>
</mods:titleInfo>
<mods:name id="4F519CDF4074564D0F37BE2645A60C86" type="personal">
<mods:role id="01957039E171C54D83B9ED945AB26F99">
<mods:roleTerm id="34FC988DCD15BC47165772F94FE68E77">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2A77A73E988083D1C59624090DB81ACA">Garrison, Rosser W.</mods:namePart>
</mods:name>
<mods:name id="4BB9E921EFB1FEB4FB2AECE91F9B1EBC" type="personal">
<mods:role id="C222EF17E9123FDF066CDA7399B86889">
<mods:roleTerm id="CDACC5568EBDB1974A2E3320FAD6F1C0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7403F87104394043DDC65CEAF1BF38B0">Ellenrieder, Natalia Von</mods:namePart>
</mods:name>
<mods:typeOfResource id="491F97791A892586DE3A226FCCA610BD">text</mods:typeOfResource>
<mods:relatedItem id="23EC3AC1B7BD227391CA062EE05F84A2" type="host">
<mods:titleInfo id="8AF7B72232743213CD51B6AE647003ED">
<mods:title id="86138DC3A4EEFF8EC03C62E018423B21">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4B7B58C2469EB5A06905A2168502A459">
<mods:date id="EFE36D4A0E6C0F417984DF49D6F71457">2018</mods:date>
<mods:detail id="DA6FABEA3B03EB935F6DBBAA18594653" type="pubDate">
<mods:number id="3E9E0044E1E0098D7AD6A89C0EF0AEFF">2018-09-03</mods:number>
</mods:detail>
<mods:detail id="47806158510DF9EE49B00D9D1D13AD8C" type="volume">
<mods:number id="F0DA3C11E2A7C8707465AAC04DB387AC">4470</mods:number>
</mods:detail>
<mods:detail id="C40B11CEDCA1574D352A8CBA22A17BD4" type="issue">
<mods:number id="CFCCBD151A73EA7A88ACD39FD33F9B40">1</mods:number>
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<mods:start id="C5662D91F4E3773671CD04F68FABC579">1</mods:start>
<mods:end id="5D3CE5283698033442EFEE4FFA0EE65E">69</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BC5153C0AB0DC68DFA035DACFBF02CB0">journal article</mods:classification>
<mods:identifier id="9B72DDB6AAB091530F1D74664E4C4A82" type="CLB-Dataset">22649</mods:identifier>
<mods:identifier id="A6B22036E3F72BF8B427C3AA02EE4BAF" type="DOI">10.11646/zootaxa.4470.1.1</mods:identifier>
<mods:identifier id="D10E7F9EA9F3345D6C948CDB57626D70" type="GBIF-Dataset">615d5c4a-3b8d-4ba6-a05d-f37e432a862b</mods:identifier>
<mods:identifier id="08C75586AF0C26A3214CD73B6FA06AB5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="96B942CC2EE20B6E06C19259000D6F21" type="Zenodo-Dep">3770039</mods:identifier>
<mods:identifier id="A45BA22B3D7E176EB06AB8E995899896" type="ZooBank">60766083-99E8-48E9-86CE-161A9EEC066B</mods:identifier>
</mods:mods>
<treatment id="03AE6457792C9F7F4A9AFCCCFD5B1A96" ID-DOI="http://doi.org/10.5281/zenodo.3799123" ID-GBIF-Taxon="163754280" ID-Zenodo-Dep="3799123" LSID="urn:lsid:plazi:treatment:03AE6457792C9F7F4A9AFCCCFD5B1A96" httpUri="http://treatment.plazi.org/id/03AE6457792C9F7F4A9AFCCCFD5B1A96" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="C31D86CA792C9F784A9AFCCCFC9F1F4F" box="[151,775,799,826]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BB8D541792C9F784A9AFCCCFC9F1F4F" blockId="11.[151,1436,799,895]" box="[151,775,799,826]" pageId="11" pageNumber="12">
<heading id="D0F0622D792C9F784A9AFCCCFC9F1F4F" bold="true" box="[151,775,799,826]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B9730953792C9F784A9AFCCCFC9F1F4F" bold="true" box="[151,775,799,826]" pageId="11" pageNumber="12">
<taxonomicName id="4C07AEC2792C9F784A9AFCCCFD201F4F" ID-CoL="9X22K" authority="Garrison &amp; von Ellenrieder" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[151,696,799,826]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cuspidata" status="sp. nov.">
<emphasis id="B9730953792C9F784A9AFCCCFECE1F4C" bold="true" box="[151,342,799,825]" italics="true" pageId="11" pageNumber="12">Argia cuspidata</emphasis>
Garrison &amp; von Ellenrieder
</taxonomicName>
,
<taxonomicNameLabel id="A240B428792C9F7848C8FCF3FC9F1F4F" box="[709,775,800,826]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA792C9F784A9AFC91FCC51F0A" pageId="11" pageNumber="12" type="description">
<paragraph id="8BB8D541792C9F784A9AFC91FCC51F0A" blockId="11.[151,1436,799,895]" pageId="11" pageNumber="12">
<figureCitation id="133CC9C4792C9F784A9AFC91FF7B1F2E" box="[151,227,834,859]" captionStart="FIGURES 1–8" captionStartId="39.[151,264,1966,1988]" captionTargetBox="[186,1424,239,1889]" captionTargetId="figure@39.[151,1437,193,1945]" captionTargetPageId="39" captionText="FIGURES 1–8. Color pattern of head. Head, anterior view (1); head posterior view (2–5); labrum, anterior view (6–8)." figureDoi="http://doi.org/10.5281/zenodo.3770041" httpUri="https://zenodo.org/record/3770041/files/figure.png" pageId="11" pageNumber="12">Figs. 1</figureCitation>
(head
<specimenCount id="9D011EC8792C9F784B22FC90FEDB1F2E" box="[303,323,835,859]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
), 13 (head, thorax, S1–
<specimenCount id="9D011EC8792C9F784841FC90FDEF1F2E" box="[588,631,835,859]" pageId="11" pageNumber="12" type="male">3 ♂</specimenCount>
), 24 (head, thorax, S1–
<specimenCount id="9D011EC8792C9F784970FC90FC3E1F2E" box="[893,934,835,859]" pageId="11" pageNumber="12" type="female">3 ♀</specimenCount>
), 45 (S1–
<specimenCount id="9D011EC8792C9F784E1BFC90FBDA1F2E" box="[1046,1090,835,859]" pageId="11" pageNumber="12" type="male">3 ♂</specimenCount>
), 46, 47 (S7–
<specimenCount id="9D011EC8792C9F784ED4FC90FA8B1F2E" box="[1241,1299,835,859]" pageId="11" pageNumber="12" type="male">10 ♂</specimenCount>
), 60 (S7–
<specimenCount id="9D011EC8792C9F784F8DFC90FF331F0A" pageId="11" pageNumber="12" type="female">10 ♀</specimenCount>
), 72 (mesostigmal plates
<specimenCount id="9D011EC8792C9F784BC1FCB4FE781F0A" box="[460,480,871,895]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
), 106 (appendages
<specimenCount id="9D011EC8792C9F7848B0FCB4FD4C1F0A" box="[701,724,871,895]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
), 136 (map)
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA792C9F784A9AFC7DFC851902" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BB8D541792C9F784A9AFC7DFEE81F9F" blockId="11.[151,1437,942,2014]" pageId="11" pageNumber="12">
<emphasis id="B9730953792C9F784A9AFC7DFE8E1FB2" bold="true" box="[151,278,942,967]" pageId="11" pageNumber="12">Etymology</emphasis>
. Named
<taxonomicName id="4C07AEC2792C9F784B70FC7DFE721FB3" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[381,490,942,966]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792C9F784B70FC7DFE721FB3" box="[381,490,942,966]" italics="true" pageId="11" pageNumber="12">cuspidata</emphasis>
</taxonomicName>
(L.
<taxonomicName id="4C07AEC2792C9F784816FC7DFD101FB3" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[539,648,942,966]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792C9F784816FC7DFD101FB3" box="[539,648,942,966]" italics="true" pageId="11" pageNumber="12">cuspidata</emphasis>
</taxonomicName>
, adj.) in reference to the large ventrally directed blunt cusp-like tooth on the male cercus.
</paragraph>
<paragraph id="8BB8D541792C9F784ACAFC25FB561846" blockId="11.[151,1437,942,2014]" pageId="11" pageNumber="12">
<emphasis id="B9730953792C9F784ACAFC25FE94187A" bold="true" box="[199,268,1014,1039]" pageId="11" pageNumber="12">Types</emphasis>
.
<materialsCitation id="3B6FDF1C792C9F784B16FC25FB561846" ID-GBIF-Occurrence="2608962915" collectingDate="1975-05-07" collectionCode="USNM" collectorName="A. Langley &amp; J. Cohen" country="Ecuador" elevation="740" latitude="-0.31361112" location="Colorados" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Santo Domingo de los Tsachilas" typeStatus="holotype">
<typeStatus id="54BC6BE3792C9F784B16FC25FE1B187A" box="[283,387,1014,1039]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
<specimenCount id="9D011EC8792C9F784B86FC24FE3B187B" box="[395,419,1015,1038]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
:
<collectingCountry id="F31095D1792C9F784BA2FC24FDA1187A" box="[431,569,1015,1039]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
,
<collectingRegion id="49C31BA3792C9F784848FC24FC6D187B" box="[581,1013,1014,1039]" country="Ecuador" name="Santo Domingo de los Tsachilas" pageId="11" pageNumber="12">Santo Domingo de los Tsáchilas Prov.</collectingRegion>
:
<quantity id="4CFF78A4792C9F784E0DFC24FBD0187A" box="[1024,1096,1014,1039]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="km" value="19.0">19 km</quantity>
east of
<collectingMunicipality id="6BDC4F3B792C9F784EACFC24FACA187A" box="[1185,1362,1014,1039]" pageId="11" pageNumber="12">Santo Domingo</collectingMunicipality>
de los
<location id="8ED8839A792C9F784A9AFBC9FE921846" LSID="urn:lsid:plazi:treatment:03AE6457792C9F7F4A9AFCCCFD5B1A96:8ED8839A792C9F784A9AFBC9FE921846" box="[151,266,1050,1075]" country="Ecuador" latitude="-0.31361112" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" name="Colorados" pageId="11" pageNumber="12" stateProvince="Santo Domingo de los Tsachilas">Colorados</location>
{
<geoCoordinate id="EE33B386792C9F784B17FBC9FE0F1846" box="[282,407,1050,1075]" degrees="0" direction="south" minutes="18" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="49" value="-0.31361112">0°18'49'' S</geoCoordinate>
,
<geoCoordinate id="EE33B386792C9F784BAEFBC8FDBE1847" box="[419,550,1051,1075]" degrees="79" direction="west" minutes="1" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="44" value="-79.02889">79°1'44'' W</geoCoordinate>
,
<quantity id="4CFF78A4792C9F78483DFBC8FDE51846" box="[560,637,1050,1075]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="12" unit="m" value="740.0">
<elevation id="002A3272792C9F78483DFBC8FDE51846" box="[560,637,1050,1075]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="12" unit="m" value="740.0">740 m</elevation>
</quantity>
},
<date id="FFB9F381792C9F78489EFBC8FC851846" box="[659,797,1050,1075]" pageId="11" pageNumber="12" value="1975-05-07">
<collectingDate id="EFFD0A69792C9F78489EFBC8FC851846" box="[659,797,1050,1075]" pageId="11" pageNumber="12" value="1975-05-07">7 May 1975</collectingDate>
</date>
,
<collectorName id="26F2B097792C9F784925FBC8FC3E1846" box="[808,934,1050,1075]" pageId="11" pageNumber="12">A. Langley</collectorName>
&amp;
<collectorName id="26F2B097792C9F7849C7FBC8FBB41846" box="[970,1068,1050,1075]" pageId="11" pageNumber="12">J. Cohen</collectorName>
leg. [
<collectionCode id="ED164D84792C9F784E67FBC8FB5C1846" box="[1130,1220,1051,1075]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12" type="Museum">USNM</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="8BB8D541792C9F784ACAFBEDFCD6195A" blockId="11.[151,1437,942,2014]" pageId="11" pageNumber="12">
<emphasis id="B9730953792C9F784ACAFBEDFDEB1822" bold="true" box="[199,627,1086,1111]" pageId="11" pageNumber="12">
Specimens examined (all
<typeStatus id="54BC6BE3792C9F784BFEFBEDFDF31822" box="[499,619,1086,1111]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
)
</emphasis>
.
<materialsCitation id="3B6FDF1C792C9F78488DFBECFBC31823" ID-GBIF-Occurrence="2608962962" box="[640,1115,1086,1111]" country="Ecuador" location="Esmeraldas Prov." pageId="11" pageNumber="12" specimenCount="9" specimenCount-female="1" specimenCount-male="8" stateProvince="Esmeraldas">
<specimenCount id="9D011EC8792C9F78488DFBECFD581823" box="[640,704,1087,1111]" pageId="11" pageNumber="12" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="9D011EC8792C9F7848C1FBECFD6F1823" box="[716,759,1087,1111]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
.
<collectingCountry id="F31095D1792C9F784909FBECFC161822" box="[772,910,1087,1111]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
,
<collectingRegion id="49C31BA3792C9F784995FBEDFBC31823" box="[920,1115,1086,1111]" country="Ecuador" name="Esmeraldas" pageId="11" pageNumber="12">Esmeraldas Prov.</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B6FDF1C792C9F784E65FBECFF4E18EB" ID-GBIF-Occurrence="2608962932" collectingDate="1997-02-02" collectionCode="JJD" collectorName="Jerrell J. Daigle" country="Ecuador" elevation="470" latitude="0.885" location="Lita" longLatPrecision="21" longitude="-78.475555" pageId="11" pageNumber="12" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D011EC8792C9F784E65FBECFB0C1823" box="[1128,1172,1087,1111]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D011EC8792C9F784E92FBECFB511823" box="[1183,1225,1087,1111]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
(in tandem), seeps
<quantity id="4CFF78A4792C9F784A9AFBB0FF7F180E" box="[151,231,1122,1147]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="12" unit="km" value="1.1">1.1 km</quantity>
NW of
<location id="8ED8839A792C9F784B4AFBB1FEEC180E" LSID="urn:lsid:plazi:treatment:03AE6457792C9F7F4A9AFCCCFD5B1A96:8ED8839A792C9F784B4AFBB1FEEC180E" box="[327,372,1122,1147]" country="Ecuador" latitude="0.885" longLatPrecision="21" longitude="-78.475555" name="Lita" pageId="11" pageNumber="12">Lita</location>
at bridge, 5:00 pm {
<geoCoordinate id="EE33B386792C9F78486CFBB1FD40180F" box="[609,728,1122,1147]" degrees="0" direction="north" minutes="53" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="6" value="0.885">0°53'6'' N</geoCoordinate>
,
<geoCoordinate id="EE33B386792C9F7848E8FBB0FCEF180F" box="[741,887,1123,1147]" degrees="78" direction="west" minutes="28" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="32" value="-78.475555">78°28'32'' W</geoCoordinate>
,
<quantity id="4CFF78A4792C9F784989FBB0FC4B180E" box="[900,979,1122,1147]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="12" unit="m" value="470.0">
<elevation id="002A3272792C9F784989FBB0FC4B180E" box="[900,979,1122,1147]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="12" unit="m" value="470.0">470 m</elevation>
</quantity>
},
<date id="FFB9F381792C9F7849E4FBB0FB31180E" box="[1001,1193,1122,1147]" pageId="11" pageNumber="12" value="1997-02-02">
<collectingDate id="EFFD0A69792C9F7849E4FBB0FB31180E" box="[1001,1193,1122,1147]" pageId="11" pageNumber="12" value="1997-02-02">2 February 1997</collectingDate>
</date>
,
<collectorName id="26F2B097792C9F784EBBFBB1FAF3180E" box="[1206,1387,1122,1147]" pageId="11" pageNumber="12">Jerrell J. Daigle</collectorName>
leg. [
<collectionCode id="ED164D84792C9F784A93FB54FF5718EB" box="[158,207,1159,1182]" pageId="11" pageNumber="12">JJD</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C792C9F784AE8FB54FA0018B6" ID-GBIF-Occurrence="2608962936" collectingDate="1997-02-05" collectionCode="RWG" collectorName="Jerrell J. Daigle" country="Ecuador" elevation="514" latitude="0.8825" location="Lita" longLatPrecision="21" longitude="-78.46" pageId="11" pageNumber="12" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D011EC8792C9F784AE8FB54FEBE18EB" box="[229,294,1159,1183]" pageId="11" pageNumber="12" type="male">2 ♂♂</specimenCount>
(
<specimenCount id="9D011EC8792C9F784B36FB54FEFE18EB" box="[315,358,1159,1183]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
in tandem with
<specimenCount id="9D011EC8792C9F78482AFB54FDA418EB" box="[551,572,1159,1182]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
of
<taxonomicName id="4C07AEC2792C9F78486AFB54FC2718EA" authority="Hagen in Selys" authorityName="Hagen in Selys" authorityYear="1865" box="[615,959,1158,1183]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="B9730953792C9F78486AFB54FC9B18EB" box="[615,771,1158,1182]" italics="true" pageId="11" pageNumber="12">Argia oculata</emphasis>
Hagen
<emphasis id="B9730953792C9F78496DFB54FCEE18EB" box="[864,886,1159,1182]" italics="true" pageId="11" pageNumber="12">in</emphasis>
Selys
</taxonomicName>
), same data but, seepage pools along RR tracks,
<quantity id="4CFF78A4792C9F784AE5FB78FEB918B6" box="[232,289,1194,1219]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="km" value="1.0">1 km</quantity>
north west of
<location id="8ED8839A792C9F784BC8FB79FE6A18B6" LSID="urn:lsid:plazi:treatment:03AE6457792C9F7F4A9AFCCCFD5B1A96:8ED8839A792C9F784BC8FB79FE6A18B6" box="[453,498,1194,1219]" country="Ecuador" latitude="0.8825" longLatPrecision="21" longitude="-78.46" name="Lita" pageId="11" pageNumber="12">Lita</location>
{
<geoCoordinate id="EE33B386792C9F78480FFB79FD1D18B7" box="[514,645,1194,1219]" degrees="0" direction="north" minutes="52" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="57" value="0.8825">0°52'57'' N</geoCoordinate>
,
<geoCoordinate id="EE33B386792C9F78489CFB79FCBA18B7" box="[657,802,1194,1219]" degrees="78" direction="west" minutes="27" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="36" value="-78.46">78°27'36'' W</geoCoordinate>
,
<quantity id="4CFF78A4792C9F784920FB79FCE318B6" box="[813,891,1194,1219]" metricMagnitude="2" metricUnit="m" metricValue="5.14" pageId="11" pageNumber="12" unit="m" value="514.0">
<elevation id="002A3272792C9F784920FB79FCE318B6" box="[813,891,1194,1219]" metricMagnitude="2" metricUnit="m" metricValue="5.14" pageId="11" pageNumber="12" unit="m" value="514.0">514 m</elevation>
</quantity>
},
<date id="FFB9F381792C9F78499DFB79FBD518B6" box="[912,1101,1194,1219]" pageId="11" pageNumber="12" value="1997-02-05">
<collectingDate id="EFFD0A69792C9F78499DFB79FBD518B6" box="[912,1101,1194,1219]" pageId="11" pageNumber="12" value="1997-02-05">5 February 1997</collectingDate>
</date>
,
<collectorName id="26F2B097792C9F784E54FB79FA9218B6" box="[1113,1290,1194,1219]" pageId="11" pageNumber="12">Jerrell J. Daigle</collectorName>
leg. [
<collectionCode id="ED164D84792C9F784F44FB78FA0A18B6" box="[1353,1426,1195,1219]" pageId="11" pageNumber="12">RWG</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C792C9F784A9AFB1CFD751892" ID-GBIF-Occurrence="2608962969" box="[151,749,1230,1255]" collectingDate="1997-02-05" collectionCode="RWG" collectorName="Kenneth J. Tennessen" country="Ecuador" elevation="514" latitude="0.8825" location="Lita" longLatPrecision="21" longitude="-78.46" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D011EC8792C9F784A9AFB1CFF5B1893" box="[151,195,1231,1255]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
, same data but
<collectorName id="26F2B097792C9F784B63FB1DFDF91893" box="[366,609,1230,1255]" pageId="11" pageNumber="12">Kenneth J. Tennessen</collectorName>
leg. [
<collectionCode id="ED164D84792C9F784890FB1CFD7E1892" box="[669,742,1231,1255]" pageId="11" pageNumber="12">RWG</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C792C9F7848FAFB1CFF49197F" ID-GBIF-Occurrence="2608962957" collectingDate="1997-02-05" collectionCode="FSCA" collectorName="Kenneth J. Tennessen" country="Ecuador" elevation="514" latitude="0.8825" location="Lita" longLatPrecision="21" longitude="-78.46" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Santo Domingo de los Tsachilas">
<specimenCount id="9D011EC8792C9F7848FAFB1CFCBB1893" box="[759,803,1231,1255]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
, same data but [
<collectionCode id="ED164D84792C9F7849D8FB1CFBBB1892" box="[981,1059,1231,1255]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="11" pageNumber="12" type="Museum">FSCA</collectionCode>
];
<collectingRegion id="49C31BA3792C9F784E38FB1CFF49197F" country="Ecuador" name="Santo Domingo de los Tsachilas" pageId="11" pageNumber="12">Santo Domingo de los Tsáchilas Prov.</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B6FDF1C792C9F784AECFB20FD9E195A" ID-GBIF-Occurrence="2608962930" collectingDate="1975-07-05" collectionCode="RWG" collectorName="A. Langley &amp; J. Cohen" country="Ecuador" elevation="744" latitude="-0.31361112" location="Colorados" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" pageId="11" pageNumber="12" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D011EC8792C9F784AECFB20FEBD197F" box="[225,293,1267,1291]" pageId="11" pageNumber="12" type="male">2 ♂♂</specimenCount>
,
<quantity id="4CFF78A4792C9F784B3FFB20FEE3197E" box="[306,379,1266,1291]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="km" value="19.0">19 km</quantity>
east of
<collectingMunicipality id="6BDC4F3B792C9F784BD5FB20FD13197E" box="[472,651,1266,1291]" pageId="11" pageNumber="12">Santo Domingo</collectingMunicipality>
de los
<location id="8ED8839A792C9F7848ECFB21FCCC197E" LSID="urn:lsid:plazi:treatment:03AE6457792C9F7F4A9AFCCCFD5B1A96:8ED8839A792C9F7848ECFB21FCCC197E" box="[737,852,1266,1291]" country="Ecuador" latitude="-0.31361112" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" name="Colorados" pageId="11" pageNumber="12">Colorados</location>
{
<geoCoordinate id="EE33B386792C9F78496BFB21FC7D197E" box="[870,997,1266,1291]" degrees="0" direction="south" minutes="18" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="49" value="-0.31361112">0°18'49'' S</geoCoordinate>
,
<geoCoordinate id="EE33B386792C9F7849FEFB20FBEF197F" box="[1011,1143,1267,1291]" degrees="79" direction="west" minutes="1" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="44" value="-79.02889">79°1'44'' W</geoCoordinate>
,
<quantity id="4CFF78A4792C9F784E89FB20FB4B197E" box="[1156,1235,1266,1291]" metricMagnitude="2" metricUnit="m" metricValue="7.44" pageId="11" pageNumber="12" unit="m" value="744.0">
<elevation id="002A3272792C9F784E89FB20FB4B197E" box="[1156,1235,1266,1291]" metricMagnitude="2" metricUnit="m" metricValue="7.44" pageId="11" pageNumber="12" unit="m" value="744.0">744 m</elevation>
</quantity>
},
<date id="FFB9F381792C9F784EE7FB21FAEC197E" box="[1258,1396,1266,1291]" pageId="11" pageNumber="12" value="1975-07-05">
<collectingDate id="EFFD0A69792C9F784EE7FB21FAEC197E" box="[1258,1396,1266,1291]" pageId="11" pageNumber="12" value="1975-07-05">5 July 1975</collectingDate>
</date>
,
<collectorName id="26F2B097792C9F784F8CFB20FF6B195A" pageId="11" pageNumber="12">A. Langley</collectorName>
&amp;
<collectorName id="26F2B097792C9F784B1BFAC4FEE0195A" box="[278,376,1302,1327]" pageId="11" pageNumber="12">J. Cohen</collectorName>
leg. [
<collectionCode id="ED164D84792C9F784BBBFAC4FE67195A" box="[438,511,1303,1327]" pageId="11" pageNumber="12">RWG</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C792C9F78481CFAC4FCD2195A" ID-GBIF-Occurrence="2608962911" box="[529,842,1302,1327]" collectingDate="1975-07-05" collectionCode="USNM" collectorName="A. Langley &amp; J. Cohen" country="Ecuador" elevation="744" latitude="-0.31361112" location="Colorados" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D011EC8792C9F78481CFAC4FDA3195B" box="[529,571,1303,1327]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
same data but [
<collectionCode id="ED164D84792C9F7848E4FAC4FCDB195A" box="[745,835,1303,1327]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BB8D541792C9F784ACAFAE8FC851902" blockId="11.[151,1437,942,2014]" pageId="11" pageNumber="12">
A medium-sized largely dark species with male cercus armed with a blunt anteapical ventral tooth (
<figureCitation id="133CC9C4792C9F784F3EFAE9FA0C1926" box="[1331,1428,1338,1363]" captionStart="FIGURES 103–106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103–106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="11" pageNumber="12">Fig. 106</figureCitation>
) and unmodified mesostigmal lobe in the female (
<figureCitation id="133CC9C4792C9F7848B1FA8DFC881902" box="[700,784,1374,1399]" captionStart="FIGURES 69–72" captionStartId="48.[151,264,1557,1579]" captionTargetBox="[264,1313,208,1530]" captionTargetId="figure@48.[264,1323,193,1535]" captionTargetPageId="48" captionText="FIGURES 69–72. Female mesostigmal plates. (a) dorsal view; (b) detail of mesostigmal lobe, dorsal view; (c) frontal (anterior) view; (d) detail of mesostigmal lobe, posterodorsal view (on edge)." figureDoi="http://doi.org/10.5281/zenodo.3770053" httpUri="https://zenodo.org/record/3770053/files/figure.png" pageId="11" pageNumber="12">Fig. 72</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA792C9F7F4ACAFA51FCD7190E" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8BB8D541792C9F784ACAFA51FE571A5E" blockId="11.[151,1437,942,2014]" pageId="11" pageNumber="12">
<emphasis id="B9730953792C9F784ACAFA51FDB919EE" bold="true" box="[199,545,1410,1435]" pageId="11" pageNumber="12">
Description of male
<typeStatus id="54BC6BE3792C9F784BB7FA51FDB919EE" box="[442,545,1410,1435]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
</emphasis>
. Head: labrum, base of mandibles, genae and frons entirely pale (ivory green), anteclypeus brown, postclypeus dark brown with a well-defined medial pale spot, epicranium black with small pale spot anterolateral to lateral ocellus, large violaceous postocular spot laterally confluent with eye, occipital bar black (as in
<figureCitation id="133CC9C4792C9F784AD6FA3DFEB41A72" box="[219,300,1518,1543]" captionStart="FIGURES 9–13" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 9–13. Color pattern of male head, thorax, and S1–4. Head, dorsal view, thorax and S1–4, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
), dorsal surface of postocular lobe black; antennae black, rear of head mostly black with broad lateral margin bordering eye ivory.
</paragraph>
<paragraph id="8BB8D541792C9F784ACAF9E5FB011B76" blockId="11.[151,1437,942,2014]" pageId="11" pageNumber="12">
Prothorax black with following areas pale: obscure pale rim on anterior lobe, dorsolateral spot on middle lobe confluent posterolaterally with pale propleuron, lateral third of hind lobe of prothorax. Pale areas of synthorax purple laterally, olive dorsally, with broad black middorsal stripe about twice as wide as olive antehumeral stripe, the latter slightly narrowing dorsally; broad black parallel humeral stripe (as in
<figureCitation id="133CC9C4792C9F784E20F971FB181ACE" box="[1069,1152,1698,1723]" captionStart="FIGURES 9–13" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 9–13. Color pattern of male head, thorax, and S1–4. Head, dorsal view, thorax and S1–4, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
) extending from base of mesinfraepisternum and connecting below antealar crest with middorsal stripe above and with dorsal portion of obsolete interpleural suture; metapleural stripe narrow; pale colors on side of thorax purple.
</paragraph>
<paragraph id="8BB8D541792C9F784ACAF8DDFC9E1BE3" blockId="11.[151,1437,942,2014]" pageId="11" pageNumber="12">
Wings hyaline with venation black; pterostigma dark brown, surmounting 1 cell in all wings; postnodals Fw 11/12, Hw 10/11; postquadrangular cells Fw 3/3, Hw 3/3; RP
<subScript id="1783D704792C9F784957F892FCFA1B3A" attach="left" box="[858,866,1857,1871]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
at Fw 6/5.5, Hw 4/4. Coxae and trochanters pale except for black on ventral and anterior portions of coxae and dorsum of trochanters; base of metafemora and external surface of tibiae pale, tarsi and armature black.
</paragraph>
<paragraph id="8BB8D541792C9F7F4ACAF872FDC71DFD" blockId="11.[151,1437,942,2014]" lastBlockId="12.[151,1437,151,1763]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Abdomen (as in Figs. 45, 46) mostly black; S1 with a black basal ring, remainder purple; S2 violaceous with a lateral black stripe interrupted medially with latter part extending to apical fourth of segment, broadening dorsally with a narrow dorsal offshoot thus restricting pale dorsum of segment; S3 black with a pale basal ring and a broad parallel pale dorsal stripe extending to and abruptly ending at apical 0.5 of segment, a vestigial obscure pale ventrolateral stripe at along medial half of segment; S4–7 similar to S3 but entirely black dorsally and with medial pale stripe longer and paler thus giving the appearance of a dark, pale, dark pattern when viewed laterally; S8 black with an elongate pale (purple) spot, about twice as high posteriorly as basally and anteriorly almost connecting with incomplete dorsal pale basal ring; S9 black laterally with a large pale spot dorsally, apical fourth black dorsally; S10 black; torus pale, appendages black.</paragraph>
<paragraph id="8BB8D541792B9F7F4ACAFE47FED91EF1" blockId="12.[151,1437,151,1763]" pageId="12" pageNumber="13">
Genital ligula as in
<taxonomicName id="4C07AEC2792B9F7F4BBFFE47FDAD1DD9" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[434,565,404,428]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9730953792B9F7F4BBFFE47FDAD1DD9" box="[434,565,404,428]" italics="true" pageId="12" pageNumber="13">A. acridens</emphasis>
</taxonomicName>
(as in
<figureCitation id="133CC9C4792B9F7F4880FE47FD7B1DD9" box="[653,739,404,429]" captionStart="FIGURES 89–93" captionStartId="52.[151,264,1914,1936]" captionTargetBox="[156,1432,207,1889]" captionTargetId="figure@52.[151,1436,193,1892]" captionTargetPageId="52" captionText="FIGURES 89–93. Genital ligula. (a) lateral view; (b) ectal view; (c) entolateral view." figureDoi="http://doi.org/10.5281/zenodo.3770062" httpUri="https://zenodo.org/record/3770062/files/figure.png" pageId="12" pageNumber="13">Fig. 89</figureCitation>
), lacking microspinulate patch on ental surface proximal to flexure on sclerotized area; distal segment largely membranous armed basally with a curved sclerotized flagellum. Torus small, oval, swollen, occupying outer half of torifer (
<figureCitation id="133CC9C4792B9F7F4943FE0FFC591D80" box="[846,961,476,501]" captionStart="FIGURES 103–106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103–106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="12" pageNumber="13">Fig. 106a</figureCitation>
); area around epiproct and base of same black; cercus (
<figureCitation id="133CC9C4792B9F7F4B35FE2CFE3D1E6D" box="[312,421,511,536]" captionStart="FIGURES 103–106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103–106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="12" pageNumber="13">Fig. 106b</figureCitation>
) slightly surpassing paraproct, slightly longer than wide, swollen basally with medial basal margin linear and carinate narrowing distally and armed with a large ventrally directed subapical tooth whose tip is bluntly rounded (
<figureCitation id="133CC9C4792B9F7F4B54FD94FD901E15" box="[345,520,583,608]" captionStart="FIGURES 103–106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103–106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="12" pageNumber="13">Figs. 106a, c, e</figureCitation>
); paraproct unilobate, almost quadrate in lateral view with a small tuberculate tip (
<figureCitation id="133CC9C4792B9F7F4A92FDBFFEAC1EF1" box="[159,308,620,645]" captionStart="FIGURES 103–106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103–106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="12" pageNumber="13">Figs. 106b, c</figureCitation>
).
</paragraph>
<paragraph id="8BB8D541792B9F7F4ACAFD5CFCAC1EDD" blockId="12.[151,1437,151,1763]" box="[199,820,655,680]" pageId="12" pageNumber="13">Dimensions. Hw 19.5, abdomen 27.2, total length 34.8.</paragraph>
<paragraph id="8BB8D541792B9F7F4ACAFD67FE611E85" blockId="12.[151,1437,151,1763]" pageId="12" pageNumber="13">
<emphasis id="B9730953792B9F7F4ACAFD67FDD01EB8" bold="true" box="[199,584,692,717]" pageId="12" pageNumber="13">
Description of female
<typeStatus id="54BC6BE3792B9F7F4BD0FD67FDD01EB8" box="[477,584,692,717]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
</emphasis>
(
<collectingCountry id="F31095D1792B9F7F4850FD67FD581EB8" box="[605,704,692,717]" name="Ecuador" pageId="12" pageNumber="13">Ecuador</collectingCountry>
:
<collectingRegion id="49C31BA3792B9F7F48DDFD67FC001EB9" box="[720,920,692,717]" country="Ecuador" name="Esmeraldas" pageId="12" pageNumber="13">Esmeraldas Prov.</collectingRegion>
, seeps
<quantity id="4CFF78A4792B9F7F49F8FD67FBD01EB8" box="[1013,1096,692,717]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="13" unit="km" value="1.1">1.1 km</quantity>
northwest of Lita at bridge, collected in tandem with male).
</paragraph>
<paragraph id="8BB8D541792B9F7F4ACAFD2FFDA6182D" blockId="12.[151,1437,151,1763]" pageId="12" pageNumber="13">
Head (
<figureCitation id="133CC9C4792B9F7F4B1CFD2FFECA1F61" box="[273,338,764,789]" captionStart="FIGURES 1–8" captionStartId="39.[151,264,1966,1988]" captionTargetBox="[186,1424,239,1889]" captionTargetId="figure@39.[151,1437,193,1945]" captionTargetPageId="39" captionText="FIGURES 1–8. Color pattern of head. Head, anterior view (1); head posterior view (2–5); labrum, anterior view (6–8)." figureDoi="http://doi.org/10.5281/zenodo.3770041" httpUri="https://zenodo.org/record/3770041/files/figure.png" pageId="12" pageNumber="13">Fig. 1</figureCitation>
), pro- and synthorax and S1, 2 as in male (
<figureCitation id="133CC9C4792B9F7F493EFD2FFC1E1F61" box="[819,902,764,789]" captionStart="FIGURES 23–27" captionStartId="43.[151,264,1912,1934]" captionTargetBox="[165,1412,210,1878]" captionTargetId="figure@43.[150,1436,193,1891]" captionTargetPageId="43" captionText="FIGURES 23–27. Color pattern of female. Head, thorax, and S1–4 (23, 24, 26, 27); pterothorax (25). Head, dorsal view, thorax and S1–4, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770049" httpUri="https://zenodo.org/record/3770049/files/figure.png" pageId="12" pageNumber="13">Fig. 24</figureCitation>
), but pale areas light brown and more extensive and dark humeral stripe parallel, as black as middorsal stripe; S1 as in male; S3 tan dorsally, with a elongate lateral black stripe with apical 0.20 enlarged and sending a dorsal offshoot thus constricting pale dorsal stripe above followed ventrally by an irregular pale lateral stripe and margined ventrally by black; S4 entirely black with following areas pale: a complete irregular pale basal ring connecting with a thin pale middorsal stripe that extends to apical 0.2 of segment, and an elongate pale ventrolateral spot along medial half of segment thus giving the appearance of a dark, pale, dark pattern when viewed laterally; S4–7 similar to S3; S8 black with pale light olive green spot dorsally and laterally, this spot constricted laterally by black and posteriorly by black apical ring; S9 black except for small ill-defined pale lateral spot at apical third; S10 black; ovipositor black with dorsoapical margin brown, cercus black (Fig. 60).
</paragraph>
<paragraph id="8BB8D541792B9F7F4ACAFBB7FBBB18D5" blockId="12.[151,1437,151,1763]" pageId="12" pageNumber="13">
Mesostigmal lobe lacking, area immediately behind obsolete posterior margin of mesostigmal plate with a slight depression (
<figureCitation id="133CC9C4792B9F7F4B6EFB54FE2D18D5" box="[355,437,1159,1184]" captionStart="FIGURES 69–72" captionStartId="48.[151,264,1557,1579]" captionTargetBox="[264,1313,208,1530]" captionTargetId="figure@48.[264,1323,193,1535]" captionTargetPageId="48" captionText="FIGURES 69–72. Female mesostigmal plates. (a) dorsal view; (b) detail of mesostigmal lobe, dorsal view; (c) frontal (anterior) view; (d) detail of mesostigmal lobe, posterodorsal view (on edge)." figureDoi="http://doi.org/10.5281/zenodo.3770053" httpUri="https://zenodo.org/record/3770053/files/figure.png" pageId="12" pageNumber="13">Fig. 72</figureCitation>
) medial arm of mesostigmal plate prominent, glabrous.
</paragraph>
<paragraph id="8BB8D541792B9F7F4ACAFB7FFCD7190E" blockId="12.[151,1437,151,1763]" pageId="12" pageNumber="13">
<emphasis id="B9730953792B9F7F4ACAFB7FFE4018B0" bold="true" box="[199,472,1196,1221]" pageId="12" pageNumber="13">
Variation in
<typeStatus id="54BC6BE3792B9F7F4B6FFB7FFE4018B0" box="[354,472,1196,1221]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
</emphasis>
. Little variation was observed in the
<typeStatus id="54BC6BE3792B9F7F4987FB7EFC7318B0" box="[906,1003,1197,1221]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
series. The pale lateral spot on S
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male can range from moderately large as in
<typeStatus id="54BC6BE3792B9F7F4887FB1CFD74189D" box="[650,748,1231,1256]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(Fig. 46) to very small (Fig. 47). Pterostigma surmounting 1 cell in males (N=9),
<quantity id="4CFF78A4792B9F7F4B8EFB27FE371978" box="[387,431,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="13" unit="in" value="1.0">1 in</quantity>
female (N =1); postnodals: Fw
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males,
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female, Hw
<quantity id="4CFF78A4792B9F7F4EABFB27FA921978" box="[1190,1290,1268,1293]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="12" pageNumber="13" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
males,
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female; postquadrangular cells Fw 3, Hw
<quantity id="4CFF78A4792B9F7F4862FACBFD031945" box="[623,667,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="13" unit="in" value="3.0">3 in</quantity>
males, Fw 3, Hw
<quantity id="4CFF78A4792B9F7F4967FACBFC0E1945" box="[874,918,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="13" unit="in" value="3.0">3 in</quantity>
female; RP
<subScript id="1783D704792B9F7F4E17FAF5FBBA1941" attach="left" box="[1050,1058,1318,1332]" fontSize="6" pageId="12" pageNumber="13">2</subScript>
at Fw 5–7, Hw
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males, Fw 5, Hw
<quantity id="4CFF78A4792B9F7F4AECFAECFE941922" box="[225,268,1342,1367]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="12" pageNumber="13" unit="in" value="4.0">4 in</quantity>
female.
<emphasis id="B9730953792B9F7F4B61FAEDFE6E1922" bold="true" box="[364,502,1342,1367]" pageId="12" pageNumber="13">Dimensions</emphasis>
.
<specimenCount id="9D011EC8792B9F7F480EFA93FD831922" box="[515,539,1344,1367]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
: Hw 19.3 ± 0.5 [18.6–20.2], abdomen 27.5 ± 0.6 [26.4–28.3], total length 35.1 ± 0.8 [33.9–36.5];
<specimenCount id="9D011EC8792B9F7F4B5FFAB0FEF1190F" box="[338,361,1379,1402]" pageId="12" pageNumber="13" type="female">♀</specimenCount>
: Hw 20.5, abdomen 25.2, total length 32.8.
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA792B9F7F4ACAFA55FA8319EA" box="[199,1307,1414,1439]" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BB8D541792B9F7F4ACAFA55FA8319EA" blockId="12.[151,1437,151,1763]" box="[199,1307,1414,1439]" pageId="12" pageNumber="13">
<emphasis id="B9730953792B9F7F4ACAFA55FEA219EA" bold="true" box="[199,314,1414,1439]" pageId="12" pageNumber="13">Diagnosis</emphasis>
. This species is similar to the sympatric
<taxonomicName id="4C07AEC2792B9F7F490FFA54FC1A19EA" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[770,898,1415,1439]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9730953792B9F7F490FFA54FC1A19EA" box="[770,898,1415,1439]" italics="true" pageId="12" pageNumber="13">A. acridens</emphasis>
</taxonomicName>
and is diagnosed under that species.
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA792B9F7F4ACAFA79FBFE1AEE" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BB8D541792B9F7F4ACAFA79FC4A1A02" blockId="12.[151,1437,151,1763]" pageId="12" pageNumber="13">
<emphasis id="B9730953792B9F7F4ACAFA79FEA319B6" bold="true" box="[199,315,1450,1475]" pageId="12" pageNumber="13">Remarks.</emphasis>
A male of
<taxonomicName id="4C07AEC2792B9F7F4BB3FA78FDD419B6" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[446,588,1451,1475]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792B9F7F4BB3FA78FDD419B6" box="[446,588,1451,1475]" italics="true" pageId="12" pageNumber="13">A. cuspidata</emphasis>
</taxonomicName>
from the same locality as the female described above as
<taxonomicName id="4C07AEC2792B9F7F4ED7FA78FAF019B6" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[1242,1384,1451,1475]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792B9F7F4ED7FA78FAF019B6" box="[1242,1384,1451,1475]" italics="true" pageId="12" pageNumber="13">A. cuspidata</emphasis>
</taxonomicName>
was collected in tandem with a female of another species. That pair, collected by J. Daigle, presented a problem as to which of these two females, the one described above, or the one in tandem, represented the true female of
<taxonomicName id="4C07AEC2792B9F7F4F89FA20FE9C1A5A" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792B9F7F4F89FA20FE9C1A5A" italics="true" pageId="12" pageNumber="13">A. cuspidata</emphasis>
</taxonomicName>
. Close examination of the latter female mesostigmal plates and body markings matched those for
<taxonomicName id="4C07AEC2792B9F7F4F89F9C4FF731A26" authorityName="Hagen in Selys" authorityYear="1865" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="B9730953792B9F7F4F89F9C4FF731A26" italics="true" pageId="12" pageNumber="13">A. oculata</emphasis>
</taxonomicName>
and some males of that species were collected at the same locality. We are, therefore, fairly certain that the single female described above truly represents the female of
<taxonomicName id="4C07AEC2792B9F7F4932F98CFC531A02" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[831,971,1631,1655]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792B9F7F4932F98CFC531A02" box="[831,971,1631,1655]" italics="true" pageId="12" pageNumber="13">A. cuspidata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB8D541792B9F7F4ACAF951FBFE1AEE" blockId="12.[151,1437,151,1763]" box="[199,1126,1666,1691]" pageId="12" pageNumber="13">
<emphasis id="B9730953792B9F7F4ACAF951FEBB1AEE" bold="true" box="[199,291,1666,1691]" pageId="12" pageNumber="13">Habitat</emphasis>
. Apparently confined to seepages associated with larger rivers and creeks.
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA792B9F7F4ACAF975FD5B1A96" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BB8D541792B9F7F4ACAF975FD5B1A96" blockId="12.[151,1437,151,1763]" pageId="12" pageNumber="13">
<emphasis id="B9730953792B9F7F4ACAF975FEC11ACA" bold="true" box="[199,345,1702,1727]" pageId="12" pageNumber="13">Distribution</emphasis>
. As far as known endemic to
<collectingCountry id="F31095D1792B9F7F48C1F975FCB31ACA" box="[716,811,1702,1727]" name="Ecuador" pageId="12" pageNumber="13">Ecuador</collectingCountry>
, restricted to
<collectingRegion id="49C31BA3792B9F7F49DCF975FBCA1ACA" box="[977,1106,1702,1727]" country="Ecuador" name="Esmeraldas" pageId="12" pageNumber="13">Esmeraldas</collectingRegion>
and
<collectingRegion id="49C31BA3792B9F7F4E9EF974FF671A96" country="Ecuador" name="Santo Domingo de los Tsachilas" pageId="12" pageNumber="13">Santo Domingo de los Tsáchilas</collectingRegion>
Provinces west of the Andes (
<figureCitation id="133CC9C4792B9F7F4859F919FD2E1A96" box="[596,694,1738,1763]" captionStart="FIGURES 136–137" captionStartId="63.[151,264,1901,1923]" captionTargetBox="[170,1246,208,1864]" captionTargetId="figure@63.[151,1437,193,1880]" captionTargetPageId="63" captionText="FIGURES 136–137. Distribution of new Argia species and closest congeners in Ecuador." figureDoi="http://doi.org/10.5281/zenodo.3770088" httpUri="https://zenodo.org/record/3770088/files/figure.png" pageId="12" pageNumber="13">Fig. 136</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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