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<document id="FCDCDC9EF1B418E89EE6BBFB2DC15909" ID-CLB-Dataset="22649" ID-DOI="10.11646/zootaxa.4470.1.1" ID-GBIF-Dataset="615d5c4a-3b8d-4ba6-a05d-f37e432a862b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3770039" ID-ZooBank="60766083-99E8-48E9-86CE-161A9EEC066B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588000738706" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2018" docId="03AE645779219F7B4A9AFC24FD721EF0" docLanguage="en" docName="zootaxa.4470.1.1.pdf" docOrigin="Zootaxa 4470 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argia acridens Garrison &amp; von Ellenrieder 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="FF971C2F79279F734A0DFFD3FF981C75" masterDocTitle="Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species" masterLastPageNumber="69" masterPageNumber="1" pageNumber="7" updateTime="1698837067205" updateUser="ExternalLinkService">
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<mods:title id="029C8FDA446B29E994E0EDFBE2010BF9">Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species</mods:title>
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<mods:namePart id="B5615AE199CBBB93B00F5756236CF713">Garrison, Rosser W.</mods:namePart>
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<mods:namePart id="18F3FDFC0BA2B68D6630AD7938B2660C">Ellenrieder, Natalia Von</mods:namePart>
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<mods:title id="F8A2ED87D89C606432B059603911E1A7">Zootaxa</mods:title>
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<mods:part id="C5F5E0CBCE642932AA543ACB538BE69A">
<mods:date id="A60E0E1A17768DBA084BD919916E2825">2018</mods:date>
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<mods:number id="FC1F8E64F3324CF3D9656A48723C25A7">2018-09-03</mods:number>
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<treatment id="03AE645779219F7B4A9AFC24FD721EF0" ID-DOI="http://doi.org/10.5281/zenodo.3799117" ID-GBIF-Taxon="163754283" ID-Zenodo-Dep="3799117" LSID="urn:lsid:plazi:treatment:03AE645779219F7B4A9AFC24FD721EF0" httpUri="http://treatment.plazi.org/id/03AE645779219F7B4A9AFC24FD721EF0" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C31D86CA79219F754A9AFC24FD621867" box="[151,762,1015,1042]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BB8D54179219F754A9AFC24FD621867" blockId="6.[151,1436,1015,1111]" box="[151,762,1015,1042]" pageId="6" pageNumber="7">
<heading id="D0F0622D79219F754A9AFC24FD621867" bold="true" box="[151,762,1015,1042]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B973095379219F754A9AFC24FD621867" bold="true" box="[151,762,1015,1042]" pageId="6" pageNumber="7">
<taxonomicName id="4C07AEC279219F754A9AFC24FD331867" ID-CoL="9X22B" authority="Garrison &amp; von Ellenrieder" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[151,683,1015,1042]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="acridens" status="sp. nov.">
<emphasis id="B973095379219F754A9AFC24FED11864" bold="true" box="[151,329,1015,1041]" italics="true" pageId="6" pageNumber="7">Argia acridens</emphasis>
Garrison &amp; von Ellenrieder
</taxonomicName>
,
<taxonomicNameLabel id="A240B42879219F7548B5FC2BFD621867" box="[696,762,1016,1042]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA79219F754A9AFBC9FBE51822" pageId="6" pageNumber="7" type="description">
<paragraph id="8BB8D54179219F754A9AFBC9FBE51822" blockId="6.[151,1436,1015,1111]" pageId="6" pageNumber="7">
<figureCitation id="133CC9C479219F754A9AFBC9FF711846" box="[151,233,1050,1075]" captionStart="FIGURES 913" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 913. Color pattern of male head, thorax, and S14. Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="6" pageNumber="7">Figs. 9</figureCitation>
(head, thorax, S1
<specimenCount id="9D011EC879219F754BC0FBC8FE671846" box="[461,511,1051,1075]" pageId="6" pageNumber="7" type="male">3 ♂</specimenCount>
), 23 (head, thorax, S1
<specimenCount id="9D011EC879219F754916FBC8FCD11846" box="[795,841,1051,1075]" pageId="6" pageNumber="7" type="female">3 ♀</specimenCount>
), 34 (S1
<specimenCount id="9D011EC879219F7549C8FBC8FC6E1846" box="[965,1014,1051,1075]" pageId="6" pageNumber="7" type="male">3 ♂</specimenCount>
), 35 (S7
<specimenCount id="9D011EC879219F754E63FBC8FB361846" box="[1134,1198,1051,1075]" pageId="6" pageNumber="7" type="male">10 ♂</specimenCount>
), 59 (S7
<specimenCount id="9D011EC879219F754F2BFBC8FAFA1846" box="[1318,1378,1051,1075]" pageId="6" pageNumber="7" type="female">10 ♀</specimenCount>
), 69 (mesostigmal plates
<specimenCount id="9D011EC879219F754B70FBECFE081822" box="[381,400,1087,1111]" pageId="6" pageNumber="7" type="female"></specimenCount>
), 89 (genital ligula), 103 (appendages
<specimenCount id="9D011EC879219F754949FBECFCC31822" box="[836,859,1087,1111]" pageId="6" pageNumber="7" type="male"></specimenCount>
), 136 (map), 142 (photo).
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA79219F744A9AFB55FD261A47" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BB8D54179219F754A9AFB55FF7E18B7" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="7">
<emphasis id="B973095379219F754A9AFB55FE8618EA" bold="true" box="[151,286,1158,1183]" pageId="6" pageNumber="7">Etymology.</emphasis>
Named
<taxonomicName id="4C07AEC279219F754B76FB55FE4318EB" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[379,475,1158,1182]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B973095379219F754B76FB55FE4318EB" box="[379,475,1158,1182]" italics="true" pageId="6" pageNumber="7">acridens</emphasis>
</taxonomicName>
(L. adj.) in reference to the large medioventrally directed acuminate tooth on the male cercus.
</paragraph>
<paragraph id="8BB8D54179219F754ACAFB1DFC02197E" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="7">
<emphasis id="B973095379219F754ACAFB1DFE8B1892" bold="true" box="[199,275,1230,1255]" pageId="6" pageNumber="7">Types.</emphasis>
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<typeStatus id="54BC6BE379219F754B10FB1DFE1D1892" box="[285,389,1230,1255]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
<specimenCount id="9D011EC879219F754B82FB1CFE3F1893" box="[399,423,1231,1254]" pageId="6" pageNumber="7" type="male"></specimenCount>
:
<collectingCountry id="F31095D179219F754BBBFB1CFDD81892" box="[438,576,1231,1255]" name="Ecuador" pageId="6" pageNumber="7">ECUADOR</collectingCountry>
,
<collectingRegion id="49C31BA379219F754843FB1DFD731893" box="[590,747,1230,1255]" country="Ecuador" name="Manabi" pageId="6" pageNumber="7">Manabí Prov.</collectingRegion>
,
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west of
<collectingMunicipality id="6BDC4F3B79219F7549A6FB1CFBC61892" box="[939,1118,1230,1255]" pageId="6" pageNumber="7">Santo Domingo</collectingMunicipality>
de los
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,
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,
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},
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,
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<emphasis id="B973095379219F7548B2FB20FD68197E" box="[703,752,1267,1291]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. leg. [
<collectionCode id="ED164D8479219F754938FB20FC17197E" box="[821,911,1267,1291]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7" type="Museum">USNM</collectionCode>
].
</materialsCitation>
</paragraph>
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<emphasis id="B973095379219F754ACAFAC5FDEC195A" bold="true" box="[199,628,1302,1327]" pageId="6" pageNumber="7">
Specimens examined (all
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)
</emphasis>
.
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,
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.
</paragraph>
<paragraph id="8BB8D54179219F754ACAFAE8FA121AE2" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="7">
<materialsCitation id="3B6FDF1C79219F754ACAFAE8FD0D1902" ID-GBIF-Occurrence="2608960776" collectingDate="2015-05-12" collectingDateMax="2015-05-15" collectingDateMin="2015-05-12" collectionCode="RWG" collectorName="Jessica Ware &amp; Melissa Sanchez" country="Ecuador" elevation="470" latitude="0.52027774" location="Canande Reserve" longLatPrecision="21" longitude="-79.08667" pageId="6" pageNumber="7" specimenCount="2" specimenCount-male="2" stateProvince="Esmeraldas">
<collectingCountry id="F31095D179219F754ACAFAE8FEC91926" box="[199,337,1339,1363]" name="Ecuador" pageId="6" pageNumber="7">ECUADOR</collectingCountry>
,
<collectingRegion id="49C31BA379219F754B50FAE9FDB91927" box="[349,545,1338,1363]" country="Ecuador" name="Esmeraldas" pageId="6" pageNumber="7">Esmeraldas Prov.</collectingRegion>
:
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,
<location id="8ED8839A79219F75488DFAE9FCDD1927" LSID="urn:lsid:plazi:treatment:03AE645779219F7B4A9AFC24FD721EF0:8ED8839A79219F75488DFAE9FCDD1927" box="[640,837,1338,1363]" country="Ecuador" latitude="0.52027774" longLatPrecision="21" longitude="-79.08667" name="Canande Reserve" pageId="6" pageNumber="7" stateProvince="Esmeraldas">Canandé Reserve</location>
(
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,
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,
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),
<date id="FFB9F38179219F754EDEFAE9FA001926" box="[1235,1432,1338,1363]" pageId="6" pageNumber="7" value="2015-05-12" valueMax="2015-05-15" valueMin="2015-05-12">
<collectingDate id="EFFD0A6979219F754EDEFAE9FA001926" box="[1235,1432,1338,1363]" pageId="6" pageNumber="7" value="2015-05-12" valueMax="2015-05-15" valueMin="2015-05-12">1215 May 2015</collectingDate>
</date>
,
<collectorName id="26F2B09779219F754A9AFA8DFEBF1903" box="[151,295,1374,1399]" pageId="6" pageNumber="7">Jessica Ware</collectorName>
&amp;
<collectorName id="26F2B09779219F754B46FA8DFD9E1902" box="[331,518,1374,1399]" pageId="6" pageNumber="7">Melissa Sanchez</collectorName>
leg. [
<collectionCode id="ED164D8479219F754848FA8CFD161902" box="[581,654,1375,1399]" pageId="6" pageNumber="7">RWG</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C79219F7548ADFA8DFBD919EE" ID-GBIF-Occurrence="2608960750" collectingDate="1975-05-07" collectionCode="RWG" collectorName="Paul J. Spangler" country="Ecuador" elevation="260" latitude="-0.3413889" location="Colorados" longLatPrecision="21" longitude="-79.77028" municipality="Santo Domingo" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Manabi">
<collectingRegion id="49C31BA379219F7548ADFA8DFCA01903" box="[672,824,1374,1399]" country="Ecuador" name="Manabi" pageId="6" pageNumber="7">Manabí Prov.</collectingRegion>
:
<specimenCount id="9D011EC879219F75494BFA8CFCEB1903" box="[838,883,1375,1399]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
,
<quantity id="4CFF78A479219F754972FA8CFC5E1902" box="[895,966,1374,1399]" metricMagnitude="4" metricUnit="m" metricValue="7.9" pageId="6" pageNumber="7" unit="km" value="79.0">79 km</quantity>
west of
<collectingMunicipality id="6BDC4F3B79219F754E2AFA8CFB401902" box="[1063,1240,1374,1399]" pageId="6" pageNumber="7">Santo Domingo</collectingMunicipality>
de los
<location id="8ED8839A79219F754F24FA8DFA041902" LSID="urn:lsid:plazi:treatment:03AE645779219F7B4A9AFC24FD721EF0:8ED8839A79219F754F24FA8DFA041902" box="[1321,1436,1374,1399]" country="Ecuador" latitude="-0.3413889" longLatPrecision="21" longitude="-79.77028" municipality="Santo Domingo" name="Colorados" pageId="6" pageNumber="7" stateProvince="Manabi">Colorados</location>
{
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,
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,
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</quantity>
},
<date id="FFB9F38179219F754825FA50FD2B19EE" box="[552,691,1410,1435]" pageId="6" pageNumber="7" value="1975-05-07">
<collectingDate id="EFFD0A6979219F754825FA50FD2B19EE" box="[552,691,1410,1435]" pageId="6" pageNumber="7" value="1975-05-07">7 May 1975</collectingDate>
</date>
,
<collectorName id="26F2B09779219F7548B3FA51FCEC19EE" box="[702,884,1410,1435]" pageId="6" pageNumber="7">Paul J. Spangler</collectorName>
<emphasis id="B973095379219F754976FA50FC2C19EE" box="[891,948,1411,1435]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
leg. [
<collectionCode id="ED164D8479219F7549FFFA50FBA319EE" box="[1010,1083,1411,1435]" pageId="6" pageNumber="7">RWG</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C79219F754E40FA50FA1B19CA" ID-GBIF-Occurrence="2608960790" collectingDate="2010-04-14" collectionCode="RWG" collectorName="Jerrell J. Daigle" country="Ecuador" elevation="420" latitude="-1.0644444" location="Solano" longLatPrecision="21" longitude="-79.976944" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Manabi Prov.">
<specimenCount id="9D011EC879219F754E40FA50FBE119EF" box="[1101,1145,1411,1435]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
, highway E40, 5 km west of
<location id="8ED8839A79219F754AB8FA75FE9E19CA" LSID="urn:lsid:plazi:treatment:03AE645779219F7B4A9AFC24FD721EF0:8ED8839A79219F754AB8FA75FE9E19CA" box="[181,262,1446,1471]" country="Ecuador" latitude="-1.0644444" longLatPrecision="21" longitude="-79.976944" name="Solano" pageId="6" pageNumber="7" stateProvince="Manabi Prov.">Solano</location>
, stream/waterfall, noon {
<geoCoordinate id="EE33B38679219F754813FA75FD1519CA" box="[542,653,1446,1471]" degrees="1" direction="south" minutes="3" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="52" value="-1.0644444">1°3'52'' S</geoCoordinate>
,
<geoCoordinate id="EE33B38679219F754897FA75FCB219CB" box="[666,810,1446,1471]" degrees="79" direction="west" minutes="58" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="37" value="-79.976944">79°58'37'' W</geoCoordinate>
,
<quantity id="4CFF78A479219F754938FA74FC1B19CA" box="[821,899,1446,1471]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="7" unit="m" value="420.0">
<elevation id="002A327279219F754938FA74FC1B19CA" box="[821,899,1446,1471]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="7" unit="m" value="420.0">420 m</elevation>
</quantity>
},
<date id="FFB9F38179219F754995FA74FBA119CA" box="[920,1081,1446,1471]" pageId="6" pageNumber="7" value="2010-04-14">
<collectingDate id="EFFD0A6979219F754995FA74FBA119CA" box="[920,1081,1446,1471]" pageId="6" pageNumber="7" value="2010-04-14">14 April 2010</collectingDate>
</date>
,
<collectorName id="26F2B09779219F754E49FA75FB6D19CA" box="[1092,1269,1446,1471]" pageId="6" pageNumber="7">Jerrell J. Daigle</collectorName>
leg. [
<collectionCode id="ED164D8479219F754F3EFA74FAE419CA" box="[1331,1404,1447,1471]" pageId="6" pageNumber="7">RWG</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C79219F754F83FA74FCBA1997" ID-GBIF-Occurrence="2608962810" collectingDate="2010-04-14" collectionCode="JJ" collectorName="Jim Johnson" country="Ecuador" elevation="420" latitude="-1.0644444" location="Solano" longLatPrecision="21" longitude="-79.976944" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Manabi Prov.">
<specimenCount id="9D011EC879219F754F83FA74FF341997" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
same data but trickle, sheet flow,
<collectorName id="26F2B09779219F75483CFA19FD251996" box="[561,701,1482,1507]" pageId="6" pageNumber="7">Jim Johnson</collectorName>
leg. [
<collectionCode id="ED164D8479219F7548F3FA18FC811997" box="[766,793,1483,1506]" pageId="6" pageNumber="7">JJ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C79219F754922FA18FAD61A72" ID-GBIF-Occurrence="2608960746" collectingDate="2010-03-04" collectionCode="FSCA" collectorName="Kenneth J. Tennessen" country="Ecuador" elevation="350" latitude="-1.0644444" location="Solano" longLatPrecision="21" longitude="-79.976944" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Manabi Prov.">
<specimenCount id="9D011EC879219F754922FA18FCC21997" box="[815,858,1483,1507]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
same data but highway E40, 5 km west of
<location id="8ED8839A79219F754F4AFA19FA001996" LSID="urn:lsid:plazi:treatment:03AE645779219F7B4A9AFC24FD721EF0:8ED8839A79219F754F4AFA19FA001996" box="[1351,1432,1482,1507]" country="Ecuador" latitude="-1.0644444" longLatPrecision="21" longitude="-79.976944" name="Solano" pageId="6" pageNumber="7" stateProvince="Manabi Prov.">Solano</location>
, trickle, sheet flow {
<geoCoordinate id="EE33B38679219F754B77FA3DFE741A72" box="[378,492,1518,1543]" degrees="1" direction="south" minutes="3" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="52" value="-1.0644444">1°3'52'' S</geoCoordinate>
,
<geoCoordinate id="EE33B38679219F754BF1FA3DFD171A73" box="[508,655,1518,1543]" degrees="79" direction="west" minutes="58" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="37" value="-79.976944">79°58'37'' W</geoCoordinate>
,
<quantity id="4CFF78A479219F754890FA3DFD751A72" box="[669,749,1518,1543]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="m" value="350.0">
<elevation id="002A327279219F754890FA3DFD751A72" box="[669,749,1518,1543]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="m" value="350.0">350 m</elevation>
</quantity>
},
<date id="FFB9F38179219F75490BFA3CFC341A72" box="[774,940,1518,1543]" pageId="6" pageNumber="7" value="2010-03-04">
<collectingDate id="EFFD0A6979219F75490BFA3CFC341A72" box="[774,940,1518,1543]" pageId="6" pageNumber="7" value="2010-03-04">4 March 2010</collectingDate>
</date>
,
<collectorName id="26F2B09779219F7549B6FA3DFB2C1A73" box="[955,1204,1518,1543]" pageId="6" pageNumber="7">Kenneth J. Tennessen</collectorName>
leg. [
<collectionCode id="ED164D8479219F754EF4FA3CFADF1A72" box="[1273,1351,1519,1543]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="7" type="Museum">FSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C79219F754F51FA3CFC811A3A" ID-GBIF-Occurrence="2608960795" collectingDate="1975-07-05" collectionCode="RWG" collectorName="A. Langley &amp; J. Cohen" country="Ecuador" elevation="744" latitude="-0.31361112" location="Colorados" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" pageId="6" pageNumber="7" specimenCount="2" specimenCount-male="2" stateProvince="Santo Domingo de los Tsachilas">
<collectingRegion id="49C31BA379219F754F51FA3CFE671A5F" country="Ecuador" name="Santo Domingo de los Tsachilas" pageId="6" pageNumber="7">Santo Domingo de los Tsáchilas Prov.</collectingRegion>
:
<specimenCount id="9D011EC879219F754803F9C0FDC81A5F" box="[526,592,1555,1579]" pageId="6" pageNumber="7" type="male">2 ♂♂</specimenCount>
,
<quantity id="4CFF78A479219F754850F9C0FD3D1A5E" box="[605,677,1554,1579]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="7" unit="km" value="19.0">19 km</quantity>
east of
<collectingMunicipality id="6BDC4F3B79219F75490DF9C0FC2A1A5E" box="[768,946,1554,1579]" pageId="6" pageNumber="7">Santo Domingo</collectingMunicipality>
de los
<location id="8ED8839A79219F754E08F9C1FBE01A5E" LSID="urn:lsid:plazi:treatment:03AE645779219F7B4A9AFC24FD721EF0:8ED8839A79219F754E08F9C1FBE01A5E" box="[1029,1144,1554,1579]" country="Ecuador" latitude="-0.31361112" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" name="Colorados" pageId="6" pageNumber="7" stateProvince="Santo Domingo de los Tsachilas">Colorados</location>
{
<geoCoordinate id="EE33B38679219F754E84F9C1FA9F1A5E" box="[1161,1287,1554,1579]" degrees="0" direction="south" minutes="18" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="49" value="-0.31361112">0°18'49'' S</geoCoordinate>
,
<geoCoordinate id="EE33B38679219F754F18F9C0FA001A5F" box="[1301,1432,1555,1579]" degrees="79" direction="west" minutes="1" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="44" value="-79.02889">79°1'44'' W</geoCoordinate>
,
<quantity id="4CFF78A479219F754A9AF9E4FF7C1A3A" box="[151,228,1590,1615]" metricMagnitude="2" metricUnit="m" metricValue="7.44" pageId="6" pageNumber="7" unit="m" value="744.0">
<elevation id="002A327279219F754A9AF9E4FF7C1A3A" box="[151,228,1590,1615]" metricMagnitude="2" metricUnit="m" metricValue="7.44" pageId="6" pageNumber="7" unit="m" value="744.0">744 m</elevation>
</quantity>
},
<date id="FFB9F38179219F754AF4F9E5FEE61A3A" box="[249,382,1590,1615]" pageId="6" pageNumber="7" value="1975-07-05">
<collectingDate id="EFFD0A6979219F754AF4F9E5FEE61A3A" box="[249,382,1590,1615]" pageId="6" pageNumber="7" value="1975-07-05">5 July 1975</collectingDate>
</date>
,
<collectorName id="26F2B09779219F754B84F9E4FD9F1A3A" box="[393,519,1590,1615]" pageId="6" pageNumber="7">A. Langley</collectorName>
&amp;
<collectorName id="26F2B09779219F754827F9E4FD141A3A" box="[554,652,1590,1615]" pageId="6" pageNumber="7">J. Cohen</collectorName>
leg. [
<collectionCode id="ED164D8479219F7548C4F9E4FC8A1A3A" box="[713,786,1591,1615]" pageId="6" pageNumber="7">RWG</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C79219F754929F9E4FB3A1A3A" ID-GBIF-Occurrence="2608960763" box="[804,1186,1590,1615]" collectingDate="1975-07-05" collectionCode="USNM" collectorName="A. Langley &amp; J. Cohen" country="Ecuador" elevation="744" latitude="-0.31361112" location="Colorados" longLatPrecision="21" longitude="-79.02889" municipality="Santo Domingo" pageId="6" pageNumber="7" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Santo Domingo de los Tsachilas Prov.">
<specimenCount id="9D011EC879219F754929F9E4FCFC1A3B" box="[804,868,1591,1615]" pageId="6" pageNumber="7" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D011EC879219F75497CF9E4FC0B1A3A" box="[881,915,1591,1615]" pageId="6" pageNumber="7" type="female">1♀</specimenCount>
same data but [
<collectionCode id="ED164D8479219F754E4CF9E4FB031A3A" box="[1089,1179,1591,1615]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7" type="Museum">USNM</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B6FDF1C79219F754EA0F9E4FA1E1AE2" ID-GBIF-Occurrence="2608962801" collectingDate="1988-10-06" collectionCode="RWG" collectorName="Sidney W. Dunkle" country="Ecuador" county="Tinalandia" elevation="2000" latitude="-0.28388888" location="Colorados" longLatPrecision="21" longitude="-79.05" municipality="Santo Domingo" pageId="6" pageNumber="7" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Santo Domingo de los Tsachilas Prov.">
<specimenCount id="9D011EC879219F754EA0F9E4FB761A3B" box="[1197,1262,1591,1615]" pageId="6" pageNumber="7" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D011EC879219F754EF7F9E4FAA01A3B" box="[1274,1336,1591,1615]" pageId="6" pageNumber="7" type="female">2 ♀♀</specimenCount>
[
<specimenCount id="9D011EC879219F754F45F9E4FAF41A3A" box="[1352,1388,1591,1615]" pageId="6" pageNumber="7" type="male">1♂</specimenCount>
,
<specimenCount id="9D011EC879219F754F74F9E4FA041A3A" box="[1401,1436,1591,1615]" pageId="6" pageNumber="7" type="female">1♀</specimenCount>
in copula],
<collectingCounty id="62D9ADCD79219F754B1BF989FE081A06" box="[278,400,1626,1651]" pageId="6" pageNumber="7">Tinalandia</collectingCounty>
,
<quantity id="4CFF78A479219F754B91F988FE7B1A06" box="[412,483,1626,1651]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="km" value="12.0">12 km</quantity>
east of
<collectingMunicipality id="6BDC4F3B79219F754831F988FD751A06" box="[572,749,1626,1651]" pageId="6" pageNumber="7">Santo Domingo</collectingMunicipality>
de los
<location id="8ED8839A79219F754932F989FC2D1A06" LSID="urn:lsid:plazi:treatment:03AE645779219F7B4A9AFC24FD721EF0:8ED8839A79219F754932F989FC2D1A06" box="[831,949,1626,1651]" country="Ecuador" county="Tinalandia" latitude="-0.28388888" longLatPrecision="21" longitude="-79.05" municipality="Santo Domingo" name="Colorados" pageId="6" pageNumber="7" stateProvince="Santo Domingo de los Tsachilas Prov.">Colorados</location>
, stream and ditch near golf course, shallow water flowing over a rock face {
<geoCoordinate id="EE33B38679219F754BF3F9ADFDF41AE2" box="[510,620,1662,1687]" degrees="0" direction="south" minutes="17" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="2" value="-0.28388888">0°17'2'' S</geoCoordinate>
,
<geoCoordinate id="EE33B38679219F754874F9ACFD751AE3" box="[633,749,1663,1687]" degrees="79" direction="west" minutes="3" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="0" value="-79.05">79°3'0'' W</geoCoordinate>
,
<quantity id="4CFF78A479219F7548F5F9ACFCC21AE2" box="[760,858,1662,1687]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="2000.0">
<elevation id="002A327279219F7548F5F9ACFCC21AE2" box="[760,858,1662,1687]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="2000.0">2,000 m</elevation>
</quantity>
},
<date id="FFB9F38179219F754962F9ADFBB91AE2" box="[879,1057,1662,1687]" pageId="6" pageNumber="7" value="1988-10-06">
<collectingDate id="EFFD0A6979219F754962F9ADFBB91AE2" box="[879,1057,1662,1687]" pageId="6" pageNumber="7" value="1988-10-06">6 October 1988</collectingDate>
</date>
,
<collectorName id="26F2B09779219F754E21F9ADFB601AE2" box="[1068,1272,1662,1687]" pageId="6" pageNumber="7">Sidney W. Dunkle</collectorName>
leg. [
<collectionCode id="ED164D8479219F754F3AF9ACFA181AE2" box="[1335,1408,1663,1687]" pageId="6" pageNumber="7">RWG</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BB8D54179219F754ACAF970FCB71AAA" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="7">
A medium-sized largely dark species with male cercus armed with a sharp medioventral tooth (
<figureCitation id="133CC9C479219F754F0EF971FAFB1ACE" box="[1283,1379,1698,1723]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="6" pageNumber="7">Fig. 103</figureCitation>
) and poorly developed mesostigmal lobe in the female (
<figureCitation id="133CC9C479219F7548C3F915FCBB1AAA" box="[718,803,1734,1759]" captionStart="FIGURES 6972" captionStartId="48.[151,264,1557,1579]" captionTargetBox="[264,1313,208,1530]" captionTargetId="figure@48.[264,1323,193,1535]" captionTargetPageId="48" captionText="FIGURES 6972. Female mesostigmal plates. (a) dorsal view; (b) detail of mesostigmal lobe, dorsal view; (c) frontal (anterior) view; (d) detail of mesostigmal lobe, posterodorsal view (on edge)." figureDoi="http://doi.org/10.5281/zenodo.3770053" httpUri="https://zenodo.org/record/3770053/files/figure.png" pageId="6" pageNumber="7">Fig. 69</figureCitation>
).
</paragraph>
<paragraph id="8BB8D54179219F754ACAF939FC151BE6" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="7">
Description of male
<typeStatus id="54BC6BE379219F754BBFF939FD801B76" box="[434,536,1770,1795]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
. Head: labrum, base of mandibles, genae and frons entirely pale (ivory green), anteclypeus brown, postclypeus pale with a well-defined midlateral spot on each side, epicranium black with small pale spot anterolateral to lateral ocellus, large violaceous postocular spot laterally confluent with eye, occipital bar black (as in
<figureCitation id="133CC9C479219F754B28F885FEF21B1A" box="[293,362,1878,1903]" captionStart="FIGURES 913" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 913. Color pattern of male head, thorax, and S14. Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
), dorsal surface of postocular lobe black; antennae black, rear of head black above narrowing ventrally toward occipital foramen, remainder of rear of head ivory.
</paragraph>
<paragraph id="8BB8D54179219F744ACAF84DFC771D35" blockId="6.[151,1437,1158,2011]" lastBlockId="7.[151,1437,151,2018]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Prothorax black with following areas pale: diffuse rim on anterior lobe, dorsolateral spot on middle lobe confluent posterolaterally with pale propleuron, lateral third of hind lobe of prothorax. Pale areas of synthorax purple laterally, olive dorsally, with broad black middorsal stripe about twice as wide as olive antehumeral stripe, the latter slightly narrowing dorsally; broad black parallel-sided humeral stripe along humeral suture with posterior half merging into lighter shades of brown (as in
<figureCitation id="133CC9C479209F7448A3FF0CFD691C8D" box="[686,753,223,248]" captionStart="FIGURES 913" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 913. Color pattern of male head, thorax, and S14. Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
), extending from base of mesinfraepisternum and connecting below antealar crest with middorsal stripe above and with dorsal portion of obsolete interpleural suture; metapleural stripe narrow; pale colors on side of thorax purple (as in
<figureCitation id="133CC9C479209F744990FEF4FC791D35" box="[925,993,295,320]" captionStart="FIGURES 913" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 913. Color pattern of male head, thorax, and S14. Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BB8D54179209F744ACAFE9FFD3C1DA6" blockId="7.[151,1437,151,2018]" pageId="7" pageNumber="8">
Wings hyaline with venation black; pterostigma dark brown, surmounting 1 cell in all wings; postnodals Fw 11/11, Hw 9/9; postquadrangular cells Fw 3/3, Hw 3/3; RP
<subScript id="1783D70479209F744923FEADFCAE1DF9" attach="left" box="[814,822,382,396]" fontSize="6" pageId="7" pageNumber="8">2</subScript>
at Fw 5/5, Hw 4/4. Coxae and trochanters pale except for black on ventral and anterior portions of coxae and dorsum of trochanters; base of metafemora and external surface of tibiae pale, tarsi and armature black.
</paragraph>
<paragraph id="8BB8D54179209F744ACAFE0DFB0A1EDE" blockId="7.[151,1437,151,2018]" pageId="7" pageNumber="8">
Abdomen (as in
<figureCitation id="133CC9C479209F744B72FE0DFE551D82" box="[383,461,478,503]" captionStart="FIGURES 913" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 913. Color pattern of male head, thorax, and S14. Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="7" pageNumber="8">Figs. 9</figureCitation>
, 34, 35) mostly black; S1 with a black basal ring, remainder purple; S2 violaceous with a black stripe laterally extending to apical fourth of segment, broadening dorsally with a narrow dorsal offshoot thus restricting pale dorsum of segment; S3 black with a pale basal ring and a narrowing pale dorsal stripe ending at apical half of segment, a diffuse pale ventrolateral stripe at along medial half of segment thus giving the appearance of a dark, pale, dark pattern when viewed laterally; S46 similar to S3; S7 black with small pale basal spot; S89 pale (purple) dorsolaterally, black ventrolaterally; S10 black; torus pale, appendages black.
</paragraph>
<paragraph id="8BB8D54179209F744ACAFD65FBF91E86" blockId="7.[151,1437,151,2018]" pageId="7" pageNumber="8">
Genital ligula (
<figureCitation id="133CC9C479209F744B7FFD65FE5C1EBA" box="[370,452,694,719]" captionStart="FIGURES 8993" captionStartId="52.[151,264,1914,1936]" captionTargetBox="[156,1432,207,1889]" captionTargetId="figure@52.[151,1436,193,1892]" captionTargetPageId="52" captionText="FIGURES 8993. Genital ligula. (a) lateral view; (b) ectal view; (c) entolateral view." figureDoi="http://doi.org/10.5281/zenodo.3770062" httpUri="https://zenodo.org/record/3770062/files/figure.png" pageId="7" pageNumber="8">Fig. 89</figureCitation>
) lacking microspinulate patch on ental surface proximal to flexure on sclerotized area; distal segment largely membranous armed basally with a curved sclerotized flagellum.
</paragraph>
<paragraph id="8BB8D54179209F744ACAFCD3FAF51FD2" blockId="7.[151,1437,151,2018]" pageId="7" pageNumber="8">
Torus transversely oval, swollen, occupying entire ventral margin of torifer and partially overlapping bilobed epiproct (
<figureCitation id="133CC9C479209F744B08FCF1FEE21F4E" box="[261,378,802,827]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Fig. 103b</figureCitation>
); area around epiproct and base of same black; cercus (
<figureCitation id="133CC9C479209F744E07FCF1FBF31F4E" box="[1034,1131,802,827]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Fig. 103</figureCitation>
ae) subequal in length to paraproct, longer than wide, swollen basally with medial basal margin strongly convex but narrowing distally and with a strongly acuminate mediolaterally directed tooth; mediodorsal surface of cercus depressed, slightly concave near basal lobe; paraproct (
<figureCitation id="133CC9C479209F744BCAFC5DFDD61FD2" box="[455,590,910,935]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Fig. 103d, e</figureCitation>
) unilobate, almost quadrate in lateral view with a small tuberculate tip.
</paragraph>
<paragraph id="8BB8D54179209F744ACAFC61FCAC1FBE" blockId="7.[151,1437,151,2018]" box="[199,820,946,971]" pageId="7" pageNumber="8">Dimensions. Hw 19.5, abdomen 27.2, total length 34.8.</paragraph>
<paragraph id="8BB8D54179209F744ACAFC05FC821946" blockId="7.[151,1437,151,2018]" pageId="7" pageNumber="8">
Description of female
<typeStatus id="54BC6BE379209F744BCDFC0BFDB91F85" box="[448,545,984,1008]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
(
<collectingCountry id="F31095D179209F744822FC05FD0A1F9A" box="[559,658,982,1007]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
:
<collectingRegion id="49C31BA379209F744891FC04FBDE1F9A" box="[668,1094,982,1007]" country="Ecuador" name="Santo Domingo de los Tsachilas" pageId="7" pageNumber="8">Santo Domingo de los Tsáchilas Prov.</collectingRegion>
,
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east of Santo Domingo de los Colorados, streams south of river): Head, pro- and synthorax and S12 as in male (
<figureCitation id="133CC9C479209F744E98FC29FB721866" box="[1173,1258,1018,1043]" captionStart="FIGURES 2327" captionStartId="43.[151,264,1912,1934]" captionTargetBox="[165,1412,210,1878]" captionTargetId="figure@43.[150,1436,193,1891]" captionTargetPageId="43" captionText="FIGURES 2327. Color pattern of female. Head, thorax, and S14 (23, 24, 26, 27); pterothorax (25). Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770049" httpUri="https://zenodo.org/record/3770049/files/figure.png" pageId="7" pageNumber="8">Fig. 23</figureCitation>
), but pale areas more extensive and dark humeral stripe brown, not as black as middorsal stripe and blue gray; S3 black with an incomplete irregular pale basal ring not connected dorsally, a diffuse pale ventrolateral stripe at along medial half of segment thus giving the appearance of a dark, pale, dark pattern when viewed laterally; S45 similar to S3; S7 black with narrow pale basal ring connected to pale lateral stripe ending at apical fourth; S8 black laterally with a narrow basal ring and broader apical ring surrounding pale green dorsolateral spot; S9 black except for small pale dorsolateral spot at apical third; S10 black; ovipositor black with dorsal margin slightly pale, cercus black (Fig. 59). Mesostigmal lobe (as in
<figureCitation id="133CC9C479209F744BA3FB25FD98197A" box="[430,512,1270,1295]" captionStart="FIGURES 6972" captionStartId="48.[151,264,1557,1579]" captionTargetBox="[264,1313,208,1530]" captionTargetId="figure@48.[264,1323,193,1535]" captionTargetPageId="48" captionText="FIGURES 6972. Female mesostigmal plates. (a) dorsal view; (b) detail of mesostigmal lobe, dorsal view; (c) frontal (anterior) view; (d) detail of mesostigmal lobe, posterodorsal view (on edge)." figureDoi="http://doi.org/10.5281/zenodo.3770053" httpUri="https://zenodo.org/record/3770053/files/figure.png" pageId="7" pageNumber="8">Fig. 69</figureCitation>
) vestigial with posteromesal margin forming an elongate dorsoanteriorly directed glabrous tubercle slightly elevated above mesepisternum.
</paragraph>
<paragraph id="8BB8D54179209F744ACAFAEDFD261A47" blockId="7.[151,1437,151,2018]" pageId="7" pageNumber="8">
Variation in
<typeStatus id="54BC6BE379209F744B5BFA93FE5D192D" box="[342,453,1344,1368]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
. Little variation was observed in the
<typeStatus id="54BC6BE379209F744979FA93FC4D192D" box="[884,981,1344,1368]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
series. The humeral stripe can be paler than the one figured for the male and old males have a dusting of pruinosity on sides of thorax. Pterostigma surmounting 1 cell in males (N = 9),
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females (N = 2); postnodals: Fw
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males,
<quantity id="4CFF78A479209F744EDEFA54FAAE19EA" box="[1235,1334,1414,1439]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.302" metricValueMin="3.048" pageId="7" pageNumber="8" unit="in" value="12.5" valueMax="13.0" valueMin="12.0">1213 in</quantity>
females, Hw
<quantity id="4CFF78A479209F744AC5FA78FE8719B6" box="[200,287,1450,1475]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="7" pageNumber="8" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">912 in</quantity>
males,
<quantity id="4CFF78A479209F744B75FA78FE4419B6" box="[376,476,1450,1475]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="7" pageNumber="8" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">1011 in</quantity>
females; postquadrangular cells Fw 23, Hw
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males, Fw 3, Hw
<quantity id="4CFF78A479209F744F05FA78FAAD19B6" box="[1288,1333,1450,1475]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="8" unit="in" value="3.0">3 in</quantity>
females; RP
<subScript id="1783D70479209F744AB4FA0DFF591999" attach="left" box="[185,193,1502,1516]" fontSize="6" pageId="7" pageNumber="8">2</subScript>
at Fw 57, Hw
<quantity id="4CFF78A479209F744B79FA1DFE231992" box="[372,443,1486,1511]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="7" pageNumber="8" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">45 in</quantity>
males, Fw 6, Hw
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females. Dimensions.
<specimenCount id="9D011EC879209F7449C1FA03FC7C1992" box="[972,996,1488,1511]" pageId="7" pageNumber="8" type="male"></specimenCount>
: Hw 21.1 ± 0.06 [19.921.9], abdomen 28.8 ± 0.09 [19.929.9], total length 36.9 ± 1.1 [35.238.7];
<specimenCount id="9D011EC879209F744944FA25FCF81A78" box="[841,864,1526,1549]" pageId="7" pageNumber="8" type="female"></specimenCount>
: Hw 21.3 ± 1.0 [20.722.5], abdomen 25.0 ± 0.03 [24.725.2], total length 32.8 ± 0.05 [32.333.2].
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA79209F7B4ACAF9EEFB911E48" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BB8D54179209F7B4ACAF9EEFAAA1CA1" blockId="7.[151,1437,151,2018]" lastBlockId="8.[151,1437,151,645]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B973095379209F744ACAF9EEFED91A23" bold="true" box="[199,321,1597,1622]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
This species is superficially similar to the sympatric
<taxonomicName id="4C07AEC279209F744994F9EDFBBD1A20" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[921,1061,1597,1621]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B973095379209F744994F9EDFBBD1A20" box="[921,1061,1597,1621]" italics="true" pageId="7" pageNumber="8">A. cuspidata</emphasis>
</taxonomicName>
but differs by morphology of the caudal appendages, thoracic and abdominal markings in the male, and morphology of the mesostigmal lamina in the female. Male cercus of
<taxonomicName id="4C07AEC279209F744BD6F955FDC51AE8" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[475,605,1669,1693]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B973095379209F744BD6F955FDC51AE8" box="[475,605,1669,1693]" italics="true" pageId="7" pageNumber="8">A. acridens</emphasis>
</taxonomicName>
is broadly convex at basal half (
<figureCitation id="133CC9C479209F7449E9F956FB031AEB" box="[996,1179,1669,1694]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Figs. 103a, e, c</figureCitation>
) and terminates in an acuminate ventromesally directed tooth (
<figureCitation id="133CC9C479209F744869F97AFC951AB7" box="[612,781,1705,1730]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Fig. 103b, c, d</figureCitation>
). Male cercus in
<taxonomicName id="4C07AEC279209F7449DDF979FBC51AB4" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[976,1117,1705,1729]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B973095379209F7449DDF979FBC51AB4" box="[976,1117,1705,1729]" italics="true" pageId="7" pageNumber="8">A. cuspidata</emphasis>
</taxonomicName>
has a largely linear carinate ridge basomedially (
<figureCitation id="133CC9C479209F744B8FF91EFD801A93" box="[386,536,1741,1766]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Figs. 106a, c</figureCitation>
) and the terminal tooth is subapical, broadly rounded apically and is directed ventrally (
<figureCitation id="133CC9C479209F744B07F922FE361B7F" box="[266,430,1777,1802]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Fig. 106b, c, e</figureCitation>
). The torus in
<taxonomicName id="4C07AEC279209F74485CF921FD481B7C" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[593,720,1777,1801]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B973095379209F74485CF921FD481B7C" box="[593,720,1777,1801]" italics="true" pageId="7" pageNumber="8">A. acridens</emphasis>
</taxonomicName>
is elongate occupying the entire ventral margin (
<figureCitation id="133CC9C479209F744EFBF922FAFC1B7F" box="[1270,1380,1777,1802]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Fig. 103a</figureCitation>
), not confined to the apicoventral margin as in
<taxonomicName id="4C07AEC279209F744860F8C5FD621B58" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[621,762,1813,1837]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B973095379209F744860F8C5FD621B58" box="[621,762,1813,1837]" italics="true" pageId="7" pageNumber="8">A. cuspidata</emphasis>
</taxonomicName>
(
<figureCitation id="133CC9C479209F744907F8C6FCE21B5B" box="[778,890,1813,1838]" captionStart="FIGURES 103106" captionStartId="55.[151,264,1911,1933]" captionTargetBox="[168,1425,204,1877]" captionTargetId="figure@55.[151,1436,193,1890]" captionTargetPageId="55" captionText="FIGURES 103106. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view; (e) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3770070" httpUri="https://zenodo.org/record/3770070/files/figure.png" pageId="7" pageNumber="8">Fig. 106a</figureCitation>
). Thoracic patterns in
<taxonomicName id="4C07AEC279209F744E7AF8C5FB6F1B58" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[1143,1271,1813,1837]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B973095379209F744E7AF8C5FB6F1B58" box="[1143,1271,1813,1837]" italics="true" pageId="7" pageNumber="8">A. acridens</emphasis>
</taxonomicName>
tend to mirror those of
<taxonomicName id="4C07AEC279209F744AF5F8E9FE1D1B24" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[248,389,1849,1873]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B973095379209F744AF5F8E9FE1D1B24" box="[248,389,1849,1873]" italics="true" pageId="7" pageNumber="8">A. cuspidata</emphasis>
</taxonomicName>
but the dark humeral stripe in
<taxonomicName id="4C07AEC279209F7448ECF8E9FCFA1B24" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[737,866,1849,1873]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B973095379209F7448ECF8E9FCFA1B24" box="[737,866,1849,1873]" italics="true" pageId="7" pageNumber="8">A. acridens</emphasis>
</taxonomicName>
(
<figureCitation id="133CC9C479209F74497CF8EAFC2C1B27" box="[881,948,1849,1874]" captionStart="FIGURES 913" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 913. Color pattern of male head, thorax, and S14. Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
) tends to be lighter and in some specimens is distinctly paler than the black middorsal stripe; in
<taxonomicName id="4C07AEC279209F7448E9F88DFCE81B00" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[740,880,1885,1909]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B973095379209F7448E9F88DFCE81B00" box="[740,880,1885,1909]" italics="true" pageId="7" pageNumber="8">A. cuspidata</emphasis>
</taxonomicName>
, the humeral stripe is usually as dark as the black middorsal stripe (
<figureCitation id="133CC9C479209F744B50F852FE291BEF" box="[349,433,1921,1946]" captionStart="FIGURES 913" captionStartId="40.[151,264,1879,1901]" captionTargetBox="[173,1407,210,1846]" captionTargetId="figure@40.[150,1436,193,1858]" captionTargetPageId="40" captionText="FIGURES 913. Color pattern of male head, thorax, and S14. Head, dorsal view, thorax and S14, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770043" httpUri="https://zenodo.org/record/3770043/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
). Greater differences are found on abdominal S3, 89 as follows: Dorsal pale area on S3 of
<taxonomicName id="4C07AEC279209F744ABBF875FEAE1BC8" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[182,310,1957,1981]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B973095379209F744ABBF875FEAE1BC8" box="[182,310,1957,1981]" italics="true" pageId="7" pageNumber="8">A. acridens</emphasis>
</taxonomicName>
is restricted basally as a small narrow posteriorly directed stripe (Fig. 34); in
<taxonomicName id="4C07AEC279209F744EA5F875FAAD1BC8" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[1192,1333,1957,1981]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B973095379209F744EA5F875FAAD1BC8" box="[1192,1333,1957,1981]" italics="true" pageId="7" pageNumber="8">A. cuspidata</emphasis>
</taxonomicName>
, the pale dorsal stripe is broader, largely parallel and ends abruptly at apical fourth (Fig. 45). S
<quantity id="4CFF78A479209F744E58F819FB061B97" box="[1109,1182,1993,2018]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="7" pageNumber="8" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">89 in</quantity>
<taxonomicName id="4C07AEC279209F744EA8F819FABC1B97" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[1189,1316,1994,2018]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B973095379209F744EA8F819FABC1B97" box="[1189,1316,1994,2018]" italics="true" pageId="7" pageNumber="8">A. acridens</emphasis>
</taxonomicName>
is entirely pale dorsolaterally (Fig. 35); pale areas in
<taxonomicName id="4C07AEC2792F9F7B4879FF4BFC991CC5" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[628,769,152,176]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792F9F7B4879FF4BFC991CC5" box="[628,769,152,176]" italics="true" pageId="8" pageNumber="9">A. cuspidata</emphasis>
</taxonomicName>
are restricted (Fig. 46) or entirely (Fig. 47) black with pale areas limited to lateral portions of S8 and dorsal blue spot not reaching apical fourth (Figs. 4647) on S9.
</paragraph>
<paragraph id="8BB8D541792F9F7B4ACAFF0CFAE11D11" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="9">
Females of both species are more similar in terms of body markings and they differ in the morphology of the mesostigmal plate. An elongate raised anteriorly directed tubercle confined to the medial half of the mesostigmal lamina characterizes
<taxonomicName id="4C07AEC2792F9F7B4B88FEFBFD9D1D35" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[389,517,296,320]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9730953792F9F7B4B88FEFBFD9D1D35" box="[389,517,296,320]" italics="true" pageId="8" pageNumber="9">A. acridens</emphasis>
</taxonomicName>
(
<figureCitation id="133CC9C4792F9F7B481BFEF4FDF31D35" box="[534,619,295,320]" captionStart="FIGURES 6972" captionStartId="48.[151,264,1557,1579]" captionTargetBox="[264,1313,208,1530]" captionTargetId="figure@48.[264,1323,193,1535]" captionTargetPageId="48" captionText="FIGURES 6972. Female mesostigmal plates. (a) dorsal view; (b) detail of mesostigmal lobe, dorsal view; (c) frontal (anterior) view; (d) detail of mesostigmal lobe, posterodorsal view (on edge)." figureDoi="http://doi.org/10.5281/zenodo.3770053" httpUri="https://zenodo.org/record/3770053/files/figure.png" pageId="8" pageNumber="9">Fig. 69</figureCitation>
); no such tubercle is present in
<taxonomicName id="4C07AEC2792F9F7B49DEFEFBFBF81D35" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[979,1120,296,320]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9730953792F9F7B49DEFEFBFBF81D35" box="[979,1120,296,320]" italics="true" pageId="8" pageNumber="9">A. cuspidata</emphasis>
</taxonomicName>
, this area instead forming a slight concavity with a gently raised area directly posterior to medial margin of the mesostigmal plate (
<figureCitation id="133CC9C4792F9F7B4F1AFE9FFAF41D11" box="[1303,1388,332,357]" captionStart="FIGURES 6972" captionStartId="48.[151,264,1557,1579]" captionTargetBox="[264,1313,208,1530]" captionTargetId="figure@48.[264,1323,193,1535]" captionTargetPageId="48" captionText="FIGURES 6972. Female mesostigmal plates. (a) dorsal view; (b) detail of mesostigmal lobe, dorsal view; (c) frontal (anterior) view; (d) detail of mesostigmal lobe, posterodorsal view (on edge)." figureDoi="http://doi.org/10.5281/zenodo.3770053" httpUri="https://zenodo.org/record/3770053/files/figure.png" pageId="8" pageNumber="9">Fig. 72</figureCitation>
).
</paragraph>
<paragraph id="8BB8D541792F9F7B4ACAFEBCFB911E48" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="9">
<emphasis id="B9730953792F9F7B4ACAFEBCFEB21DFD" bold="true" box="[199,298,367,392]" pageId="8" pageNumber="9">Habitat.</emphasis>
Apparently confined to seepages associated with larger rivers and creeks. A pair collected by S. W. Dunkle was found at a &quot;stream and ditch near a golf course&quot; with the habitat characterized as &quot;shallow water flowing over a rock face.&quot; Jim Johnson (pers. comm.
<date id="FFB9F381792F9F7B48E2FE6BFCE21DA5" box="[751,890,440,464]" pageId="8" pageNumber="9" value="2017-12-09">9 Dec. 2017</date>
) photographed and collected this species at &quot;....a vegetated cliff with water dripping off of it. This was along one side of a road, and close to a high gradient stream that flowed under the road. The adults perched on vegetation along that cliff, and my recollection is that it was in shade when we found them, although I am not completely positive about that.&quot;
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA792F9F7B4ACAFD94FD721EF0" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BB8D541792F9F7B4ACAFD94FD721EF0" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="9">
<emphasis id="B9730953792F9F7B4ACAFD94FEF81E15" bold="true" box="[199,352,583,608]" pageId="8" pageNumber="9">Distribution.</emphasis>
As far as known endemic to
<collectingCountry id="F31095D1792F9F7B48A2FD94FC961E15" box="[687,782,583,608]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
, restricted to
<collectingRegion id="49C31BA3792F9F7B49A4FD94FBB51E15" box="[937,1069,583,608]" country="Ecuador" name="Esmeraldas" pageId="8" pageNumber="9">Esmeraldas</collectingRegion>
,
<collectingRegion id="49C31BA3792F9F7B4E34FD94FB161E15" box="[1081,1166,583,608]" country="Ecuador" name="Manabi" pageId="8" pageNumber="9">Manabí</collectingRegion>
and
<collectingRegion id="49C31BA3792F9F7B4EC5FD9BFEBE1EF0" country="Ecuador" name="Santo Domingo de los Tsachilas" pageId="8" pageNumber="9">Santo Domingo de los Tsáchilas</collectingRegion>
Provinces west of the Andes (
<figureCitation id="133CC9C4792F9F7B4876FDBFFD461EF0" box="[635,734,620,645]" captionStart="FIGURES 136137" captionStartId="63.[151,264,1901,1923]" captionTargetBox="[170,1246,208,1864]" captionTargetId="figure@63.[151,1437,193,1880]" captionTargetPageId="63" captionText="FIGURES 136137. Distribution of new Argia species and closest congeners in Ecuador." figureDoi="http://doi.org/10.5281/zenodo.3770088" httpUri="https://zenodo.org/record/3770088/files/figure.png" pageId="8" pageNumber="9">Fig. 136</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>