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<document id="49D313D573CA03513063EEDA37CC417F" ID-DOI="10.11646/zootaxa.5027.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5448029" ID-ZooBank="CCF10500-01A3-476C-A4BE-57161E164EBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630464259914" checkinUser="plazi" docAuthor="Kompier, Tom, Karube, Haruki, Futahashi, Ryo &amp; Phan, Quoc Toan" docDate="2021" docId="03A687CBFFC7FFF9E0E770D1FCDEF8D5" docLanguage="en" docName="zootaxa.5027.1.1.pdf" docOrigin="Zootaxa 5027 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Planaeschna tomokunii : Asahina 1996" docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="FF9FFFB3FFC0FFFEE0707207FFC3FFCA" masterDocTitle="The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="8" updateTime="1699280300883" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D40FACC9515BEA2611A371F7D42C2CD4">The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)</mods:title>
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<mods:namePart id="D29E9A26801F191E1B0E0DB3B9CD26CC">Kompier, Tom</mods:namePart>
<mods:affiliation id="6FD34512251EA93B8C1F751D5F3AE90E">Schoutenstraat 69, 2596 SK Den Haag, the Netherlands.</mods:affiliation>
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<mods:namePart id="B108927BD43FCC99D3272458791FF735">Karube, Haruki</mods:namePart>
<mods:affiliation id="4F0DFFC860468B1074B9759604C239DF">Kanagawa Prefectural Museum of Natural History, Odawara, Japan.</mods:affiliation>
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<mods:namePart id="F1C0C438F1514CC09F5E7DCFDD8677B3">Futahashi, Ryo</mods:namePart>
<mods:affiliation id="05B81BC83A7B48D9A40310E06A33D332">National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan.</mods:affiliation>
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<mods:namePart id="E4C2E0D0B2C95A996F61E59D97334803">Phan, Quoc Toan</mods:namePart>
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<mods:affiliation id="E848076C0E0E8F317C5D39623BDC6A9D">The Center for Entomology &amp; Parasitology Research, College of Medicine &amp; Pharmacy, Duy Tan University, 550000, Da Nang, Vietnam. pqtoan 84 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3154 - 6546</mods:affiliation>
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<paragraph id="8BB036DDFFC7FFF9E0E770D1FD49FD3A" blockId="7.[151,650,726,789]" box="[151,650,726,753]" pageId="7" pageNumber="8">
<heading id="D0F881B1FFC7FFF9E0E770D1FD49FD3A" bold="true" box="[151,650,726,753]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B97BEACFFFC7FFF9E0E770D1FD49FD3A" bold="true" box="[151,650,726,753]" pageId="7" pageNumber="8">
2.
<taxonomicName id="4C0F4D5EFFC7FFF9E0C570D1FD49FD3A" ID-CoL="6VN7P" authority="Asahina, 1996" authorityName=": Asahina" authorityYear="1996" box="[181,650,726,753]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E0C570D1FE17FD3B" bold="true" box="[181,468,726,753]" italics="true" pageId="7" pageNumber="8">Planaeschna tomokunii</emphasis>
<bibRefCitation id="EF9E4B2CFFC7FFF9E1AA70D1FD49FD3A" author="Asahina, S." box="[474,650,726,752]" pageId="7" pageNumber="8" pagination="69 - 80" refId="ref16318" refString="Asahina, S. (1996) Records of the northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo 3. Aeschnidae, Corduliidae and Libellulidae. Bulletin National Science Museum Tokyo, Series A, 22, 69 - 80." type="journal article" year="1996">Asahina, 1996</bibRefCitation>
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<paragraph id="8BB036DDFFC7FFF9E0E770FCFF28FCDF" blockId="7.[151,650,726,789]" box="[151,235,763,789]" pageId="7" pageNumber="8">
(
<figureCitation id="13342A58FFC7FFF9E0D070FCFF20FCDF" box="[160,227,763,789]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1968]" captionTargetBox="[247,1338,181,1920]" captionTargetId="figure-22@8.[247,1339,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Planaeschna tomokunii [AC] lateral habitus alive, including face, [DE] scans of dorsal habitus, [FG] anal appendages of male in lateral (F) and dorsal (G) view. (A, FG) male, Tam Dao NP, Vinh Phuc Prov., 3-VII-2016, (B) immature male, same location, 4-VI-2016, (C, E) female, same location, 24-IX-2015, (D) male, same location, 16-VI-2016." figureDoi="http://doi.org/10.5281/zenodo.5448039" httpUri="https://zenodo.org/record/5448039/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
)
</paragraph>
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<paragraph id="8BB036DDFFC7FFF9E0E77138FC07FCBC" blockId="7.[151,1437,831,886]" pageId="7" pageNumber="8">
<taxonomicName id="4C0F4D5EFFC7FFF9E0E77138FDFAFC9D" ID-CoL="6VN7P" authority=": Asahina (1996)" authorityName=": Asahina" authorityYear="1996" box="[151,569,831,855]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E0E77138FE4AFC9D" box="[151,393,831,855]" italics="true" pageId="7" pageNumber="8">Planaeschna tomokunii</emphasis>
:
<bibRefCitation id="EF9E4B2CFFC7FFF9E1E97138FDFAFC9D" author="Asahina, S." box="[409,569,831,855]" pageId="7" pageNumber="8" pagination="69 - 80" refId="ref16318" refString="Asahina, S. (1996) Records of the northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo 3. Aeschnidae, Corduliidae and Libellulidae. Bulletin National Science Museum Tokyo, Series A, 22, 69 - 80." type="journal article" year="1996">Asahina (1996)</bibRefCitation>
</taxonomicName>
pp. 7072,
<figureCitation id="13342A58FFC7FFF9E2CC7138FCD4FC9D" box="[700,791,831,855]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="8.[151,250,1944,1968]" captionStartId-1="9.[151,250,1943,1967]" captionStartId-2="11.[151,250,1801,1825]" captionTargetBox-0="[247,1338,181,1920]" captionTargetBox-1="[225,1363,181,1920]" captionTargetBox-2="[151,1436,921,1778]" captionTargetId-0="figure-22@8.[247,1339,181,1920]" captionTargetId-1="figure-17@9.[225,1363,181,1920]" captionTargetId-2="figure-362@11.[151,1436,921,1778]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="11" captionText-0="FIGURE 4. Planaeschna tomokunii [AC] lateral habitus alive, including face, [DE] scans of dorsal habitus, [FG] anal appendages of male in lateral (F) and dorsal (G) view. (A, FG) male, Tam Dao NP, Vinh Phuc Prov., 3-VII-2016, (B) immature male, same location, 4-VI-2016, (C, E) female, same location, 24-IX-2015, (D) male, same location, 16-VI-2016." captionText-1="FIGURE 5. Planaeschna tsuchi sp. nov. [AB] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [EF] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, EF) holotype, (D) female paratype, 24-X-2015." captionText-2="FIGURE 6. Planaeschna viridis male, Bach Ma NP. Thus Thien-Hue Prov., 7-VIII-2016. (A) lateral habitus alive, including face, [BC] anal appendages in lateral (B) and dorsal (C) view, (D) dorsal pattern of S2." figureDoi-0="http://doi.org/10.5281/zenodo.5448039" figureDoi-1="http://doi.org/10.5281/zenodo.5448041" figureDoi-2="http://doi.org/10.5281/zenodo.5448043" httpUri-0="https://zenodo.org/record/5448039/files/figure.png" httpUri-1="https://zenodo.org/record/5448041/files/figure.png" httpUri-2="https://zenodo.org/record/5448043/files/figure.png" pageId="7" pageNumber="8">Figs 46</figureCitation>
(description of female
<typeStatus id="54B4887FFFC7FFF9E4617138FBADFC9D" box="[1041,1134,831,855]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
);
<bibRefCitation id="EF9E4B2CFFC7FFF9E4F37138FAB0FC9D" author="Sasamoto, A. &amp; Do, C. &amp; Vu, L. V." box="[1155,1395,831,855]" pageId="7" pageNumber="8" pagination="587 - 594" refId="ref17356" refString="Sasamoto, A., Do, C. &amp; Vu, L. V. (2013) Discovery of a new species of the genus Planaeschna from Northern Vietnam, with a first description of male P. tomokunii. Zootaxa, 3652 (5), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3652.5.8" type="journal article" year="2013">
Sasamoto
<emphasis id="B97BEACFFFC7FFF9E49F7147FAE5FC9D" box="[1263,1318,831,855]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2013)
</bibRefCitation>
pp. 590593,
<figureCitation id="13342A58FFC7FFF9E15D7159FEB7FCBC" box="[301,372,862,886]" captionStart="FIGURE 11" captionStartId="20.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1647]" captionTargetId="figure-22@20.[151,1436,181,1647]" captionTargetPageId="20" captionText="FIGURE 11. Comparison of details of Planaeschna crux sp. nov. and P. cucphuongensis [AB, HI] dorsal pattern of S2, [C, J] dorsal view of thorax showing antealar sinus, [D, F] male anal appendages in dorsal view, [E, G] male anal appendages in lateral view. (A, CE) holotype P. crux, (B) paratype female P. crux used for description, (FH, J) male P. cucphuongensis, Xuan Son NP, Phu Tho Prov., 22-X-2016, (I) female P. cucphuongensis, Cuc Phuong NP, Ninh Binh Prov., 16-VI-2015." figureDoi="http://doi.org/10.5281/zenodo.5448056" httpUri="https://zenodo.org/record/5448056/files/figure.png" pageId="7" pageNumber="8">Figs 11</figureCitation>
22 (first description of male, photos of female
<typeStatus id="54B4887FFFC7FFF9E32C7159FC7AFCBC" box="[860,953,862,886]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
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<paragraph id="8BB036DDFFC7FFF9E0E771A3FAF9FBCC" blockId="7.[151,1437,932,1823]" pageId="7" pageNumber="8">
<emphasis id="B97BEACFFFC7FFF9E0E771A3FE4EFC74" bold="true" box="[151,397,932,958]" pageId="7" pageNumber="8">Materials examined.</emphasis>
<materialsCitation id="3B673C80FFC7FFF9E1E571A3FBEAFC75" box="[405,1065,932,959]" collectingDate="2015-09-24" collectorName="Tam Dao NP" country="Vietnam" location="Vinh Phuc Prov." pageId="7" pageNumber="8" specimenCount="1" specimenCount-female="1" stateProvince="Vinh Phuc">
<specimenCount id="9D09FD54FFC7FFF9E1E571A3FE03FC75" box="[405,448,932,959]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
,
<collectorName id="26FA530BFFC7FFF9E1BE71A2FDA1FC75" box="[462,610,933,959]" pageId="7" pageNumber="8">Tam Dao NP</collectorName>
,
<collectingRegion id="49CBF83FFFC7FFF9E21E71A3FCE9FC75" box="[622,810,932,959]" country="Vietnam" name="Vinh Phuc" pageId="7" pageNumber="8">Vinh Phuc Prov.</collectingRegion>
,
<collectingCountry id="F318764DFFC7FFF9E34571A3FC54FC74" box="[821,919,932,958]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
,
<date id="FFB1101DFFC7FFF9E3D371A3FBEAFC75" box="[931,1065,932,959]" pageId="7" pageNumber="8" value="2015-09-24">
<collectingDate id="EFF5E9F5FFC7FFF9E3D371A3FBEAFC75" box="[931,1065,932,959]" pageId="7" pageNumber="8" value="2015-09-24">24-IX-2015</collectingDate>
</date>
</materialsCitation>
, TK leg.;
<materialsCitation id="3B673C80FFC7FFF9E4EC71A3FE40FC28" collectingDate="2016-06-04" collectorName="Tam Dao NP" country="Vietnam" location="Vinh Phuc Prov." pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="9D09FD54FFC7FFF9E4EC71A3FB0AFC75" box="[1180,1225,932,959]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
, same locality and collector,
<date id="FFB1101DFFC7FFF9E17971CFFE40FC28" box="[265,387,968,994]" pageId="7" pageNumber="8" value="2016-06-04">
<collectingDate id="EFF5E9F5FFC7FFF9E17971CFFE40FC28" box="[265,387,968,994]" pageId="7" pageNumber="8" value="2016-06-04">4-VI-2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673C80FFC7FFF9E1E071CFFC58FC28" box="[400,923,968,995]" collectingDate="2016-07-03" collectorName="Tam Dao NP" country="Vietnam" location="Vinh Phuc Prov." pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="9D09FD54FFC7FFF9E1E071CFFE7CFC29" box="[400,447,968,995]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
, same location and collector,
<date id="FFB1101DFFC7FFF9E36871CFFC58FC28" box="[792,923,968,994]" pageId="7" pageNumber="8" value="2016-07-03">
<collectingDate id="EFF5E9F5FFC7FFF9E36871CFFC58FC28" box="[792,923,968,994]" pageId="7" pageNumber="8" value="2016-07-03">3-VII-2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673C80FFC7FFF9E3D971CFFF10FBCC" collectingDate="2016-07-16" collectorName="Tam Dao NP" country="Vietnam" location="Vinh Phuc Prov." pageId="7" pageNumber="8" specimenCount="2" specimenCount-male="2" stateProvince="Vinh Phuc">
<specimenCount id="9D09FD54FFC7FFF9E3D971CFFC2EFC29" box="[937,1005,968,995]" pageId="7" pageNumber="8" type="male">2 ♂♂</specimenCount>
, same location and collector,
<date id="FFB1101DFFC7FFF9E53771CFFF10FBCC" pageId="7" pageNumber="8" value="2016-07-16">
<collectingDate id="EFF5E9F5FFC7FFF9E53771CFFF10FBCC" pageId="7" pageNumber="8" value="2016-07-16">16-VII- 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673C80FFC7FFF9E0AE71EBFCDBFBCC" box="[222,792,1004,1031]" collectingDate="2016-07-17" collectorName="Tam Dao NP" country="Vietnam" location="Vinh Phuc Prov." pageId="7" pageNumber="8" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="9D09FD54FFC7FFF9E0AE71EBFECBFBCD" box="[222,264,1004,1031]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
<specimenCount id="9D09FD54FFC7FFF9E17F71EBFEFAFBCD" box="[271,313,1004,1031]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
, same location and collector,
<date id="FFB1101DFFC7FFF9E2F771EBFCDBFBCC" box="[647,792,1004,1030]" pageId="7" pageNumber="8" value="2016-07-17">
<collectingDate id="EFF5E9F5FFC7FFF9E2F771EBFCDBFBCC" box="[647,792,1004,1030]" pageId="7" pageNumber="8" value="2016-07-17">17-VII-2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673C80FFC7FFF9E35371EBFAF5FBCC" box="[803,1334,1004,1031]" collectingDate="2016-08-27" collectorName="Tam Dao NP" country="Vietnam" location="Vinh Phuc Prov." pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="9D09FD54FFC7FFF9E35371EBFC8CFBCD" box="[803,847,1004,1031]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
, same location and collector,
<date id="FFB1101DFFC7FFF9E4EB71EBFAF5FBCC" box="[1179,1334,1004,1030]" pageId="7" pageNumber="8" value="2016-08-27">
<collectingDate id="EFF5E9F5FFC7FFF9E4EB71EBFAF5FBCC" box="[1179,1334,1004,1030]" pageId="7" pageNumber="8" value="2016-08-27">27-VIII-2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036DDFFC7FFF9E0B77617FA4EFAED" blockId="7.[151,1437,932,1823]" pageId="7" pageNumber="8">
<emphasis id="B97BEACFFFC7FFF9E0B77617FED2FBE0" bold="true" box="[199,273,1040,1066]" pageId="7" pageNumber="8">Notes.</emphasis>
<bibRefCitation id="EF9E4B2CFFC7FFF9E16B7617FE0EFBE1" author="Asahina, S." box="[283,461,1040,1067]" pageId="7" pageNumber="8" pagination="69 - 80" refId="ref16318" refString="Asahina, S. (1996) Records of the northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo 3. Aeschnidae, Corduliidae and Libellulidae. Bulletin National Science Museum Tokyo, Series A, 22, 69 - 80." type="journal article" year="1996">Asahina (1996)</bibRefCitation>
described
<taxonomicName id="4C0F4D5EFFC7FFF9E23F7616FD1CFBE0" authorityName=": Asahina" authorityYear="1996" box="[591,735,1040,1066]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E23F7616FD1CFBE0" box="[591,735,1040,1066]" italics="true" pageId="7" pageNumber="8">P. tomokunii</emphasis>
</taxonomicName>
based on a single female. Without detailed explanation, he considered
<taxonomicName id="4C0F4D5EFFC7FFF9E16A7632FE6AFB84" authorityName=": Asahina" authorityYear="1996" box="[282,425,1076,1102]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E16A7632FE6AFB84" box="[282,425,1076,1102]" italics="true" pageId="7" pageNumber="8">P. tomokunii</emphasis>
</taxonomicName>
to be closely related to
<taxonomicName id="4C0F4D5EFFC7FFF9E2B57632FCFDFB84" authorityName="Asahina" authorityYear="1951" box="[709,830,1077,1102]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis id="B97BEACFFFC7FFF9E2B57632FCFDFB84" box="[709,830,1077,1102]" italics="true" pageId="7" pageNumber="8">P. taiwana</emphasis>
</taxonomicName>
. The description of
<taxonomicName id="4C0F4D5EFFC7FFF9E45B7633FAD4FB84" box="[1067,1303,1076,1102]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BEACFFFC7FFF9E45B7633FB7FFB84" box="[1067,1212,1076,1102]" italics="true" pageId="7" pageNumber="8">Planaeschna</emphasis>
species
</taxonomicName>
based on a single female is problematic, because there is little interspecific structural variation in females. For the identification of
<taxonomicName id="4C0F4D5EFFC7FFF9E0C8767BFE8AFB5C" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[184,329,1148,1174]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEACFFFC7FFF9E0C8767BFE8AFB5C" box="[184,329,1148,1174]" italics="true" pageId="7" pageNumber="8">Planaeschna</emphasis>
</taxonomicName>
, the combination of the morphology of the male anal appendages and the maculation pattern is effective, but in 1996 there were no
<taxonomicName id="4C0F4D5EFFC7FFF9E25976A7FD79FB70" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[553,698,1184,1210]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEACFFFC7FFF9E25976A7FD79FB70" box="[553,698,1184,1210]" italics="true" pageId="7" pageNumber="8">Planaeschna</emphasis>
</taxonomicName>
known from
<collectingCountry id="F318764DFFC7FFF9E33F76A7FC71FB70" box="[847,946,1184,1210]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
.
<taxonomicName id="4C0F4D5EFFC7FFF9E3CC76A7FB1CFB70" authorityName=": Asahina" authorityYear="1996" box="[956,1247,1184,1210]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B97BEACFFFC7FFF9E3CC76A7FB1CFB70" box="[956,1247,1184,1210]" italics="true" pageId="7" pageNumber="8">Planaeschna tamdaoensis</emphasis>
</taxonomicName>
with a black and yellow face and plain-faced
<taxonomicName id="4C0F4D5EFFC7FFF9E1AE76C2FDAFFB14" authorityName=": Asahina" authorityYear="1996" box="[478,620,1220,1246]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E1AE76C2FDAFFB14" box="[478,620,1220,1246]" italics="true" pageId="7" pageNumber="8">P. tomokunii</emphasis>
</taxonomicName>
were the first two species described.
<taxonomicName id="4C0F4D5EFFC7FFF9E45176C3FAE8FB14" authorityName=": Asahina" authorityYear="1996" box="[1057,1323,1220,1246]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E45176C3FAE8FB14" box="[1057,1323,1220,1246]" italics="true" pageId="7" pageNumber="8">Planaeschna tomokunii</emphasis>
</taxonomicName>
remained enigmatic until
<bibRefCitation id="EF9E4B2CFFC7FFF9E13476EFFDFDFAC8" author="Sasamoto, A. &amp; Do, C. &amp; Vu, L. V." box="[324,574,1256,1282]" pageId="7" pageNumber="8" pagination="587 - 594" refId="ref17356" refString="Sasamoto, A., Do, C. &amp; Vu, L. V. (2013) Discovery of a new species of the genus Planaeschna from Northern Vietnam, with a first description of male P. tomokunii. Zootaxa, 3652 (5), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3652.5.8" type="journal article" year="2013">
Sasamoto
<emphasis id="B97BEACFFFC7FFF9E1C776EEFE2DFAC8" box="[439,494,1256,1282]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2013)
</bibRefCitation>
described the male based on specimens collected at the
<typeStatus id="54B4887FFFC7FFF9E4D676EEFB15FAC9" box="[1190,1238,1257,1283]" pageId="7" pageNumber="8">type</typeStatus>
locality, allowing the species to be correctly identified. So far,
<taxonomicName id="4C0F4D5EFFC7FFF9E2FB770AFCD5FAEC" authorityName=": Asahina" authorityYear="1996" box="[651,790,1292,1318]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E2FB770AFCD5FAEC" box="[651,790,1292,1318]" italics="true" pageId="7" pageNumber="8">P. tomokunii</emphasis>
</taxonomicName>
may be a true endemic of the Tam Dao mountain range.
</paragraph>
<paragraph id="8BB036DDFFC7FFF9E0B77737FB2CF9A1" blockId="7.[151,1437,932,1823]" pageId="7" pageNumber="8">
Of all
<taxonomicName id="4C0F4D5EFFC7FFF9E1647737FE3CFA80" box="[276,511,1328,1354]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BEACFFFC7FFF9E1647737FE66FA80" box="[276,421,1328,1354]" italics="true" pageId="7" pageNumber="8">Planaeschna</emphasis>
species
</taxonomicName>
known from
<collectingCountry id="F318764DFFC7FFF9E2ED7737FD3FFA80" box="[669,764,1328,1354]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
or
<collectingCountry id="F318764DFFC7FFF9E3567737FCAFFA80" box="[806,876,1328,1354]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
, none have the combination of slender cerci and dorsal pattern of S2 of male
<taxonomicName id="4C0F4D5EFFC7FFF9E1A97752FDA6FAA4" authorityName=": Asahina" authorityYear="1996" box="[473,613,1364,1390]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E1A97752FDA6FAA4" box="[473,613,1364,1390]" italics="true" pageId="7" pageNumber="8">P. tomokunii</emphasis>
</taxonomicName>
. The dorsal and lateral pattern of S2 is somewhat like that of
<taxonomicName id="4C0F4D5EFFC7FFF9E56C7752FAA9FAA4" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[1308,1386,1365,1390]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crux">
<emphasis id="B97BEACFFFC7FFF9E56C7752FAA9FAA4" box="[1308,1386,1365,1390]" italics="true" pageId="7" pageNumber="8">P. crux</emphasis>
</taxonomicName>
, but that species has clearly paddle-shaped cerci (see
<figureCitation id="13342A58FFC7FFF9E2BF777FFC9BFA58" box="[719,856,1400,1426]" captionStart="FIGURE 11" captionStartId="20.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1647]" captionTargetId="figure-22@20.[151,1436,181,1647]" captionTargetPageId="20" captionText="FIGURE 11. Comparison of details of Planaeschna crux sp. nov. and P. cucphuongensis [AB, HI] dorsal pattern of S2, [C, J] dorsal view of thorax showing antealar sinus, [D, F] male anal appendages in dorsal view, [E, G] male anal appendages in lateral view. (A, CE) holotype P. crux, (B) paratype female P. crux used for description, (FH, J) male P. cucphuongensis, Xuan Son NP, Phu Tho Prov., 22-X-2016, (I) female P. cucphuongensis, Cuc Phuong NP, Ninh Binh Prov., 16-VI-2015." figureDoi="http://doi.org/10.5281/zenodo.5448056" httpUri="https://zenodo.org/record/5448056/files/figure.png" pageId="7" pageNumber="8">Fig. 11AE</figureCitation>
). The anal appendages of
<taxonomicName id="4C0F4D5EFFC7FFF9E4FA777EFADBFA58" authorityName=": Asahina" authorityYear="1996" box="[1162,1304,1400,1426]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E4FA777EFADBFA58" box="[1162,1304,1400,1426]" italics="true" pageId="7" pageNumber="8">P. tomokunii</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C0F4D5EFFC7FFF9E16E779AFE5BFA7C" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[286,408,1437,1462]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="samurai">
<emphasis id="B97BEACFFFC7FFF9E16E779AFE5BFA7C" box="[286,408,1437,1462]" italics="true" pageId="7" pageNumber="8">P. samurai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D5EFFC7FFF9E1A5779AFDF8FA7C" authorityName=": Karube" authorityYear="2004" box="[469,571,1436,1462]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B97BEACFFFC7FFF9E1A5779AFDF8FA7C" box="[469,571,1436,1462]" italics="true" pageId="7" pageNumber="8">P. viridis</emphasis>
</taxonomicName>
(compare
<figureCitation id="13342A58FFC7FFF9E2C9779BFCECFA7C" box="[697,815,1436,1462]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1968]" captionTargetBox="[247,1338,181,1920]" captionTargetId="figure-22@8.[247,1339,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Planaeschna tomokunii [AC] lateral habitus alive, including face, [DE] scans of dorsal habitus, [FG] anal appendages of male in lateral (F) and dorsal (G) view. (A, FG) male, Tam Dao NP, Vinh Phuc Prov., 3-VII-2016, (B) immature male, same location, 4-VI-2016, (C, E) female, same location, 24-IX-2015, (D) male, same location, 16-VI-2016." figureDoi="http://doi.org/10.5281/zenodo.5448039" httpUri="https://zenodo.org/record/5448039/files/figure.png" pageId="7" pageNumber="8">Fig. 4FG</figureCitation>
with
<figureCitation id="13342A58FFC7FFF9E305779BFC37FA7C" box="[885,1012,1436,1462]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1966]" captionTargetBox="[242,1345,181,1919]" captionTargetId="figure-22@6.[242,1345,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Planaeschna samurai sp. nov. [A, C] face, [B, D] lateral habitus alive, [E, F] scan of dorsal habitus, [GH] anal appendages in dorsal (G) and lateral (H) view. (A) paratype male, Pia Oac NP, Cao Bang Prov., 18-X-2014, (B, E, GH) holotype, (CD, F) paratype female, Tu Le, Yen Bai Prov., 16-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448037" httpUri="https://zenodo.org/record/5448037/files/figure.png" pageId="7" pageNumber="8">Figs 3GH</figureCitation>
and
<figureCitation id="13342A58FFC7FFF9E441779BFBB5FA7D" box="[1073,1142,1436,1463]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1825]" captionTargetBox="[151,1436,921,1778]" captionTargetId="figure-362@11.[151,1436,921,1778]" captionTargetPageId="11" captionText="FIGURE 6. Planaeschna viridis male, Bach Ma NP. Thus Thien-Hue Prov., 7-VIII-2016. (A) lateral habitus alive, including face, [BC] anal appendages in lateral (B) and dorsal (C) view, (D) dorsal pattern of S2." figureDoi="http://doi.org/10.5281/zenodo.5448043" httpUri="https://zenodo.org/record/5448043/files/figure.png" pageId="7" pageNumber="8">6BC</figureCitation>
), but differ in detail. The cerci are more parallel-sided in dorsal view compared to
<taxonomicName id="4C0F4D5EFFC7FFF9E36D77C6FC55FA10" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[797,918,1473,1498]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="samurai">
<emphasis id="B97BEACFFFC7FFF9E36D77C6FC55FA10" box="[797,918,1473,1498]" italics="true" pageId="7" pageNumber="8">P. samurai</emphasis>
</taxonomicName>
, with a slightly longer epiproct, and in lateral view not waved and curving downward as in
<taxonomicName id="4C0F4D5EFFC7FFF9E2EA77E2FD3CFA34" authorityName=": Karube" authorityYear="2004" box="[666,767,1508,1534]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B97BEACFFFC7FFF9E2EA77E2FD3CFA34" box="[666,767,1508,1534]" italics="true" pageId="7" pageNumber="8">P. viridis</emphasis>
</taxonomicName>
, but gently curving upwards. The dorsal and lateral pattern of S2 distinguishes it clearly from both
<taxonomicName id="4C0F4D5EFFC7FFF9E215740EFD1CF9E8" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[613,735,1545,1570]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="samurai">
<emphasis id="B97BEACFFFC7FFF9E215740EFD1CF9E8" box="[613,735,1545,1570]" italics="true" pageId="7" pageNumber="8">P. samurai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D5EFFC7FFF9E36A740EFC43F9E8" authorityName=": Karube" authorityYear="2004" box="[794,896,1544,1570]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B97BEACFFFC7FFF9E36A740EFC43F9E8" box="[794,896,1544,1570]" italics="true" pageId="7" pageNumber="8">P. viridis</emphasis>
</taxonomicName>
(compare
<figureCitation id="13342A58FFC7FFF9E38C740FFBB0F9E8" box="[1020,1139,1544,1570]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1968]" captionTargetBox="[247,1338,181,1920]" captionTargetId="figure-22@8.[247,1339,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Planaeschna tomokunii [AC] lateral habitus alive, including face, [DE] scans of dorsal habitus, [FG] anal appendages of male in lateral (F) and dorsal (G) view. (A, FG) male, Tam Dao NP, Vinh Phuc Prov., 3-VII-2016, (B) immature male, same location, 4-VI-2016, (C, E) female, same location, 24-IX-2015, (D) male, same location, 16-VI-2016." figureDoi="http://doi.org/10.5281/zenodo.5448039" httpUri="https://zenodo.org/record/5448039/files/figure.png" pageId="7" pageNumber="8">Fig. 4DE</figureCitation>
with
<figureCitation id="13342A58FFC7FFF9E4C8740FFAEDF9E8" box="[1208,1326,1544,1570]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1966]" captionTargetBox="[242,1345,181,1919]" captionTargetId="figure-22@6.[242,1345,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Planaeschna samurai sp. nov. [A, C] face, [B, D] lateral habitus alive, [E, F] scan of dorsal habitus, [GH] anal appendages in dorsal (G) and lateral (H) view. (A) paratype male, Pia Oac NP, Cao Bang Prov., 18-X-2014, (B, E, GH) holotype, (CD, F) paratype female, Tu Le, Yen Bai Prov., 16-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448037" httpUri="https://zenodo.org/record/5448037/files/figure.png" pageId="7" pageNumber="8">Figs 3EF</figureCitation>
and
<figureCitation id="13342A58FFC7FFF9E51A740FFA53F9E9" box="[1386,1424,1544,1571]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1825]" captionTargetBox="[151,1436,921,1778]" captionTargetId="figure-362@11.[151,1436,921,1778]" captionTargetPageId="11" captionText="FIGURE 6. Planaeschna viridis male, Bach Ma NP. Thus Thien-Hue Prov., 7-VIII-2016. (A) lateral habitus alive, including face, [BC] anal appendages in lateral (B) and dorsal (C) view, (D) dorsal pattern of S2." figureDoi="http://doi.org/10.5281/zenodo.5448043" httpUri="https://zenodo.org/record/5448043/files/figure.png" pageId="7" pageNumber="8">6D</figureCitation>
). Furthermore, the triangular spot on the metepisternum is confluent with the lateral stripe on the mesepimeron in
<taxonomicName id="4C0F4D5EFFC7FFF9E5F8742AFF30F9A0" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="samurai">
<emphasis id="B97BEACFFFC7FFF9E5F8742AFF30F9A0" italics="true" pageId="7" pageNumber="8">P. samurai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D5EFFC7FFF9E1597456FE4EF9A0" authorityName=": Karube" authorityYear="2004" box="[297,397,1616,1642]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B97BEACFFFC7FFF9E1597456FE4EF9A0" box="[297,397,1616,1642]" italics="true" pageId="7" pageNumber="8">P. viridis</emphasis>
</taxonomicName>
, but separated in
<taxonomicName id="4C0F4D5EFFC7FFF9E2237456FD1DF9A0" authorityName=": Asahina" authorityYear="1996" box="[595,734,1616,1642]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E2237456FD1DF9A0" box="[595,734,1616,1642]" italics="true" pageId="7" pageNumber="8">P. tomokunii</emphasis>
</taxonomicName>
(compare
<figureCitation id="13342A58FFC7FFF9E3267457FC0FF9A0" box="[854,972,1616,1642]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1968]" captionTargetBox="[247,1338,181,1920]" captionTargetId="figure-22@8.[247,1339,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Planaeschna tomokunii [AC] lateral habitus alive, including face, [DE] scans of dorsal habitus, [FG] anal appendages of male in lateral (F) and dorsal (G) view. (A, FG) male, Tam Dao NP, Vinh Phuc Prov., 3-VII-2016, (B) immature male, same location, 4-VI-2016, (C, E) female, same location, 24-IX-2015, (D) male, same location, 16-VI-2016." figureDoi="http://doi.org/10.5281/zenodo.5448039" httpUri="https://zenodo.org/record/5448039/files/figure.png" pageId="7" pageNumber="8">Fig. 4AC</figureCitation>
with
<figureCitation id="13342A58FFC7FFF9E47C7457FB45F9A1" box="[1036,1158,1616,1643]" captionStart="FIGURE 3" captionStartId="6.[151,250,1942,1966]" captionTargetBox="[242,1345,181,1919]" captionTargetId="figure-22@6.[242,1345,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Planaeschna samurai sp. nov. [A, C] face, [B, D] lateral habitus alive, [E, F] scan of dorsal habitus, [GH] anal appendages in dorsal (G) and lateral (H) view. (A) paratype male, Pia Oac NP, Cao Bang Prov., 18-X-2014, (B, E, GH) holotype, (CD, F) paratype female, Tu Le, Yen Bai Prov., 16-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448037" httpUri="https://zenodo.org/record/5448037/files/figure.png" pageId="7" pageNumber="8">Figs 3B, D</figureCitation>
and
<figureCitation id="13342A58FFC7FFF9E4CD7457FB21F9A1" box="[1213,1250,1616,1643]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1825]" captionTargetBox="[151,1436,921,1778]" captionTargetId="figure-362@11.[151,1436,921,1778]" captionTargetPageId="11" captionText="FIGURE 6. Planaeschna viridis male, Bach Ma NP. Thus Thien-Hue Prov., 7-VIII-2016. (A) lateral habitus alive, including face, [BC] anal appendages in lateral (B) and dorsal (C) view, (D) dorsal pattern of S2." figureDoi="http://doi.org/10.5281/zenodo.5448043" httpUri="https://zenodo.org/record/5448043/files/figure.png" pageId="7" pageNumber="8">6A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFC7FFF9E0B77473FD6BF931" pageId="7" pageNumber="8" type="biology_ecology">
<paragraph id="8BB036DDFFC7FFF9E0B77473FD6BF931" blockId="7.[151,1437,932,1823]" pageId="7" pageNumber="8">
<emphasis id="B97BEACFFFC7FFF9E0B77473FEEFF944" bold="true" box="[199,300,1652,1678]" pageId="7" pageNumber="8">Ecology.</emphasis>
<taxonomicName id="4C0F4D5EFFC7FFF9E1417473FDFBF944" authorityName=": Asahina" authorityYear="1996" box="[305,568,1652,1678]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC7FFF9E1417473FDFBF944" box="[305,568,1652,1678]" italics="true" pageId="7" pageNumber="8">Planaeschna tomokunii</emphasis>
</taxonomicName>
is found around
<quantity id="4CF79B38FFC7FFF9E2817473FC86F945" box="[753,837,1652,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="m" value="1000.0">1000 m</quantity>
asl. on the densely forested slopes of Tam Dao, where it apparently breeds in small brooks and seeps. Field observations were done of specimens hawking at dusk over a car track through the forest on the mountainside. In one instance, a male successfully grabbed a female over the track. It occurred together with
<taxonomicName id="4C0F4D5EFFC7FFF9E18A74E6FD62F930" authorityName=": Asahina" authorityYear="1996" box="[506,673,1760,1786]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B97BEACFFFC7FFF9E18A74E6FD62F930" box="[506,673,1760,1786]" italics="true" pageId="7" pageNumber="8">P. tamdaoensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFC7FFF9E0B77503FCDEF8D5" box="[199,797,1796,1823]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BB036DDFFC7FFF9E0B77503FCDEF8D5" blockId="7.[151,1437,932,1823]" box="[199,797,1796,1823]" pageId="7" pageNumber="8">
<emphasis id="B97BEACFFFC7FFF9E0B77503FEA2F8D4" bold="true" box="[199,353,1796,1822]" pageId="7" pageNumber="8">Distribution.</emphasis>
Endemic to
<collectingCountry id="F318764DFFC7FFF9E19E7503FD8EF8D4" box="[494,589,1796,1822]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
(
<collectingRegion id="49CBF83FFFC7FFF9E22B7503FCD5F8D5" box="[603,790,1796,1823]" country="Vietnam" name="Vinh Phuc" pageId="7" pageNumber="8">Vinh Phuc Prov.</collectingRegion>
)
</paragraph>
</subSubSection>
</treatment>
</document>