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<document id="D9C7BA0BDA17F862FACF024DE420F856" ID-CLB-Dataset="29741" ID-DOI="10.11646/zootaxa.4443.1.1" ID-GBIF-Dataset="fb772898-fe24-438e-9a4a-1a98c5b393a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1305628" ID-ZooBank="urn:lsid:zoobank.org:pub:8CF7D069-783C-4D20-8A10-6987529AB4BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530775745819" checkinUser="plazi" docAuthor="Chris J. Hodgson &amp; Douglas J. Williams" docDate="2018" docId="03A687D07BF80384EEA8FF34FAF3FDC0" docLanguage="en" docName="zootaxa.4443.1.1.pdf.imf" docOrigin="Zootaxa 4443 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Paralecanium palawanense Hodgson, spec. n." docType="treatment" docVersion="7" lastPageNumber="131" masterDocId="FF9FFFA87B780306EE3FFFACFFC6FFFD" masterDocTitle="Revision of the soft scale genus Paralecanium (Hemiptera: Coccomorpha: Coccidae) with the introduction of three new genera and twenty new species" masterLastPageNumber="162" masterPageNumber="1" pageNumber="129" updateTime="1698552844880" updateUser="ExternalLinkService">
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<mods:title id="839B2CFEAB4D09D337CBAD0D89EBAAA9">Revision of the soft scale genus Paralecanium (Hemiptera: Coccomorpha: Coccidae) with the introduction of three new genera and twenty new species</mods:title>
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<mods:namePart id="CA4867B33E5F7457148C9AF5992FC537">Chris J. Hodgson</mods:namePart>
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<mods:namePart id="846A5B56AA52432CDAE402AD005EC3DA">Douglas J. Williams</mods:namePart>
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<mods:title id="2AB55AE9D655E5C8C089C63A574BE6E2">Zootaxa</mods:title>
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<mods:date id="D900D8AF6DF17BDA330007CFA3BE3F07">2018</mods:date>
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<mods:number id="6824BDC0B00F31ECCB6EFE4B5BDF2C0B">2018-07-05</mods:number>
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<mods:number id="22F66CA44003565C46DB01BFD7D1E3C7">4443</mods:number>
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<treatment id="03A687D07BF80384EEA8FF34FAF3FDC0" ID-DOI="http://doi.org/10.5281/zenodo.5981546" ID-GBIF-Taxon="146071261" ID-Zenodo-Dep="5981546" LSID="urn:lsid:plazi:treatment:03A687D07BF80384EEA8FF34FAF3FDC0" httpUri="http://treatment.plazi.org/id/03A687D07BF80384EEA8FF34FAF3FDC0" lastPageId="130" lastPageNumber="131" pageId="128" pageNumber="129">
<subSubSection id="C315654D7BF80386EEA8FF34FF3FFF2E" pageId="128" pageNumber="129" type="nomenclature">
<paragraph id="8BB036C67BF80386EEA8FF34FD79FF4F" blockId="128.[151,703,152,211]" box="[151,703,152,178]" pageId="128" pageNumber="129">
<heading id="D0F881AA7BF80386EEA8FF34FD79FF4F" bold="true" box="[151,703,152,178]" fontSize="11" level="1" pageId="128" pageNumber="129" reason="1">
<emphasis id="B97BEAD47BF80386EEA8FF34FD79FF4F" bold="true" box="[151,703,152,178]" pageId="128" pageNumber="129">
<taxonomicName id="4C0F4D457BF80386EEA8FF34FD9DFF4F" authority="Hodgson" authorityName="Hodgson" box="[151,603,152,178]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="palawanense" status="sp. nov.">
<emphasis id="B97BEAD47BF80386EEA8FF34FE23FF4F" bold="true" box="[151,485,152,178]" italics="true" pageId="128" pageNumber="129">Paralecanium palawanense</emphasis>
Hodgson
</taxonomicName>
<taxonomicNameLabel id="A24857AF7BF80386EC5DFF34FD79FF4F" box="[610,703,152,178]" pageId="128" pageNumber="129" rank="species">spec. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB036C67BF80386EEA8FF16FF3FFF2E" blockId="128.[151,703,152,211]" box="[151,249,186,211]" pageId="128" pageNumber="129">
(
<figureCitation id="13342A437BF80386EEA0FF16FF37FF2E" box="[159,241,186,211]" captionStart="FIGURE 54" captionStartId="129.[151,250,1947,1969]" captionTargetBox="[216,1416,259,1883]" captionTargetId="figure@129.[151,1436,193,1925]" captionTargetPageId="129" captionText="FIGURE 54. Paralecanium palawanense Hodgson, spec. n., adult female." httpUri="https://zenodo.org/record/1305738/files/figure.png" pageId="128" pageNumber="129">Fig. 54</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C315654D7BF80386EEA8FEAEFB05FDA3" pageId="128" pageNumber="129" type="materials_examined">
<paragraph id="8BB036C67BF80386EEA8FEAEFC28FE0E" blockId="128.[151,1437,258,2011]" pageId="128" pageNumber="129">
<emphasis id="B97BEAD47BF80386EEA8FEAEFE38FEE6" bold="true" box="[151,510,258,283]" pageId="128" pageNumber="129">Material examined. Holotype f</emphasis>
and
<emphasis id="B97BEAD47BF80386EC0BFEAEFC28FEE6" bold="true" box="[564,1006,258,283]" pageId="128" pageNumber="129">paratype ff. PHILIPPINE ISLANDS:</emphasis>
left label:
<taxonomicName id="4C0F4D457BF80386EA5CFEAEFB3CFEE6" box="[1123,1274,258,283]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="128" pageNumber="162" phylum="Arthropoda" rank="genus">Paralecanium</taxonomicName>
/
<taxonomicName id="4C0F4D457BF80386EB31FEAEFA5DFEE6" box="[1294,1435,258,283]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="palawanense">palawanense</taxonomicName>
/ Hodgson / holotype and paratype ff; right label: Calophyllum / upper surfaces of lvs / PHILIPPINE IS: / PALAWAN / Irawan│Iwahig / forest, 2000,
<date id="FFB110067BF80386ECAFFEE7FCE5FE9E" box="[656,803,330,355]" pageId="128" pageNumber="129" value="1988-02-11">11 Feb. 1988</date>
, J.H. Martin coll #5266 / B.M. 1988-2 (BMNH): 1/6adff (fg, but heavily sclerotised;
<emphasis id="B97BEAD47BF80386EFEEFEC2FDF1FE7A" bold="true" box="[465,567,366,391]" pageId="128" pageNumber="129">holotype</emphasis>
female bottom righthand specimen, clearly marked).
<emphasis id="B97BEAD47BF80386EAAEFEC2FAE6FE7A" bold="true" box="[1169,1312,366,391]" pageId="128" pageNumber="129">Paratype ff:</emphasis>
Left label:
<taxonomicName id="4C0F4D457BF80386EEA8FE3EFEF7FE56" box="[151,305,402,427]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="128" pageNumber="162" phylum="Arthropoda" rank="genus">Paralecanium</taxonomicName>
/
<taxonomicName id="4C0F4D457BF80386EF68FE3EFE2EFE56" box="[343,488,402,427]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="palawanense">palawanense</taxonomicName>
/ Hodgson / paratype ff; right label: Calophyllum / upper surfaces of lvs / PHILIPPINE IS: / PALAWAN / Irawan│ Iwahig / forest, 2000,
<date id="FFB110067BF80386EDB2FE1BFBECFE32" box="[909,1066,438,463]" pageId="128" pageNumber="129" value="1988-02-11">11 Feb. 1988</date>
, J.H. Martin coll #5266 / B.M. 1988-2 (BMNH): 2/12adff (f, but mature and with some damage to venter).
</paragraph>
<paragraph id="8BB036C67BF80386EEF8FE53FC5DFDEA" blockId="128.[151,1437,258,2011]" box="[199,923,510,535]" pageId="128" pageNumber="129">Note. The description is based on the least-sclerotised specimen.</paragraph>
<paragraph id="8BB036C67BF80386EEF8FD8EFC9FFDC6" blockId="128.[151,1437,258,2011]" box="[199,857,546,571]" pageId="128" pageNumber="129">
<emphasis id="B97BEAD47BF80386EEF8FD8EFE01FDC6" bold="true" box="[199,455,546,571]" pageId="128" pageNumber="129">Unmounted material.</emphasis>
Derm black (J. Martin,
<emphasis id="B97BEAD47BF80386ECEAFD88FC84FDC6" box="[725,834,548,571]" italics="true" pageId="128" pageNumber="129">pers. com</emphasis>
.).
</paragraph>
<paragraph id="8BB036C67BF80386EEF8FDEAFB05FDA3" blockId="128.[151,1437,258,2011]" box="[199,1219,582,607]" pageId="128" pageNumber="129">
<emphasis id="B97BEAD47BF80386EEF8FDEAFDDEFDA2" bold="true" box="[199,536,582,607]" pageId="128" pageNumber="129">Slide-mounted adult female.</emphasis>
Body almost round. Length 2.0
<quantity id="4CF79B237BF80386EDB4FDEAFC24FDA3" box="[907,994,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="128" pageNumber="129" unit="mm" value="2.5">2.5 mm</quantity>
, width 1.82.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C315654D7BF80384EEF8FDC0FABFFF28" lastPageId="130" lastPageNumber="131" pageId="128" pageNumber="129" type="description">
<paragraph id="8BB036C67BF80386EEF8FDC0FDCBFAF6" blockId="128.[151,1437,258,2011]" pageId="128" pageNumber="129">
<emphasis id="B97BEAD47BF80386EEF8FDC0FEEFFD7E" box="[199,297,620,643]" italics="true" pageId="128" pageNumber="129">Dorsum.</emphasis>
Derm rather uniformly sclerotised at maturity, with round to oval areolations, each with darker borders, mostly in radial bands submarginally and submedially; areolations particularly obvious on older specimens, when most of derm also with minute small pale spots, each less than 1 µm wide. Marginal radial lines clear and distinct, extending well onto dorsum; alternate rays significantly longer and with many converging and coalescing more medially on older specimens, present as follows: 11 lines anteriorly between anterior stigmatic clefts, no lines from each cleft, each side with 3 lines between stigmatic clefts and 11 on abdomen. Abdominal clear areas generally absent but occasionally possibly present but obscure. Dorsal setae: hard to detect; basal sockets extremely hard to separate from smaller pores (due to sclerotisation of dorsum) but each seta straight to slightly curved, parallel-sided with a blunt apex, 45 µm long; apparently restricted to marginal and submarginal areas near radial lines; absent more medially. Large dorsal pores each small, 45 µm wide, possibly with visible micropores, surrounded by slightly more heavily sclerotised derm; frequent in broad submedial bands of 1835 pores on each side; pores not forming a clear polygonal pattern. Small simple pores of perhaps of 2 sizes present, each represented by small clear spot in derm, smaller pores each about 1 µm wide, larger pores each perhaps 2 µm wide; also not in an obvious polygonal pattern. Anal plates each with a rounded outer margin; each plate with 1 or 2 small pores medially plus 1 seta near middle of inner margin; also with 4 small setae near apex; length of plates 150160 µm, combined width 105115 µm. Anogenital fold with 1 pair of small setae along anterior margin, each lateral margin with (possibly) 1 seta at anterior end, another near middle and possibly another posteriorly. On more mature specimens, anal plate apodeme (lateral margin sclerotisation) extending anteriorly and forming 2 “flamelike” extensions into body cavity.
</paragraph>
<paragraph id="8BB036C67BF80386EEF8FABBFCDCF9D6" blockId="128.[151,1437,258,2011]" pageId="128" pageNumber="129">
<emphasis id="B97BEAD47BF80386EEF8FABBFEE4FAD3" box="[199,290,1303,1326]" italics="true" pageId="128" pageNumber="129">Margin.</emphasis>
Margins without marginal ornamentation apart from a castellation for each seta. Marginal setae oval, fan-like; width of each fan mostly 2027 µm, length about 1823 µm; with about 8095 setae anteriorly between anterior stigmatic clefts, each side with 3036 between clefts and 7588 along abdominal margin. Stigmatic clefts quite deep, each with a strongly sclerotised inner margin and 03 stigmatic spines; some clefts (about a quarter) appear to lack setae and setal sockets; about 20% have a single large median spine, and the remainder each have 2 or 3 spines; when spines present, median spine always present and placed towards anterior end of cleft; median spine much the longest, each 3035 µm long, each lateral spine 1317 µm long. Eyespots each more or less round, width of socket 5065 µm, diameter of lens about 20 µm.
</paragraph>
<paragraph id="8BB036C67BF80384EEF8F99BFABFFF28" blockId="128.[151,1437,258,2011]" lastBlockId="130.[151,1436,151,573]" lastPageId="130" lastPageNumber="131" pageId="128" pageNumber="129">
<emphasis id="B97BEAD47BF80386EEF8F99BFED6F9B3" box="[199,272,1591,1614]" italics="true" pageId="128" pageNumber="129">Venter.</emphasis>
Membranous, but with a complete submarginal band about 70115 µm wide on more mature specimens; also with a dense darker area just posterior to each anal plate. Multilocular disc-pores present around genital opening and on preceding 2 segments only, each side with 59 on abdominal segment VII, 1113 on segment VI and 14 on segment V. Spiracular disc-pores sparsely present in bands 1 pore wide between margin and each spiracle, with 1014 pores in each band. Ventral microducts present at least near mouthparts. Ventral setae: with 1 pair of interantennal setae; 1 pair of slightly longer pregenital setae on abdominal segment VII, each seta about 90 µm long, plus other long setae, each about 60 µm long, in bands across abdominal segments as follows (on each side): segment VII 02;
<date id="FFB110067BF80386EC2AF89FFDAEF8B6" box="[533,616,1843,1867]" pageId="128" pageNumber="129">VI 14</date>
;
<date id="FFB110067BF80386EC4AF89FFD78F8B6" box="[629,702,1842,1867]" pageId="128" pageNumber="129">V 36</date>
;
<date id="FFB110067BF80386ECF4F89FFD39F8B6" box="[715,767,1842,1867]" pageId="128" pageNumber="129">IV 6</date>
or 7,
<date id="FFB110067BF80386ED7BF89FFC54F8B6" box="[836,914,1842,1867]" pageId="128" pageNumber="129">III 59</date>
and
<date id="FFB110067BF80386EDF4F89FFBD5F8B6" box="[971,1043,1842,1867]" pageId="128" pageNumber="129">II 58</date>
; other setae very sparse. Antennae reduced, each with 6 segments, total length 90100 µm; apical segment short, 1012 µm long, apical seta short, each about 812 µm long; antennal setae apparently entirely absent from scape, pedicel and segment III; only fleshy setae present on segments IV and V, and 5 or 6 setae on apical segment. Clypeolabral shield about 9095 µm long. Spiracles small, width of each peritreme 1518 µm. Legs very reduced but with most segments still distinct, each with no tibio-tarsal segmental line; total length of hind legs (µm) 4050, claw 2.53.0; tarsal digitules absent; claw digitules possibly dissimilar, 1 rather longer than other with small capitate apices; leg setae mainly absent.
</paragraph>
</subSubSection>
<caption id="DF70664E7BF90387EEA8F837FC49F84C" box="[151,911,1947,1969]" httpUri="https://zenodo.org/record/1305738/files/figure.png" pageId="129" pageNumber="130" startId="129.[151,250,1947,1969]" targetBox="[216,1416,259,1883]" targetPageId="129">
<paragraph id="8BB036C67BF90387EEA8F837FC49F84C" blockId="129.[151,911,1947,1969]" box="[151,911,1947,1969]" pageId="129" pageNumber="130">
<emphasis id="B97BEAD47BF90387EEA8F837FEE6F84C" bold="true" box="[151,288,1947,1969]" pageId="129" pageNumber="130">FIGURE 54.</emphasis>
<taxonomicName id="4C0F4D457BF90387EF1AF837FD62F84C" authority="Hodgson" authorityName="Hodgson" box="[293,676,1947,1969]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="palawanense" status="sp. nov.">
<emphasis id="B97BEAD47BF90387EF1AF837FDF9F84C" box="[293,575,1947,1969]" italics="true" pageId="129" pageNumber="130">Paralecanium palawanense</emphasis>
Hodgson
</taxonomicName>
,
<emphasis id="B97BEAD47BF90387EC91F837FD3BF84C" bold="true" box="[686,765,1947,1969]" pageId="129" pageNumber="130">
<taxonomicNameLabel id="A24857AF7BF90387EC91F837FD3BF84C" box="[686,765,1947,1969]" pageId="129" pageNumber="130" rank="species">spec. n.</taxonomicNameLabel>
</emphasis>
, adult female.
</paragraph>
</caption>
<subSubSection id="C315654D7BFA0384EEF8FF73FADEFE08" pageId="130" pageNumber="131" type="discussion">
<paragraph id="8BB036C67BFA0384EEF8FF73FA98FE32" blockId="130.[151,1436,151,573]" pageId="130" pageNumber="131">
<emphasis id="B97BEAD47BFA0384EEF8FF73FE89FF05" bold="true" box="[199,335,223,248]" pageId="130" pageNumber="131">Comments.</emphasis>
Adult female
<taxonomicName id="4C0F4D457BFA0384EFC3FF4DFD68FF05" box="[508,686,223,248]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="palawanense" status="sp. nov.">
<emphasis id="B97BEAD47BFA0384EFC3FF4DFD68FF05" box="[508,686,223,248]" italics="true" pageId="130" pageNumber="131">P. palawanense</emphasis>
</taxonomicName>
<emphasis id="B97BEAD47BFA0384EC87FF73FCD2FF05" bold="true" box="[696,788,223,248]" pageId="130" pageNumber="131">
<taxonomicNameLabel id="A24857AF7BFA0384EC87FF73FCD2FF05" box="[696,788,223,248]" pageId="130" pageNumber="131" rank="species">spec. n.</taxonomicNameLabel>
</emphasis>
have a number of rather unique features: (i) anal plate apodemes each with “flame-like” extensions anteriorly on mature specimens; (ii) extreme reduction of legs but still with most segmentation visible; (iii) segmental bands of long setae medially on abdominal segments; (iv) possible absence of stigmatic spines from some clefts; (v) when stigmatic spines present, the larger setae are positioned anteriorly in the cleft rather than medially; (vi) absence of marginal ornamentation apart from enlarged setal sockets, and (vii) antennae without setose setae on scape, pedicel and segments IIIV.
<taxonomicName id="4C0F4D457BFA0384EA5BFE38FA5AFE50" box="[1124,1436,404,429]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="palawanense">
<emphasis id="B97BEAD47BFA0384EA5BFE38FA5AFE50" box="[1124,1436,404,429]" italics="true" pageId="130" pageNumber="131">Paralecanium palawanense</emphasis>
</taxonomicName>
is otherwise rather similar to
<taxonomicName id="4C0F4D457BFA0384EFE1FE15FDACFE2D" baseAuthorityName="Green" baseAuthorityYear="1904" box="[478,618,439,464]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="calophylli">
<emphasis id="B97BEAD47BFA0384EFE1FE15FDACFE2D" box="[478,618,439,464]" italics="true" pageId="130" pageNumber="131">P. calophylli</emphasis>
</taxonomicName>
, which, as its name suggests, was also collected off
<emphasis id="B97BEAD47BFA0384EA8BFE1BFAF1FE2D" box="[1204,1335,439,464]" italics="true" pageId="130" pageNumber="131">Calophillus</emphasis>
sp.
</paragraph>
<paragraph id="8BB036C67BFA0384EEF8FE70FADEFE08" blockId="130.[151,1436,151,573]" box="[199,1304,476,501]" pageId="130" pageNumber="131">
<materialsCitation id="3B673C9B7BFA0384EEF8FE70FADEFE08" ID-GBIF-Occurrence="1872522453" box="[199,1304,476,501]" country="Philippines" location="Palawan" pageId="130" pageNumber="131" specimenCount="1" stateProvince="Palawan">
<taxonomicName id="4C0F4D457BFA0384EEF8FE70FE39FE08" box="[199,511,476,501]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="palawanense">
<emphasis id="B97BEAD47BFA0384EEF8FE70FE39FE08" box="[199,511,476,501]" italics="true" pageId="130" pageNumber="131">Paralecanium palawanense</emphasis>
</taxonomicName>
is currently only known from the island of
<collectingRegion id="49CBF8247BFA0384EDD9FE70FB83FE08" box="[998,1093,476,501]" country="Philippines" name="Palawan" pageId="130" pageNumber="131">Palawan</collectingRegion>
in the
<collectingCountry id="F31876567BFA0384EAADFE70FAD2FE08" box="[1170,1300,476,501]" name="Philippines" pageId="130" pageNumber="131">Philippines</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C315654D7BFA0384EEF8FE53FAF3FDC0" pageId="130" pageNumber="131" type="description">
<paragraph id="8BB036C67BFA0384EEF8FE53FD60FDE5" blockId="130.[151,1436,151,573]" box="[199,678,511,536]" pageId="130" pageNumber="131">
<emphasis id="B97BEAD47BFA0384EEF8FE53FE88FDE5" bold="true" box="[199,334,511,536]" pageId="130" pageNumber="131">Host-plant.</emphasis>
<emphasis id="B97BEAD47BFA0384EF6BFE53FE22FDE5" box="[340,484,511,536]" italics="true" pageId="130" pageNumber="131">Calophyllum</emphasis>
sp. (
<taxonomicName id="4C0F4D457BFA0384EC25FE53FD5CFDE5" box="[538,666,511,536]" family="Clusiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="130" pageNumber="131" phylum="Magnoliopsida" rank="family">Clusiaceae</taxonomicName>
).
</paragraph>
<paragraph id="8BB036C67BFA0384EEF8FD88FAF3FDC0" blockId="130.[151,1436,151,573]" box="[199,1333,548,573]" pageId="130" pageNumber="131">
<emphasis id="B97BEAD47BFA0384EEF8FD88FE50FDC0" bold="true" box="[199,406,548,573]" pageId="130" pageNumber="131">Name derivation.</emphasis>
This species is named after the island in the Philippines on which it was collected.
</paragraph>
</subSubSection>
</treatment>
</document>