186 lines
55 KiB
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186 lines
55 KiB
XML
<document id="1E9F2C65F336A03D1B07B7E1A50D1D90" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. & Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFB3076A5637FE3E7DCAFC76" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Holthuija Ng & Forges 2015, n. gen." docType="treatment" docUuid="8DB52511-0A93-47A6-9382-6DFAF32A962C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="160" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="160" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3997581A9ED3EA7BAAAAB229C4251B75">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="88D5100B4940645E39D344AA8B5197EB">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="0925B3297DDCEA948E121CBD2FBEF5F3">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="75A7DDE9FBD5D88065C65F4AA4DCCD65">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="DFCC2DB36737BCD92349A46DFF556337">2015</mods:date>
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<mods:number id="DAC397DF1996A88ACFA3B48E5C78BF7C">63</mods:number>
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<treatment id="03A9654BFFB3076A5637FE3E7DCAFC76" ID-DOI="http://doi.org/10.5281/zenodo.5466807" ID-Zenodo-Dep="5466807" LSID="urn:lsid:zoobank.org:act:8DB52511-0A93-47A6-9382-6DFAF32A962C" httpUri="http://treatment.plazi.org/id/03A9654BFFB3076A5637FE3E7DCAFC76" lastPageNumber="160" pageId="50" pageNumber="160">
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<subSubSection id="C31A87D6FFB3076A5637FE3E7B73FE76" box="[336,529,445,469]" pageId="50" pageNumber="160" type="nomenclature">
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<paragraph id="8BBFD45DFFB3076A5637FE3E7B73FE76" blockId="50.[336,529,445,469]" box="[336,529,445,469]" pageId="50" pageNumber="160">
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<heading id="D0F76331FFB3076A5637FE3E7B73FE76" bold="true" box="[336,529,445,469]" centered="true" fontSize="10" level="2" pageId="50" pageNumber="160" reason="2">
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<emphasis id="B974084FFFB3076A5637FE3E7B73FE76" bold="true" box="[336,529,445,469]" pageId="50" pageNumber="160">
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<taxonomicName id="4C00AFDEFFB3076A5637FE3E78D9FE76" ID-CoL="7PKL9" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[336,443,445,469]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B974084FFFB3076A5637FE3E78D9FE76" bold="true" box="[336,443,445,469]" italics="true" pageId="50" pageNumber="160">Holthuija</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B534FFB3076A56A3FE3E7B73FE76" box="[452,529,445,469]" pageId="50" pageNumber="160" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C31A87D6FFB3076A5716FE7E7C1EFEB6" pageId="50" pageNumber="160" type="diagnosis">
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<paragraph id="8BBFD45DFFB3076A5716FE7E7C1EFEB6" blockId="50.[111,754,509,2069]" lastBlockId="50.[811,1452,189,277]" pageId="50" pageNumber="160">
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<emphasis id="B974084FFFB3076A5716FE7E7984FDB6" bold="true" box="[113,230,509,533]" pageId="50" pageNumber="160">Diagnosis.</emphasis>
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Carapace ovate; dorsal surface inflated, regions distinct, covered by granules or spines; gastric and branchial regions clearly delimited by grooves (
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<figureCitation id="133BC8D8FFB3076A569CFDBE7B1EFDF6" box="[507,636,573,597]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="54.[113,148,2005,2027]" captionStartId-1="55.[136,171,1961,1983]" captionStartId-2="56.[113,148,1208,1230]" captionTargetBox-0="[113,1451,561,1990]" captionTargetBox-1="[136,1474,234,1945]" captionTargetBox-2="[113,1451,190,1192]" captionTargetId-0="figure-15@54.[113,1451,561,1990]" captionTargetId-1="figure-8@55.[136,1474,232,1947]" captionTargetId-2="figure-15@56.[113,1451,190,1193]" captionTargetPageId-0="54" captionTargetPageId-1="55" captionTargetPageId-2="56" captionText-0="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." captionText-1="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." captionText-2="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi-0="http://doi.org/10.5281/zenodo.5384644" figureDoi-1="http://doi.org/10.5281/zenodo.5384646" figureDoi-2="http://doi.org/10.5281/zenodo.5384648" httpUri-0="https://zenodo.org/record/5384644/files/figure.png" httpUri-1="https://zenodo.org/record/5384646/files/figure.png" httpUri-2="https://zenodo.org/record/5384648/files/figure.png" pageId="50" pageNumber="160">Figs. 24–26</figureCitation>
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). Intestinal region without median spine (
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<figureCitation id="133BC8D8FFB3076A56D8FDDE7B25FDD6" box="[447,583,605,629]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="54.[113,148,2005,2027]" captionStartId-1="55.[136,171,1961,1983]" captionStartId-2="56.[113,148,1208,1230]" captionTargetBox-0="[113,1451,561,1990]" captionTargetBox-1="[136,1474,234,1945]" captionTargetBox-2="[113,1451,190,1192]" captionTargetId-0="figure-15@54.[113,1451,561,1990]" captionTargetId-1="figure-8@55.[136,1474,232,1947]" captionTargetId-2="figure-15@56.[113,1451,190,1193]" captionTargetPageId-0="54" captionTargetPageId-1="55" captionTargetPageId-2="56" captionText-0="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." captionText-1="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." captionText-2="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi-0="http://doi.org/10.5281/zenodo.5384644" figureDoi-1="http://doi.org/10.5281/zenodo.5384646" figureDoi-2="http://doi.org/10.5281/zenodo.5384648" httpUri-0="https://zenodo.org/record/5384644/files/figure.png" httpUri-1="https://zenodo.org/record/5384646/files/figure.png" httpUri-2="https://zenodo.org/record/5384648/files/figure.png" pageId="50" pageNumber="160">Figs. 24–26</figureCitation>
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). Pseudorostral spines relatively long, diverging, forming a V (
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<figureCitation id="133BC8D8FFB3076A55EBFDFE7B89FD36" box="[652,747,637,661]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Figs. 24</figureCitation>
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,
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<figureCitation id="133BC8D8FFB3076A5716FD1E79ECFD16" box="[113,142,669,693]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">25</figureCitation>
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,
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<figureCitation id="133BC8D8FFB3076A57FBFD1E7984FD16" box="[156,230,669,693]" captionStart="Fig" captionStartId="73.[135,170,1833,1855]" captionTargetBox="[138,1471,189,1817]" captionTargetId="figure-8@73.[138,1471,188,1818]" captionTargetPageId="73" captionText="Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384672" httpUri="https://zenodo.org/record/5384672/files/figure.png" pageId="50" pageNumber="160">37D–J</figureCitation>
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). Supraorbital eave large, anterior part strongly expanded, foliaceous, margin convex; spine; antorbital spine long, sharp (
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<figureCitation id="133BC8D8FFB3076A5788FD5E7805FD56" box="[239,359,733,757]" captionStart="Fig" captionStartId="73.[135,170,1833,1855]" captionTargetBox="[138,1471,189,1817]" captionTargetId="figure-8@73.[138,1471,188,1818]" captionTargetPageId="73" captionText="Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384672" httpUri="https://zenodo.org/record/5384672/files/figure.png" pageId="50" pageNumber="160">Fig. 37D–J</figureCitation>
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). Intercalated spine distinct, separated from supraorbital eave and postorbital spine by distinct gaps; postorbital spine strong, lobiform; hepatic region with 1 strong spine, much shorter than postorbital spine; 1 tubercle below (
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<figureCitation id="133BC8D8FFB3076A5644FCDE78C3FCD6" box="[291,417,861,885]" captionStart="Fig" captionStartId="73.[135,170,1833,1855]" captionTargetBox="[138,1471,189,1817]" captionTargetId="figure-8@73.[138,1471,188,1818]" captionTargetPageId="73" captionText="Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384672" httpUri="https://zenodo.org/record/5384672/files/figure.png" pageId="50" pageNumber="160">Fig. 37D–J</figureCitation>
|
||
). Lateral carapace margin with 3 large spines, 1 branchial spine (
|
||
<figureCitation id="133BC8D8FFB3076A5688FCFE7B2FFC36" box="[495,589,893,917]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Figs. 24</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A553CFCFE7B1BFC36" box="[603,633,893,917]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">25</figureCitation>
|
||
). Posterior carapace margin with 2 distinct median spines (
|
||
<figureCitation id="133BC8D8FFB3076A55EAFC1E7B89FC16" box="[653,747,925,949]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Figs. 24</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A5716FC3E79EDFC76" box="[113,143,957,981]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">25</figureCitation>
|
||
). Eyes relatively long, slender, with ovoid cornea (
|
||
<figureCitation id="133BC8D8FFB3076A55A2FC3E79D9FC56" captionStart="Fig" captionStartId="73.[135,170,1833,1855]" captionTargetBox="[138,1471,189,1817]" captionTargetId="figure-8@73.[138,1471,188,1818]" captionTargetPageId="73" captionText="Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384672" httpUri="https://zenodo.org/record/5384672/files/figure.png" pageId="50" pageNumber="160">Fig. 37D–J</figureCitation>
|
||
). Antennal flagellum short, slender. Basal antennal article longer than broad, rectangular; surface with several tubercles, with 2 spines distally; inner and outer lateral margins with low granules; proximal outer angle rounded; antero-external crested rim of antennular fossa overlaps distal part of basal antennal article by about a third of its width (
|
||
<figureCitation id="133BC8D8FFB3076A57A3FB1E782AFB16" box="[196,328,1181,1205]" captionStart="Fig" captionStartId="76.[113,148,1692,1714]" captionTargetBox="[113,1451,331,1676]" captionTargetId="figure-15@76.[113,1451,331,1677]" captionTargetPageId="76" captionText="Fig. 40. Antennae, antennules and epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; L, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; M, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384678" httpUri="https://zenodo.org/record/5384678/files/figure.png" pageId="50" pageNumber="160">Fig. 40D–I</figureCitation>
|
||
). Epistome slightly wider than long, anterior margin with 2 low lobes and granular extension below each; posterior margin composed of 4 rectangular plates separated by shallow fissures (
|
||
<figureCitation id="133BC8D8FFB3076A5562FB7E7BE8FAB6" box="[517,650,1277,1301]" captionStart="Fig" captionStartId="76.[113,148,1692,1714]" captionTargetBox="[113,1451,331,1676]" captionTargetId="figure-15@76.[113,1451,331,1677]" captionTargetPageId="76" captionText="Fig. 40. Antennae, antennules and epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; L, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; M, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384678" httpUri="https://zenodo.org/record/5384678/files/figure.png" pageId="50" pageNumber="160">Figs. 40D–I</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A55F1FB7E7B87FAB6" box="[662,741,1277,1301]" captionStart="Fig" captionStartId="78.[113,148,1966,1988]" captionTargetBox="[112,1451,654,1951]" captionTargetId="figure-15@78.[112,1451,652,1952]" captionTargetPageId="78" captionText="Fig. 42. Epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; C, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; E, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; F, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; G, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; H, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384682" httpUri="https://zenodo.org/record/5384682/files/figure.png" pageId="50" pageNumber="160">42C–E</figureCitation>
|
||
). Suborbital margin separated from basal antennal article and margin of postorbital tooth by fissures (
|
||
<figureCitation id="133BC8D8FFB3076A5544FABE7BC2FAF6" box="[547,672,1341,1365]" captionStart="Fig" captionStartId="76.[113,148,1692,1714]" captionTargetBox="[113,1451,331,1676]" captionTargetId="figure-15@76.[113,1451,331,1677]" captionTargetPageId="76" captionText="Fig. 40. Antennae, antennules and epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; L, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; M, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384678" httpUri="https://zenodo.org/record/5384678/files/figure.png" pageId="50" pageNumber="160">Fig. 40D–I</figureCitation>
|
||
). Outer surface of third maxilliped covered by short setae; ischium subrectangular, slightly longer than broad; postero-external angle of merus relatively broad, “inserted” into shallower concavity on outer margin of ischium; antero-internal part of ischium rounded, auriculiform (
|
||
<figureCitation id="133BC8D8FFB3076A5546FA5E7BC7FA56" box="[545,677,1501,1525]" captionStart="Fig" captionStartId="82.[113,148,1833,1855]" captionTargetBox="[170,1394,189,1818]" captionTargetId="figure-234@82.[170,1394,189,1819]" captionTargetPageId="82" captionText="Fig. 46. Third maxillipeds. A, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; B, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; C, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; D, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; E, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; F, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; G, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; H, Sakaija sakaii, male (9.0 × 6.6 mm) (NSMT-Cr 8094), Japan; I, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; J, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; K, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; L, Sakaija longispinosa n. sp., paratype ovigerous female (14.2 × 10.7 mm) (NMV J63197), Australia; M, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; N, Planaja plana n. gen. n. sp., paratype male (38.3 × 30.7 mm) (ZRC 2013.1370), Philippines; O, Ovimaja compressipes, holotype female (51.2 × 40.5 mm) (NHM 1860.15), China; P, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384690" httpUri="https://zenodo.org/record/5384690/files/figure.png" pageId="50" pageNumber="160">Fig. 46A–F</figureCitation>
|
||
). Male chelipeds relatively short in adult males, surfaces of merus and carpus almost smooth; carpus elongate; propodus of palm elongated, not inflated, curved, smooth, palm longer than fingers; fingers long, slender, gently curved, with small basal gape when closed (
|
||
<figureCitation id="133BC8D8FFB3076A5620F9FE78C1F936" box="[327,419,1661,1685]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Figs. 24</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A56D6F9FE78ACF936" box="[433,462,1661,1685]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">25</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A56BAF9FE7B47F936" box="[477,549,1661,1685]" captionStart="Fig" captionStartId="90.[113,148,1833,1855]" captionTargetBox="[151,1414,189,1818]" captionTargetId="figure-229@90.[151,1414,189,1821]" captionTargetPageId="90" captionText="Fig. 54. Chelae. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., paratype male (76.5 × 61.3 mm) (ZRC 2001.0577), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; G, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; L, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; M, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, holotype female (51.2 × 40.5 mm) (NHM 1860.15), China; P, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384710" httpUri="https://zenodo.org/record/5384710/files/figure.png" pageId="50" pageNumber="160">54D–I</figureCitation>
|
||
). Ambulatory legs relatively short, slender; merus without dorsal subdistal spine; dactylus elongate, curved, covered with long setae except for corneous tip (
|
||
<figureCitation id="133BC8D8FFB3076A5666F95E7839F956" box="[257,347,1757,1781]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Figs. 24</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A5601F95E78E1F956" box="[358,387,1757,1781]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">25</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A56E9F95E78BDF956" box="[398,479,1757,1781]" captionStart="Fig" captionStartId="92.[113,148,1891,1913]" captionTargetBox="[160,1404,189,1876]" captionTargetId="figure-189@92.[160,1406,189,1876]" captionTargetPageId="92" captionText="Fig. 56. First ambulatory dactyli and propodi. A, Paramaya spinigera, dried female (70.1 × 58.8 mm) (KPM NH4195), Japan; B, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; C, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; E, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan [figure laterally inverted]; F, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; G, Holthuija poorei, n. sp. paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; H, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; I, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines [figure laterally inverted]; J, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384714" httpUri="https://zenodo.org/record/5384714/files/figure.png" pageId="50" pageNumber="160">56C–G</figureCitation>
|
||
). Thoracic sternum wide; surfaces of somites 5–8 with numerous rounded tubercles and granules; sternites 3 and 4 with obliquely longitudinal depressions; margin between sternites 2 and 3 demarcated by deep notch; anterior margin of sterno-abdominal cavity forming complete rim (
|
||
<figureCitation id="133BC8D8FFB3076A5610F8FE789DF836" box="[375,511,1917,1941]" captionStart="Fig" captionStartId="86.[113,148,1750,1772]" captionTargetBox="[113,1452,388,1735]" captionTargetId="figure-138@86.[113,1452,388,1735]" captionTargetPageId="86" captionText="Fig. 50. Male anterior thoracic sternums and abdomens. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea." figureDoi="http://doi.org/10.5281/zenodo.5384702" httpUri="https://zenodo.org/record/5384702/files/figure.png" pageId="50" pageNumber="160">Figs. 50D–I</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A5569F8FE7B54F836" box="[526,566,1917,1941]" captionStart="Fig" captionStartId="88.[113,148,1712,1734]" captionTargetBox="[113,1452,369,1696]" captionTargetId="figure-170@88.[113,1452,369,1696]" captionTargetPageId="88" captionText="Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384706" httpUri="https://zenodo.org/record/5384706/files/figure.png" pageId="50" pageNumber="160">52I</figureCitation>
|
||
). Male abdomen subrectangular, with 6 free somites and telson; somites 3 and 4 subequal to or wider than somites 6 and telson (
|
||
<figureCitation id="133BC8D8FFB3076A55A2F83E79DEF856" captionStart="Fig" captionStartId="86.[113,148,1750,1772]" captionTargetBox="[113,1452,388,1735]" captionTargetId="figure-138@86.[113,1452,388,1735]" captionTargetPageId="86" captionText="Fig. 50. Male anterior thoracic sternums and abdomens. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea." figureDoi="http://doi.org/10.5281/zenodo.5384702" httpUri="https://zenodo.org/record/5384702/files/figure.png" pageId="50" pageNumber="160">Fig. 50D–I</figureCitation>
|
||
). Male press-button abdominal locking mechanism submedian in position on sterno-abdominal cavity (
|
||
<figureCitation id="133BC8D8FFB3076A55EBF87E7B86F7B6" box="[652,740,2045,2069]" captionStart="Fig" captionStartId="88.[113,148,1712,1734]" captionTargetBox="[113,1452,369,1696]" captionTargetId="figure-170@88.[113,1452,369,1696]" captionTargetPageId="88" captionText="Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384706" httpUri="https://zenodo.org/record/5384706/files/figure.png" pageId="50" pageNumber="160">Fig. 52I</figureCitation>
|
||
). Female abdomen dome-shaped, covering most of thoracic sternum. G1 long, relatively stout, almost straight or gently curved, distal part with scattered short setae (
|
||
<figureCitation id="133BC8D8FFB3076A527AFF7E7C0DFEB6" box="[1309,1391,253,277]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. A–C, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; F–H, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; I–K, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; L–O, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [A–F, I, L]; 0.5 mm [D, G, H, J, K, M–O]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="50" pageNumber="160">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A87D6FFB3076A544AFEBE7ACDFED6" pageId="50" pageNumber="160" type="type_taxon">
|
||
<paragraph id="8BBFD45DFFB3076A544AFEBE7ACDFED6" blockId="50.[813,1451,317,373]" pageId="50" pageNumber="160">
|
||
<emphasis id="B974084FFFB3076A544AFEBE7AB3FEF6" bold="true" box="[813,977,317,341]" pageId="50" pageNumber="160">
|
||
<typeStatus id="54BB6AFFFFB3076A544AFEBE7A08FEF6" box="[813,874,317,341]" pageId="50" pageNumber="160">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C00AFDEFFB3076A54BAFEBD7C43FEF6" ID-CoL="84FZ3" authority="Walker, 1887" authorityName="Walker" authorityYear="1887" box="[989,1313,317,341]" class="Malacostraca" family="Majidae" genus="Maia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="miersii">
|
||
<emphasis id="B974084FFFB3076A54BAFEBD7D16FEF6" box="[989,1140,318,341]" italics="true" pageId="50" pageNumber="160">Maia miersii</emphasis>
|
||
<bibRefCitation id="EF91A9ACFFB3076A5318FEBE7C43FEF6" author="Walker AO" box="[1151,1313,317,341]" pageId="50" pageNumber="160" pagination="107 - 117" refId="ref89616" refString="Walker AO (1887) Notes on a collection of Crustacea from Singapore. Journal of the Linnean Society of London, 20 (118): 107 - 117, pls. 6 - 9." type="journal article" year="1887">Walker, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
, by present designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A87D6FFB3076A544AFE1E7D55FD96" pageId="50" pageNumber="160" type="etymology">
|
||
<paragraph id="8BBFD45DFFB3076A544AFE1E7D55FD96" blockId="50.[812,1451,413,566]" pageId="50" pageNumber="160">
|
||
<emphasis id="B974084FFFB3076A544AFE1E7ADEFE16" bold="true" box="[813,956,413,437]" pageId="50" pageNumber="160">Etymology.</emphasis>
|
||
The name is derived from an arbitrary combination of the name Holthuis, with the genus name
|
||
<taxonomicName id="4C00AFDEFFB3076A544AFE5D7A07FE56" authorityName="Lamarck" authorityYear="1801" box="[813,869,478,501]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B974084FFFB3076A544AFE5D7A07FE56" box="[813,869,478,501]" italics="true" pageId="50" pageNumber="160">Maja</emphasis>
|
||
</taxonomicName>
|
||
. This honours the major contribution by the late Lipke B. Holthuis in resolving the convoluted nomenclature of the genus. Gender feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A87D6FFB3076A544AFDDE7DCAFC76" pageId="50" pageNumber="160" type="discussion">
|
||
<paragraph id="8BBFD45DFFB3076A544AFDDE7DCAFC76" blockId="50.[813,1452,605,981]" pageId="50" pageNumber="160">
|
||
<emphasis id="B974084FFFB3076A544AFDDE7AFCFDD6" bold="true" box="[813,926,605,629]" pageId="50" pageNumber="160">Remarks.</emphasis>
|
||
Six species are included in
|
||
<taxonomicName id="4C00AFDEFFB3076A538EFDDE7C31FDD6" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[1257,1363,605,629]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B974084FFFB3076A538EFDDE7C31FDD6" box="[1257,1363,605,629]" italics="true" pageId="50" pageNumber="160">Holthuija</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A5239FDDD7CCEFDD5" box="[1374,1452,606,630]" pageId="50" pageNumber="160" rank="genus">n. gen.</taxonomicNameLabel>
|
||
<taxonomicName id="4C00AFDEFFB3076A544AFDFD7D33FD36" authority="(Walker, 1887)" baseAuthorityName="Walker" baseAuthorityYear="1887" box="[813,1105,637,661]" class="Malacostraca" family="Majidae" genus="Maia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="miersii">
|
||
<emphasis id="B974084FFFB3076A544AFDFD7AFEFD36" box="[813,924,638,661]" italics="true" pageId="50" pageNumber="160">H. miersii</emphasis>
|
||
(
|
||
<bibRefCitation id="EF91A9ACFFB3076A54CAFDFE7D2AFD36" author="Walker AO" box="[941,1096,637,661]" pageId="50" pageNumber="160" pagination="107 - 117" refId="ref89616" refString="Walker AO (1887) Notes on a collection of Crustacea from Singapore. Journal of the Linnean Society of London, 20 (118): 107 - 117, pls. 6 - 9." type="journal article" year="1887">Walker, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C00AFDEFFB3076A5338FDFD7CC5FD36" authority="(Rathbun, 1916)" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[1119,1447,637,661]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="suluensis">
|
||
<emphasis id="B974084FFFB3076A5338FDFD7D84FD36" box="[1119,1254,637,661]" italics="true" pageId="50" pageNumber="160">H. suluensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF91A9ACFFB3076A539FFDFE7CFDFD36" author="Rathbun MJ" box="[1272,1439,637,661]" pageId="50" pageNumber="160" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer " Albatross, " 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C00AFDEFFB3076A544AFD1D7AE5FD16" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[813,903,669,693]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="pauli" status="sp. nov.">
|
||
<emphasis id="B974084FFFB3076A544AFD1D7AE5FD16" box="[813,903,669,693]" italics="true" pageId="50" pageNumber="160">H. pauli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A54F6FD1D7AACFD16" box="[913,974,670,693]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C00AFDEFFB3076A54BAFD1D7D34FD16" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[989,1110,670,693]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="cognata" status="sp. nov.">
|
||
<emphasis id="B974084FFFB3076A54BAFD1D7D34FD16" box="[989,1110,670,693]" italics="true" pageId="50" pageNumber="160">H. cognata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A5307FD1D7DFCFD16" box="[1120,1182,670,693]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C00AFDEFFB3076A53CBFD1D7C71FD16" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[1196,1299,670,693]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="aussie" status="sp. nov.">
|
||
<emphasis id="B974084FFFB3076A53CBFD1D7C71FD16" box="[1196,1299,670,693]" italics="true" pageId="50" pageNumber="160">H. aussie</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A527BFD1D7C3AFD16" box="[1308,1368,670,693]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C00AFDEFFB3076A52F6FD1D7A11FD76" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="poorei" status="sp. nov.">
|
||
<emphasis id="B974084FFFB3076A52F6FD1D7A11FD76" italics="true" pageId="50" pageNumber="160">H. poorei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A541BFD3D7ADAFD76" box="[892,952,702,725]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
These species are all easily distinguished by their prominently foliate supraorbital eave, which is the most expanded in all the genera treated here. All species also have a strongly granulated carapace surface, epistome, suborbital and pterygostomial regions, third maxilliped, thoracic sternum and abdomen. The suborbital tooth is also sharply separated from the postorbital tooth by a distinct fissure; and the G1 is generally simple, with a simple distal part and only scattered short setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |