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<document id="018E34B44241F2E9CC9E7F9FFF8F3CDB" ID="10.11646/zootaxa.4126.3.1" ID-CLB-Dataset="38782" ID-DOI="10.11646/zootaxa.4126.3.1" ID-GBIF-Dataset="1a5e6006-4597-44a3-93a6-4a082b070c7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="262862" ID-ZooBank="F0A10548-915F-4125-9E55-7569E07CCBAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466403114188" checkinUser="plazi" docAuthor="Enushchenko, Ilya V. &amp; Semenov, Viktor B." docDate="2016" docId="03A687EBFFD89624FF059F52FB45FDE6" docLanguage="en" docName="zootaxa.4126.3.1.pdf" docOrigin="Zootaxa 4126 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gyrophaena (Gyrophaena) nana Paykull 1800" docType="treatment" docVersion="9" lastPageNumber="324" masterDocId="FF9FFF93FFCC9633FF92986DFFCEFFCA" masterDocTitle="A review of the genus Gyrophaena Mannerheim 1830 (Coleoptera: Staphylinidae: Aleocharinae: Gyrophaenina) of the Caucasus and adjacent territories" masterLastPageNumber="337" masterPageNumber="301" pageNumber="321" updateTime="1698663063067" updateUser="ExternalLinkService">
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<mods:title id="A43918DB4D92D9EF565A64C9507A3E6A">A review of the genus Gyrophaena Mannerheim 1830 (Coleoptera: Staphylinidae: Aleocharinae: Gyrophaenina) of the Caucasus and adjacent territories</mods:title>
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<mods:namePart id="A1654DA4729F50FC71ED0D7AA02D4CFA">Enushchenko, Ilya V.</mods:namePart>
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<mods:namePart id="6597670E50889AF4A821563076F17FD5">Semenov, Viktor B.</mods:namePart>
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<mods:date id="3A0A6E52AA6FADDF66AB04BDC5CD4143">2016</mods:date>
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<treatment id="03A687EBFFD89624FF059F52FB45FDE6" ID-DOI="http://doi.org/10.5281/zenodo.6057796" ID-GBIF-Taxon="120828941" ID-Zenodo-Dep="6057796" LSID="urn:lsid:plazi:treatment:03A687EBFFD89624FF059F52FB45FDE6" httpUri="http://treatment.plazi.org/id/03A687EBFFD89624FF059F52FB45FDE6" lastPageId="23" lastPageNumber="324" pageId="20" pageNumber="321">
<subSubSection id="C3156576FFD89627FF059F52FEE0F8B1" pageId="20" pageNumber="321" type="nomenclature">
<paragraph id="8BB036FDFFD89627FF059F52FD28F890" blockId="20.[151,742,1855,1915]" box="[151,742,1855,1882]" pageId="20" pageNumber="321">
<heading id="D0F88191FFD89627FF059F52FD28F890" bold="true" box="[151,742,1855,1882]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
<taxonomicName id="4C0F4D7EFFD89627FF059F52FD28F890" ID-CoL="3J24Q" authority="Paykull, 1800" authorityName="Paykull" authorityYear="1800" box="[151,742,1855,1882]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="nana" subGenus="Gyrophaena">
<emphasis id="B97BEAEFFFD89627FF059F52FD28F890" bold="true" box="[151,742,1855,1882]" pageId="20" pageNumber="321">
<emphasis id="B97BEAEFFFD89627FF059F52FEE3F893" bold="true" box="[151,301,1855,1881]" italics="true" pageId="20" pageNumber="321">Gyrophaena</emphasis>
(
<emphasis id="B97BEAEFFFD89627FEAF9F52FE1DF893" bold="true" box="[317,467,1855,1881]" italics="true" pageId="20" pageNumber="321">Gyrophaena</emphasis>
)
<emphasis id="B97BEAEFFFD89627FE769F52FDECF893" bold="true" box="[484,546,1855,1881]" italics="true" pageId="20" pageNumber="321">nana</emphasis>
(
<bibRefCitation id="EF9E4B0CFFD89627FDA09F2DFD10F890" author="Paykull" box="[562,734,1856,1882]" pageId="20" pageNumber="321" refString="Paykull, G. de (1800) Fauna Suecica. Insecta. Tomus III. Upsaliae (Uppsala), J. F. Edman, 459 pp." type="book" year="1800">Paykull, 1800</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB036FDFFD89627FF059F0FFEE0F8B1" blockId="20.[151,742,1855,1915]" box="[151,302,1890,1915]" pageId="20" pageNumber="321">(Figs. 46–48)</paragraph>
</subSubSection>
<subSubSection id="C3156576FFD89624FF059FC7FCE2FF21" lastPageId="23" lastPageNumber="324" pageId="20" pageNumber="321" type="reference_group">
<paragraph id="8BB036FDFFD89627FF059FC7FDDBF80A" blockId="20.[151,579,1962,2015]" box="[151,533,1962,1984]" pageId="20" pageNumber="321">
<treatmentCitationGroup id="AB1F11D3FFD89627FF059FC7FDDBF80A" box="[151,533,1962,1984]" pageId="20" pageNumber="321">
<taxonomicName id="4C0F4D7EFFD89627FF059FC7FDDBF80A" ID-CoL="4ZK7M" authority="Paykull, 1800: 408" authorityName="Paykull" authorityPageNumber="408" authorityYear="1800" box="[151,533,1962,1984]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B97BEAEFFFD89627FF059FC7FE81F80A" box="[151,335,1962,1984]" italics="true" pageId="20" pageNumber="321">Staphylinus nanus</emphasis>
<treatmentCitation id="0AAE10ECFFD89627FEC79FC7FDDBF80A" author="Paykull" box="[341,533,1962,1984]" page="408" pageId="20" pageNumber="321" year="1800">
<bibRefCitation id="EF9E4B0CFFD89627FEC79FC7FE28F80A" author="Paykull" box="[341,486,1962,1984]" pageId="20" pageNumber="321" refString="Paykull, G. de (1800) Fauna Suecica. Insecta. Tomus III. Upsaliae (Uppsala), J. F. Edman, 459 pp." type="book" year="1800">Paykull, 1800</bibRefCitation>
: 408
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB036FDFFD89626FF059FA4FE9DF87C" blockId="20.[151,579,1962,2015]" lastBlockId="21.[151,339,1953,1976]" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321">
<treatmentCitationGroup id="AB1F11D3FFD89626FF059FA4FE9DF87C" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321">
<taxonomicName id="4C0F4D7EFFD89627FF059FA4FD8DF815" ID-CoL="5TVKV" authority="Stephens, 1832: 153" authorityName="Stephens" authorityPageNumber="153" authorityYear="1832" box="[151,579,1993,2015]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B97BEAEFFFD89627FF059FA4FEA1F815" box="[151,367,1993,2015]" italics="true" pageId="20" pageNumber="321">Aleochara marginata</emphasis>
<treatmentCitation id="0AAE10ECFFD89627FEE79FA4FD8DF815" author="Stephens" box="[373,579,1993,2015]" page="153" pageId="20" pageNumber="321" year="1832">
<bibRefCitation id="EF9E4B0CFFD89627FEE79FA4FDDDF815" author="Stephens" box="[373,531,1993,2015]" pageId="20" pageNumber="321" refString="Stephens, J. F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, 5, 448 pp. [Baldwin &amp; Cradock, London]" type="book" year="1832">Stephens, 1832</bibRefCitation>
: 153
</treatmentCitation>
</taxonomicName>
<emphasis id="B97BEAEFFFD99626FF059FCCFE9DF87C" bold="true" box="[151,339,1953,1976]" pageId="21" pageNumber="322">FIGURES 40–45.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB036FDFFDA9625FF059FCCFE9DF87C" blockId="22.[151,339,1953,1976]" box="[151,339,1953,1976]" pageId="22" pageNumber="323">
<emphasis id="B97BEAEFFFDA9625FF059FCCFE9DF87C" bold="true" box="[151,339,1953,1976]" pageId="22" pageNumber="323">FIGURES 46–51.</emphasis>
</paragraph>
<paragraph id="8BB036FDFFDB9624FF0598F5FD4AFF64" blockId="23.[151,812,152,235]" box="[151,644,152,174]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB1F11D3FFDB9624FF0598F5FD4AFF64" box="[151,644,152,174]" pageId="23" pageNumber="324">
<taxonomicName id="4C0F4D7EFFDB9624FF0598F5FD4AFF64" ID-CoL="6L96M" authority="A. Fleischer, 1909: 332" authorityName="A. Fleischer" authorityPageNumber="332" authorityYear="1909" box="[151,644,152,174]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="B97BEAEFFFDB9624FF0598F5FE5FFF64" box="[151,401,152,174]" italics="true" pageId="23" pageNumber="324">Gyrophaena nigriventris</emphasis>
<treatmentCitation id="0AAE10ECFFDB9624FE0598F5FD4AFF64" author="Fleischer" box="[407,644,152,174]" page="332" pageId="23" pageNumber="324" year="1909">
<bibRefCitation id="EF9E4B0CFFDB9624FE0598F5FD9BFF64" author="Fleischer" box="[407,597,152,174]" pageId="23" pageNumber="324" refString="Fleischer, A. (1909) Eine neue Varietat der Gyrophaena nana Payk. Wiener Entomologische Zeitung, 28, 322." type="journal article" year="1909">A. Fleischer, 1909</bibRefCitation>
: 332
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB036FDFFDB9624FF0598DBFDE3FF06" blockId="23.[151,812,152,235]" box="[151,557,182,204]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB1F11D3FFDB9624FF0598DBFDE3FF06" box="[151,557,182,204]" pageId="23" pageNumber="324">
<taxonomicName id="4C0F4D7EFFDB9624FF0598DBFDE3FF06" ID-CoL="6L96G" authority="Casey, 1906: 301" authorityName="Casey" authorityPageNumber="301" authorityYear="1906" box="[151,557,182,204]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="perpolita">
<emphasis id="B97BEAEFFFDB9624FF0598DBFEB6FF06" box="[151,376,182,204]" italics="true" pageId="23" pageNumber="324">Gyrophaena perpolita</emphasis>
<treatmentCitation id="0AAE10ECFFDB9624FEEC98DBFDE3FF06" author="Casey" box="[382,557,182,204]" page="301" pageId="23" pageNumber="324" year="1906">
<bibRefCitation id="EF9E4B0CFFDB9624FEEC98DBFE33FF06" author="Casey" box="[382,509,182,204]" pageId="23" pageNumber="324" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
: 301
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB036FDFFDB9624FF0598B8FCE2FF21" blockId="23.[151,812,152,235]" box="[151,812,213,235]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB1F11D3FFDB9624FF0598B8FCE2FF21" box="[151,812,213,235]" pageId="23" pageNumber="324">
<taxonomicName id="4C0F4D7EFFDB9624FF0598B8FE80FF21" ID-CoL="3J24Q" box="[151,334,213,235]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="B97BEAEFFFDB9624FF0598B8FE80FF21" box="[151,334,213,235]" italics="true" pageId="23" pageNumber="324">Gyrophaena nana</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AAE10ECFFDB9624FEC898B8FDD7FF21" author="Roubal" box="[346,537,213,235]" page="484" pageId="23" pageNumber="324" year="1913">
<bibRefCitation id="EF9E4B0CFFDB9624FEC898B8FE29FF21" author="Roubal" box="[346,487,213,235]" pageId="23" pageNumber="324" refString="Roubal, J. (1913) Przyczynek do fauny rodziny kusokrywkow (Staphylinidae) polnocnego Kaukazu (Beitrag zur Staphylinenfauna des nordlichen Kaukasus). Kosmos, 38, 477 - 487." type="journal article" year="1913">Roubal, 1913</bibRefCitation>
: 484
</treatmentCitation>
;
<treatmentCitation id="0AAE10ECFFDB9624FDB198B8FCE2FF21" author="Nikitskiy" box="[547,812,213,235]" page="113" pageId="23" pageNumber="324" year="2008">
<bibRefCitation id="EF9E4B0CFFDB9624FDB198B8FD33FF21" author="Nikitskiy" box="[547,765,213,235]" pageId="23" pageNumber="324" refString="Nikitskiy, N. B., Bibin, A. R. &amp; Dolgin, M. M. (2008) Ksilotrofnye zhestkokrylye (Coleoptera) Kavkazskogo gosudarstvennogo prirodnogo biosfernogo zapovednika i sopredelnykh territoriy [Xylophilous beetles (Coleoptera) of the Caucasian State Biospheric Natural Reserve and adjacent territories]. Syktyvkar, 452 pp. [in Russian]" type="book" year="2008">Nikitskiy &amp; al., 2008</bibRefCitation>
: 113
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3156576FFDB9624FF05997AFDE6FE0A" pageId="23" pageNumber="324" type="materials_examined">
<paragraph id="8BB036FDFFDB9624FF05997AFDE6FE0A" blockId="23.[151,1436,279,557]" pageId="23" pageNumber="324">
<emphasis id="B97BEAEFFFDB9624FF05997AFEB6FEFA" bold="true" box="[151,376,279,304]" pageId="23" pageNumber="324">Material examined</emphasis>
:
<emphasis id="B97BEAEFFFDB9624FE15997AFDA2FEFA" bold="true" box="[391,620,279,304]" pageId="23" pageNumber="324">
<collectingCountry id="F318766DFFDB9624FE15997AFE3CFEFA" box="[391,498,279,304]" name="Russia" pageId="23" pageNumber="324">RUSSIA</collectingCountry>
: ADYGEA:
</emphasis>
Maikopskiy District: 1 Ƌ, 2 ♀: Caucasian Biosphere Reserve, Guzeripl´,
<date id="FFB1103DFFDB9624FF059951FED6FE9E" box="[151,280,316,340]" pageId="23" pageNumber="324" value="1990-06-21">21.VI.1990</date>
, V.V. Grebennikov leg. (cVS,
<collectionCode id="ED1EAE38FFDB9624FD129951FD06FE9E" box="[640,712,316,340]" pageId="23" pageNumber="324">ZMM</collectionCode>
); 1 Ƌ: Guzeripl´, Abago Mountain,
<quantity id="4CF79B18FFDB9624FBE09951FB03FE9E" box="[1138,1229,316,340]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="324" unit="m" value="1800.0">1800 m</quantity>
, window trap,
<date id="FFB1103DFFDB9624FA129951FED7FEB2" pageId="23" pageNumber="324" value="2000-06-01" valueMax="2000-06-22" valueMin="2000-06-01">1– 22.VI.2000</date>
, A.R. Bibin leg. (cVS); 1 Ƌ, 2 ♀: ibid., window trap,
<date id="FFB1103DFFDB9624FCFF990DFB88FEB2" box="[877,1094,352,376]" pageId="23" pageNumber="324" value="2000-06-22" valueMax="2000-07-14" valueMin="2000-06-22">22.VI–14.VII.2000</date>
, A.R. Bibin leg. (cVS,
<collectionCode id="ED1EAE38FFDB9624FADA990DFA5EFEB2" box="[1352,1424,352,376]" pageId="23" pageNumber="324">ZMM</collectionCode>
); 1 ♀: ibid.,
<quantity id="4CF79B18FFDB9624FE8599E9FEBFFE56" box="[279,369,388,412]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="324" unit="m" value="1400.0">1400 m</quantity>
, window trap,
<date id="FFB1103DFFDB9624FD8D99E9FD73FE57" box="[543,701,388,413]" pageId="23" pageNumber="324" value="2001-06-09" valueMax="2001-06-26" valueMin="2001-06-09">9–26.VI.2001</date>
, A.R. Bibin leg. (cVS);
<emphasis id="B97BEAEFFFDB9624FC4599E9FB2FFE57" bold="true" box="[983,1249,388,413]" pageId="23" pageNumber="324">
<collectingCountry id="F318766DFFDB9624FC4599E9FBADFE57" box="[983,1123,388,413]" name="Georgia" pageId="23" pageNumber="324">GEORGIA</collectingCountry>
: SVANETI:
</emphasis>
1 Ƌ: ‘Caucasus Swanetien Leder Reitter’ (
<collectionCode id="ED1EAE38FFDB9624FE5299C5FDD2FE0A" LSID="urn:lsid:biocol.org:col:34991" box="[448,540,424,448]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="23" pageNumber="324">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3156576FFDB9624FF5599A1FB19FE2E" box="[199,1239,460,485]" pageId="23" pageNumber="324" type="distribution">
<paragraph id="8BB036FDFFDB9624FF5599A1FB19FE2E" blockId="23.[151,1436,279,557]" box="[199,1239,460,485]" pageId="23" pageNumber="324">
<emphasis id="B97BEAEFFFDB9624FF5599A1FEAEFE2F" bold="true" box="[199,352,460,485]" pageId="23" pageNumber="324">Distribution.</emphasis>
Europe, Caucasus, Far East, N
<collectingCountry id="F318766DFFDB9624FD5199A1FCEAFE2F" box="[707,804,460,485]" name="United States of America" pageId="23" pageNumber="324">America</collectingCountry>
(
<bibRefCitation id="EF9E4B0CFFDB9624FCA199A1FC03FE2F" author="Seevers" box="[819,973,460,485]" pageId="23" pageNumber="324" refString="Seevers, C. H. (1951) A revision of the North American and European staphylinid beetles of the subtribe Gyrophaenina (Aleocharinae, Bolitocharini). Fieldiana: Zoology, 32 (10), 1 - 762." type="journal article" year="1951">Seevers 1951</bibRefCitation>
),
<collectingCountry id="F318766DFFDB9624FC7399A1FBFFFE2F" box="[993,1073,460,485]" name="Turkey" pageId="23" pageNumber="324">Turkey</collectingCountry>
(
<bibRefCitation id="EF9E4B0CFFDB9624FBD299A1FB04FE2E" author="Korge" box="[1088,1226,460,484]" pageId="23" pageNumber="324" refString="Korge, H. (1971) Beitrage zur Kenntnis der Koleopterenfauna Kleinasiens. Annotationes Zoologicae et Botanicae, 67: 1 - 68." type="journal article" year="1971">Korge 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3156576FFDB9624FF559982FB45FDE6" pageId="23" pageNumber="324" type="discussion">
<paragraph id="8BB036FDFFDB9624FF559982FB45FDE6" blockId="23.[151,1436,279,557]" pageId="23" pageNumber="324">
<emphasis id="B97BEAEFFFDB9624FF559982FEF5FDC2" bold="true" box="[199,315,495,520]" pageId="23" pageNumber="324">Remarks.</emphasis>
The first record of
<taxonomicName id="4C0F4D7EFFDB9624FD8C999DFDB8FDC2" box="[542,630,496,520]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="B97BEAEFFFDB9624FD8C999DFDB8FDC2" box="[542,630,496,520]" italics="true" pageId="23" pageNumber="324">G. nana</emphasis>
</taxonomicName>
for Caucasus (Karachay-Cherkessia, Teberda River) was published by
<bibRefCitation id="EF9E4B0CFFDB9624FF059A79FEF3FDE6" author="Roubal" box="[151,317,532,557]" pageId="23" pageNumber="324" refString="Roubal, J. (1913) Przyczynek do fauny rodziny kusokrywkow (Staphylinidae) polnocnego Kaukazu (Beitrag zur Staphylinenfauna des nordlichen Kaukasus). Kosmos, 38, 477 - 487." type="journal article" year="1913">Roubal (1913)</bibRefCitation>
. It has also been recorded from Abago Mountain by
<bibRefCitation id="EF9E4B0CFFDB9624FC199A79FB49FDE6" box="[907,1159,532,557]" pageId="23" pageNumber="324" refString="Nikitskiy, N. B., Bibin, A. R. &amp; Dolgin, M. M. (2008) Ksilotrofnye zhestkokrylye (Coleoptera) Kavkazskogo gosudarstvennogo prirodnogo biosfernogo zapovednika i sopredelnykh territoriy [Xylophilous beetles (Coleoptera) of the Caucasian State Biospheric Natural Reserve and adjacent territories]. Syktyvkar, 452 pp. [in Russian]" type="book">Nikitskiy &amp; al. (2008)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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