256 lines
37 KiB
XML
256 lines
37 KiB
XML
<document id="BE63EE2D618EB594DD2CDAD77E08E586" ID-CLB-Dataset="32079" ID-DOI="10.11646/zootaxa.4320.3.12" ID-GBIF-Dataset="6132c40b-b273-4e7a-9718-ef97cf2280c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="893728" ID-ZooBank="Fcc8Fd2C-C644-43Fb-B28E-1Cd75De38C30" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712289128" checkinUser="plazi" docAuthor="Nel, André" docDate="2017" docId="03AA8781FFFFFFAE87E7FED7FE65E018" docLanguage="en" docName="zootaxa.4320.3.12.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Burmacompsocus coniugans Nel 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="598" masterDocId="FF93FFF9FFFEFFAF8770FFB0FFEDE749" masterDocTitle="New species of Compsocidae (Insecta, Psocodea) from Cretaceous Burmese amber" masterLastPageNumber="600" masterPageNumber="597" pageNumber="598" updateTime="1698468571971" updateUser="ExternalLinkService">
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<mods:titleInfo id="4DCA7CE185BFC1BB2E8287ACFA96B396">
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<mods:title id="6CEF5B3332C065876E1C1A7F99D0B52E">New species of Compsocidae (Insecta, Psocodea) from Cretaceous Burmese amber</mods:title>
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</mods:titleInfo>
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<mods:name id="38D762B0187F0E35B7173636029F555A" type="personal">
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<mods:namePart id="6EFEC009707258C547FEF3D2C10BFC54">Nel, André</mods:namePart>
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<mods:typeOfResource id="67B6EFBA2817BA3E966DFD2839391A42">text</mods:typeOfResource>
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<mods:title id="90196CAE56F240301AEBA5A439ADE688">Zootaxa</mods:title>
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<mods:part id="19D00847FB7E15D7668284E915AFCB72">
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<mods:date id="D050B0A6EB426172BFC92CC962C82DD7">2017</mods:date>
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<mods:number id="3D5E1F945A9ABDC0B205D2A4FF70F34E">2017-09-18</mods:number>
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<mods:number id="2D7AEC23F50153F3EA7AAB3B0A4E241C">4320</mods:number>
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<treatment id="03AA8781FFFFFFAE87E7FED7FE65E018" ID-DOI="http://doi.org/10.5281/zenodo.6008861" ID-GBIF-Taxon="134090539" ID-Zenodo-Dep="6008861" LSID="urn:lsid:plazi:treatment:03AA8781FFFFFFAE87E7FED7FE65E018" httpUri="http://treatment.plazi.org/id/03AA8781FFFFFFAE87E7FED7FE65E018" lastPageNumber="598" pageId="1" pageNumber="598">
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<subSubSection id="C319651CFFFFFFAE87E7FED7FC78E6CB" box="[151,917,359,386]" pageId="1" pageNumber="598" type="nomenclature">
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<paragraph id="8BBC3697FFFFFFAE87E7FED7FC78E6CB" blockId="1.[151,917,359,386]" box="[151,917,359,386]" pageId="1" pageNumber="598">
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<heading id="D0F481FBFFFFFFAE87E7FED7FC78E6CB" bold="true" box="[151,917,359,386]" fontSize="11" level="1" pageId="1" pageNumber="598" reason="1">
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<emphasis id="B977EA85FFFFFFAE87E7FED7FC78E6CB" bold="true" box="[151,917,359,386]" pageId="1" pageNumber="598">
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<taxonomicName id="4C034D14FFFFFFAE87E7FED7FD74E6CB" ID-CoL="NXLR" authority="Sroka & Nel" authorityName="Nel" authorityYear="2017" box="[151,665,359,386]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans" status="sp. nov.">
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<emphasis id="B977EA85FFFFFFAE87E7FED7FE19E6C8" bold="true" box="[151,500,359,385]" italics="true" pageId="1" pageNumber="598">Burmacompsocus coniugans</emphasis>
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Sroka & Nel
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24457FEFFFFFFAE85D6FED8FCEEE6CB" box="[678,771,360,386]" pageId="1" pageNumber="598" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="13382A12FFFFFFAE8463FED8FC66E6CB" box="[787,907,360,386]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Figs 1–10</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C319651CFFFFFFAE87E7FE1DFB3AE6AE" pageId="1" pageNumber="598" type="etymology">
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<paragraph id="8BBC3697FFFFFFAE87E7FE1DFB3AE6AE" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
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<emphasis id="B977EA85FFFFFFAE87E7FE1DFEFAE68D" bold="true" box="[151,279,429,452]" pageId="1" pageNumber="598">Etymology.</emphasis>
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The name of the new species is derived from the Latin expression for "fusing" and refers to the fusion of veins Rs and M in the forewing, which probably constitutes the most reliable distinguishing character.
