366 lines
51 KiB
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366 lines
51 KiB
XML
<document id="68B4DE20F7B761E20A77A52FB245D4AC" ID-DOI="10.11646/zootaxa.5027.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5448029" ID-ZooBank="CCF10500-01A3-476C-A4BE-57161E164EBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630464259914" checkinUser="plazi" docAuthor="Kompier, Tom, Karube, Haruki, Futahashi, Ryo & Phan, Quoc Toan" docDate="2021" docId="03A687CBFFDAFFE4E0E77290FC90F91D" docLanguage="en" docName="zootaxa.5027.1.1.pdf" docOrigin="Zootaxa 5027 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Planaeschna ishigakiana Asahina 1951" docType="treatment" docVersion="4" lastPageNumber="27" masterDocId="FF9FFFB3FFC0FFFEE0707207FFC3FFCA" masterDocTitle="The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="27" updateTime="1699280300883" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1404DE5D35A524AF5F2CF4B1C70C5777">The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)</mods:title>
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<mods:namePart id="2764F89633B102D0CE7372F31DC27805">Kompier, Tom</mods:namePart>
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<mods:affiliation id="E5BE14F48DD90357FC8245C2AC93DE2F">Schoutenstraat 69, 2596 SK Den Haag, the Netherlands.</mods:affiliation>
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<mods:namePart id="8815B9E32BFD3DFA2E313E8D0B091DE9">Karube, Haruki</mods:namePart>
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<mods:affiliation id="2652ADF23F01FC2BB1901F223F92DCE6">Kanagawa Prefectural Museum of Natural History, Odawara, Japan.</mods:affiliation>
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<mods:namePart id="455D2CB30A4E17C751D59D851612F72D">Futahashi, Ryo</mods:namePart>
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<mods:affiliation id="2AC10B71C134D490344904944FBB10FB">National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan.</mods:affiliation>
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<mods:namePart id="CFE9C65A666C41E93EC73ECC3DFB4B53">Phan, Quoc Toan</mods:namePart>
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<mods:affiliation id="BA6417250FF6BEBED9CDB17D916AC305">The Center for Entomology & Parasitology Research, College of Medicine & Pharmacy, Duy Tan University, 550000, Da Nang, Vietnam. pqtoan 84 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3154 - 6546</mods:affiliation>
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<mods:nameIdentifier id="CF704C8B34E1E558B77EFCC2BADD7674" type="email">pqtoan84@gmail.com</mods:nameIdentifier>
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<mods:title id="E3812669391397EEA86CF6FC1351D2C0">Zootaxa</mods:title>
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<treatment id="03A687CBFFDAFFE4E0E77290FC90F91D" ID-DOI="http://doi.org/10.5281/zenodo.5492692" ID-Zenodo-Dep="5492692" LSID="urn:lsid:plazi:treatment:03A687CBFFDAFFE4E0E77290FC90F91D" httpUri="http://treatment.plazi.org/id/03A687CBFFDAFFE4E0E77290FC90F91D" lastPageNumber="27" pageId="26" pageNumber="27">
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<paragraph id="8BB036DDFFDAFFE4E0E77290FD6FFF7B" blockId="26.[151,684,151,213]" box="[151,684,151,178]" pageId="26" pageNumber="27">
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<heading id="D0F881B1FFDAFFE4E0E77290FD6FFF7B" bold="true" box="[151,684,151,178]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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<emphasis id="B97BEACFFFDAFFE4E0E77290FD6FFF7B" bold="true" box="[151,684,151,178]" pageId="26" pageNumber="27">
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10.
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<taxonomicName id="4C0F4D5EFFDAFFE4E0B37290FD6FFF7B" ID-CoL="4JJ3T" authority="Asahina, 1951" authorityName="Asahina" authorityYear="1951" box="[195,684,151,178]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
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<emphasis id="B97BEACFFFDAFFE4E0B37290FE36FF78" bold="true" box="[195,501,151,178]" italics="true" pageId="26" pageNumber="27">Planaeschna ishigakiana</emphasis>
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<bibRefCitation id="EF9E4B2CFFDAFFE4E18C7290FD6FFF7B" author="Asahina, S." box="[508,684,151,177]" pageId="26" pageNumber="27" pagination="15 - 22" refId="ref16213" refString="Asahina, S. (1951) New dragonflies from the north-eastern Asia (Odonata). Kontyu, 19, 15 - 22." type="journal article" year="1951">Asahina, 1951</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3156556FFDAFFE4E0E772BCFEE4FF1F" box="[151,295,187,213]" pageId="26" pageNumber="27" type="description">
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<paragraph id="8BB036DDFFDAFFE4E0E772BCFEE4FF1F" blockId="26.[151,684,151,213]" box="[151,295,187,213]" pageId="26" pageNumber="27">
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(
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<figureCitation id="13342A58FFDAFFE4E0EF72BCFF35FF1F" box="[159,246,187,213]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Figs 16</figureCitation>
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,
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<figureCitation id="13342A58FFDAFFE4E17272BCFEDCFF1F" box="[258,287,187,213]" captionStart="FIGURE 17" captionStartId="28.[151,250,1349,1373]" captionTargetBox="[156,1436,194,1325]" captionTargetId="figure-22@28.[151,1436,181,1325]" captionTargetPageId="28" captionText="FIGURE 17. Planaeschna ishigakiana spp. anal appendages. [A–C] after Asahina (1988), lateral view (top) and dorsal view (bottom), [D] Xuan Son NP, Phu Tho Prov., 26-X-2014 (same as in Fig. 7). (A) P. i. ishigakiana, Ishigakijima (1988: p. 28, Figs 2–3), (B) P. i. nagaminei, Amami Oshima (1988: p. 29, Figs 13–14), (C) P. i. ishigakiana, Iriomote Island (1988: p. 28, Figs 4–5), (D) P. i. guentherpetersi stat. nov." figureDoi="http://doi.org/10.5281/zenodo.5448068" httpUri="https://zenodo.org/record/5448068/files/figure.png" pageId="26" pageNumber="27">17</figureCitation>
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)
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</paragraph>
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<subSubSection id="C3156556FFDAFFE4E0E77307FC6CFEFC" pageId="26" pageNumber="27" type="reference_group">
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<paragraph id="8BB036DDFFDAFFE4E0E77307FC70FED2" blockId="26.[151,947,256,310]" box="[151,947,256,280]" pageId="26" pageNumber="27">
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<taxonomicName id="4C0F4D5EFFDAFFE4E0E77307FC8EFED2" authority="Sasamoto, Do & Vu, 2013" authorityName="Sasamoto, Do & Vu" authorityYear="2013" baseAuthorityName="Kompier & Karube & Futahashi & Phan" baseAuthorityYear="2013" box="[151,845,256,280]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="ishigakiana" status="stat. nov." subSpecies="guentherpetersi">
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<emphasis id="B97BEACFFFDAFFE4E0E77307FDFFFED2" box="[151,572,256,280]" italics="true" pageId="26" pageNumber="27">Planaeschna ishigakiana guentherpetersi</emphasis>
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<bibRefCitation id="EF9E4B2CFFDAFFE4E2327307FC8EFED2" author="Sasamoto, A. & Do, C. & Vu, L. V." box="[578,845,256,280]" pageId="26" pageNumber="27" pagination="587 - 594" refId="ref17356" refString="Sasamoto, A., Do, C. & Vu, L. V. (2013) Discovery of a new species of the genus Planaeschna from Northern Vietnam, with a first description of male P. tomokunii. Zootaxa, 3652 (5), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3652.5.8" type="journal article" year="2013">Sasamoto, Do & Vu, 2013</bibRefCitation>
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</taxonomicName>
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<emphasis id="B97BEACFFFDAFFE4E3237307FC70FED2" bold="true" box="[851,947,256,280]" pageId="26" pageNumber="27">
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<taxonomicNameLabel id="A24857B4FFDAFFE4E3237307FC70FED2" box="[851,947,256,280]" pageId="26" pageNumber="27">stat. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
<paragraph id="8BB036DDFFDAFFE4E0E77319FC6CFEFC" blockId="26.[151,947,256,310]" box="[151,943,286,310]" pageId="26" pageNumber="27">
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<taxonomicName id="4C0F4D5EFFDAFFE4E0E77319FD71FEFC" ID-CoL="9WVXY" authority=": Sasamoto et al. (2013)" authorityName="Sasamoto" authorityYear="2013" box="[151,690,286,310]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi">
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<emphasis id="B97BEACFFFDAFFE4E0E77319FE7CFEFC" box="[151,447,286,310]" italics="true" pageId="26" pageNumber="27">Planaeschna guentherpetersi</emphasis>
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||
:
|
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<bibRefCitation id="EF9E4B2CFFDAFFE4E1BD7319FD71FEFC" author="Sasamoto, A. & Do, C. & Vu, L. V." box="[461,690,286,310]" pageId="26" pageNumber="27" pagination="587 - 594" refId="ref17356" refString="Sasamoto, A., Do, C. & Vu, L. V. (2013) Discovery of a new species of the genus Planaeschna from Northern Vietnam, with a first description of male P. tomokunii. Zootaxa, 3652 (5), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3652.5.8" type="journal article" year="2013">
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||
Sasamoto
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<emphasis id="B97BEACFFFDAFFE4E2467318FDAAFEFC" box="[566,617,286,310]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
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(2013)
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</bibRefCitation>
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</taxonomicName>
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pp. 587–590,
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<figureCitation id="13342A58FFDAFFE4E3347319FC68FEFC" box="[836,939,286,310]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="2.[151,250,1910,1934]" captionStartId-1="3.[151,250,1867,1891]" captionStartId-2="6.[151,250,1942,1966]" captionStartId-3="8.[151,250,1944,1968]" captionStartId-4="9.[151,250,1943,1967]" captionStartId-5="11.[151,250,1801,1825]" captionStartId-6="13.[151,250,1943,1967]" captionStartId-7="14.[151,250,1943,1967]" captionStartId-8="15.[151,250,1874,1898]" captionStartId-9="17.[151,250,1943,1967]" captionTargetBox-0="[219,1354,194,1883]" captionTargetBox-1="[200,1398,187,1839]" captionTargetBox-2="[242,1345,181,1919]" captionTargetBox-3="[247,1338,181,1920]" captionTargetBox-4="[225,1363,181,1920]" captionTargetBox-5="[151,1436,921,1778]" captionTargetBox-6="[225,1362,181,1920]" captionTargetBox-7="[201,1386,181,1920]" captionTargetBox-8="[187,1401,181,1850]" captionTargetBox-9="[267,1321,181,1920]" captionTargetId-0="figure-22@2.[196,1391,181,1886]" captionTargetId-1="figure-17@3.[170,1416,181,1843]" captionTargetId-2="figure-22@6.[242,1345,181,1919]" captionTargetId-3="figure-22@8.[247,1339,181,1920]" captionTargetId-4="figure-17@9.[225,1363,181,1920]" captionTargetId-5="figure-362@11.[151,1436,921,1778]" captionTargetId-6="figure-17@13.[225,1362,181,1920]" captionTargetId-7="figure-22@14.[201,1386,181,1920]" captionTargetId-8="figure-17@15.[187,1401,181,1850]" captionTargetId-9="figure-17@17.[267,1321,181,1920]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="11" captionTargetPageId-6="13" captionTargetPageId-7="14" captionTargetPageId-8="15" captionTargetPageId-9="17" captionText-0="FIGURE 1. A neighbour-joining phylogeny of Vietnamese Planaeschna species based on nuclear genes (ITS1-5.8S-ITS2, 1,081 bp). Maximum likelihood and Bayesian phylogenies exhibited substantially similar topologies. On each node, statistical support values are indicated in the order of [bootstrap value of neighbour-joining]/[bootstrap value of maximum likelihood]/ posterior probability (percentages) of Bayesian]. Accession numbers are shown in parentheses." captionText-1="FIGURE 2. A neighbour-joining phylogeny of Vietnamese Planaeschna species based on mitochondrial genes (16SrRNA and COI, 965 bp). Maximum likelihood and Bayesian phylogenies exhibited substantially similar topologies. On each node, statistical support values are indicated in the order of [bootstrap value of neighbour-joining]/[bootstrap value of maximum likelihood]/[posterior probability (percentages) of Bayesian]. Asterisks indicate support values lower than 50%. Accession numbers are shown in parentheses." captionText-2="FIGURE 3. Planaeschna samurai sp. nov. [A, C] face, [B, D] lateral habitus alive, [E, F] scan of dorsal habitus, [G–H] anal appendages in dorsal (G) and lateral (H) view. (A) paratype male, Pia Oac NP, Cao Bang Prov., 18-X-2014, (B, E, G–H) holotype, (C–D, F) paratype female, Tu Le, Yen Bai Prov., 16-X-2015." captionText-3="FIGURE 4. Planaeschna tomokunii [A–C] lateral habitus alive, including face, [D–E] scans of dorsal habitus, [F–G] anal appendages of male in lateral (F) and dorsal (G) view. (A, F–G) male, Tam Dao NP, Vinh Phuc Prov., 3-VII-2016, (B) immature male, same location, 4-VI-2016, (C, E) female, same location, 24-IX-2015, (D) male, same location, 16-VI-2016." captionText-4="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." captionText-5="FIGURE 6. Planaeschna viridis male, Bach Ma NP. Thus Thien-Hue Prov., 7-VIII-2016. (A) lateral habitus alive, including face, [B–C] anal appendages in lateral (B) and dorsal (C) view, (D) dorsal pattern of S2." captionText-6="FIGURE 7. Planaeschna asahinai alive in hand [A, B] habitus, [C, D] face. (A, C) male, 7-IX-2016, Sao La NR, Thua Thien- Hue Prov., (B, D) female, 6-VIII-2016, Bach Ma NP, Thua Thien-Hue Prov." captionText-7="FIGURE 8. Planaeschna celia males [A] scan of dorsal habitus, [B] face, [C–D] lateral habitus alive. (A) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (B–D) same location, 26-X-2014 (two individuals)." captionText-8="FIGURE 9. Comparison of details of male Planaeschna tamdaoensis and male P. celia [A, B] facial pattern, [C, D] anal appendages lateral, [E, F] anal appendages dorsal, [G, H] dorsal pattern of S2. [A, C, E, G] P. tamdaoensis, [B, D, F, H] P. celia. (A) Pia Oac NP, Cao Bang Prov., 12-IX-2015, (B) Xuan Son NP, Phu Tho Prov., 26-X-2014 (different specimen than Fig. 2B), (C, E) Tam Dao NP, Vinh Phuc Prov., 27-VIII-2016, (D, F, H) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (G) Pia Oac NP, Cao Bang Prov., 27-VI-2015." captionText-9="FIGURE 10. Planaeschna crux sp. nov. alive in hand [A–C] lateral habitus, [D–F] face. (A, D) male, Pia Oac NP, Cao Bang Prov., 3-X-2015, (B, E), immature male, Tu Le, Yen Bai Prov., 30-VI-2015, (C) female, Tu Le, Yen Bai Prov., 15-V-2015, (F) female, Pia Oac NP, Cao Bang Prov., 3-X-2015." figureDoi-0="http://doi.org/10.5281/zenodo.5448031" figureDoi-1="http://doi.org/10.5281/zenodo.5448035" figureDoi-2="http://doi.org/10.5281/zenodo.5448037" figureDoi-3="http://doi.org/10.5281/zenodo.5448039" figureDoi-4="http://doi.org/10.5281/zenodo.5448041" figureDoi-5="http://doi.org/10.5281/zenodo.5448043" figureDoi-6="http://doi.org/10.5281/zenodo.5448045" figureDoi-7="http://doi.org/10.5281/zenodo.5448047" figureDoi-8="http://doi.org/10.5281/zenodo.5448049" figureDoi-9="http://doi.org/10.5281/zenodo.5448053" httpUri-0="https://zenodo.org/record/5448031/files/figure.png" httpUri-1="https://zenodo.org/record/5448035/files/figure.png" httpUri-2="https://zenodo.org/record/5448037/files/figure.png" httpUri-3="https://zenodo.org/record/5448039/files/figure.png" httpUri-4="https://zenodo.org/record/5448041/files/figure.png" httpUri-5="https://zenodo.org/record/5448043/files/figure.png" httpUri-6="https://zenodo.org/record/5448045/files/figure.png" httpUri-7="https://zenodo.org/record/5448047/files/figure.png" httpUri-8="https://zenodo.org/record/5448049/files/figure.png" httpUri-9="https://zenodo.org/record/5448053/files/figure.png" pageId="26" pageNumber="27">Figs 1–10</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156556FFDAFFE4E0E77363FC15FA58" pageId="26" pageNumber="27" type="materials_examined">
|
||
<paragraph id="8BB036DDFFDAFFE4E0E77363FD9BFE0D" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">
|
||
<emphasis id="B97BEACFFFDAFFE4E0E77363FE48FEB4" bold="true" box="[151,395,356,382]" pageId="26" pageNumber="27">Materials examined.</emphasis>
|
||
<materialsCitation id="3B673C80FFDAFFE4E1E17362FC54FEB4" box="[401,919,356,383]" collectingDate="2013-10-26" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="26" pageNumber="27" specimenCount="1" specimenCount-female="1" stateProvince="Phu Tho">
|
||
<specimenCount id="9D09FD54FFDAFFE4E1E17362FE78FEB5" box="[401,443,357,383]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectorName id="26FA530BFFDAFFE4E1B87362FDA1FEB5" box="[456,610,357,383]" pageId="26" pageNumber="27">Xuan Son NP</collectorName>
|
||
,
|
||
<collectingRegion id="49CBF83FFFDAFFE4E21C7363FCCDFEB5" box="[620,782,356,383]" country="Vietnam" name="Phu Tho" pageId="26" pageNumber="27">Phu Tho Prov.</collectingRegion>
|
||
,
|
||
<date id="FFB1101DFFDAFFE4E3697363FC54FEB4" box="[793,919,356,382]" pageId="26" pageNumber="27" value="2013-10-26">
|
||
<collectingDate id="EFF5E9F5FFDAFFE4E3697363FC54FEB4" box="[793,919,356,382]" pageId="26" pageNumber="27" value="2013-10-26">26-X-2013</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, TK leg.;
|
||
<materialsCitation id="3B673C80FFDAFFE4E4747362FED5FE68" collectingDate="2014-10-26" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Phu Tho">
|
||
<specimenCount id="9D09FD54FFDAFFE4E4747362FBEDFEB5" box="[1028,1070,357,383]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
|
||
<specimenCount id="9D09FD54FFDAFFE4E4447362FB94FEB5" box="[1076,1111,357,383]" pageId="26" pageNumber="27" type="female">1♀</specimenCount>
|
||
, same location and collector,
|
||
<date id="FFB1101DFFDAFFE4E0E7738FFED5FE68" box="[151,278,392,418]" pageId="26" pageNumber="27" value="2014-10-26">
|
||
<collectingDate id="EFF5E9F5FFDAFFE4E0E7738FFED5FE68" box="[151,278,392,418]" pageId="26" pageNumber="27" value="2014-10-26">26-X-2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B673C80FFDAFFE4E151738EFCF9FE69" box="[289,826,392,419]" collectingDate="2014-11-23" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="2" stateProvince="Phu Tho">
|
||
<specimenCount id="9D09FD54FFDAFFE4E151738EFEA1FE69" box="[289,354,393,419]" pageId="26" pageNumber="27" type="female">2 ♀♀</specimenCount>
|
||
, same location and collector,
|
||
<date id="FFB1101DFFDAFFE4E2C2738EFCF9FE69" box="[690,826,393,419]" pageId="26" pageNumber="27" value="2014-11-23">
|
||
<collectingDate id="EFF5E9F5FFDAFFE4E2C2738EFCF9FE69" box="[690,826,393,419]" pageId="26" pageNumber="27" value="2014-11-23">23-XI-2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B673C80FFDAFFE4E336738EFA9DFE69" box="[838,1374,392,419]" collectingDate="2014-11-29" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="2" stateProvince="Phu Tho">
|
||
<specimenCount id="9D09FD54FFDAFFE4E336738EFC44FE69" box="[838,903,393,419]" pageId="26" pageNumber="27" type="female">2 ♀♀</specimenCount>
|
||
, same location and collector,
|
||
<date id="FFB1101DFFDAFFE4E4A7738EFA9DFE69" box="[1239,1374,393,419]" pageId="26" pageNumber="27" value="2014-11-29">
|
||
<collectingDate id="EFF5E9F5FFDAFFE4E4A7738EFA9DFE69" box="[1239,1374,393,419]" pageId="26" pageNumber="27" value="2014-11-29">29-XI-2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B673C80FFDAFFE4E51A738EFD96FE0D" collectingDate="2015-10-11" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="26" pageNumber="27" specimenCount="1" specimenCount-female="1" stateProvince="Phu Tho">
|
||
<specimenCount id="9D09FD54FFDAFFE4E51A738EFA56FE69" box="[1386,1429,393,419]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
|
||
, same location and collector,
|
||
<date id="FFB1101DFFDAFFE4E1A773ABFD96FE0D" box="[471,597,428,455]" pageId="26" pageNumber="27" value="2015-10-11">
|
||
<collectingDate id="EFF5E9F5FFDAFFE4E1A773ABFD96FE0D" box="[471,597,428,455]" pageId="26" pageNumber="27" value="2015-10-11">11-X-2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB036DDFFDAFFE4E0B773D6FC42FCBD" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">
|
||
<emphasis id="B97BEACFFFDAFFE4E0B773D6FED2FE21" bold="true" box="[199,273,465,491]" pageId="26" pageNumber="27">Notes.</emphasis>
|
||
<bibRefCitation id="EF9E4B2CFFDAFFE4E16A73D6FDDDFE21" author="Sasamoto, A. & Do, C. & Vu, L. V." box="[282,542,464,491]" pageId="26" pageNumber="27" pagination="587 - 594" refId="ref17356" refString="Sasamoto, A., Do, C. & Vu, L. V. (2013) Discovery of a new species of the genus Planaeschna from Northern Vietnam, with a first description of male P. tomokunii. Zootaxa, 3652 (5), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3652.5.8" type="journal article" year="2013">
|
||
Sasamoto
|
||
<emphasis id="B97BEACFFFDAFFE4E1E073D6FE08FE20" box="[400,459,464,490]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
compared
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E2EF73D6FCAEFE20" box="[671,877,464,490]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi">
|
||
<emphasis id="B97BEACFFFDAFFE4E2EF73D6FCAEFE20" box="[671,877,464,490]" italics="true" pageId="26" pageNumber="27">P. guentherpetersi</emphasis>
|
||
</taxonomicName>
|
||
with other species with rich brown facial colour (
|
||
<figureCitation id="13342A58FFDAFFE4E0D073F3FEE6FDC5" box="[160,293,500,527]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Fig. 16A, C</figureCitation>
|
||
) in their diagnosis of the species, but not with species with more colourful faces. The dorsal pattern on S2 (
|
||
<figureCitation id="13342A58FFDAFFE4E0B3701FFE81FDF9" box="[195,322,536,563]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Fig. 16E–F</figureCitation>
|
||
) is unique among Vietnamese
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E2E4701FFCB8FDF8" box="[660,891,536,562]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97BEACFFFDAFFE4E2E4701FFCE6FDF8" box="[660,805,536,562]" italics="true" pageId="26" pageNumber="27">Planaeschna</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(but see
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E3AE701FFB6BFDF9" box="[990,1192,536,563]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="undefined-2">
|
||
<emphasis id="B97BEACFFFDAFFE4E3AE701FFBACFDF8" box="[990,1135,536,562]" italics="true" pageId="26" pageNumber="27">Planaeschna</emphasis>
|
||
sp. 2
|
||
</taxonomicName>
|
||
below, and
|
||
<figureCitation id="13342A58FFDAFFE4E55B701FFA4CFDF9" box="[1323,1423,536,563]" captionStart="FIGURE 18" captionStartId="30.[151,250,1944,1968]" captionTargetBox="[160,1427,181,1920]" captionTargetId="figure-22@30.[159,1428,181,1920]" captionTargetPageId="30" captionText="FIGURE 18. Planaeschna sp. 2. [A–B] female lateral habitus including face, alive, [C–D] dorsolateral habitus. (A) Huu Lien NR, Lang Son Prov., 1-XII-2013, (B–C) Xuan Son NP, Phu Tho Prov., 20-X-2013, (D) probable male, Huu Lien NR, Lang Son Prov., 11-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448070" httpUri="https://zenodo.org/record/5448070/files/figure.png" pageId="26" pageNumber="27">Fig. 18C</figureCitation>
|
||
). However, this pattern occurs in several species recently described from continental
|
||
<collectingCountry id="F318764DFFDAFFE4E43D703BFB50FD9C" box="[1101,1171,572,598]" name="China" pageId="26" pageNumber="27">China</collectingCountry>
|
||
, and is also found in
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E5F8703AFED9FDB0" authorityName="Asahina" authorityYear="1951" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E5F8703AFED9FDB0" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F318764DFFDAFFE4E1107066FE63FDB1" box="[352,416,609,635]" name="Japan" pageId="26" pageNumber="27">Japan</collectingCountry>
|
||
and
|
||
<collectingCountry id="F318764DFFDAFFE4E1A77067FDEEFDB0" box="[471,557,608,634]" name="Taiwan" pageId="26" pageNumber="27">Taiwan</collectingCountry>
|
||
. Three subspecies are known for
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E3D67066FB86FDB0" authorityName="Asahina" authorityYear="1951" box="[934,1093,608,634]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E3D67066FB86FDB0" box="[934,1093,608,634]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
: nominotypical
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E48D7066FA5EFDB0" authorityName="Asahina" authorityYear="1951" box="[1277,1437,608,634]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E48D7066FA5EFDB0" box="[1277,1437,608,634]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
was described from Ishigaki-jima, and later recorded also from nearby Iriomote-jima, of the Yaeyama Islands in the Ryukyu archipelago of
|
||
<collectingCountry id="F318764DFFDAFFE4E1B870AEFDC8FD09" box="[456,523,681,707]" name="Japan" pageId="26" pageNumber="27">Japan</collectingCountry>
|
||
, just east of
|
||
<collectingCountry id="F318764DFFDAFFE4E2E670AFFD2FFD08" box="[662,748,680,706]" name="Taiwan" pageId="26" pageNumber="27">Taiwan</collectingCountry>
|
||
. The subspecies
|
||
<emphasis id="B97BEACFFFDAFFE4E3D770AEFB8CFD08" box="[935,1103,681,706]" italics="true" pageId="26" pageNumber="27">P. i. nagaminei</emphasis>
|
||
<bibRefCitation id="EF9E4B2CFFDAFFE4E42570AFFB34FD09" author="Asahina, S." box="[1109,1271,680,707]" pageId="26" pageNumber="27" pagination="27 - 34" refId="ref16291" refString="Asahina, S. (1988) A problematical race of Planaeschna ishigakiana from Amami-Oshima, Middle Ryukyus. Tombo, 31, 27 - 34." type="journal article" year="1988">Asahina, 1988</bibRefCitation>
|
||
was described from Amami Oshima, Ryukyu archipelago of
|
||
<collectingCountry id="F318764DFFDAFFE4E2EB70CAFD1DFD2D" box="[667,734,717,743]" name="Japan" pageId="26" pageNumber="27">Japan</collectingCountry>
|
||
, followed by the description of
|
||
<emphasis id="B97BEACFFFDAFFE4E43270CAFB1CFD2C" box="[1090,1247,716,742]" italics="true" pageId="26" pageNumber="27">P. i. flavostria</emphasis>
|
||
<bibRefCitation id="EF9E4B2CFFDAFFE4E49670CBFA98FD2C" author="Yeh, W." box="[1254,1371,716,742]" pageId="26" pageNumber="27" pagination="26 - 28" refId="ref17758" refString="Yeh, W. (1996) New records of the Odonata taken in Taiwan. Tombo, 39, 26 - 28." type="journal article" year="1996">Yeh, 1996</bibRefCitation>
|
||
, from
|
||
<collectingCountry id="F318764DFFDAFFE4E0E770F7FF2EFCC0" box="[151,237,752,778]" name="Taiwan" pageId="26" pageNumber="27">Taiwan</collectingCountry>
|
||
. The three subspecies of
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E27270F6FD63FCC0" authorityName="Asahina" authorityYear="1951" box="[514,672,752,778]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E27270F6FD63FCC0" box="[514,672,752,778]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
are different in patterning, but structurally identical. All three have a more colourful facial pattern.
|
||
<bibRefCitation id="EF9E4B2CFFDAFFE4E1987112FD25FCE5" author="Sasamoto, A. & Do, C. & Vu, L. V." box="[488,742,788,815]" pageId="26" pageNumber="27" pagination="587 - 594" refId="ref17356" refString="Sasamoto, A., Do, C. & Vu, L. V. (2013) Discovery of a new species of the genus Planaeschna from Northern Vietnam, with a first description of male P. tomokunii. Zootaxa, 3652 (5), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3652.5.8" type="journal article" year="2013">
|
||
Sasamoto
|
||
<emphasis id="B97BEACFFFDAFFE4E22C7112FD57FCE4" box="[604,660,788,814]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
did not compare their
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E3967112FB71FCE4" box="[998,1202,788,814]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi">
|
||
<emphasis id="B97BEACFFFDAFFE4E3967112FB71FCE4" box="[998,1202,788,814]" italics="true" pageId="26" pageNumber="27">P. guentherpetersi</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E4A67112FAB6FCE4" authorityName="Asahina" authorityYear="1951" box="[1238,1397,788,814]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E4A67112FAB6FCE4" box="[1238,1397,788,814]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
for their diagnosis, which was reasonable in view of the different facial pattern and in view of the large distributional gap between these species, even if the pattern on S2 was the same.
|
||
</paragraph>
|
||
<paragraph id="8BB036DDFFDAFFE4E0B77186FE3CFC28" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">
|
||
Our DNA analysis suggested that the three known subspecies of
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E3F87186FBE7FC50" authorityName="Asahina" authorityYear="1951" box="[904,1060,896,922]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E3F87186FBE7FC50" box="[904,1060,896,922]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
not only clustered well with slight differences, but surprisingly, that specimens of
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E2DF71A2FCAEFC74" box="[687,877,933,958]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guenterpetersi">
|
||
<emphasis id="B97BEACFFFDAFFE4E2DF71A2FCAEFC74" box="[687,877,933,958]" italics="true" pageId="26" pageNumber="27">P. guenterpetersi</emphasis>
|
||
</taxonomicName>
|
||
were also included in the same clade, suggesting that they were the same species.
|
||
</paragraph>
|
||
<paragraph id="8BB036DDFFDAFFE4E0B771EAFADFFA81" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">
|
||
In fact,
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E15371EAFE37FBCC" box="[291,500,1004,1031]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi">
|
||
<emphasis id="B97BEACFFFDAFFE4E15371EAFEF7FBCC" box="[291,308,1005,1030]" italics="true" pageId="26" pageNumber="27">P</emphasis>
|
||
.
|
||
<emphasis id="B97BEACFFFDAFFE4E13471EBFE37FBCC" box="[324,500,1004,1030]" italics="true" pageId="26" pageNumber="27">guentherpetersi</emphasis>
|
||
</taxonomicName>
|
||
not only has the same pattern on S2, but also identical anal appendages to those of
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E0C97616FE99FBE0" authorityName="Asahina" authorityYear="1951" box="[185,346,1040,1066]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E0C97616FE99FBE0" box="[185,346,1040,1066]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="13342A58FFDAFFE4E1EB7617FE2DFBE1" box="[411,494,1040,1067]" captionStart="FIGURE 17" captionStartId="28.[151,250,1349,1373]" captionTargetBox="[156,1436,194,1325]" captionTargetId="figure-22@28.[151,1436,181,1325]" captionTargetPageId="28" captionText="FIGURE 17. Planaeschna ishigakiana spp. anal appendages. [A–C] after Asahina (1988), lateral view (top) and dorsal view (bottom), [D] Xuan Son NP, Phu Tho Prov., 26-X-2014 (same as in Fig. 7). (A) P. i. ishigakiana, Ishigakijima (1988: p. 28, Figs 2–3), (B) P. i. nagaminei, Amami Oshima (1988: p. 29, Figs 13–14), (C) P. i. ishigakiana, Iriomote Island (1988: p. 28, Figs 4–5), (D) P. i. guentherpetersi stat. nov." figureDoi="http://doi.org/10.5281/zenodo.5448068" httpUri="https://zenodo.org/record/5448068/files/figure.png" pageId="26" pageNumber="27">Fig. 17</figureCitation>
|
||
for a comparison of
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E2987617FBF0FBE0" box="[744,1075,1040,1066]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi">
|
||
<emphasis id="B97BEACFFFDAFFE4E2987617FBF0FBE0" box="[744,1075,1040,1066]" italics="true" pageId="26" pageNumber="27">Planaeschna guentherpetersi</emphasis>
|
||
</taxonomicName>
|
||
from Xuan Son with drawings by
|
||
<bibRefCitation id="EF9E4B2CFFDAFFE4E0CC7633FEAEFB85" author="Asahina, S." box="[188,365,1076,1103]" pageId="26" pageNumber="27" pagination="27 - 34" refId="ref16291" refString="Asahina, S. (1988) A problematical race of Planaeschna ishigakiana from Amami-Oshima, Middle Ryukyus. Tombo, 31, 27 - 34." type="journal article" year="1988">Asahina (1988)</bibRefCitation>
|
||
of
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E1E97632FD96FB84" box="[409,597,1076,1102]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="ishigakiana" subSpecies="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E1E97632FD96FB84" box="[409,597,1076,1102]" italics="true" pageId="26" pageNumber="27">P. i. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B97BEACFFFDAFFE4E2E27632FC83FB84" box="[658,832,1077,1102]" italics="true" pageId="26" pageNumber="27">P. i. nagaminei</emphasis>
|
||
). The three known subspecies of
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E4BE7632FAB3FB84" authorityName="Asahina" authorityYear="1951" box="[1230,1392,1076,1102]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E4BE7632FAB3FB84" box="[1230,1392,1076,1102]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
are geographically isolated: the Yaeyama Islands are about
|
||
<quantity id="4CF79B38FFDAFFE4E28D765EFC92FBB8" box="[765,849,1112,1139]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="km" value="200.0">200 km</quantity>
|
||
from
|
||
<collectingCountry id="F318764DFFDAFFE4E3E2765FFC27FBB8" box="[914,996,1112,1138]" name="Taiwan" pageId="26" pageNumber="27">Taiwan</collectingCountry>
|
||
and over
|
||
<quantity id="4CF79B38FFDAFFE4E420765FFB67FBB8" box="[1104,1188,1112,1139]" metricMagnitude="5" metricUnit="m" metricValue="6.5" pageId="26" pageNumber="27" unit="km" value="650.0">650 km</quantity>
|
||
from Amami Oshima. It is therefore not surprising that these populations display some differences in colouration. Although specimens currently identified as
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E1EC76A6FDA8FB70" box="[412,619,1184,1210]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi">
|
||
<emphasis id="B97BEACFFFDAFFE4E1EC76A6FDA8FB70" box="[412,619,1184,1210]" italics="true" pageId="26" pageNumber="27">P. guentherpetersi</emphasis>
|
||
</taxonomicName>
|
||
likewise shows some differences in colouration compared to the other three subspecies (see diagnosis below), the DNA and morphological evidence presented here shows these are in fact conspecific with specimens belonging to
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E2D276EEFC80FAC8" authorityName="Asahina" authorityYear="1951" box="[674,835,1256,1282]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E2D276EEFC80FAC8" box="[674,835,1256,1282]" italics="true" pageId="26" pageNumber="27">P. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF9E4B2CFFDAFFE4E32176EFFBC3FAC9" author="Asahina, S." box="[849,1024,1256,1283]" pageId="26" pageNumber="27" pagination="27 - 34" refId="ref16291" refString="Asahina, S. (1988) A problematical race of Planaeschna ishigakiana from Amami-Oshima, Middle Ryukyus. Tombo, 31, 27 - 34." type="journal article" year="1988">Asahina (1988)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF9E4B2CFFDAFFE4E44976EFFB7AFAC9" author="Yeh, W." box="[1081,1209,1256,1283]" pageId="26" pageNumber="27" pagination="26 - 28" refId="ref17758" refString="Yeh, W. (1996) New records of the Odonata taken in Taiwan. Tombo, 39, 26 - 28." type="journal article" year="1996">Yeh (1996)</bibRefCitation>
|
||
gave precedence to structural similarities over differences in coloration between these taxa when determining specific status. We concur with their approach and therefore treat this taxon as a fourth subspecies:
|
||
<emphasis id="B97BEACFFFDAFFE4E3B47736FADFFA81" bold="true" box="[964,1308,1328,1355]" pageId="26" pageNumber="27">
|
||
<emphasis id="B97BEACFFFDAFFE4E3B47736FB69FA81" bold="true" box="[964,1194,1328,1355]" italics="true" pageId="26" pageNumber="27">
|
||
P. i.
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E3867737FB69FA81" authorityName="Kompier & Karube & Futahashi & Phan" authorityYear="2021" baseAuthorityName="Sasamoto, Do & Vu" baseAuthorityYear="2013" box="[1014,1194,1328,1355]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi" status="stat. nov.">guentherpetersi</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A24857B4FFDAFFE4E4C17736FADFFA81" box="[1201,1308,1329,1355]" pageId="26" pageNumber="27">stat. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB036DDFFDAFFE4E0B77752FC15FA58" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">To our knowledge, this is the first known example of a species within this genus with such a large disjunctive range consisting of isolated populations that differ distinctly in patterning.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156556FFDAFFE4E0B7779BFBD2F9E9" pageId="26" pageNumber="27" type="diagnosis">
|
||
<paragraph id="8BB036DDFFDAFFE4E0B7779BFBD2F9E9" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">
|
||
<emphasis id="B97BEACFFFDAFFE4E0B7779BFE16FA7C" bold="true" box="[199,469,1436,1462]" pageId="26" pageNumber="27">Differential Diagnosis.</emphasis>
|
||
The male of this subspecies differs from the other subspecies by the largely brown facial pattern lacking clear black and yellow markings (
|
||
<figureCitation id="13342A58FFDAFFE4E36277C7FCBFFA11" box="[786,892,1472,1499]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Fig. 16A</figureCitation>
|
||
), the more brown rather than black tones of the dark parts of the pterothorax (
|
||
<figureCitation id="13342A58FFDAFFE4E19D77E3FD91FA35" box="[493,594,1508,1535]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Fig. 16B</figureCitation>
|
||
) and from
|
||
<emphasis id="B97BEACFFFDAFFE4E2A377E2FCB1FA34" box="[723,882,1508,1534]" italics="true" pageId="26" pageNumber="27">P. i. flavostria</emphasis>
|
||
by its darker maculation (
|
||
<figureCitation id="13342A58FFDAFFE4E4EF77E3FAC4FA35" box="[1183,1287,1508,1535]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Fig. 16B</figureCitation>
|
||
). The female differs by the all brown face (
|
||
<figureCitation id="13342A58FFDAFFE4E192740FFD84F9E9" box="[482,583,1544,1571]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Fig. 16C</figureCitation>
|
||
) and the lateral pattern of S2 (
|
||
<figureCitation id="13342A58FFDAFFE4E3EB740FFBC7F9E9" box="[923,1028,1544,1571]" captionStart="FIGURE 16" captionStartId="27.[151,250,1942,1966]" captionTargetBox="[236,1352,181,1919]" captionTargetId="figure-17@27.[236,1352,181,1919]" captionTargetPageId="27" captionText="FIGURE 16. Planaeschna ishigakiana guentherpetersi [A, C] face alive, [B, D] lateral habitus alive, [E–F] scan of dorsal habitus, [G–H] anal appendages in lateral (G) and dorsal (H) view. Note that epiproct actually reaches halfway length of cerci. (A–B, E, G–H) male, Xuan Son NP, Phu Tho Prov., 26-X-2014, (C–D, F) female, same location, 23-XI-2014." figureDoi="http://doi.org/10.5281/zenodo.5448066" httpUri="https://zenodo.org/record/5448066/files/figure.png" pageId="26" pageNumber="27">Fig. 16D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156556FFDAFFE4E0B7742AFB7FF944" pageId="26" pageNumber="27" type="biology_ecology">
|
||
<paragraph id="8BB036DDFFDAFFE4E0B7742AFB7FF944" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">
|
||
<emphasis id="B97BEACFFFDAFFE4E0B7742AFEEFF98D" bold="true" box="[199,300,1581,1607]" pageId="26" pageNumber="27">Ecology.</emphasis>
|
||
This is the most common
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E23C742BFD1EF98C" authorityName="Kompier & Karube & Futahashi & Phan" authorityYear="2021" box="[588,733,1580,1606]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BEACFFFDAFFE4E23C742BFD1EF98C" box="[588,733,1580,1606]" italics="true" pageId="26" pageNumber="27">Planaeschna</emphasis>
|
||
</taxonomicName>
|
||
in Xuan Son NP, where it inhabits shallow streams under heavy forest cover. Females were observed during the day ovipositing in rotting wood just above the water surface, or hung up in nearby trees. Males were rarely observed. All records were from October and November.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156556FFDAFFE4E0B7749FFC90F91D" pageId="26" pageNumber="27" type="distribution">
|
||
<paragraph id="8BB036DDFFDAFFE4E0B7749FFC90F91D" blockId="26.[151,1437,356,1751]" pageId="26" pageNumber="27">
|
||
<emphasis id="B97BEACFFFDAFFE4E0B7749FFEA2F978" bold="true" box="[199,353,1688,1714]" pageId="26" pageNumber="27">Distribution.</emphasis>
|
||
<collectingCountry id="F318764DFFDAFFE4E117749EFE64F979" box="[359,423,1689,1715]" name="Japan" pageId="26" pageNumber="27">Japan</collectingCountry>
|
||
(Yaeyama Islands (
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E2F3749EFCF5F978" box="[643,822,1688,1714]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="ishigakiana" subSpecies="ishigakiana">
|
||
<emphasis id="B97BEACFFFDAFFE4E2F3749EFCF5F978" box="[643,822,1688,1714]" italics="true" pageId="26" pageNumber="27">P. i. ishigakiana</emphasis>
|
||
</taxonomicName>
|
||
), Amami Oshima (
|
||
<emphasis id="B97BEACFFFDAFFE4E47D749EFB70F978" box="[1037,1203,1689,1714]" italics="true" pageId="26" pageNumber="27">P. i. nagaminei</emphasis>
|
||
)),
|
||
<collectingCountry id="F318764DFFDAFFE4E4A3749FFAD5F978" box="[1235,1302,1688,1714]" name="China" pageId="26" pageNumber="27">China</collectingCountry>
|
||
(
|
||
<collectingCountry id="F318764DFFDAFFE4E554749FFABBF978" box="[1316,1400,1688,1714]" name="Taiwan" pageId="26" pageNumber="27">Taiwan</collectingCountry>
|
||
(
|
||
<emphasis id="B97BEACFFFDAFFE4E5F8749EFEDBF91C" italics="true" pageId="26" pageNumber="27">P. i. flavostria</emphasis>
|
||
)),
|
||
<collectingCountry id="F318764DFFDAFFE4E14874BBFE54F91C" box="[312,407,1724,1750]" name="Vietnam" pageId="26" pageNumber="27">Vietnam</collectingCountry>
|
||
(
|
||
<collectingRegion id="49CBF83FFFDAFFE4E1D674BBFD8BF91D" box="[422,584,1724,1751]" country="Vietnam" name="Phu Tho" pageId="26" pageNumber="27">Phu Tho Prov.</collectingRegion>
|
||
(
|
||
<emphasis id="B97BEACFFFDAFFE4E22874BAFCFAF91C" box="[600,825,1724,1750]" italics="true" pageId="26" pageNumber="27">
|
||
P. i.
|
||
<taxonomicName id="4C0F4D5EFFDAFFE4E2F974BBFCFAF91C" authorityName="Kompier & Karube & Futahashi & Phan" authorityYear="2021" baseAuthorityName="Sasamoto, Do & Vu" baseAuthorityYear="2013" box="[649,825,1724,1750]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="guentherpetersi">guentherpetersi</taxonomicName>
|
||
</emphasis>
|
||
)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |