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<document id="2E059BF5464CF5F38ADEF01DFCF084A8" ID-DOI="10.11646/zootaxa.5311.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8094233" ID-ZooBank="F32BB695-1524-495D-9109-62ECC079F367" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1688030177252" checkinUser="plazi" docAuthor="Matias, Rafael" docDate="2023" docId="03A687F84524275BFF4EA4A5FB7F36C0" docLanguage="en" docName="zootaxa.5311.2.3.pdf" docOrigin="Zootaxa 5311 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5311.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ephutomma Ashmead 1899" docType="treatment" docVersion="3" lastPageNumber="210" masterDocId="FF9FFF804523275CFFD9A432FF89322D" masterDocTitle="Sex-association experiments reveal the true identity of Physetopoda lampedusia (Invrea, 1957) (Hymenoptera, Mutillidae, Smicromyrmini)" masterLastPageNumber="231" masterPageNumber="203" pageNumber="210" updateTime="1688148932863" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="725DF6B95096785FE1521FE1832EB3F8">Sex-association experiments reveal the true identity of Physetopoda lampedusia (Invrea, 1957) (Hymenoptera, Mutillidae, Smicromyrmini)</mods:title>
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<mods:namePart id="80993F2EDC86312DB1C98AB1FB75DAF8">Matias, Rafael</mods:namePart>
<mods:nameIdentifier id="C72A8CCF56227F9B9ACCEF18A62C6C65" type="ORCID">0000-0002-5072-0538</mods:nameIdentifier>
<mods:affiliation id="D6C168B6AFEF7D23742CCBFA4987BDCE">MNHNC - Museu Nacional de História Natural e da Ciência, Universidade de Lisboa, 1250 - 102 Lisboa, Portugal &amp; cE 3 c - Centre for Ecology, Evolution and Environmental Change, Faculdade de Ciências, Universidade de Lisboa, 1749 - 016 Lisboa, Portugal. rfsmatias @ gmail. com; https: // orcid. org / 0000 - 0002 - 5072 - 0538</mods:affiliation>
<mods:nameIdentifier id="CF7F4EBCA1E80E4E744B5C3ECB6B9B9B" type="email">rfsmatias@gmail.com</mods:nameIdentifier>
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<mods:title id="BE1D7D5C526DF6EB13468E737C5AE46E">Zootaxa</mods:title>
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<mods:date id="B5E2E5B2EEAA2DBA79C3B3D40B050805">2023</mods:date>
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<mods:identifier id="F412FF6535C76188728DBD2707161CC7" type="DOI">10.11646/zootaxa.5311.2.3</mods:identifier>
<mods:identifier id="7F5889AD48B958CFA8E0E334BB7A0D7C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A687F84524275BFF4EA4A5FB7F36C0" LSID="urn:lsid:plazi:treatment:03A687F84524275BFF4EA4A5FB7F36C0" httpUri="http://treatment.plazi.org/id/03A687F84524275BFF4EA4A5FB7F36C0" lastPageNumber="210" pageId="7" pageNumber="210">
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<emphasis id="B97BEAFC4524275BFF4EA4A5FDC0329C" bold="true" box="[151,585,151,178]" pageId="7" pageNumber="210">
Genus
<taxonomicName id="4C0F4D6D4524275BFF37A4A5FDC0329C" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[238,585,151,178]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAFC4524275BFF37A4A5FE0D329F" bold="true" box="[238,388,151,178]" italics="true" pageId="7" pageNumber="210">Ephutomma</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFE53A4A5FDC0329C" author="Ashmead, W. H." box="[394,585,151,177]" pageId="7" pageNumber="210" pagination="45 - 60" refId="ref19011" refString="Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
</taxonomicName>
</emphasis>
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<treatmentCitationGroup id="AB1F11C04524275BFF4EA4EEFF79331C" pageId="7" pageNumber="210">
<treatmentCitation id="0AAE10FF4524275BFF4EA4EEFE5632DE" author="Ashmead, W. H." box="[151,479,219,244]" page="52" pageId="7" pageNumber="210" year="1899">
<taxonomicName id="4C0F4D6D4524275BFF4EA4EEFE5632DE" authority="Ashmead, 1899: 52" authorityName="Ashmead" authorityPageNumber="52" authorityYear="1899" box="[151,479,219,244]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAFC4524275BFF4EA4EEFE9832D9" box="[151,273,220,244]" italics="true" pageId="7" pageNumber="210">Ephutomma</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFECEA4EEFE5632DE" author="Ashmead, W. H." box="[279,479,219,244]" pageId="7" pageNumber="210" pagination="45 - 60" refId="ref19011" refString="Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899: 52</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, ♁ non
<specimenCount id="9D09FD674524275BFDF7A4EEFDCB32DE" box="[558,578,220,243]" pageId="7" pageNumber="210" type="female">♀</specimenCount>
;