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</paragraph>
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</subSubSection>
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<subSubSection id="C319651CFFFFFFAE87B7FE42FD7AE565" pageId="1" pageNumber="598" type="diagnosis">
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<paragraph id="8BBC3697FFFFFFAE87B7FE42FD7AE565" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
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<emphasis id="B977EA85FFFFFFAE87B7FE42FED1E540" bold="true" box="[199,316,498,521]" pageId="1" pageNumber="598">Diagnosis.</emphasis>
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Forewing with Rs fused with M for a short distance, row of denticles on anterior carina of forefemur absent, lacinia with pronounced subapical teeth.
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</paragraph>
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</subSubSection>
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<subSubSection id="C319651CFFFFFFAE87B7FD88FB4CE225" pageId="1" pageNumber="598" type="description">
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<paragraph id="8BBC3697FFFFFFAE87B7FD88FE38E538" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
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<emphasis id="B977EA85FFFFFFAE87B7FD88FEBFE506" bold="true" box="[199,338,568,591]" pageId="1" pageNumber="598">Description.</emphasis>
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<typeStatus id="54B88835FFFFFFAE8629FD88FE51E506" box="[345,444,568,591]" pageId="1" pageNumber="598" type="holotype">Holotype</typeStatus>
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specimen represents male. Body surface strongly corrugated. No flattened scales present on surface of body and/or wings.
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</paragraph>
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<paragraph id="8BBC3697FFFFFFAE87B7FDCDFAC9E408" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
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Head
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<quantity id="4CFB9B72FFFFFFAE8677FDCDFE85E5DD" box="[263,360,637,660]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="598" unit="mm" value="0.52">0.52 mm</quantity>
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long,
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<quantity id="4CFB9B72FFFFFFAE86DFFDCDFDE2E5DD" box="[431,527,637,660]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="598" unit="mm" value="0.58">0.58 mm</quantity>
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wide. Three ocelli situated on slightly elevated tubercle (
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<figureCitation id="13382A12FFFFFFAE83F1FDCDFB09E5DD" box="[1153,1252,637,660]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Figs 1, 3</figureCitation>
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). Lacinia curved outwards apically, with several distinct subapical denticles on inner side (
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<figureCitation id="13382A12FFFFFFAE84C3FD10FC18E5FE" box="[947,1013,672,695]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
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). Denticles not observable from lateral view, from where lacinia appears rod-like (see
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<figureCitation id="13382A12FFFFFFAE85F8FD72FD2AE590" box="[648,711,706,729]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
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). Maxillary palpi (
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<figureCitation id="13382A12FFFFFFAE84FEFD72FC20E590" box="[910,973,706,729]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
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) with three distinguishable segments, basal segment not visible. Apical segment
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<quantity id="4CFB9B72FFFFFFAE8531FD55FD4AE5B5" box="[577,679,741,764]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="598" unit="mm" value="0.13">0.13 mm</quantity>
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long, slightly widened distally, subapical segment
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<quantity id="4CFB9B72FFFFFFAE8383FD55FAB5E5B5" box="[1267,1368,741,764]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="598" unit="mm" value="0.08">0.08 mm</quantity>
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long, cylindrical. Labial palpi short, rounded apically, bearing numerous long hair-like setae (
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<figureCitation id="13382A12FFFFFFAE8301FCB8FB5AE456" box="[1137,1207,776,799]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
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). Antennae filiform, composed of 13 segments, equipped with sparse long hair-like setae. Length of segments decreasing apically.
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</paragraph>
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<paragraph id="8BBC3697FFFFFFAE87B7FCFDFD4EE4A6" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
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Wings hyaline, covered with numerous short microtrichia, denser along wing veins. No long setae on veins and/or wing margins. Forewing (
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<figureCitation id="13382A12FFFFFFAE86DFFCC0FDFDE4CE" box="[431,528,880,903]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Figs 1, 3</figureCitation>
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)
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<quantity id="4CFB9B72FFFFFFAE8552FCC0FD6EE4CE" box="[546,643,880,903]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" pageId="1" pageNumber="598" unit="mm" value="1.87">1.87 mm</quantity>
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long,
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<quantity id="4CFB9B72FFFFFFAE85B9FCC0FCC7E4CE" box="[713,810,880,903]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="598" unit="mm" value="0.73">0.73 mm</quantity>
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wide. Pterostigma
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<quantity id="4CFB9B72FFFFFFAE848DFCC0FBB0E4CE" box="[1021,1117,880,903]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="598" unit="mm" value="0.34">0.34 mm</quantity>
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long,
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||
<quantity id="4CFB9B72FFFFFFAE83D4FCC0FAE8E4CE" box="[1188,1285,880,903]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="598" unit="mm" value="0.13">0.13 mm</quantity>
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wide, basally closed, not sclerotized. Stems of M and Rs fused for short distance. Areola postica
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<quantity id="4CFB9B72FFFFFFAE835FFC22FB6EE4E0" box="[1071,1155,914,937]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="598" unit="mm" value="0.4">0.4 mm</quantity>
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long,
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<quantity id="4CFB9B72FFFFFFAE83BCFC22FAC0E4E0" box="[1228,1325,914,937]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="598" unit="mm" value="0.12">0.12 mm</quantity>
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wide, not connected to M by crossvein. Nodulus present. A2 not joining A1. Hind wing (
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<figureCitation id="13382A12FFFFFFAE84AFFC05FBA6E485" box="[991,1099,949,972]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Figs 9, 10</figureCitation>
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)
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<quantity id="4CFB9B72FFFFFFAE832AFC05FB55E485" box="[1114,1208,949,972]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="598" unit="mm" value="1.55">1.55 mm</quantity>
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||
long,
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<quantity id="4CFB9B72FFFFFFAE838CFC05FAB6E485" box="[1276,1371,949,972]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="598" unit="mm" value="0.44">0.44 mm</quantity>
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||
wide. First segment of Rs not visible. M two-branched.
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</paragraph>
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<paragraph id="8BBC3697FFFFFFAE87B7FC4AFED6E330" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
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Tibiae with rows of long spine-like setae. Group of thicker setae apically. Tarsi (
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<figureCitation id="13382A12FFFFFFAE8327FC4AFB76E358" box="[1111,1179,1018,1041]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 7</figureCitation>
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) three-segmented, first segment longest, equipped with rows of spine-like setae, shorter than on tibiae. Second and third segments approximately equal in length, tiny spine-like setae situated apically. Claws with two subapical denticles, pulvillus not visible (
|
||
<figureCitation id="13382A12FFFFFFAE879CFBD2FEC0E330" box="[236,301,1122,1145]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC3697FFFFFFAE87B7FB35FBF6E3F6" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
|
||
Abdomen
|
||
<quantity id="4CFB9B72FFFFFFAE8647FB35FE99E3D5" box="[311,372,1157,1180]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="598" unit="mm" value="1.0">1 mm</quantity>
|
||
long, slightly deformed. Hair-like setae observable in posterior part. Genitalia partly obscured, parameres and aedeagus partially observable (
|
||
<figureCitation id="13382A12FFFFFFAE85F2FB18FD29E3F6" box="[642,708,1192,1215]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 6</figureCitation>
|
||
). Hypandrium broadly rounded.
|
||
</paragraph>
|
||
<paragraph id="8BBC3697FFFFFFAE87B7FB7AFB4CE225" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
|
||
<emphasis id="B977EA85FFFFFFAE87B7FB7AFEDBE3A8" bold="true" box="[199,310,1226,1249]" pageId="1" pageNumber="598">Affinities.</emphasis>
|
||
Following
|
||
<bibRefCitation id="EF924B66FFFFFFAE86B0FB7AFD97E3A8" author="Mockford" box="[448,634,1226,1249]" pageId="1" pageNumber="598" refString="Mockford, E. L. (1967) The electrentomoid psocids (Psocoptera). Psyche, 74, 109 - 165." type="journal article" year="1967">Mockford (1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF924B66FFFFFFAE85FCFB7AFD2FE3A8" author="Mockford" box="[652,706,1226,1249]" pageId="1" pageNumber="598" refString="Mockford, E. L. (1993) North American Psocoptera (Insecta). Flora; Fauna Handbook 10. Sandhill Crane Press, Gainesville, Florida, 455 pp." type="book" year="1993">1993</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="EF924B66FFFFFFAE847DFB7AFC2EE3A8" author="Smithers" box="[781,963,1226,1249]" pageId="1" pageNumber="598" refString="Smithers, C. N. (1990) Keys to the families and genera of Psocoptera (Arthropoda, Insecta). Technical Reports of the Australian Museum, 1990 (2), 1 - 82. https: // doi. org / 10.3853 / j. 1031 - 8062.2.1990.77" type="journal article" year="1990">Smithers (1990)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C034D14FFFFFFAE84A5FB7AFB8AE3A8" authorityName="Nel" authorityYear="2017" box="[981,1127,1226,1249]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans">
|
||
<emphasis id="B977EA85FFFFFFAE84A5FB7AFB8AE3A8" box="[981,1127,1226,1249]" italics="true" pageId="1" pageNumber="598">B. coniugans</emphasis>
|
||
</taxonomicName>
|
||
exhibits a combination of morphological traits diagnostic for the recent family
|
||
<taxonomicName id="4C034D14FFFFFFAE8592FB5DFC0AE24D" authority="Mockford" authorityName="Mockford" box="[738,999,1261,1284]" class="Insecta" family="Compsocidae" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="family">Compsocidae Mockford</taxonomicName>
|
||
of the suborder Troctomorpha, namely (only characters visible in the holotype of
|
||
<taxonomicName id="4C034D14FFFFFFAE85F0FAA0FCFCE26E" authorityName="Nel" authorityYear="2017" box="[640,785,1296,1319]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans">
|
||
<emphasis id="B977EA85FFFFFFAE85F0FAA0FCFCE26E" box="[640,785,1296,1319]" italics="true" pageId="1" pageNumber="598">B. coniugans</emphasis>
|
||
</taxonomicName>
|
||
listed): antenna with 13 segments, tarsi three-segmented, macroptery with complex forewing venation, pterostigmal area not sclerotized or more opaque than rest of membrane, forewing with nodulus, hind wing with M forked, absence of flattened scales on wings and body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319651CFFFFFFAE87B7FAC8FE65E018" pageId="1" pageNumber="598" type="reference_group">
|
||
<paragraph id="8BBC3697FFFFFFAE87B7FAC8FBF3E1ED" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
|
||
<taxonomicName id="4C034D14FFFFFFAE87B7FAC8FEBFE2C6" ID-CoL="NXLR" authorityName="Nel" authorityYear="2017" box="[199,338,1400,1423]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans">
|
||
<emphasis id="B977EA85FFFFFFAE87B7FAC8FEBFE2C6" box="[199,338,1400,1423]" italics="true" pageId="1" pageNumber="598">B. coniugans</emphasis>
|
||
</taxonomicName>
|
||
also follows most of the additional characteristics of the genus
|
||
<taxonomicName id="4C034D14FFFFFFAE8373FAC8FB2DE2C6" ID-CoL="8MQPK" authorityName="Nel & Waller" authorityYear="2007" box="[1027,1216,1400,1423]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B977EA85FFFFFFAE8373FAC8FB2DE2C6" box="[1027,1216,1400,1423]" italics="true" pageId="1" pageNumber="598">Burmacompsocus</emphasis>
|
||
</taxonomicName>
|
||
as described by
|
||
<bibRefCitation id="EF924B66FFFFFFAE8205FAC8FE8DE2F8" author="Nel" pageId="1" pageNumber="598" refString="Nel, A. & Waller, A. (2007) The first fossil Compsocidae from Cretaceous Burmese amber (Insecta, Psocoptera, Troctomorpha). Cretaceous Research, 28, 1039 - 1041. https: // doi. org / 10.1016 / j. cretres. 2007.02.002" type="journal article" year="2007">Nel and Waller (2007)</bibRefCitation>
|
||
: lacinia tip with lateral cusp curved outward, claw with two preapical denticles, pterostigma closed basally, forewing veins 1A and 2A not joined, first segment of Rs in hind wing absent. At the same time,
|
||
<taxonomicName id="4C034D14FFFFFFAE8260FA0DFA76E29D" ID-CoL="NXLR" authorityName="Nel" authorityYear="2017" box="[1296,1435,1469,1492]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans">
|
||
<emphasis id="B977EA85FFFFFFAE8260FA0DFA76E29D" box="[1296,1435,1469,1492]" italics="true" pageId="1" pageNumber="598">B. coniugans</emphasis>
|
||
</taxonomicName>
|
||
differs in several aspects from
|
||
<taxonomicName id="4C034D14FFFFFFAE869FFA50FD86E2BE" ID-CoL="NXLS" box="[495,619,1504,1527]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="perreaui">
|
||
<emphasis id="B977EA85FFFFFFAE869FFA50FD86E2BE" box="[495,619,1504,1527]" italics="true" pageId="1" pageNumber="598">B. perreaui</emphasis>
|
||
</taxonomicName>
|
||
, the type species of
|
||
<taxonomicName id="4C034D14FFFFFFAE8425FA50FBFFE2BE" ID-CoL="8MQPK" authorityName="Nel & Waller" authorityYear="2007" box="[853,1042,1504,1527]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B977EA85FFFFFFAE8425FA50FBFFE2BE" box="[853,1042,1504,1527]" italics="true" pageId="1" pageNumber="598">Burmacompsocus</emphasis>
|
||
</taxonomicName>
|
||
. The forewing veins Rs and M are connected by a crossvein in
|
||
<taxonomicName id="4C034D14FFFFFFAE86A2F9B2FDA3E150" ID-CoL="NXLS" box="[466,590,1538,1561]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="perreaui">
|
||
<emphasis id="B977EA85FFFFFFAE86A2F9B2FDA3E150" box="[466,590,1538,1561]" italics="true" pageId="1" pageNumber="598">B. perreaui</emphasis>
|
||
</taxonomicName>
|
||
(see fig. 1D in
|
||
<bibRefCitation id="EF924B66FFFFFFAE8472F9B2FC34E150" author="Nel" box="[770,985,1538,1561]" pageId="1" pageNumber="598" refString="Nel, A. & Waller, A. (2007) The first fossil Compsocidae from Cretaceous Burmese amber (Insecta, Psocoptera, Troctomorpha). Cretaceous Research, 28, 1039 - 1041. https: // doi. org / 10.1016 / j. cretres. 2007.02.002" type="journal article" year="2007">Nel & Waller 2007</bibRefCitation>
|
||
), whilst these veins are fused for a short distance in
|
||
<taxonomicName id="4C034D14FFFFFFAE8666F995FE4EE175" ID-CoL="NXLR" authorityName="Nel" authorityYear="2017" box="[278,419,1573,1596]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans">
|
||
<emphasis id="B977EA85FFFFFFAE8666F995FE4EE175" box="[278,419,1573,1596]" italics="true" pageId="1" pageNumber="598">B. coniugans</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13382A12FFFFFFAE86C5F995FE14E175" box="[437,505,1573,1596]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 3</figureCitation>
|
||
). Another distinguishing character is represented by the row of denticles on anterior carina of the forefemur, mentioned by
|
||
<bibRefCitation id="EF924B66FFFFFFAE8533F9F8FCD4E116" author="Nel" box="[579,825,1608,1631]" pageId="1" pageNumber="598" refString="Nel, A. & Waller, A. (2007) The first fossil Compsocidae from Cretaceous Burmese amber (Insecta, Psocoptera, Troctomorpha). Cretaceous Research, 28, 1039 - 1041. https: // doi. org / 10.1016 / j. cretres. 2007.02.002" type="journal article" year="2007">Nel and Waller (2007)</bibRefCitation>
|
||
as present in
|
||
<taxonomicName id="4C034D14FFFFFFAE84A8F9F8FBBFE116" ID-CoL="NXLS" box="[984,1106,1608,1631]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="perreaui">
|
||
<emphasis id="B977EA85FFFFFFAE84A8F9F8FBBFE116" box="[984,1106,1608,1631]" italics="true" pageId="1" pageNumber="598">B. perreaui</emphasis>
|
||
</taxonomicName>
|
||
and missing in
|
||
<taxonomicName id="4C034D14FFFFFFAE8277F9F8FA78E116" ID-CoL="NXLR" authorityName="Nel" authorityYear="2017" box="[1287,1429,1608,1631]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans">
|
||
<emphasis id="B977EA85FFFFFFAE8277F9F8FA78E116" box="[1287,1429,1608,1631]" italics="true" pageId="1" pageNumber="598">B. coniugans</emphasis>
|
||
</taxonomicName>
|
||
. Moreover,
|
||
<bibRefCitation id="EF924B66FFFFFFAE867DF9DAFE13E1C8" author="Nel" box="[269,510,1642,1665]" pageId="1" pageNumber="598" refString="Nel, A. & Waller, A. (2007) The first fossil Compsocidae from Cretaceous Burmese amber (Insecta, Psocoptera, Troctomorpha). Cretaceous Research, 28, 1039 - 1041. https: // doi. org / 10.1016 / j. cretres. 2007.02.002" type="journal article" year="2007">Nel and Waller (2007)</bibRefCitation>
|
||
described lacinia tip of
|
||
<taxonomicName id="4C034D14FFFFFFAE8475F9DAFC92E1C8" ID-CoL="NXLS" box="[773,895,1642,1665]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="perreaui">
|
||
<emphasis id="B977EA85FFFFFFAE8475F9DAFC92E1C8" box="[773,895,1642,1665]" italics="true" pageId="1" pageNumber="598">B. perreaui</emphasis>
|
||
</taxonomicName>
|
||
bearing few indistinct rounded denticles, whereas
|
||
<taxonomicName id="4C034D14FFFFFFAE87E7F93DFECCE1ED" ID-CoL="NXLR" authorityName="Nel" authorityYear="2017" box="[151,289,1677,1700]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="coniugans">
|
||
<emphasis id="B977EA85FFFFFFAE87E7F93DFECCE1ED" box="[151,289,1677,1700]" italics="true" pageId="1" pageNumber="598">B. coniugans</emphasis>
|
||
</taxonomicName>
|
||
exhibits relatively pronounced denticles subapically on lacinia (
|
||
<figureCitation id="13382A12FFFFFFAE84A0F93DFBFCE1ED" box="[976,1041,1677,1700]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1877,1899]" captionTargetBox="[159,1428,194,1856]" captionTargetId="figure@2.[159,1428,194,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Burmacompsocus coniugans sp. nov.: (1) photograph of holotype (dorsal view); (2) photograph of holotype (ventral view); (3) drawing of holotype (dorsal view) with marked forewing venation pattern; (4) drawing of holotype (ventral view); (5) detail of mouthparts (ventral view); (6) detail of genitalia (ventral view); (7) tarsus of left hind leg; (8) detail of claw; (9) right hind wing, visible part; (10) left hind wing, visible part." httpUri="https://zenodo.org/record/893730/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC3697FFFFFFAE87B7F900FE65E018" blockId="1.[151,1436,429,1873]" pageId="1" pageNumber="598">
|
||
In addition to
|
||
<taxonomicName id="4C034D14FFFFFFAE862EF900FE3AE18E" ID-CoL="NXLS" box="[350,471,1712,1735]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="perreaui">
|
||
<emphasis id="B977EA85FFFFFFAE862EF900FE3AE18E" box="[350,471,1712,1735]" italics="true" pageId="1" pageNumber="598">B. perreaui</emphasis>
|
||
</taxonomicName>
|
||
, the genus also encompases
|
||
<taxonomicName id="4C034D14FFFFFFAE847CF900FB30E18E" ID-CoL="NXLQ" authority="(Cockerell, 1916)" authorityName="Cockerell" authorityYear="1916" baseAuthorityName="Cockerell" baseAuthorityYear="1916" box="[780,1245,1711,1735]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="banksi">
|
||
<emphasis id="B977EA85FFFFFFAE847CF900FBFBE18E" box="[780,1046,1711,1735]" italics="true" pageId="1" pageNumber="598">Burmacompsocus banksi</emphasis>
|
||
(Cockerell, 1916)
|
||
</taxonomicName>
|
||
. This species was only tentatively placed in the genus, based on its similar wing venation (
|
||
<bibRefCitation id="EF924B66FFFFFFAE84CBF962FB40E1A0" author="Mockford" box="[955,1197,1745,1769]" pageId="1" pageNumber="598" refString="Mockford, E. L., Lienhard, Ch. & Yoshizawa, K. (2013) Revised classification of " Psocoptera " from Cretaceous amber, a reassessment of published information. Insecta Matsumurana, 69, 1 - 26." type="journal article" year="2013">
|
||
Mockford
|
||
<emphasis id="B977EA85FFFFFFAE8341F962FB8EE1A0" box="[1073,1123,1745,1769]" italics="true" pageId="1" pageNumber="598">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
). This attribution was confirmed by
|
||
<bibRefCitation id="EF924B66FFFFFFAE865CF945FE00E045" box="[300,493,1780,1804]" pageId="1" pageNumber="598" refString="Azar, D., Hakim, M. & Huang, D. (2016) A new compsocid booklouse from the Cretaceous amber of Myanmar (Psocodea: Troctomorpha: Amphientometae: Compsocidae). Cretaceous Research, 68, 28 - 33. https: // doi. org / 10.1016 / j. cretres. 2016.08.003" type="journal article">
|
||
Azar
|
||
<emphasis id="B977EA85FFFFFFAE8617F945FE78E045" box="[359,405,1780,1804]" italics="true" pageId="1" pageNumber="598">et al</emphasis>
|
||
. (2016)
|
||
</bibRefCitation>
|
||
, however a resolution of the specific placement of
|
||
<taxonomicName id="4C034D14FFFFFFAE837EF945FB9CE045" ID-CoL="NXLQ" baseAuthorityName="Cockerell" baseAuthorityYear="1916" box="[1038,1137,1780,1804]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="banksi">
|
||
<emphasis id="B977EA85FFFFFFAE837EF945FB9CE045" box="[1038,1137,1780,1804]" italics="true" pageId="1" pageNumber="598">B. banksi</emphasis>
|
||
</taxonomicName>
|
||
was seen as impossible due to the fragmentary preservation of the fossil (
|
||
<bibRefCitation id="EF924B66FFFFFFAE850BF8A8FCC3E066" author="Azar" box="[635,814,1815,1839]" pageId="1" pageNumber="598" refString="Azar, D., Hakim, M. & Huang, D. (2016) A new compsocid booklouse from the Cretaceous amber of Myanmar (Psocodea: Troctomorpha: Amphientometae: Compsocidae). Cretaceous Research, 68, 28 - 33. https: // doi. org / 10.1016 / j. cretres. 2016.08.003" type="journal article" year="2016">
|
||
Azar
|
||
<emphasis id="B977EA85FFFFFFAE85C7F8A8FD0BE066" box="[695,742,1815,1839]" italics="true" pageId="1" pageNumber="598">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). We concur with this opinion and exclude
|
||
<taxonomicName id="4C034D14FFFFFFAE838DF8A8FA8CE066" ID-CoL="NXLQ" baseAuthorityName="Cockerell" baseAuthorityYear="1916" box="[1277,1377,1815,1839]" class="Insecta" family="Compsocidae" genus="Burmacompsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="banksi">
|
||
<emphasis id="B977EA85FFFFFFAE838DF8A8FA8CE066" box="[1277,1377,1815,1839]" italics="true" pageId="1" pageNumber="598">B. banksi</emphasis>
|
||
</taxonomicName>
|
||
from the comparative study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |