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<document ID-DOI="http://doi.org/10.5281/zenodo.6458594" ID-GBIF-Dataset="adeeb71f-7f8d-4e00-bc9f-35089363f76e" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6458594" checkinTime="1600878147105" checkinUser="plazi" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03A687BCFFA2FFA213B5F37BF7D6FBF0" docLanguage="en" docName="hbmw_9_Phyllostomidae_444.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Lophostoma brasiliense Peters 1867" docType="treatment" docVersion="11" lastPageNumber="503" masterDocId="FF9FFFC4FFB1FFB1133CFFBAFFE0F244" masterDocTitle="Phyllostomidae" masterLastPageNumber="583" masterPageNumber="444" pageNumber="503" updateTime="1658329166853" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phyllostomidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>444</mods:start>
<mods:end>583</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6458594</mods:identifier>
<mods:identifier type="GBIF-Dataset">adeeb71f-7f8d-4e00-bc9f-35089363f76e</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">6458594</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6803095" ID-GBIF-Taxon="196400479" ID-Zenodo-Dep="6803095" LSID="urn:lsid:plazi:treatment:03A687BCFFA2FFA213B5F37BF7D6FBF0" httpUri="http://treatment.plazi.org/id/03A687BCFFA2FFA213B5F37BF7D6FBF0" lastPageNumber="503" pageId="19" pageNumber="503">
<subSubSection box="[137,194,3265,3311]" pageId="19" pageNumber="503" type="multiple">
<paragraph blockId="19.[134,1196,3265,3390]" box="[137,194,3265,3311]" pageId="19" pageNumber="503">
<heading box="[137,194,3265,3311]" pageId="19" pageNumber="503">
<figureCitation box="[137,194,3265,3311]" captionStart="Plate 36: Phyllostomidae" captionStartId="17.[137,167,3288,3313]" captionTargetBox="[23,2765,17,3655]" captionTargetPageId="16" captionText="24. Long-legged Bat (Macrophyllum macrophyllum), 25. Fringe-lipped Bat (Trachops cirrhosus), 26. Striped Hairy-nosed Bat (Gardnerycteris crenulatum), 27. Keenan's Hairy-nosed Bat (Gardnerycteris keenani), 28. Koepckes Hairy-nosed Bat (Gardnerycteris koepckeae), 29. Kalkos Round-eared Bat (Lophostoma kalkoae), 30. Pygmy Round-eared Bat (Lophostoma brasiliense), 31. Carrikers Round-eared Bat (Lophostoma carrikeri), 32. Schulzs Round-eared Bat (Lophostoma schulzi), 33. Western Round-eared Bat (Lophostoma occidentale), 34. Davis's Round-eared Bat (Lophostoma evotis), 35. White-throated Round-eared Bat (Lophostoma silvicola), 36. Greater Round-eared Bat (Tonatia bidens), 37. Stripe-headed Round-eared Bat (Tonatia saurophila), 38. Pale-faced Bat (Phylloderma stenops), 39. Pale Spear-nosed Bat (Phyllostomus discolor), 40. Lesser Spear-nosed Bat (Phyllostomus elongatus), 41. Greater Spear-nosed Bat (Phyllostomus hastatus), 42. Guianan Spear-nosed Bat (Phyllostomus latifolius), 43. Woolly False Vampire Bat (Chrotopterus auritus), 44. Southern Golden Bat (Mimon bennettii), 45. Cozumelan Golden Bat (Mimon cozumelae), 46. Spectral Bat (Vampyrum spectrum)" figureDoi="http://doi.org/10.5281/zenodo.6458675" httpUri="https://zenodo.org/record/6458675/files/figure.png" pageId="19" pageNumber="503">30.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[210,714,3265,3311]" pageId="19" pageNumber="503" type="vernacular_names">
<paragraph blockId="19.[134,1196,3265,3390]" box="[210,714,3265,3311]" pageId="19" pageNumber="503">
<heading box="[210,714,3265,3311]" pageId="19" pageNumber="503">
<vernacularName box="[210,714,3265,3311]" pageId="19" pageNumber="503">Pygmy Round-eared Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[785,1195,3265,3311]" pageId="19" pageNumber="503" type="nomenclature">
<paragraph blockId="19.[134,1196,3265,3390]" box="[785,1195,3265,3311]" pageId="19" pageNumber="503">
<heading box="[785,1195,3265,3311]" pageId="19" pageNumber="503">
<taxonomicName authority="Peters, 1867" authorityName="Peters" authorityYear="1867" box="[785,1195,3265,3311]" class="Mammalia" family="Phyllostomidae" genus="Lophostoma" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="brasiliense">
<emphasis box="[785,1195,3265,3311]" italics="true" pageId="19" pageNumber="503">Lophostoma brasiliense</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="vernacular_names">
<paragraph blockId="19.[134,1196,3265,3390]" box="[136,1137,3330,3351]" pageId="19" pageNumber="503">
<heading box="[136,1137,3330,3351]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[136,212,3330,3351]" pageId="19" pageNumber="503">French:</emphasis>
<vernacularName box="[221,462,3330,3351]" pageId="19" pageNumber="503">Lophostome des marais</vernacularName>
/
<emphasis bold="true" box="[484,574,3330,3351]" pageId="19" pageNumber="503">German:</emphasis>
<vernacularName box="[584,826,3330,3351]" pageId="19" pageNumber="503">Kleine Rundohrblattnase</vernacularName>
/
<emphasis bold="true" box="[847,939,3330,3351]" pageId="19" pageNumber="503">Spanish:</emphasis>
<vernacularName box="[948,1053,3330,3351]" pageId="19" pageNumber="503">Lofostoma</vernacularName>
pigmeo
</heading>
</paragraph>
<paragraph blockId="19.[134,1196,3265,3390]" box="[135,870,3368,3389]" pageId="19" pageNumber="503">
<heading box="[135,870,3368,3389]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[135,383,3368,3389]" pageId="19" pageNumber="503">Other common names:</emphasis>
<vernacularName box="[392,634,3368,3389]" pageId="19" pageNumber="503">Lesser Round-eared Bat</vernacularName>
,
<vernacularName box="[649,870,3368,3389]" pageId="19" pageNumber="503">Little Round-eared Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[135,775,3444,3469]" pageId="19" pageNumber="503" type="reference_group">
<paragraph blockId="19.[134,1258,3441,3475]" box="[135,775,3444,3469]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[135,290,3444,3469]" pageId="19" pageNumber="503">Taxonomy.</emphasis>
<taxonomicName authority="Peters, 1867" authorityName="Peters" authorityYear="1867" box="[302,770,3444,3469]" class="Mammalia" family="Phyllostomidae" genus="Lophostoma" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="brasiliense">Lophostoma brasiliense Peters, 1867</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection box="[787,1257,3444,3469]" pageId="19" pageNumber="503" type="materials_examined">
<paragraph blockId="19.[134,1258,3441,3475]" box="[787,1257,3444,3469]" pageId="19" pageNumber="503">
<materialsCitation ID-GBIF-Occurrence="3819564490" box="[787,1257,3444,3469]" pageId="19" pageNumber="503">
“Baia” (=
<collectingCountry box="[922,1048,3444,3469]" name="El Salvador" pageId="19" pageNumber="503">Salvador</collectingCountry>
),
<collectingRegion box="[1074,1155,3444,3469]" country="Brazil" name="Bahia" pageId="19" pageNumber="503">Bahia</collectingRegion>
,
<collectingCountry box="[1172,1254,3444,3469]" name="Brazil" pageId="19" pageNumber="503">Brazil</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="discussion">
<paragraph blockId="19.[2025,2620,285,708]" pageId="19" pageNumber="503">
<taxonomicName authority="Peters, 1867" authorityName="Peters" authorityYear="1867" box="[2026,2322,285,314]" class="Mammalia" family="Phyllostomidae" genus="Lophostoma" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="brasiliense">Lophostoma brasiliense</taxonomicName>
was formerly in the genus
<taxonomicName box="[2120,2217,320,353]" class="Mammalia" family="Phyllostomidae" genus="Tonatia" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="genus">Tonatia</taxonomicName>
, but T. E. Lee and collaborators in 2002 showed that
<taxonomicName box="[2385,2482,360,393]" class="Mammalia" family="Phyllostomidae" genus="Tonatia" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="genus">Tonatia</taxonomicName>
was paraphyletic and recommended restricting it to
<taxonomicName baseAuthorityName="Spix" baseAuthorityYear="1823" box="[2066,2176,438,471]" class="Mammalia" family="Phyllostomidae" genus="Tonatia" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="bidens">T. bidens</taxonomicName>
and
<taxonomicName authorityName="Koopman &amp; E. E. Williams" authorityYear="1951" box="[2247,2414,438,471]" class="Mammalia" family="Phyllostomidae" genus="Tonatia" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="saurophila">T. saurophila</taxonomicName>
, while reclassifying remaining taxa to
<taxonomicName authorityName="d'Orbigny" authorityYear="1836" box="[2364,2517,478,511]" class="Mammalia" family="Phyllostomidae" genus="Lophostoma" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="genus">Lophostoma</taxonomicName>
. Other small species of
<taxonomicName authorityName="d'Orbigny" authorityYear="1836" box="[2248,2401,526,551]" class="Mammalia" family="Phyllostomidae" genus="Lophostoma" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="genus">Lophostoma</taxonomicName>
, such as L. nicaraguae,
<taxonomicName baseAuthorityName="P. Gervais" baseAuthorityYear="1856" box="[2134,2269,561,590]" class="Mammalia" family="Phyllostomidae" genus="Micronycteris" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="minuta">L. minuta</taxonomicName>
, and L. venezuelae, once recognized as valid, are currently treated as synonyms of
<taxonomicName authority="Peters, 1867" authorityName="Peters" authorityYear="1867" box="[2243,2410,635,668]" class="Mammalia" family="Phyllostomidae" genus="Lophostoma" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="brasiliense">L. brasiliense</taxonomicName>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6458662" ID-Zenodo-Dep="6458662" httpUri="https://zenodo.org/record/6458662/files/figure.png" inLine="true" pageId="19" pageNumber="503" targetBox="[1411,2001,293,705]" targetPageId="19">
<paragraph blockId="19.[2025,2620,285,708]" lastBlockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[2025,2201,679,708]" pageId="19" pageNumber="503">Distribution.</emphasis>
From S
<collectingCountry box="[2357,2459,679,708]" name="Mexico" pageId="19" pageNumber="503">Mexico</collectingCountry>
(
<collectingRegion box="[2493,2618,679,708]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="19" pageNumber="503">Veracruz</collectingRegion>
and S
<collectingRegion box="[1500,1611,714,747]" country="Mexico" name="Yucatan" pageId="19" pageNumber="503">Yucatan</collectingRegion>
Peninsula) through Central America to N &amp; E South America, including
<collectingCountry box="[1468,1607,762,787]" name="Colombia" pageId="19" pageNumber="503">Colombia</collectingCountry>
,
<collectingCountry box="[1622,1769,762,787]" name="Venezuela" pageId="19" pageNumber="503">Venezuela</collectingCountry>
, the Guianas,
<collectingCountry box="[1972,2090,762,787]" name="Ecuador" pageId="19" pageNumber="503">Ecuador</collectingCountry>
,
<collectingCountry box="[2107,2174,762,787]" name="Peru" pageId="19" pageNumber="503">Peru</collectingCountry>
,
<collectingCountry box="[2192,2274,762,787]" name="Brazil" pageId="19" pageNumber="503">Brazil</collectingCountry>
,
<collectingCountry box="[2290,2386,762,787]" name="Bolivia" pageId="19" pageNumber="503">Bolivia</collectingCountry>
, and an isolated record from
<collectingCountry box="[1594,1723,793,826]" name="Paraguay" pageId="19" pageNumber="503">Paraguay</collectingCountry>
; also on
<collectingCountry box="[1847,1968,793,826]" name="Trinidad and Tobago" pageId="19" pageNumber="503">Trinidad</collectingCountry>
I.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="description">
<paragraph blockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1413,1660,837,866]" pageId="19" pageNumber="503">Descriptive notes.</emphasis>
Head-body 42-61 mm, tail 8-13 mm, ear 19-2-24-8 mm, hindfoot 9-11-9 mm, forearm 34—40-5 mm; weight 9-9-13-8 g. Greater lengths of skulls are 18-7-21-6 mm. There is no apparently sexual dimorphism, except in genitalia. The Pygmy Round-eared Bat is the smallest species of
<taxonomicName box="[2016,2166,950,983]" class="Magnoliopsida" family="Thymelaeaceae" genus="Lophostoma" kingdom="Plantae" order="Malvales" pageId="19" pageNumber="503" phylum="Tracheophyta" rank="genus">Lophostoma</taxonomicName>
, characterized by long rounded ears connected by band of skin across forehead. It is easily identified from other species of
<taxonomicName box="[1453,1602,1030,1063]" class="Magnoliopsida" family="Thymelaeaceae" genus="Lophostoma" kingdom="Plantae" order="Malvales" pageId="19" pageNumber="503" phylum="Tracheophyta" rank="genus">Lophostoma</taxonomicName>
by its smaller external and cranial measurements. Dorsal pelage is long and smooth but not dense; hairs are brown and slightly darker around face and have white bases. Venteris pale brown and does not contrast sharply with dorsum. Rostrum is sparsely furred, almost naked;tip of chin has rows of small rounded tubercles forming a U-shape. Shorttail is enclosed in uropatagium, exceptfortip that protrudesslightly near middle of dorsal side. In the field, Pygmy Round-eared Bats can be confused with bigeared bats (
<taxonomicName box="[1575,1741,1266,1299]" class="Mammalia" family="Phyllostomidae" genus="Micronycteris" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="genus">Micronycteris</taxonomicName>
) because they are also small and gray or brown phyllostomines with long, rounded ears. Nevertheless, generic characteristics such as two lower incisors versus four in
<taxonomicName box="[1617,1780,1345,1378]" class="Mammalia" family="Phyllostomidae" genus="Micronycteris" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="genus">Micronycteris</taxonomicName>
and pair of large tubercles forming V-shape on tip of chin distinguish the Pygmy Round-eared Bat from species of
<taxonomicName box="[2176,2340,1384,1417]" class="Mammalia" family="Phyllostomidae" genus="Micronycteris" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="genus">Micronycteris</taxonomicName>
. Skull is robust, with an undeveloped sagittal crest, and is constricted in postorbital region.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="biology_ecology">
<paragraph blockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1412,1524,1463,1496]" pageId="19" pageNumber="503">Habitat.</emphasis>
Varied habitats often secondary growth in savanna regions below elevations of
<quantity box="[1450,1538,1507,1536]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="503" unit="m" value="500.0">500 m</quantity>
, moist areas and riparian habitats, deciduous forests, forest fragments, forest edges near agricultural areas, swampy primary forests, and creek side primary forests.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="food_feeding">
<paragraph blockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1412,1681,1582,1615]" pageId="19" pageNumber="503">Food and Feeding.</emphasis>
The Pygmy Round-eared Bat is a foliage-gleaning insectivore. It prefers large arthropods such as coleopterans and lepidopterans, and it might eat fruit. In
<collectingCountry box="[1530,1612,1660,1693]" name="Brazil" pageId="19" pageNumber="503">Brazil</collectingCountry>
, pollen has been found in stomach contents.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="breeding">
<paragraph blockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1412,1547,1700,1733]" pageId="19" pageNumber="503">Breeding.</emphasis>
Reproductive patterns of Pygmy Round-eared Bats vary between Central America and South America. In
<collectingCountry box="[1863,2010,1739,1772]" name="Costa Rica" pageId="19" pageNumber="503">Costa Rica</collectingCountry>
, single peak in pregnancy occurs in late dry season in March, followed by lactation in early wet season in June. Males are reproductive for several months before peak pregnancy in April-May and at end of wet season in September—-October. In South America, bimodal reproductive patterns are more common, with one peak in pregnancy in dry seasons and another peak in wet seasons.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="activity">
<paragraph blockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1410,1654,1937,1970]" pageId="19" pageNumber="503">Activity patterns.</emphasis>
The Pygmy Round-eared Bat is nocturnal, with activity peaking shortly after dusk. It roosts in large active arboreal termite nests. In
<collectingCountry box="[2375,2442,1976,2009]" name="Peru" pageId="19" pageNumber="503">Peru</collectingCountry>
, roosts were c.60-70 cm wide, ¢.60-70 cm high, and ¢.
<quantity box="[2001,2088,2016,2049]" metricMagnitude="-1" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="503" unit="cm" value="28.0">28 cm</quantity>
deep, with entrances of ¢.
<quantity box="[2463,2533,2016,2049]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="503" unit="cm" value="6.0">6 cm</quantity>
in diameter. In
<collectingCountry box="[1563,1709,2055,2088]" name="Costa Rica" pageId="19" pageNumber="503">Costa Rica</collectingCountry>
, roosts cavities have been described as lacking any compartments or lateral chambers, being a single cylindrical chamber ¢.
<quantity box="[2214,2302,2094,2127]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="503" unit="cm" value="20.0">20 cm</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="biology_ecology">
<paragraph blockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1412,2128,2134,2167]" pageId="19" pageNumber="503">Movements, Home range and Social organization.</emphasis>
In
<collectingCountry box="[2185,2336,2134,2167]" name="Costa Rica" pageId="19" pageNumber="503">Costa Rica</collectingCountry>
, adult male and female Pygmy Round-eared Bats roosting together suggest harem-based mating system. They construct and actively maintain roost cavities to prevent disturbance by termites. Pygmy Round-eared Bats have been documented sharing roosts with Fringe-lipped Bats (
<taxonomicName box="[1491,1736,2296,2325]" class="Mammalia" family="Phyllostomidae" genus="Trachops" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="503" phylum="Chordata" rank="species" species="cirrhosus">Trachops cirrhosus</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection box="[1412,2464,2331,2364]" pageId="19" pageNumber="503" type="conservation">
<paragraph blockId="19.[1410,2619,714,2484]" box="[1412,2464,2331,2364]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1412,1762,2331,2364]" pageId="19" pageNumber="503">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="503" type="bibRefCitation_list">
<paragraph blockId="19.[1410,2619,714,2484]" pageId="19" pageNumber="503">
<emphasis bold="true" box="[1413,1566,2380,2405]" pageId="19" pageNumber="503">Bibliography.</emphasis>
Bernard &amp; Fenton (2002), Durant et al. (2013), Eisenberg &amp; Redford (1999), Genoways &amp; Williams (1984), Goodwin (1942), Kalko et al. (2006), Lee et al. (2002), Munin et al. (2012), Simmons &amp; Voss (1998), Williams &amp; Genoways (1980a), Willig (1985a), York et al. (2008).
</paragraph>
</subSubSection>
</treatment>
</document>