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<document ID-DOI="http://doi.org/10.5281/zenodo.6448815" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6448815" approvalRequired="392" approvalRequired_for_illustrations="1" approvalRequired_for_matCits="49" approvalRequired_for_taxonomicNames="120" approvalRequired_for_treatments="222" checkinTime="1635825784914" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03AD87FAFFE1F60F89663C58F98AF701" docLanguage="en" docName="hbmw_9_Pteropodidae_16.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Epomops franqueti" docType="treatment" docVersion="3" lastPageNumber="95" masterDocId="FF94FF82FFC4F62A891E341CFFA5FF9B" masterDocTitle="Pteropodidae" masterLastPageNumber="162" masterPageNumber="16" pageNumber="95" updateTime="1656519960238" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Pteropodidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>16</mods:start>
<mods:end>162</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6448815</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">6448815</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6448955" ID-Zenodo-Dep="6448955" LSID="urn:lsid:plazi:treatment:03AD87FAFFE1F60F89663C58F98AF701" httpUri="http://treatment.plazi.org/id/03AD87FAFFE1F60F89663C58F98AF701" lastPageNumber="95" pageId="37" pageNumber="95">
<subSubSection box="[120,177,2116,2162]" pageId="37" pageNumber="95" type="multiple">
<paragraph blockId="37.[119,1254,2116,2207]" box="[120,177,2116,2162]" pageId="37" pageNumber="95">
<heading box="[120,177,2116,2162]" pageId="37" pageNumber="95">
<figureCitation box="[120,177,2116,2162]" captionStart="Plate 4: Pteropodidae" captionStartId="33.[122,152,3248,3269]" captionTargetBox="[14,2755,18,3663]" captionTargetPageId="32" captionText="50. Western Woermanns Fruit Bat (Megaloglossus azagnyi), 51. Eastern Woermanns Fruit Bat (Megaloglossus woermanni), 52. Angolan Soft-furred Fruit Bat (Lissonycteris angolensis), 53. Little Collared Fruit Bat (Myonycteris torquata), 54. Sao Tome Collared Fruit Bat (Myonycteris brachycephala), 55. Sierra Leone Collared Fruit Bat (Myonycteris leptodon), 56. Bergmanss Collared Fruit Bat (Myonycteris relicta), 57. Broad-faced Fruit Bat (Plerotes anchietae), 58. Hammer-headed Fruit Bat (Hypsignathus monstrosus), 59. Franquets Epauletted Fruit Bat (Epomops franqueti), 60. Buttikofers Epauletted Fruit Bat (Epomops buettikoferi), 61. Veldkamps Epauletted Fruit Bat (Nanonycteris veldkampr), 62. Gambian Epauletted Fruit Bat (Epomophorus gambianus), 63. Peterss Epauletted Fruit Bat (Epomophorus erypturus), 64. Angolan Epauletted Fruit Bat (Epomophorus angolensis), 65. Little Epauletted Fruit Bat (Epomophorus labiatus), 66. Minor Epauletted Fruit Bat (Epomophorus minor), 67. Ansells Epauletted Fruit Bat (Epomophorus anselli), 68. Wahlberg's Epauletted Fruit Bat (Epomophorus wahlbergi), 69. Dobsons Epauletted Fruit Bat (Epomophorus dobsonu), 70. Sanborns Epauletted Fruit Bat (Epomophorus grands), 71. Lesser Epauletted Fruit Bat (Epomophorus pusillus), 72. Haymans Epauletted Fruit Bat (Epomophorus intermedius)" figureDoi="http://doi.org/10.5281/zenodo.6448849" httpUri="https://zenodo.org/record/6448849/files/figure.png" pageId="37" pageNumber="95">59.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[194,853,2116,2162]" pageId="37" pageNumber="95" type="vernacular_names">
<paragraph blockId="37.[119,1254,2116,2207]" box="[194,853,2116,2162]" pageId="37" pageNumber="95">
<heading box="[194,853,2116,2162]" pageId="37" pageNumber="95">
<vernacularName box="[194,853,2116,2162]" pageId="37" pageNumber="95">Franquets Epauletted Fruit Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[923,1254,2116,2162]" pageId="37" pageNumber="95" type="nomenclature">
<paragraph blockId="37.[119,1254,2116,2207]" box="[923,1254,2116,2162]" pageId="37" pageNumber="95">
<heading box="[923,1254,2116,2162]" pageId="37" pageNumber="95">
<taxonomicName baseAuthorityName="Tomes" baseAuthorityYear="1860" box="[923,1254,2116,2162]" class="Mammalia" family="Pteropodidae" genus="Epomops" kingdom="Animalia" order="Chiroptera" pageId="37" pageNumber="95" phylum="Chordata" rank="species" species="franqueti">
<emphasis box="[923,1254,2116,2162]" italics="true" pageId="37" pageNumber="95">Epomops franqueti</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[120,1213,2180,2201]" pageId="37" pageNumber="95" type="vernacular_names">
<paragraph blockId="37.[119,1254,2116,2207]" box="[120,1213,2180,2201]" pageId="37" pageNumber="95">
<heading box="[120,1213,2180,2201]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[120,195,2180,2201]" pageId="37" pageNumber="95">French:</emphasis>
<vernacularName box="[205,460,2180,2201]" pageId="37" pageNumber="95">Epomophore de Franquet</vernacularName>
/
<emphasis bold="true" box="[480,571,2180,2201]" pageId="37" pageNumber="95">German:</emphasis>
<vernacularName box="[581,870,2180,2201]" pageId="37" pageNumber="95">Franguet-Epaulettenflughund</vernacularName>
/
<emphasis bold="true" box="[891,982,2180,2201]" pageId="37" pageNumber="95">Spanish:</emphasis>
<vernacularName box="[992,1213,2180,2201]" pageId="37" pageNumber="95">Epomops de Franquet</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="reference_group">
<paragraph blockId="37.[732,1325,2248,2675]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[733,889,2248,2281]" pageId="37" pageNumber="95">Taxonomy.</emphasis>
<taxonomicName authority="Tomes, 1860" authorityName="Tomes" authorityYear="1860" class="Mammalia" family="Pteropodidae" genus="Epomophorus" kingdom="Animalia" order="Chiroptera" pageId="37" pageNumber="95" phylum="Chordata" rank="species" species="franqueti">Epomophorus franqueti Tomes, 1860</taxonomicName>
,
<materialsCitation box="[819,945,2287,2320]" pageId="37" pageNumber="95">
<collectingCountry box="[833,928,2287,2320]" name="Gabon" pageId="37" pageNumber="95">Gabon</collectingCountry>
.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="discussion">
<paragraph blockId="37.[732,1325,2248,2675]" pageId="37" pageNumber="95">
A described subspecies strepitans from southern
<collectingCountry box="[875,979,2366,2399]" name="Nigeria" pageId="37" pageNumber="95">Nigeria</collectingCountry>
has not been regularly accepted because there seems to be clinal differences in
<taxonomicName authorityName="Tomes" authorityYear="1860" box="[1012,1173,2445,2478]" class="Mammalia" family="Pteropodidae" genus="Epomophorus" kingdom="Animalia" order="Chiroptera" pageId="37" pageNumber="95" phylum="Chordata" rank="species" species="franqueti">E. franqueti</taxonomicName>
, with size increasing from West to Central Africa. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="distribution">
<caption inLine="true" pageId="37" pageNumber="95" targetBox="[1579,1583,2177,2202]" targetPageId="37">
<paragraph blockId="37.[732,1325,2248,2675]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[732,909,2564,2597]" pageId="37" pageNumber="95">Distribution.</emphasis>
W &amp; C Africa from SE
<collectingCountry name="Ivory Coast" pageId="37" pageNumber="95">Ivory Coast</collectingCountry>
E to
<collectingCountry box="[886,1067,2612,2637]" name="South Sudan" pageId="37" pageNumber="95">South Sudan</collectingCountry>
and
<collectingCountry box="[1141,1253,2612,2637]" name="Uganda" pageId="37" pageNumber="95">Uganda</collectingCountry>
, S to N
<collectingCountry box="[766,872,2642,2675]" name="Zambia" pageId="37" pageNumber="95">Zambia</collectingCountry>
and N
<collectingCountry box="[975,1078,2642,2675]" name="Angola" pageId="37" pageNumber="95">Angola</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="description">
<paragraph blockId="37.[117,1324,2685,3464]" lastBlockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[118,375,2685,2714]" pageId="37" pageNumber="95">Descriptive notes.</emphasis>
Head-body
<quantity box="[575,761,2685,2714]" metricMagnitude="-1" metricUnit="m" metricValue="1.375" metricValueMax="1.65" metricValueMin="1.1" pageId="37" pageNumber="95" unit="mm" value="137.5" valueMax="165.0" valueMin="110.0">110-165 mm</quantity>
(males) and
<quantity box="[976,1162,2685,2714]" metricMagnitude="-1" metricUnit="m" metricValue="1.475" metricValueMax="1.8" metricValueMin="1.15" pageId="37" pageNumber="95" unit="mm" value="147.5" valueMax="180.0" valueMin="115.0">115-180 mm</quantity>
(females), tail
<quantity box="[173,291,2725,2754]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="37" pageNumber="95" unit="mm" value="0.5" valueMax="1.0" valueMin="0.0">0-1 mm</quantity>
, ear
<quantity box="[366,516,2725,2754]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="37" pageNumber="95" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">23-27 mm</quantity>
, hindfoot
<quantity box="[672,820,2725,2754]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="37" pageNumber="95" unit="mm" value="26.0" valueMax="28.0" valueMin="24.0">24-28 mm</quantity>
(males) and
<quantity box="[1019,1168,2725,2754]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="37" pageNumber="95" unit="mm" value="23.0" valueMax="25.0" valueMin="21.0">21-25 mm</quantity>
(females), forearm
<quantity box="[243,407,2764,2793]" metricMagnitude="-2" metricUnit="m" metricValue="9.25" metricValueMax="10.1" metricValueMin="8.4" pageId="37" pageNumber="95" unit="mm" value="92.5" valueMax="101.0" valueMin="84.0">84-101 mm</quantity>
(males) and
<quantity box="[601,748,2764,2793]" metricMagnitude="-2" metricUnit="m" metricValue="8.65" metricValueMax="9.6" metricValueMin="7.7" pageId="37" pageNumber="95" unit="mm" value="86.5" valueMax="96.0" valueMin="77.0">77-96 mm</quantity>
(females); weight
<quantity box="[1015,1142,2764,2793]" metricMagnitude="-1" metricUnit="kg" metricValue="1.3199999999999998" metricValueMax="1.72" metricValueMin="0.9199999999999999" pageId="37" pageNumber="95" unit="g" value="132.0" valueMax="172.0" valueMin="92.0">92-172 g</quantity>
(males) and
<quantity box="[118,246,2799,2832]" metricMagnitude="-2" metricUnit="kg" metricValue="9.55" metricValueMax="13.0" metricValueMin="6.1" pageId="37" pageNumber="95" unit="g" value="95.5" valueMax="130.0" valueMin="61.0">61-130 g</quantity>
(females). Males average larger than females, with larger and more robust heads. Muzzle is relatively long and broad;lips and cheeks are fleshy and expandable, particularly in males; eyes are large, with brown irises; and ears are dark brown, with anterior and posterior basal ear patches. Epaulettes formed by an eversible muscular pocketfilled with long, erectile, white hairs are present on adult males and used in display; females have small, inconspicuous invagination on each shoulder with 12-mm pale brown hairs; dorsum is generally fawn, light brown to reddish brown in both sexes; sometimes paler grayish brown on shoulders and nape, with unicolored hairs; and pelage is soft, slightly fluffy, and mid-dorsally ¢.
<quantity box="[765,847,3115,3148]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="95" unit="mm" value="8.0">8 mm</quantity>
, extending about halfway on forearm dorsally and ventrally. Venter is paler brown, regularly changing into ample white patch on chest and abdomen. Wings have claw on second digits; membranes are medium to dark brown, attached to second toes, and sparsely covered in hairs for about one-half upper surface and about one-third lower surface. Skull is medium-sized, with basicranial deflection; dorsal profile is generally straight or somewhat concave in rostral and interorbital regions; rostrum is of medium relative length; zygomatic arches are sturdy and long; and palate is relatively short and weakly concave from side-to-side and anteroposteriorly. There are three thick and 5-8 thin palatal ridges; ridges 1-2 are undivided; ridge 3 is exceptionally divided; ridges 4 and onward are narrowly divided and finely serrated; and post-dental palate is almost flat. Larynx is ossified in adult males. Dental formula for all species of Epomopsis 12/2, C1/1,P 2/3, M 1/2 (x2) = 28, but I* is frequently missing, often in females. P' is separated from C' by large diastema, and P|is reduced but larger than an incisor. Chromosomal complement for four females and one male from
<collectingCountry box="[1873,2021,481,506]" name="Cameroon" pageId="37" pageNumber="95">Cameroon</collectingCountry>
had 2n = 36 and FNa = 66, with 13 pairs of metacentric or submetacentric and four pairs of submetacentric or subtelocentric autosomes. X-chromosome was medium-sized subtelocentric, and Y-chromosome was small subtelocentric. In
<collectingCountry box="[1738,1834,591,624]" name="Gabon" pageId="37" pageNumber="95">Gabon</collectingCountry>
, chromosomal complement had 2n = 35 (single male) and 36 (females) and FNa = 66; Y-chromosome was missing, leading to XX/XO sex chromosome system.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="biology_ecology">
<paragraph blockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[1399,1510,718,743]" pageId="37" pageNumber="95">Habitat.</emphasis>
Primarily rainforest and rainforest-savanna mosaic, both wetter and drier types, and marginally
<collectingCountry box="[1709,1811,748,781]" name="Guinea-Bissau" pageId="37" pageNumber="95">Guinea</collectingCountry>
Savanna and Zambezian Woodland biotic zones (in relict and riverine forests), from sea level up to elevations of ¢.
<quantity box="[2267,2380,788,821]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="37" pageNumber="95" unit="m" value="1850.0">1850 m</quantity>
. Franquets Epauletted Fruit Bat was also reported from secondary forests (
<collectingCountry box="[2266,2367,827,860]" name="Gabon" pageId="37" pageNumber="95">Gabon</collectingCountry>
), often clearings close to dense groundcover, and gardens.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="food_feeding">
<paragraph blockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[1398,1672,906,939]" pageId="37" pageNumber="95">Food and Feeding.</emphasis>
Franquets Epauletted Fruit Bat takes fruit from canopies and sometimes also fallen fruit. Fruits are stored in expandable cheeks and taken to a perch at up to
<quantity box="[1610,1696,985,1018]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="95" unit="m" value="200.0">200 m</quantity>
away. Food is held by a foot, mouth, wrists, or in pocket formed by wing membrane when a wrist is flexed (thumbs are generally only used to cling on perch or leaf). Fruits from at least 14 plant genera in ten families are consumed. In
<collectingCountry box="[1438,1534,1104,1137]" name="Gabon" pageId="37" pageNumber="95">Gabon</collectingCountry>
, females prefer fruits from
<taxonomicName box="[1923,2037,1104,1137]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="37" pageNumber="95" phylum="Tracheophyta" rank="genus">Solanum</taxonomicName>
torvum (
<taxonomicName box="[2158,2322,1104,1137]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="37" pageNumber="95" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
); males preferfigs.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="breeding">
<paragraph blockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[1397,1533,1143,1176]" pageId="37" pageNumber="95">Breeding.</emphasis>
Litter size of Franquets Epauletted Fruit Bat is one. In
<collectingCountry box="[2336,2448,1143,1176]" name="Uganda" pageId="37" pageNumber="95">Uganda</collectingCountry>
, reproductive chronology is continuous bimodal polyestry, with postpartum estrus. Gestation lasts 5-6 months, and births occur in March and September. Lactating females are found almost year-round; young are probably weaned only a few weeks before the next birth. Franquets Epauletted Fruit Bat has an “exploded” lek mating system (i.e. not tight clusters such as leks of the Hammer-headed Fruit Bat,
<taxonomicName authorityName="H. Allen" authorityYear="1861" box="[2257,2590,1341,1374]" class="Mammalia" family="Pteropodidae" genus="Hypsignathus" kingdom="Animalia" order="Chiroptera" pageId="37" pageNumber="95" phylum="Chordata" rank="species" species="monstrosus">Hypsignathus monstrosus</taxonomicName>
), where a male has a territory of ¢.
<quantity box="[1856,1947,1380,1413]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="95" unit="m" value="200.0">200 m</quantity>
in diameter. These territories contain enough resources for females to feed and roost but are probably selected for optimizing display and minimizing predation. Individual leks are
<quantity box="[2157,2262,1459,1492]" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="37" pageNumber="95" unit="km" value="3.0" valueMax="4.0" valueMin="2.0">2—4 km</quantity>
apart. During displays, males beat their wings, show off their epaulettes, and make honking sounds about once a second; this calling continues throughout the night.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="activity">
<paragraph blockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[1395,1630,1578,1611]" pageId="37" pageNumber="95">Activity patterns.</emphasis>
Franquets Epauletted Fruit Bat is nocturnal. It is slow and maneuverable, generally flying through canopies and aboveground.It roosts in dense foliage of large trees (e.g.
<taxonomicName box="[1637,1775,1657,1690]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="37" pageNumber="95" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
,
<taxonomicName box="[1794,2004,1657,1690]" class="Magnoliopsida" family="Combretaceae" kingdom="Plantae" order="Myrtales" pageId="37" pageNumber="95" phylum="Tracheophyta" rank="family">Combretaceae</taxonomicName>
), close to or over water surfaces, where it hangs from branches
<quantity box="[1704,1788,1696,1729]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="37" pageNumber="95" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4-6 m</quantity>
aboveground, rarely in low bushes.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="biology_ecology">
<paragraph blockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[1397,2125,1740,1769]" pageId="37" pageNumber="95">Movements, Home range and Social organization.</emphasis>
Franquets Epauletted Fruit Bat roosts singly, sometimes in pairs and exceptionally in trios. In
<collectingCountry box="[2301,2396,1775,1808]" name="Gabon" pageId="37" pageNumber="95">Gabon</collectingCountry>
, home ranges have radii of
<quantity box="[1581,1735,1815,1848]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="37" pageNumber="95" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600-800 m</quantity>
, with males having larger home ranges than females.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="conservation">
<paragraph blockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[1397,1745,1854,1887]" pageId="37" pageNumber="95">Status and Conservation.</emphasis>
Classified as Least Concern on The [UCN Red List. Franquets Epauletted Fruit Bat has a wide distribution and presumably large population. It is probably not declining fast enough to be assigned to a higher category. Major threats are probably habitat loss due to deforestation and recently persecution and extermination as a result of fear for their potential to transmit zoonotic diseases (e.g. Ebola virus).
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="95" type="bibRefCitation_list">
<paragraph blockId="37.[1395,2607,275,2206]" pageId="37" pageNumber="95">
<emphasis bold="true" box="[1397,1550,2099,2124]" pageId="37" pageNumber="95">Bibliography.</emphasis>
ACR (2018), Aellen (1952), Andersen (1910a), Bergmans (1989), Gembu Tungaluna (2012), Haiduk et al. (1980), Happold, M. (2013i), Kityo &amp; Nalikka (2016), Monadjem, Taylor et al. (2010), Primus et al. (2006), Simmons (2005).
</paragraph>
</subSubSection>
</treatment>
</document>