<treatmentCitation id="0AAE10FF4524275BFD96A4EEFD7032D9" author="Mickel, C. E." box="[591,761,220,244]" page="32" pageId="7" pageNumber="210" year="1928">
<bibRefCitation id="EF9E4B1F4524275BFD96A4EEFD7032D9" author="Mickel, C. E." box="[591,761,220,244]" pageId="7" pageNumber="210" pagination="1 - 351" refId="ref21270" refString="Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. Bulletin of the United States National Museum, 143, 1 - 351. https: // doi. org / 10.5479 / si. 03629236.143.1" type="journal article" year="1928">Mickel 1928: 32</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE10FF4524275BFCDAA4EEFC4732D9" author="Skorikov, A. S." box="[771,974,219,244]" page="321" pageId="7" pageNumber="210" year="1935">
<bibRefCitation id="EF9E4B1F4524275BFCDAA4EEFC4732D9" author="Skorikov, A. S." box="[771,974,219,244]" pageId="7" pageNumber="210" pagination="257 - 349" refId="ref22251" refString="Skorikov, A. S. (1935) Zur Mutilliden-Fauna Zentralasiens. Trudy Tadzhikskoi Basy Akademii Nauk SSSR [Tajik Base of the Academy of Sciences of the USSR], 5, 257 - 349, 7 pls. [in Russian]" type="journal article" year="1935">Skorikov 1935: 321</bibRefCitation>
</treatmentCitation>
, ♁ non
<specimenCount id="9D09FD674524275BFBC6A4EEFBBA32DE" box="[1055,1075,220,243]" pageId="7" pageNumber="210" type="female">♀</specimenCount>
;
<treatmentCitation id="0AAE10FF4524275BFBE6A4EEFAD732D9" author="Lelej, A. S. &amp; Kabakov, O. N." box="[1087,1374,220,244]" page="192" pageId="7" pageNumber="210" year="1980">
<bibRefCitation id="EF9E4B1F4524275BFBE6A4EEFAD732D9" author="Lelej, A. S. &amp; Kabakov, O. N." box="[1087,1374,220,244]" pageId="7" pageNumber="210" pagination="141 - 156" refId="ref20602" refString="Lelej, A. S. &amp; Kabakov, O. N. (1980) [On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan]. Entomologicheskoe Obozrenie, 59 (1), 181 - 196, 14 figs. [in Russian, English translation in Entomological Review, 59, 141 - 156]" type="journal article" year="1980">Lelej &amp; Kabakov 1980: 192</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE10FF4524275BFAB1A4EEFEAD333F" author="Lelej, A. S." page="52" pageId="7" pageNumber="210" year="2002">
<bibRefCitation id="EF9E4B1F4524275BFAB1A4EEFEAD333F" author="Lelej, A. S." pageId="7" pageNumber="210" refId="ref20508" refString="Lelej, A. S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic region. Dalnauka, Vladivostok, 171 pp." type="book" year="2002">Lelej 2002: 52</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE10FF4524275BFEF6A4C8FDB8333F" author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Palmerini, M. M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." box="[303,561,250,274]" page="172" pageId="7" pageNumber="210" year="2020">
<bibRefCitation id="EF9E4B1F4524275BFEF6A4C8FDB8333F" author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Palmerini, M. M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." box="[303,561,250,274]" pageId="7" pageNumber="210" pagination="5 - 427" refId="ref21817" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Palmerini, M. M., Scaramozzino, P. L., Williams, K. A. &amp; Romano, M. (2020 [&quot; 2018 &quot;]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bolletino del Museo Regionale di Scienze Naturali, 36 (1 - 2), 5 - 427." type="journal article" year="2020">
Pagliano
<emphasis id="B97BEAFC4524275BFE57A4C9FE4B333F" box="[398,450,250,274]" italics="true" pageId="7" pageNumber="210">et al.</emphasis>
2020: 172
</bibRefCitation>
</treatmentCitation>
. Gender: neuter (Art. 30.1.2 of the Code (ICZN 1999),
<emphasis id="B97BEAFC4524275BFBBEA4C9FB20333F" box="[1127,1193,251,274]" italics="true" pageId="7" pageNumber="210">contra</emphasis>
<treatmentCitation id="0AAE10FF4524275BFB76A4C8FF6D331C" author="Lelej, A. S. &amp; Brothers, D." page="22" pageId="7" pageNumber="210" year="2008">
<bibRefCitation id="EF9E4B1F4524275BFB76A4C8FF6D331C" author="Lelej, A. S. &amp; Brothers, D." pageId="7" pageNumber="210" pagination="1 - 79" refId="ref20536" refString="Lelej, A. S. &amp; Brothers, D. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 1889.1.1" type="journal article" year="2008">Lelej &amp; Brothers 2008: 22</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB036EE4524275BFF4EA50AFE6B3362" blockId="7.[151,1437,219,397]" box="[151,482,311,336]" pageId="7" pageNumber="210">
<treatmentCitationGroup id="AB1F11C04524275BFF4EA50AFE6B3362" box="[151,482,311,336]" pageId="7" pageNumber="210">
<treatmentCitation id="0AAE10FF4524275BFF4EA50AFE573362" author="Suarez, F. J." box="[151,478,311,336]" page="51" pageId="7" pageNumber="210" year="1965">
<taxonomicName id="4C0F4D6D4524275BFF4EA50AFE573362" authority="Suarez, 1965 b: 51" authorityName="Suarez" authorityPageNumber="51" authorityYear="1965" box="[151,478,311,336]" class="Insecta" family="Mutillidae" genus="Eremomyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAFC4524275BFF4EA50AFE963362" box="[151,287,312,335]" italics="true" pageId="7" pageNumber="210">Eremomyrme</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFEFCA50AFE573362" author="Suarez, F. J." box="[293,478,311,336]" pageId="7" pageNumber="210" pagination="51 - 52" refId="ref22466" refString="Suarez, F. J. (1965 b) Nota sinonimica preliminar y propuesta del nombre generico Eremomyrme nom. nov. (Hymenoptera, Mutillidae). Memorie della Societa Entomologica Italiana, 44, 51 - 52." type="journal article" year="1965">Suárez, 1965b: 51</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31565654524275BFE31A50AFE7233A0" pageId="7" pageNumber="210" type="multiple">
<paragraph id="8BB036EE4524275BFE31A50AFE7233A0" blockId="7.[151,1437,219,397]" pageId="7" pageNumber="210">
<typeStatus id="54B4884C4524275BFE31A50AFD93337D" box="[488,538,312,336]" pageId="7" pageNumber="210">Type</typeStatus>
species
<taxonomicName id="4C0F4D6D4524275BFDB6A50AFC7F337D" authority="Dalla Torre, 1897" authorityName="Dalla Torre" authorityYear="1897" box="[623,1014,312,336]" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="turcestanica">
<emphasis id="B97BEAFC4524275BFDB6A50AFCB53362" box="[623,828,312,336]" italics="true" pageId="7" pageNumber="210">Mutilla turcestanica</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFC9BA50AFC7F337D" author="Dalla Torre, C. G. de" box="[834,1014,312,336]" pageId="7" pageNumber="210" refId="ref19570" refString="Dalla Torre, C. G. de (1897) Catalogus Hymenopterorum hucusque Descriptorum Systematicus et Synonimicus. Volumen VIII: Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], (4) + viii + 750 pp." type="book" year="1897">Dalla Torre, 1897</bibRefCitation>
</taxonomicName>
, ♁ (replacement name for
<taxonomicName id="4C0F4D6D4524275BFADDA50AFE143343" authority="Radoszkowski, 1877" authorityName="Radoszkowski" authorityYear="1877" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B97BEAFC4524275BFADDA50AFA153362" box="[1284,1436,312,336]" italics="true" pageId="7" pageNumber="210">Mutilla incerta</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFF1EA564FE143343" author="Radoszkowski, O." box="[199,413,342,366]" pageId="7" pageNumber="210" pagination="1 - 87" refId="ref22056" refString="Radoszkowski, O. (1877) Chrysidiformis, Mutillidae et Sphegidae. Voyage au Turkestan d'Alexis Fedtschenko, Series 14, 2 (5), 1 - 87. [in Russian]" type="journal article" year="1877">Radoszkowski, 1877</bibRefCitation>
</taxonomicName>
, ♁, by automatic designation. Junior objective synonym of
<taxonomicName id="4C0F4D6D4524275BFC23A564FA913343" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[1018,1304,342,366]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAFC4524275BFC23A564FBFD3343" box="[1018,1140,342,366]" italics="true" pageId="7" pageNumber="210">Ephutomma</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFBA0A564FA913343" author="Ashmead, W. H." box="[1145,1304,342,366]" pageId="7" pageNumber="210" pagination="45 - 60" refId="ref19011" refString="Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
</taxonomicName>
according to
<treatmentCitation id="0AAE10FF4524275BFF1EA547FE7E33A0" author="Lelej, A. S. &amp; Kabakov, O. N." box="[199,503,373,397]" page="193" pageId="7" pageNumber="210" year="1980">
<bibRefCitation id="EF9E4B1F4524275BFF1EA547FE7E33A0" author="Lelej, A. S. &amp; Kabakov, O. N." box="[199,503,373,397]" pageId="7" pageNumber="210" pagination="141 - 156" refId="ref20602" refString="Lelej, A. S. &amp; Kabakov, O. N. (1980) [On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan]. Entomologicheskoe Obozrenie, 59 (1), 181 - 196, 14 figs. [in Russian, English translation in Entomological Review, 59, 141 - 156]" type="journal article" year="1980">Lelej &amp; Kabakov (1980: 193)</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31565654524275BFF4EA589FDB233D4" pageId="7" pageNumber="210" type="type_taxon">
<paragraph id="8BB036EE4524275BFF4EA589FDB233D4" blockId="7.[151,1437,443,1262]" pageId="7" pageNumber="210">
<emphasis id="B97BEAFC4524275BFF4EA589FEA233F8" bold="true" box="[151,299,443,469]" pageId="7" pageNumber="210">
<typeStatus id="54B4884C4524275BFF4EA589FF5B33F8" box="[151,210,443,469]" pageId="7" pageNumber="210">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C0F4D6D4524275BFEE0A589FD5C33F8" authority="Radoszkowski, 1877" authorityName="Radoszkowski" authorityYear="1877" box="[313,725,443,469]" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B97BEAFC4524275BFEE0A589FE6B33F8" box="[313,482,443,469]" italics="true" pageId="7" pageNumber="210">Mutilla incerta</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFE30A589FD5C33F8" author="Radoszkowski, O." box="[489,725,443,469]" pageId="7" pageNumber="210" pagination="1 - 87" refId="ref22056" refString="Radoszkowski, O. (1877) Chrysidiformis, Mutillidae et Sphegidae. Voyage au Turkestan d'Alexis Fedtschenko, Series 14, 2 (5), 1 - 87. [in Russian]" type="journal article" year="1877">Radoszkowski, 1877</bibRefCitation>
</taxonomicName>
, nom. praeocc., non
<bibRefCitation id="EF9E4B1F4524275BFC62A589FBD933F8" author="Spinola, M." box="[955,1104,443,469]" pageId="7" pageNumber="210" pagination="85 - 149" refId="ref22330" refString="Spinola, M. (1841) Hymenopteres recueillis a Cayenne en 1839 par M. Leprieur, pharmacien de la Marine Royale. Annales de la Societe entomologique de France, 10, 85 - 149." type="journal article" year="1841">Spinola 1841</bibRefCitation>
(=
<taxonomicName id="4C0F4D6D4524275BFBAFA589FE9433D4" authority="Dalla Torre, 1897" authorityName="Dalla Torre" authorityYear="1897" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="turcestanica">
<emphasis id="B97BEAFC4524275BFBAFA589FAD133F8" box="[1142,1368,443,469]" italics="true" pageId="7" pageNumber="210">Mutilla turcestanica</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFAB9A589FE9433D4" author="Dalla Torre, C. G. de" pageId="7" pageNumber="210" refId="ref19570" refString="Dalla Torre, C. G. de (1897) Catalogus Hymenopterorum hucusque Descriptorum Systematicus et Synonimicus. Volumen VIII: Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], (4) + viii + 750 pp." type="book" year="1897">Dalla Torre, 1897</bibRefCitation>
</taxonomicName>
), by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C31565654524275BFF1EA631FAE531DC" pageId="7" pageNumber="210" type="diagnosis">
<paragraph id="8BB036EE4524275BFF1EA631FE303110" blockId="7.[151,1437,443,1262]" pageId="7" pageNumber="210">
<emphasis id="B97BEAFC4524275BFF1EA631FEB33030" bold="true" box="[199,314,515,541]" pageId="7" pageNumber="210">Diagnosis</emphasis>
: MALE. Most species with mandible broad and widened to the apex, excised beneath, apically tridentate, with a curved carina on the upper side and a large lobe basally on the lower edge; preapical inner tooth equals or is larger than apical tooth (
<taxonomicName id="4C0F4D6D4524275BFDE4A67EFD653048" authorityName=": Lelej" authorityYear="1985" box="[573,748,588,613]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="montarcense">
<emphasis id="B97BEAFC4524275BFDE4A67EFD653048" box="[573,748,588,613]" italics="true" pageId="7" pageNumber="210">E. montarcense</emphasis>
</taxonomicName>
differs in most of these characteristics, see below). Clypeus concave, with the anterior border medially protruding over which there are two denticles or tubercles. Flagellomere 1 is 1.5–1.6 times as long as its maximum width (except in
<taxonomicName id="4C0F4D6D4524275BFCF6A6A6FC553080" authorityName=": Lelej" authorityYear="1985" box="[815,988,660,685]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="montarcense">
<emphasis id="B97BEAFC4524275BFCF6A6A6FC553080" box="[815,988,660,685]" italics="true" pageId="7" pageNumber="210">E. montarcense</emphasis>
</taxonomicName>
). Felt lines present laterally only on T2. Genitalia with lamellar, long and narrow volsella (adapted from
<bibRefCitation id="EF9E4B1F4524275BFCA8A685FC6430FF" author="Lelej, A. S." box="[881,1005,695,722]" pageId="7" pageNumber="210" refId="ref20456" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighboring countries]. Nauka, Leningrad, 268 pp., 64 tabs. [in Russian]" type="book" year="1985">Lelej 1985</bibRefCitation>
,
<bibRefCitation id="EF9E4B1F4524275BFC23A685FBBE30FC" author="Lelej, A. S." box="[1018,1079,695,721]" pageId="7" pageNumber="210" refId="ref20508" refString="Lelej, A. S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic region. Dalnauka, Vladivostok, 171 pp." type="book" year="2002">2002</bibRefCitation>
). Similar to genus
<taxonomicName id="4C0F4D6D4524275BFAD5A685FA1430FC" authorityName="Schuster" authorityYear="1949" box="[1292,1437,695,721]" class="Insecta" family="Mutillidae" genus="Physetopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAFC4524275BFAD5A685FA1430FC" box="[1292,1437,695,721]" italics="true" pageId="7" pageNumber="210">Physetopoda</emphasis>
</taxonomicName>
differing in mandible structure: sub-apical teeth large, as large or larger than apical tooth (small in
<taxonomicName id="4C0F4D6D4524275BFB22A6E9FA0530D8" authorityName="Schuster" authorityYear="1949" box="[1275,1420,731,757]" class="Insecta" family="Mutillidae" genus="Physetopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAFC4524275BFB22A6E9FA0530D8" box="[1275,1420,731,757]" italics="true" pageId="7" pageNumber="210">Physetopoda</emphasis>
</taxonomicName>
), external basal tooth not well developed (large and protruding in
<taxonomicName id="4C0F4D6D4524275BFC57A6CDFB963134" authorityName="Schuster" authorityYear="1949" box="[910,1055,767,793]" class="Insecta" family="Mutillidae" genus="Physetopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAFC4524275BFC57A6CDFB963134" box="[910,1055,767,793]" italics="true" pageId="7" pageNumber="210">Physetopoda</emphasis>
</taxonomicName>
, except in
<taxonomicName id="4C0F4D6D4524275BFB7CA732FE243110" authority="Klug sensu Pagliano &amp; Agnoli 2018" authorityName="Klug sensu Pagliano &amp; Agnoli" authorityYear="2018" class="Insecta" family="Mutillidae" genus="Physetopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B97BEAFC4524275BFB7CA732FA993134" box="[1189,1296,767,793]" italics="true" pageId="7" pageNumber="210">P. pusilla</emphasis>
Klug
<emphasis id="B97BEAFC4524275BFA87A732FA153134" box="[1374,1436,768,793]" italics="true" pageId="7" pageNumber="210">sensu</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFF4EA711FE243110" author="Pagliano, G. &amp; Agnoli, G. L." box="[151,429,803,829]" pageId="7" pageNumber="210" pagination="127 - 137" refId="ref21749" refString="Pagliano, G. &amp; Agnoli, G. L. (2018) Note sugli Imenotteri Mutillidi dell'Italia meridionale con chiave dicotomica dei maschi delle specie mediterranee di Physetopoda Schuster, 1949 (Hymenoptera, Vespoidea: Mutillidae). Bollettino della Societa entomologica italiana, 150 (3), 127 - 137. https: // doi. org / 10.4081 / BollettinoSEI. 2018.127" type="journal article" year="2018">Pagliano &amp; Agnoli 2018</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB036EE4524275BFF1EA775FAE531DC" blockId="7.[151,1437,443,1262]" pageId="7" pageNumber="210">
FEMALE. Eyes elliptical, closely set; minimal distance between eyes
<emphasis id="B97BEAFC4524275BFC34A77AFB8E314C" box="[1005,1031,840,865]" italics="true" pageId="7" pageNumber="210">ca</emphasis>
. 1.1–1.3 times the longitudinal eye diameter. Clypeus with a basal process and bent upper part. Mesosoma sometimes narrow, with the head somewhat wider than pronotum. Pattern of metasomal pale pubescence formed by a large basal spot on T2 with an apical band medially expanded triangularly, and another band over T3. Scutellar scale not well developed. Pygidial area developed, with more or less parallel sides, not widened or narrowed basally (adapted from
<bibRefCitation id="EF9E4B1F4524275BFB44A7E5FA9131DF" author="Lelej, A. S." box="[1181,1304,983,1010]" pageId="7" pageNumber="210" refId="ref20456" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighboring countries]. Nauka, Leningrad, 268 pp., 64 tabs. [in Russian]" type="book" year="1985">Lelej 1985</bibRefCitation>
,
<bibRefCitation id="EF9E4B1F4524275BFAFAA7E5FAE931DC" author="Lelej, A. S." box="[1315,1376,983,1009]" pageId="7" pageNumber="210" refId="ref20508" refString="Lelej, A. S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic region. Dalnauka, Vladivostok, 171 pp." type="book" year="2002">2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31565654524275BFF1EA7C9FB7F36C0" pageId="7" pageNumber="210" type="multiple">
<paragraph id="8BB036EE4524275BFF1EA7C9FB7F36C0" blockId="7.[151,1437,443,1262]" pageId="7" pageNumber="210">
<emphasis id="B97BEAFC4524275BFF1EA7C9FE073638" bold="true" box="[199,398,1019,1045]" pageId="7" pageNumber="210">Included species</emphasis>
: seventeen, distributed throughout the Palaearctic (15 species:
<taxonomicName id="4C0F4D6D4524275BFBACA7CEFF523617" authority="(Skorikov, 1935)" baseAuthorityName="Skorikov" baseAuthorityYear="1935" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="angustatum">
<emphasis id="B97BEAFC4524275BFBACA7CEFA953638" box="[1141,1308,1020,1045]" italics="true" pageId="7" pageNumber="210">E. angustatum</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFAF6A7C9FF5B3617" author="Skorikov, A. S." pageId="7" pageNumber="210" pagination="257 - 349" refId="ref22251" refString="Skorikov, A. S. (1935) Zur Mutilliden-Fauna Zentralasiens. Trudy Tadzhikskoi Basy Akademii Nauk SSSR [Tajik Base of the Academy of Sciences of the USSR], 5, 257 - 349, 7 pls. [in Russian]" type="journal article" year="1935">Skorikov, 1935</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFF30A012FDAB3614" authority="(Lelej, 1976)" baseAuthorityName="Lelej" baseAuthorityYear="1976" box="[233,546,1055,1081]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="coronatum">
<emphasis id="B97BEAFC4524275BFF30A012FE0B3614" box="[233,386,1056,1081]" italics="true" pageId="7" pageNumber="210">E. coronatum</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFE4AA02DFD923614" author="Lelej, A. S." box="[403,539,1055,1081]" pageId="7" pageNumber="210" pagination="268 - 281" refId="ref20248" refString="Lelej, A. S. (1976) Addition to the fauna of velvet ants of Mongolia (Hymenoptera, Mutillidae). Insects of Mongolia, 4, 268 - 281. [in Russian]" type="journal article" year="1976">Lelej, 1976</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFDE9A012FCCF3617" authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[560,838,1055,1082]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="desertum">
<emphasis id="B97BEAFC4524275BFDE9A012FD303614" box="[560,697,1055,1081]" italics="true" pageId="7" pageNumber="210">E. desertum</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFD1BA02DFCCF3617" author="Lelej, A. S." box="[706,838,1055,1082]" pageId="7" pageNumber="210" refId="ref20456" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighboring countries]. Nauka, Leningrad, 268 pp., 64 tabs. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFC8DA012FB6F3614" authority="Ovchinnikov, 2002" authorityName="Ovchinnikov" authorityYear="2002" box="[852,1254,1055,1081]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="ferghanicum">
<emphasis id="B97BEAFC4524275BFC8DA012FB8B3614" box="[852,1026,1055,1081]" italics="true" pageId="7" pageNumber="210">E. ferghanicum</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFBD2A02DFB6F3614" author="Ovchinnikov, S. V." box="[1035,1254,1055,1081]" pageId="7" pageNumber="210" pagination="79 - 104" refId="ref21586" refString="Ovchinnikov, S. V. (2002) Materials to the fauna of velvet ants (Hymenoptera, Mutillidae) of Kyrghizia. Entomological Investigations in Kyrgystan, 22, 79 - 104. [in Russian]" type="journal article" year="2002">Ovchinnikov, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFB2DA012FEC83670" authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="gabesianum">
<emphasis id="B97BEAFC4524275BFB2DA012FA153614" box="[1268,1436,1055,1081]" italics="true" pageId="7" pageNumber="210">E. gabesianum</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFF4EA071FEC83670" author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[151,321,1091,1117]" pageId="7" pageNumber="210" refId="ref19240" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFE94A076FD203670" authority="Lelej &amp; Ullah" authorityName="Lelej &amp; Ullah" box="[333,681,1091,1117]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="himalayanum">
<emphasis id="B97BEAFC4524275BFE94A076FD8D3670" box="[333,516,1091,1117]" italics="true" pageId="7" pageNumber="210">E. himalayanum</emphasis>
Lelej &amp; Ullah
</taxonomicName>
<emphasis id="B97BEAFC4524275BFD68A076FD4E3670" box="[689,711,1092,1117]" italics="true" pageId="7" pageNumber="210">in</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFD16A071FC053670" author="Lelej, A. S. &amp; Ullah, M. &amp; Mahmood, K." box="[719,908,1091,1117]" pageId="7" pageNumber="210" pagination="53 - 60" refId="ref20661" refString="Lelej, A. S., Ullah, M. &amp; Mahmood, K. (2007) Additions to the knowledge of the Mutillidae (Hymenoptera) of Pakistan. Zootaxa, 1444 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 1444.1.4" type="journal article" year="2007">
Lelej
<emphasis id="B97BEAFC4524275BFCC9A076FCC03670" box="[784,841,1091,1117]" italics="true" pageId="7" pageNumber="210">et al.</emphasis>
2007
</bibRefCitation>
,
<taxonomicName id="4C0F4D6D4524275BFC4EA076FB3C3670" authority="(Lelej, 1976)" baseAuthorityName="Lelej" baseAuthorityYear="1976" box="[919,1205,1091,1117]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="kazenasi">
<emphasis id="B97BEAFC4524275BFC4EA076FB9E3670" box="[919,1047,1091,1117]" italics="true" pageId="7" pageNumber="210">E. kazenasi</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFBFEA071FB243670" author="Lelej, A. S." box="[1063,1197,1091,1117]" pageId="7" pageNumber="210" pagination="268 - 281" refId="ref20248" refString="Lelej, A. S. (1976) Addition to the fauna of velvet ants of Mongolia (Hymenoptera, Mutillidae). Insects of Mongolia, 4, 268 - 281. [in Russian]" type="journal article" year="1976">Lelej, 1976</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFB18A076FF5236AC" authority="(Lelej, 1977)" baseAuthorityName="Lelej" baseAuthorityYear="1977" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="kerzhneri">
<emphasis id="B97BEAFC4524275BFB18A076FAC23670" box="[1217,1355,1091,1117]" italics="true" pageId="7" pageNumber="210">E. kerzhneri</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFA83A071FF5A36AC" author="Lelej, A. S." pageId="7" pageNumber="210" pagination="526 - 536" refId="ref20286" refString="Lelej, A. S. (1977) New data on the velvet-ant fauna of Mongolia (Hymenoptera, Mutillidae). Insects of Mongolia, 5, 526 - 536. [in Russian]" type="journal article" year="1977">Lelej, 1977</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFF3FA05AFDE236AC" authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[230,619,1127,1153]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="mesopotamicum">
<emphasis id="B97BEAFC4524275BFF3FA05AFE3236AC" box="[230,443,1128,1153]" italics="true" pageId="7" pageNumber="210">E. mesopotamicum</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFE1BA055FDE236AC" author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[450,619,1127,1153]" pageId="7" pageNumber="210" refId="ref19240" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFDAFA05AFC0936AF" authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[630,896,1127,1154]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="minutum">
<emphasis id="B97BEAFC4524275BFDAFA05AFD7F36AC" box="[630,758,1128,1153]" italics="true" pageId="7" pageNumber="210">E. minutum</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFD24A055FC0936AF" author="Lelej, A. S." box="[765,896,1127,1154]" pageId="7" pageNumber="210" refId="ref20456" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighboring countries]. Nauka, Leningrad, 268 pp., 64 tabs. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFC53A05AFB5436AC" authority="(Suarez, 1974)" baseAuthorityName="Suarez" baseAuthorityYear="1974" box="[906,1245,1127,1153]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="mongolense">
<emphasis id="B97BEAFC4524275BFC53A05AFBA636AC" box="[906,1071,1127,1153]" italics="true" pageId="7" pageNumber="210">E. mongolense</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFBE7A055FB5C36AC" author="Suarez, F. J." box="[1086,1237,1127,1153]" pageId="7" pageNumber="210" pagination="5 - 19" refId="ref22507" refString="Suarez, F. J. (1974) Nuevas especies de Mutilidos recogidos por la Mision del Dr. Z. Kaszab en Mongolia, durante las campanas de 1963 a 1968 (Hymenoptera: Myrmosidae y Mutillidae). Archivos del Instituto de Aclimatacion, Almeria, 19, 5 - 19." type="journal article" year="1974">Suárez, 1974</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFB31A05AFA1C36AC" authorityName=": Lelej" authorityYear="1985" box="[1256,1429,1128,1153]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="montarcense">
<emphasis id="B97BEAFC4524275BFB31A05AFA1C36AC" box="[1256,1429,1128,1153]" italics="true" pageId="7" pageNumber="210">E. montarcense</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFF4EA0BEFE643688" authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[151,493,1163,1189]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="semirubrum">
<emphasis id="B97BEAFC4524275BFF4EA0BEFEB43688" box="[151,317,1163,1189]" italics="true" pageId="7" pageNumber="210">E. semirubrum</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFE9DA0B9FE643688" author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[324,493,1163,1189]" pageId="7" pageNumber="210" refId="ref19240" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D6D4524275BFE21A0BEFC1A3688" authority="(Dalla Torre, 1897)" baseAuthorityName="Dalla Torre" baseAuthorityYear="1897" box="[504,915,1163,1190]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="turcestanicum">
<emphasis id="B97BEAFC4524275BFE21A0BEFD3C3688" box="[504,693,1164,1189]" italics="true" pageId="7" pageNumber="210">E. turcestanicum</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFD1AA0B9FC023688" author="Dalla Torre, C. G. de" box="[707,907,1163,1190]" pageId="7" pageNumber="210" refId="ref19570" refString="Dalla Torre, C. G. de (1897) Catalogus Hymenopterorum hucusque Descriptorum Systematicus et Synonimicus. Volumen VIII: Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], (4) + viii + 750 pp." type="book" year="1897">Dalla Torre, 1897</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C0F4D6D4524275BFC13A0BEFB65368B" authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[970,1260,1163,1190]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="turgayense">
<emphasis id="B97BEAFC4524275BFC13A0BEFBEA3688" box="[970,1123,1164,1189]" italics="true" pageId="7" pageNumber="210">E. turgayense</emphasis>
<bibRefCitation id="EF9E4B1F4524275BFBB0A0B9FB65368B" author="Lelej, A. S." box="[1129,1260,1163,1190]" pageId="7" pageNumber="210" refId="ref20456" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighboring countries]. Nauka, Leningrad, 268 pp., 64 tabs. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
), Oriental (one:
<taxonomicName id="4C0F4D6D4524275BFF4EA082FE4D36E4" authority="(Turner, 1911))" baseAuthorityName="Turner" baseAuthorityYear="1911" box="[151,452,1199,1225]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="fletcheri">
<emphasis id="B97BEAFC4524275BFF4EA082FE9A36E4" box="[151,275,1199,1225]" italics="true" pageId="7" pageNumber="210">E. fletcheri</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFEF8A09DFE3B36E4" author="Turner, R. E." box="[289,434,1199,1225]" pageId="7" pageNumber="210" pagination="141 - 159" refId="ref22652" refString="Turner, R. E. (1911) New Hymenoptera from Ceylon. Mutillidae and Scoliidae. Spolia zeylanica, 7, 141 - 159." type="journal article" year="1911">Turner, 1911</bibRefCitation>
))
</taxonomicName>
and Afrotropical (one:
<taxonomicName id="4C0F4D6D4524275BFD13A082FBA236E4" authority="(Magretti, 1892))" baseAuthorityName="Magretti" baseAuthorityYear="1892" box="[714,1067,1199,1225]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="somalicum">
<emphasis id="B97BEAFC4524275BFD13A082FCEB36E4" box="[714,866,1199,1225]" italics="true" pageId="7" pageNumber="210">E. somalicum</emphasis>
(
<bibRefCitation id="EF9E4B1F4524275BFCA8A09DFB9036E4" author="Magretti, P." box="[881,1049,1199,1225]" pageId="7" pageNumber="210" pagination="950 - 960" refId="ref21010" refString="Magretti, P. (1892) Di alcune specie d'Imenotteri raccolte dall' Ing. L. Bricchetti Robecchi nel paese dei Somali. Annali del Museo civico di storia naturale di Genova, Series 2, 10, 950 - 960." type="journal article" year="1892">Magretti, 1892</bibRefCitation>
))
</taxonomicName>
regions; two occur in Europe:
<taxonomicName id="4C0F4D6D4524275BFA5DA082FE9336C0" baseAuthorityName="Skorikov" baseAuthorityYear="1935" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="angustatum">
<emphasis id="B97BEAFC4524275BFA5DA082FE9336C0" italics="true" pageId="7" pageNumber="210">E. angustatum</emphasis>
</taxonomicName>
in European
<collectingCountry id="F318767E4524275BFE69A0E1FE7736C0" box="[432,510,1235,1261]" name="Russia" pageId="7" pageNumber="210">Russia</collectingCountry>
, and
<taxonomicName id="4C0F4D6D4524275BFDE1A0E6FD6C36C0" authorityName=": Lelej" authorityYear="1985" box="[568,741,1236,1261]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="montarcense">
<emphasis id="B97BEAFC4524275BFDE1A0E6FD6C36C0" box="[568,741,1236,1261]" italics="true" pageId="7" pageNumber="210">E. montarcense</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9E4B1F4524275BFD2DA0E1FC5136C0" author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Palmerini, M. M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." box="[756,984,1235,1261]" pageId="7" pageNumber="210" pagination="5 - 427" refId="ref21817" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Palmerini, M. M., Scaramozzino, P. L., Williams, K. A. &amp; Romano, M. (2020 [&quot; 2018 &quot;]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bolletino del Museo Regionale di Scienze Naturali, 36 (1 - 2), 5 - 427." type="journal article" year="2020">
Pagliano
<emphasis id="B97BEAFC4524275BFC87A0E6FC1E36C0" box="[862,919,1235,1261]" italics="true" pageId="7" pageNumber="210">et al.</emphasis>
2020
</bibRefCitation>
,
<bibRefCitation id="EF9E4B1F4524275BFC3AA0E1FB6336C0" author="Lelej, A. S. &amp; Williams, K. A." box="[995,1258,1235,1261]" pageId="7" pageNumber="210" pagination="101 - 140" refId="ref20716" refString="Lelej, A. S. &amp; Williams, K. A. (2023) Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae). Zootaxa, 5231 (2), 101 - 140. https: // doi. org / 10.11646 / zootaxa. 5231.2.1" type="journal article" year="2023">Lelej &amp; Williams 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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