557 lines
79 KiB
XML
557 lines
79 KiB
XML
<document id="B4F3666CBF1AE610E73AD67F4A01B497" ID-CLB-Dataset="272041" ID-DOI="10.1093/zoolinnean/zlad013" ID-GBIF-Dataset="effa6181-8326-4c7d-8c91-809f7fabb862" ID-ISSN="0024-4082" ID-Zenodo-Dep="8432389" ID-ZooBank="C3E2DFF9-A0A6-41EA-A149-0F73A2BEE5E6" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696939870042" checkinUser="plazi" docAuthor="Silva, Felipe De Vasconcelos, Pola, Marta & Cervera, Juan Lucas" docDate="2023" docId="03A6878CFF816505FD1AFBD4EAA5FB39" docLanguage="en" docName="zlad013.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad013" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Tritonia Cuvier 1798" docType="treatment" docVersion="4" lastPageNumber="463" masterDocId="FF9FFFF4FF906517FC52FF84EE67FF8B" masterDocTitle="A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)" masterLastPageNumber="476" masterPageNumber="445" pageNumber="462" updateTime="1699694776904" updateUser="plazi">
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<mods:title id="27E6754E98B040EB119F15C7EA82E79A">A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)</mods:title>
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<mods:namePart id="55A53702A92364CF9E10A70847226310">Silva, Felipe De Vasconcelos</mods:namePart>
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<mods:affiliation id="26FB9708198BE439FD97B90CA129BA26">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:nameIdentifier id="C2CD9617B88AAD5AC307E151A9D3B32E" type="email">felipe.devasconcelos@uca.es</mods:nameIdentifier>
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<mods:namePart id="D2744332612DE849A2E3CADD60123B88">Pola, Marta</mods:namePart>
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<mods:affiliation id="C6162C47B66B000D6F351A9B67711310">Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid; Campus de Excelencia Internacional UAM + CSIC, C / Darwin, 2, 28049 Madrid, Spain</mods:affiliation>
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<mods:namePart id="B4A4526D98429E706A57E95E3773C53F">Cervera, Juan Lucas</mods:namePart>
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<mods:affiliation id="D8BF558D7DD3BFD14E0F332281F6C722">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain & Instituto Universitario de Investigación Marina (INMAR), Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Universidad de Cádiz, Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:date id="3C6984114449419BE21FF24917F0B813">2023</mods:date>
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<treatment id="03A6878CFF816505FD1AFBD4EAA5FB39" ID-DOI="http://doi.org/10.5281/zenodo.8432403" ID-GBIF-Taxon="213768880" ID-Zenodo-Dep="8432403" LSID="urn:lsid:plazi:treatment:03A6878CFF816505FD1AFBD4EAA5FB39" httpUri="http://treatment.plazi.org/id/03A6878CFF816505FD1AFBD4EAA5FB39" lastPageId="18" lastPageNumber="463" pageId="17" pageNumber="462">
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<subSubSection id="C3156511FF816506FD1AFBD4EC44FBE1" box="[328,547,1104,1130]" pageId="17" pageNumber="462" type="nomenclature">
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<paragraph id="8BB0369AFF816506FD1AFBD4EC44FBE1" blockId="17.[328,547,1104,1130]" box="[328,547,1104,1130]" pageId="17" pageNumber="462">
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<heading id="D0F881F6FF816506FD1AFBD4EC44FBE1" bold="true" box="[328,547,1104,1130]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="462" reason="2">
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<taxonomicName id="4C0F4D19FF816506FD1AFBD4EC44FBE1" ID-CoL="7PJZ6" authority="Cuvier 1798" authorityName="Cuvier" authorityYear="1798" box="[328,547,1104,1130]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
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<emphasis id="B97BEA88FF816506FD1AFBD4EC44FBE1" bold="true" box="[328,547,1104,1130]" pageId="17" pageNumber="462">
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<emphasis id="B97BEA88FF816506FD1AFBD4EFFCFBE1" bold="true" box="[328,411,1104,1130]" italics="true" pageId="17" pageNumber="462">Tritonia</emphasis>
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Cuvier 1798
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156511FF816506FD14FBF2EC41FB1B" box="[326,550,1142,1169]" pageId="17" pageNumber="462" type="description">
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<paragraph id="8BB0369AFF816506FD14FBF2EC41FB1B" blockId="17.[326,550,1142,1169]" box="[326,550,1142,1169]" pageId="17" pageNumber="462">
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<emphasis id="B97BEA88FF816506FD14FBF2EC41FB1B" bold="true" box="[326,550,1142,1169]" pageId="17" pageNumber="462">
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(
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<figureCitation id="13342A1FFF816506FD02FBF2EFF9FB1B" box="[336,414,1142,1168]" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="17" pageNumber="462">Figs 3A</figureCitation>
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,
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<figureCitation id="13342A1FFF816506FDF8FBF2EFAFFB1B" box="[426,456,1142,1168]" captionStart="Figure 4" captionStartId="13.[113,178,1528,1552]" captionTargetBox="[309,1263,147,1498]" captionTargetId="figure-134@13.[306,1266,144,1501]" captionTargetPageId="13" captionText="Figure 4. Scanning electron micrographs of radulae of analysed key species of Tritoniinae and Marioniinae subfam. nov.: A, tricuspid rachidian teeth and differentiated first lateral of Tritonia hombergii (ZMBN 83471); B, tricuspid rachidian teeth and differentiated first lateral with single accessory denticle (circled) of Candiella plebeia (MNCN15.05/93393); C, unicuspid rachidian teeth and undifferentiated first lateral of Tritoniopsis elegans (MNHNC/MB28-004506); D, tricuspid rachidian teeth and differentiated first lateral of Tritonidoxa capensis (MNCN15.05/93399); E, unicuspid rachidian teeth and undifferentiated first lateral of Tochuina gigantea (NHMD-115967); F, tricuspid rachidian teeth and differentiated first lateral with several accessory denticles (circled) of Tritonicula hamnerorum (MNCN15.05/93378); G, tricuspid rachidian teeth and differentiated first lateral of Marionia blainvillea (NHMD115968); Scale bars: A, D, E = 60 μm; B, C = 22 μm; F = 20 μm; G = 110 μm." figureDoi="http://doi.org/10.5281/zenodo.8432401" httpUri="https://zenodo.org/record/8432401/files/figure.png" pageId="17" pageNumber="462">4A</figureCitation>
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,
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<figureCitation id="13342A1FFF816506FD81FBF2EF96FB1B" box="[467,497,1142,1168]" captionStart="Figure 5" captionStartId="14.[129,194,1262,1286]" captionTargetBox="[164,1437,147,1230]" captionTargetId="figure-262@14.[161,1441,144,1234]" captionTargetPageId="14" captionText="Figure 5. Digestive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view: A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (MNCN15.05/93393); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis(MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations:a, anus; aodg, anterior ODG complex; e, esophagus; i, intestine; odg, ovotestis-digestive gland complex; s, stomach; podg, posterior ODG complex." figureDoi="http://doi.org/10.5281/zenodo.8432405" httpUri="https://zenodo.org/record/8432405/files/figure.png" pageId="17" pageNumber="462">5A</figureCitation>
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,
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<figureCitation id="13342A1FFF816506FDAFFBF2EC7CFB1B" box="[509,539,1142,1168]" captionStart="Figure 6" captionStartId="15.[113,178,1771,1795]" captionTargetBox="[317,1256,144,1743]" captionTargetId="figure-8@15.[316,1256,144,1743]" captionTargetPageId="15" captionText="Figure 6. Reproductive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view:A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (ZMBN 83471); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis (MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations: ag, albumen gland; am, ampulla; bc, bursa copulatrix; fo, female opening; fgm, female gland mass; hd, hermaphrodite duct; o, oviduct; p, penis; pp, prostatic portion; ps, penial sheath; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.8432407" httpUri="https://zenodo.org/record/8432407/files/figure.png" pageId="17" pageNumber="462">6A</figureCitation>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156511FF816506FC23FB2FEF4CFAAA" pageId="17" pageNumber="462" type="reference_group">
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<paragraph id="8BB0369AFF816506FC23FB2FEF4CFAAA" blockId="17.[113,763,1194,1313]" pageId="17" pageNumber="462">
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||
<taxonomicName id="4C0F4D19FF816506FC23FB2FEFA6FB48" ID-CoL="8b640069-24d2-4c86-a76f-8712455621fc" authority="MacGillivray 1843" authorityName="MacGillivray" authorityYear="1843" box="[113,449,1195,1219]" class="Gastropoda" family="Tritoniidae" genus="Sphaerostoma" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
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<emphasis id="B97BEA88FF816506FC23FB2FEE9DFB48" box="[113,250,1195,1219]" italics="true" pageId="17" pageNumber="462">Sphaerostoma</emphasis>
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<bibRefCitation id="EF9E4B6BFF816506FD52FB2FEFA6FB48" author="MacGillivray W" box="[256,449,1195,1219]" pageId="17" pageNumber="462" refId="ref25868" refString="MacGillivray W. A history of the molluscous animals of the counties of Aberdeen, Kincardine and Banff, to which is appended an account of the cirripedal animals of the same district. London: Cunningham & Mortimer, 1843." type="book" year="1843">MacGillivray 1843</bibRefCitation>
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</taxonomicName>
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, Hist. Moll, 24: 335.
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<taxonomicName id="4C0F4D19FF816506FC23FB4EEFC4FB69" ID-CoL="a18238e2-59f6-4a34-9779-719ec5bb26a3" authority="Stearns 1873" authorityName="Stearns" authorityYear="1873" box="[113,419,1226,1250]" class="Gastropoda" family="Tritoniidae" genus="Lateribranchiaea" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
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<emphasis id="B97BEA88FF816506FC23FB4EEF70FB69" box="[113,279,1226,1250]" italics="true" pageId="17" pageNumber="462">Lateribranchiaea</emphasis>
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Stearns 1873
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</taxonomicName>
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, Proc. Calif. Acad. Nat. Sci, 5: 77.
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<taxonomicName id="4C0F4D19FF816506FC23FB6DEC4FFA8A" ID-CoL="3580609e-198e-46f2-b904-cfebe00766c3" authority="Iredale and O'Donoghue 1923" authorityName="Iredale and O'Donoghue" authorityYear="1923" box="[113,552,1257,1281]" class="Gastropoda" family="Tritoniidae" genus="Candellista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
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<emphasis id="B97BEA88FF816506FC23FB6DEEB8FA8A" box="[113,223,1257,1281]" italics="true" pageId="17" pageNumber="462">Candellista</emphasis>
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Iredale and O’Donoghue 1923
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</taxonomicName>
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, Proc. Malacol. Soc. Lond, 15(5): 197.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3156511FF816506FC23FABBEE88FAFE" pageId="17" pageNumber="462" type="type_taxon">
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<paragraph id="8BB0369AFF816506FC23FABBEE88FAFE" blockId="17.[113,762,1342,1397]" pageId="17" pageNumber="462">
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||
<emphasis id="B97BEA88FF816506FC23FABBEE93FADD" box="[113,244,1343,1366]" italics="true" pageId="17" pageNumber="462">
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||
<typeStatus id="54B48838FF816506FC23FABBEEC6FADD" box="[113,161,1343,1366]" pageId="17" pageNumber="462">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C0F4D19FF816506FD52FABBEC36FADD" ID-CoL="7TFDN" authority="Cuvier 1803" authorityName="Cuvier" authorityYear="1803" box="[256,593,1342,1366]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="hombergii">
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<emphasis id="B97BEA88FF816506FD52FABBEFDAFADD" box="[256,445,1342,1366]" italics="true" pageId="17" pageNumber="462">Tritonia hombergii</emphasis>
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Cuvier 1803
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</taxonomicName>
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, by subsequent designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156511FF816506FC23FA17EC1FFA62" pageId="17" pageNumber="462" type="diagnosis">
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<paragraph id="8BB0369AFF816506FC23FA17EC1FFA62" blockId="17.[113,762,1426,1513]" pageId="17" pageNumber="462">
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<emphasis id="B97BEA88FF816506FC23FA17EEB0FA21" box="[113,215,1427,1450]" italics="true" pageId="17" pageNumber="462">Diagnosis:</emphasis>
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Oral veil broad and bilobed with several simple or branched velar processes. Gills fully arborescent with at least seven major pairs. Penis large, elongated or conical.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156511FF816506FC23F98EEA7BFC26" pageId="17" pageNumber="462" type="description">
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<paragraph id="8BB0369AFF816506FC23F98EEB3CFD38" blockId="17.[113,763,1546,1978]" lastBlockId="17.[810,1459,604,691]" pageId="17" pageNumber="462">
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<emphasis id="B97BEA88FF816506FC23F98EEE97F9A9" box="[113,240,1546,1570]" italics="true" pageId="17" pageNumber="462">Morphology:</emphasis>
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Body broad, length up to
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(
|
||
<figureCitation id="13342A1FFF816506FE38F98EECD3F9A8" box="[618,692,1546,1571]" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="17" pageNumber="462">Fig. 3A</figureCitation>
|
||
). Oral veil broad and bilobed. Seven to 40 simple or branched velar processes. Notal border may be distinct. Fully arborescent gills with at least seven large pairs and smaller ones in between. Rachidian tooth uni- or tricuspid (
|
||
<figureCitation id="13342A1FFF816506FD3DF90CEFDDF92B" box="[367,442,1672,1696]" captionStart="Figure 4" captionStartId="13.[113,178,1528,1552]" captionTargetBox="[309,1263,147,1498]" captionTargetId="figure-134@13.[306,1266,144,1501]" captionTargetPageId="13" captionText="Figure 4. Scanning electron micrographs of radulae of analysed key species of Tritoniinae and Marioniinae subfam. nov.: A, tricuspid rachidian teeth and differentiated first lateral of Tritonia hombergii (ZMBN 83471); B, tricuspid rachidian teeth and differentiated first lateral with single accessory denticle (circled) of Candiella plebeia (MNCN15.05/93393); C, unicuspid rachidian teeth and undifferentiated first lateral of Tritoniopsis elegans (MNHNC/MB28-004506); D, tricuspid rachidian teeth and differentiated first lateral of Tritonidoxa capensis (MNCN15.05/93399); E, unicuspid rachidian teeth and undifferentiated first lateral of Tochuina gigantea (NHMD-115967); F, tricuspid rachidian teeth and differentiated first lateral with several accessory denticles (circled) of Tritonicula hamnerorum (MNCN15.05/93378); G, tricuspid rachidian teeth and differentiated first lateral of Marionia blainvillea (NHMD115968); Scale bars: A, D, E = 60 μm; B, C = 22 μm; F = 20 μm; G = 110 μm." figureDoi="http://doi.org/10.5281/zenodo.8432401" httpUri="https://zenodo.org/record/8432401/files/figure.png" pageId="17" pageNumber="462">Fig. 4A</figureCitation>
|
||
), smooth or denticulate (with accessory denticles or folds) with rectangular base. First lateral tooth differentiated. Large number of lateral teeth (up to 200 per half row). Ratio jaw/body length: 0.27–0.40. Masticatory borders of jaws denticulate, with medium-sized (up to 100 µm) denticles with conical bases and sharp cusps; large-sized (up to 300 µm) denticles with conical bases and blunt cusps; or rod-shaped denticles. Cuticular folds may be present. ODG complex holohepatic (
|
||
<figureCitation id="13342A1FFF816506FCACF806EF2CF811" box="[254,331,1922,1946]" captionStart="Figure 5" captionStartId="14.[129,194,1262,1286]" captionTargetBox="[164,1437,147,1230]" captionTargetId="figure-262@14.[161,1441,144,1234]" captionTargetPageId="14" captionText="Figure 5. Digestive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view: A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (MNCN15.05/93393); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis(MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations:a, anus; aodg, anterior ODG complex; e, esophagus; i, intestine; odg, ovotestis-digestive gland complex; s, stomach; podg, posterior ODG complex." figureDoi="http://doi.org/10.5281/zenodo.8432405" httpUri="https://zenodo.org/record/8432405/files/figure.png" pageId="17" pageNumber="462">Fig. 5A</figureCitation>
|
||
). Gonopore at end of one-third of body length, furthest from rhinophores. Anus at middle of the body length; nephroproct above anus. Bursa copulatrix oħen pyriform. Penis large, elongate or conical; distal characters may be present. Ampulla long, cylindrical and coiled (
|
||
<figureCitation id="13342A1FFF816506F950FD1FEB2DFD38" box="[1282,1354,667,691]" captionStart="Figure 6" captionStartId="15.[113,178,1771,1795]" captionTargetBox="[317,1256,144,1743]" captionTargetId="figure-8@15.[316,1256,144,1743]" captionTargetPageId="15" captionText="Figure 6. Reproductive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view:A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (ZMBN 83471); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis (MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations: ag, albumen gland; am, ampulla; bc, bursa copulatrix; fo, female opening; fgm, female gland mass; hd, hermaphrodite duct; o, oviduct; p, penis; pp, prostatic portion; ps, penial sheath; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.8432407" httpUri="https://zenodo.org/record/8432407/files/figure.png" pageId="17" pageNumber="462">Fig. 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF816506FF78FD5EEA7BFC26" blockId="17.[810,1459,730,942]" pageId="17" pageNumber="462">
|
||
<emphasis id="B97BEA88FF816506FF78FD5EED9AFD7A" box="[810,1021,730,753]" italics="true" pageId="17" pageNumber="462">Species composition:</emphasis>
|
||
<taxonomicName id="4C0F4D19FF816506F843FD5EED07FC9A" authority="Chimienti et al. 2020" authorityName="Chimienti" authorityYear="2020" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="callogorgiae">
|
||
<emphasis id="B97BEA88FF816506F843FD5EEA81FD79" box="[1041,1254,730,754]" italics="true" pageId="17" pageNumber="462">Tritonia callogorgiae</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF816506F8A8FD5EED07FC9A" author="Chimienti G & Angeleui L & Furfaro G" pageId="17" pageNumber="462" pagination="103364" refId="ref24637" refString="Chimienti G, Angeleui L, Furfaro G, et al. Habitat, morphology and trophism of Tritonia callogorgiae sp. nov., a large nudibranch inhabiting Callogorgia verticillata forests in the Mediterranean Sea. Deep Sea Research Part I: Oceanographic Research Papers 2020; 165: 103364." type="journal article" year="2020">
|
||
Chimienti
|
||
<emphasis id="B97BEA88FF816506F924FD5EEBCAFD79" box="[1398,1453,730,754]" italics="true" pageId="17" pageNumber="462">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506FF3FFD7EEA7FFC9A" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[877,1048,761,785]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="exsulans">
|
||
<emphasis id="B97BEA88FF816506FF3FFD7EEA7FFC9A" box="[877,1048,761,785]" italics="true" pageId="17" pageNumber="462">Tritonia exsulans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506F874FD7EEADCFC9A" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[1062,1211,761,785]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="festiva">
|
||
<emphasis id="B97BEA88FF816506F874FD7EEADCFC9A" box="[1062,1211,761,785]" italics="true" pageId="17" pageNumber="462">Tritonia festiva</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506F898FD7EED07FCBB" authority="Odhner 1922" authorityName="Odhner" authorityYear="1922" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="griegi">
|
||
<emphasis id="B97BEA88FF816506F898FD7EEB3FFC9A" box="[1226,1368,762,785]" italics="true" pageId="17" pageNumber="462">Tritonia griegi</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF816506F933FD7DED07FCBB" author="Odhner N" pageId="17" pageNumber="462" pagination="1 - 47" refId="ref26522" refString="Odhner N. Norwegian opisthobranchiate Mollusca in the collection of the zoological Museum of Kristiania. Nyt Magazin for Naturvidenskaberne 1922; 60: 1 - 47." type="journal article" year="1922">Odhner 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506FF27FC9DEB29FCBB" authority="Ortea and Moro 2020" authorityName="Ortea and Moro" authorityYear="2020" box="[885,1358,792,816]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="hirondelle">
|
||
<emphasis id="B97BEA88FF816506FF27FC9DEA51FCBB" box="[885,1078,792,816]" italics="true" pageId="17" pageNumber="462">Tritonia hirondelle</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF816506F81AFC9CEB29FCBB" author="Ortea J & Moro L" box="[1096,1358,792,816]" pageId="17" pageNumber="462" pagination="29 - 34" refId="ref26633" refString="Ortea J, Moro L. El FS Poseidon rinde homenaje al hirondelle: descripcion de una nueva especie de Tritonia Cuvier, 1798 (Mollusca: Nudibranchia) de las aguas profundas de las islas de Cabo Verde para celebrar las singladuras de un yate carismatico. Avicennia 2020; 27: 29 - 34." type="journal article" year="2020">Ortea and Moro 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506F931FC9DEDECFCDB" authorityName="Cuvier" authorityYear="1803" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="hombergii">
|
||
<emphasis id="B97BEA88FF816506F931FC9DEDECFCDB" italics="true" pageId="17" pageNumber="462">Tritonia hombergii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506FFF7FCBCEA81FCDB" authority="Bergh 1904" authorityName="Bergh" authorityYear="1904" box="[933,1254,824,848]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="incerta">
|
||
<emphasis id="B97BEA88FF816506FFF7FCBCEA2CFCC4" box="[933,1099,824,847]" italics="true" pageId="17" pageNumber="462">Tritonia incerta</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF816506F80DFCBCEA81FCDB" author="Bergh LSR" box="[1119,1254,824,848]" pageId="17" pageNumber="462" pagination="1 - 56" refId="ref24457" refString="Bergh LSR. Malacologische Untersuchungen. Reisen im Archipel der Philippinen von Dr. Carl Gottflied Semper. Zweiter Theil. Wissenschaflliche Resultate. Band 9, Theil 6, Lief. 1, 1904, pp 1 - 56." type="book chapter" year="1904">Bergh 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506F8AFFCBCEDC1FCE4" authority="Bergh 1907" authorityName="Bergh" authorityYear="1907" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="indecora">
|
||
<emphasis id="B97BEA88FF816506F8AFFCBCEBD4FCDB" box="[1277,1459,824,848]" italics="true" pageId="17" pageNumber="462">Tritonia indecora</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF816506FF78FCD3EDC1FCE4" author="Bergh R" box="[810,934,855,879]" pageId="17" pageNumber="462" pagination="1 - 144" refId="ref24498" refString="Bergh R. The Opisthobranchiata of South Africa. Transactions of the South Aflican Philosophical Society 1907; 17: 1 - 144." type="journal article" year="1907">Bergh 1907</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506FFE0FCDCEB30FCE4" authority="Zhang and Zhang 2021" authorityName="Zhang and Zhang" authorityYear="2021" box="[946,1367,855,879]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="iocasica">
|
||
<emphasis id="B97BEA88FF816506FFE0FCDCEA32FCE4" box="[946,1109,856,879]" italics="true" pageId="17" pageNumber="462">Tritonia iocasica</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF816506F80CFCD3EB30FCE4" author="Zhang S & Zhang S" box="[1118,1367,855,879]" pageId="17" pageNumber="462" pagination="1817 - 1829" refId="ref28172" refString="Zhang S, Zhang S. Tritonia iocasica sp. nov., a new tritoniid species from a seamount in the tropical western Pacific (Heterobranchia: Nudibranchia). Journal of Oceanology and Limnology 2021; 39: 1817 - 1829." type="journal article" year="2021">Zhang and Zhang 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506F931FCDCED13FC05" authorityName="Er. Marcus" authorityYear="1959" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="odhneri">
|
||
<emphasis id="B97BEA88FF816506F931FCDCED13FC05" italics="true" pageId="17" pageNumber="462">Tritonia odhneri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506FFD0FCF3EA9FFC05" authority="Minichev 1971" authorityName="Minichev" authorityYear="1971" box="[898,1272,886,910]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="primorjensis">
|
||
<emphasis id="B97BEA88FF816506FFD0FCF3EA35FC05" box="[898,1106,887,910]" italics="true" pageId="17" pageNumber="462">Tritonia primorjensis</emphasis>
|
||
Minichev 1971
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506F956FCF3EBD4FC05" authorityName="Aurivillius" authorityYear="1887" box="[1284,1459,886,910]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="psoloides">
|
||
<emphasis id="B97BEA88FF816506F956FCF3EBD4FC05" box="[1284,1459,886,910]" italics="true" pageId="17" pageNumber="462">Tritonia psoloides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF816506FF04FC12EA71FC26" authorityName="Pallas" authorityYear="1788" box="[854,1046,918,941]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="tetraquetra">
|
||
<emphasis id="B97BEA88FF816506FF04FC12EA71FC26" box="[854,1046,918,941]" italics="true" pageId="17" pageNumber="462">Tritonia tetraquetra</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156511FF816505FF78FC52EAA5FB39" lastPageId="18" lastPageNumber="463" pageId="17" pageNumber="462" type="discussion">
|
||
<paragraph id="8BB0369AFF816506FF78FC52EA2DF914" blockId="17.[810,1460,982,1977]" pageId="17" pageNumber="462">
|
||
<emphasis id="B97BEA88FF816506FF78FC52EDEFFC65" box="[810,904,982,1006]" italics="true" pageId="17" pageNumber="462">Remarks:</emphasis>
|
||
The recent reassessment of
|
||
<taxonomicName id="4C0F4D19FF816506F8C9FC53EA8CFC65" authorityName="Cuvier" authorityYear="1798" box="[1179,1259,983,1006]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF816506F8C9FC53EA8CFC65" box="[1179,1259,983,1006]" italics="true" pageId="17" pageNumber="462">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF9E4B6BFF816506F95EFC52ED87FB85" author="Korshunova T & Martynov A" pageId="17" pageNumber="462" pagination="0242103" refId="ref25588" refString="Korshunova T, Martynov A. Consolidated data on the phylogeny and evolution of the family Tritoniidae (Gastropoda: Nudibranchia) contribute to genera reassessment and clarify the taxonomic status of the neuroscience models Tritonia and Tochuina. PLoS One 2020; 15: e 0242103. hups: // doi. org / 10.1371 / journal. pone. 0242103" type="journal article" year="2020">Korshunova and Martynov (2020)</bibRefCitation>
|
||
resulted in several species being transferred to the genera
|
||
<taxonomicName id="4C0F4D19FF816506FFCFFB91EA6DFBA6" authorityName="Risso" authorityYear="1826" box="[925,1034,1045,1069]" class="Gastropoda" family="Tritoniidae" genus="Duvaucelia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF816506FFCFFB91EA6DFBA6" box="[925,1034,1045,1069]" italics="true" pageId="17" pageNumber="462">Duvaucelia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF816506F86CFB91EACDFBA6" authorityName="Korshunova and Martynov" authorityYear="2020" box="[1086,1194,1045,1069]" class="Gastropoda" family="Tritoniidae" genus="Tritonicula" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF816506F86CFB91EACDFBA6" box="[1086,1194,1045,1069]" italics="true" pageId="17" pageNumber="462">Tritonicula</emphasis>
|
||
</taxonomicName>
|
||
. Yet,
|
||
<taxonomicName id="4C0F4D19FF816506F8B1FB92EB54FBA6" authorityName="Cuvier" authorityYear="1798" box="[1251,1331,1046,1069]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF816506F8B1FB92EB54FBA6" box="[1251,1331,1046,1069]" italics="true" pageId="17" pageNumber="462">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
remains the most heterogeneous genus of plateless tritoniids, with a majority of the species exhibiting characters not found in the
|
||
<typeStatus id="54B48838FF816506F901FBD0EBE6FBE7" box="[1363,1409,1108,1132]" pageId="17" pageNumber="462">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C0F4D19FF816506FF0AFBF0EA77FB00" authorityName="Cuvier" authorityYear="1803" box="[856,1040,1139,1163]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="hombergii">
|
||
<emphasis id="B97BEA88FF816506FF0AFBF0EA77FB00" box="[856,1040,1139,1163]" italics="true" pageId="17" pageNumber="462">Tritonia hombergii</emphasis>
|
||
</taxonomicName>
|
||
, as well as high values of minimum
|
||
<emphasis id="B97BEA88FF816506F9D5FBF7EBD4FB01" box="[1415,1459,1139,1162]" italics="true" pageId="17" pageNumber="462">COI</emphasis>
|
||
interspecific
|
||
<emphasis id="B97BEA88FF816506FFE1FB17EDA7FB21" box="[947,960,1171,1194]" italics="true" pageId="17" pageNumber="462">p</emphasis>
|
||
-distances (Table 3). For example, the Pacific
|
||
<taxonomicName id="4C0F4D19FF816506FF78FB36EDDAFB41" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[810,957,1202,1226]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="festiva">
|
||
<emphasis id="B97BEA88FF816506FF78FB36EDDAFB41" box="[810,957,1202,1226]" italics="true" pageId="17" pageNumber="462">Tritonia festiva</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF816506FFA0FB36EAFCFB41" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[1010,1179,1202,1226]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="exsulans">
|
||
<emphasis id="B97BEA88FF816506FFA0FB36EAFCFB41" box="[1010,1179,1202,1226]" italics="true" pageId="17" pageNumber="462">Tritonia exsulans</emphasis>
|
||
</taxonomicName>
|
||
, recovered as sister-species in our phylogenetic analyses (
|
||
<figureCitation id="13342A1FFF816506F80BFB55EAF6FB62" box="[1113,1169,1233,1257]" captionStart="Figure 2" captionStartId="10.[129,194,1719,1743]" captionTargetBox="[325,1278,148,1689]" captionTargetId="figure-6@10.[321,1281,144,1692]" captionTargetPageId="10" captionText="Figure 2. Bayesian phylogenetic tree based on the concatenated molecular data (COI + 16S + H3) of Tritoniidae (sensu Odhner 1963). Outgroup taxa are depicted in grey.Tritoniidae genera are independently coloured, and branches with several replicates for each species hypothesis were collapsed for clarity.Names in bold are species which sequences were generated for this study with the number of sequenced specimens showed in brackets. PP are shown right/top and BS leħ/boưom.Unsupported branches are not labelled. Green circles indicate maximum support by both analyses. Solid vertical bars indicate specimens grouped as species entities based on COI dataset. Empty vertical bars indicate the specimens grouped as species based on the 16S dataset." figureDoi="http://doi.org/10.5281/zenodo.8432395" httpUri="https://zenodo.org/record/8432395/files/figure.png" pageId="17" pageNumber="462">Fig. 2</figureCitation>
|
||
), have a similar penial shape (boưle-shaped penis with distal tip in
|
||
<taxonomicName id="4C0F4D19FF816506F8FFFB75EB59FA83" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[1197,1342,1264,1288]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="festiva">
|
||
<emphasis id="B97BEA88FF816506F8FFFB75EB59FA83" box="[1197,1342,1264,1288]" italics="true" pageId="17" pageNumber="462">Tritonia festiva</emphasis>
|
||
</taxonomicName>
|
||
and boưle-shaped penis with distal ring in
|
||
<taxonomicName id="4C0F4D19FF816506F821FA94EB7CFAA3" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[1139,1307,1296,1320]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="exsulans">
|
||
<emphasis id="B97BEA88FF816506F821FA94EB7CFAA3" box="[1139,1307,1296,1320]" italics="true" pageId="17" pageNumber="462">Tritonia exsulans</emphasis>
|
||
</taxonomicName>
|
||
) and cuticular folds in the stomach (
|
||
<bibRefCitation id="EF9E4B6BFF816506F840FAABEAAFFACC" author="MacFarland FM" box="[1042,1224,1327,1351]" pageId="17" pageNumber="462" pagination="1 - 72" refId="ref25838" refString="MacFarland FM. Studies of opisthobranchiate mollusks of the Pacific coast of North America. Memoirs of the California Academy of Sciences 1966; 6: 1 - 72." type="journal article" year="1966">MacFarland 1966</bibRefCitation>
|
||
). These characters are also found in the Pacific
|
||
<taxonomicName id="4C0F4D19FF816506F876FACBEB0EFAED" authority="(Schrodl 2003)" baseAuthorityName="Schrodl" baseAuthorityYear="2003" box="[1060,1385,1358,1382]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="odhneri">
|
||
<emphasis id="B97BEA88FF816506F876FACBEAA3FAED" box="[1060,1220,1358,1382]" italics="true" pageId="17" pageNumber="462">Tritonia odhneri</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9E4B6BFF816506F887FACAEB39FAED" author="Schrodl M" box="[1237,1374,1358,1382]" pageId="17" pageNumber="462" refId="ref27482" refString="Schrodl M. Sea slugs of southern South America - systematics, biogeography and biology of Chilean and Magellanic Nudipleura (Mollusca: Opisthobranchia). Hackenheim: ConchBooks, 2003." type="book" year="2003">Schrödl 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which was recovered clustered with
|
||
<taxonomicName id="4C0F4D19FF816506F81DFAEAEAB8FA0D" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[1103,1247,1390,1414]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="festiva">
|
||
<emphasis id="B97BEA88FF816506F81DFAEAEAB8FA0D" box="[1103,1247,1390,1414]" italics="true" pageId="17" pageNumber="462">Tritonia festiva</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF816506F95FFAEAEBD4FA0D" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[1293,1459,1390,1414]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="exsulans">
|
||
<emphasis id="B97BEA88FF816506F95FFAEAEBD4FA0D" box="[1293,1459,1390,1414]" italics="true" pageId="17" pageNumber="462">Tritonia exsulans</emphasis>
|
||
</taxonomicName>
|
||
in a recent molecular study (
|
||
<bibRefCitation id="EF9E4B6BFF816506F806FA09EB63FA2E" author="Moles J & Berning MI & Hooker Y" box="[1108,1284,1421,1445]" pageId="17" pageNumber="462" pagination="107209" refId="ref26342" refString="Moles J, Berning MI, Hooker Y, et al. Due South: the evolutionary history of sub-Antarctic and Antarctic Tritoniidae nudibranchs. Molecular Phylogenetics and Evolution 2021; 162: 107209." type="journal article" year="2021">
|
||
Moles
|
||
<emphasis id="B97BEA88FF816506F8C8FA0AEAA4FA2E" box="[1178,1219,1421,1445]" italics="true" pageId="17" pageNumber="462">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
). However, such characters are not reported for Atlantic
|
||
<emphasis id="B97BEA88FF816506F8BDFA29EBD3FA4F" box="[1263,1460,1452,1476]" italics="true" pageId="17" pageNumber="462">
|
||
<taxonomicName id="4C0F4D19FF816506F8BDFA29EBD7FA4F" authorityName="Cuvier" authorityYear="1803" box="[1263,1456,1452,1476]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="hombergii">Tritonia hombergii</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Furthermore, in recent molecular phylogenies (
|
||
<bibRefCitation id="EF9E4B6BFF816506F967FA4FED90F988" author="Korshunova T & Martynov A" pageId="17" pageNumber="462" pagination="0242103" refId="ref25588" refString="Korshunova T, Martynov A. Consolidated data on the phylogeny and evolution of the family Tritoniidae (Gastropoda: Nudibranchia) contribute to genera reassessment and clarify the taxonomic status of the neuroscience models Tritonia and Tochuina. PLoS One 2020; 15: e 0242103. hups: // doi. org / 10.1371 / journal. pone. 0242103" type="journal article" year="2020">Korshunova and Martynov 2020</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9E4B6BFF816506F851FA6FEAD6F988" author="Moles J & Berning MI & Hooker Y" box="[1027,1201,1515,1539]" pageId="17" pageNumber="462" pagination="107209" refId="ref26342" refString="Moles J, Berning MI, Hooker Y, et al. Due South: the evolutionary history of sub-Antarctic and Antarctic Tritoniidae nudibranchs. Molecular Phylogenetics and Evolution 2021; 162: 107209." type="journal article" year="2021">
|
||
Moles
|
||
<emphasis id="B97BEA88FF816506F81AFA68EA16F988" box="[1096,1137,1515,1539]" italics="true" pageId="17" pageNumber="462">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
; present study), the
|
||
<typeStatus id="54B48838FF816506F9D7FA6FEBD4F988" box="[1413,1459,1515,1539]" pageId="17" pageNumber="462">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C0F4D19FF816506FF2BF98FEA56F9A9" authorityName="Cuvier" authorityYear="1803" box="[889,1073,1546,1570]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="hombergii">
|
||
<emphasis id="B97BEA88FF816506FF2BF98FEA56F9A9" box="[889,1073,1546,1570]" italics="true" pageId="17" pageNumber="462">Tritonia hombergii</emphasis>
|
||
</taxonomicName>
|
||
was recovered as sister-species to the Pacific
|
||
<taxonomicName id="4C0F4D19FF816506FF26F9AEEA60F9C9" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[884,1031,1578,1602]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="festiva">
|
||
<emphasis id="B97BEA88FF816506FF26F9AEEA60F9C9" box="[884,1031,1578,1602]" italics="true" pageId="17" pageNumber="462">Tritonia festiva</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF816506F846F9AEEADAF9CA" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[1044,1213,1577,1601]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="exsulans">
|
||
<emphasis id="B97BEA88FF816506F846F9AEEADAF9CA" box="[1044,1213,1577,1601]" italics="true" pageId="17" pageNumber="462">Tritonia exsulans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF816506F8A3F9AEEBD4F9CA" authorityName="Pallas" authorityYear="1788" box="[1265,1459,1578,1601]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="tetraquetra">
|
||
<emphasis id="B97BEA88FF816506F8A3F9AEEBD4F9CA" box="[1265,1459,1578,1601]" italics="true" pageId="17" pageNumber="462">Tritonia tetraquetra</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<figureCitation id="13342A1FFF816506FF2AF9CDEDD6F9EA" box="[888,945,1609,1633]" captionStart="Figure 2" captionStartId="10.[129,194,1719,1743]" captionTargetBox="[325,1278,148,1689]" captionTargetId="figure-6@10.[321,1281,144,1692]" captionTargetPageId="10" captionText="Figure 2. Bayesian phylogenetic tree based on the concatenated molecular data (COI + 16S + H3) of Tritoniidae (sensu Odhner 1963). Outgroup taxa are depicted in grey.Tritoniidae genera are independently coloured, and branches with several replicates for each species hypothesis were collapsed for clarity.Names in bold are species which sequences were generated for this study with the number of sequenced specimens showed in brackets. PP are shown right/top and BS leħ/boưom.Unsupported branches are not labelled. Green circles indicate maximum support by both analyses. Solid vertical bars indicate specimens grouped as species entities based on COI dataset. Empty vertical bars indicate the specimens grouped as species based on the 16S dataset." figureDoi="http://doi.org/10.5281/zenodo.8432395" httpUri="https://zenodo.org/record/8432395/files/figure.png" pageId="17" pageNumber="462">Fig. 2</figureCitation>
|
||
). This molecular evidence is consistent with morphological data that further distinguish the Pacific
|
||
<taxonomicName id="4C0F4D19FF816506F960F9EDED36F914" pageId="17" pageNumber="462">
|
||
<emphasis id="B97BEA88FF816506F960F9EDEBE5F90B" box="[1330,1410,1641,1664]" italics="true" pageId="17" pageNumber="462">Tritonia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C0F4D19FF816506FFDFF90CEA24F914" authorityName="Cuvier" authorityYear="1803" box="[909,1091,1671,1695]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="species" species="hombergii">
|
||
<emphasis id="B97BEA88FF816506FFDFF90CEA24F914" box="[909,1091,1671,1695]" italics="true" pageId="17" pageNumber="462">Tritonia hombergii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF816505FF14F923EA19FDD4" blockId="17.[810,1460,982,1977]" lastBlockId="18.[825,1474,144,1202]" lastPageId="18" lastPageNumber="463" pageId="17" pageNumber="462">
|
||
Although our molecular data support the monophyly of
|
||
<taxonomicName id="4C0F4D19FF816506FF78F943ED1DF955" authorityName="Cuvier" authorityYear="1798" box="[810,890,1735,1758]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF816506FF78F943ED1DF955" box="[810,890,1735,1758]" italics="true" pageId="17" pageNumber="462">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
, it is difficult to establish synapomorphies for such a heterogeneous group.
|
||
<bibRefCitation id="EF9E4B6BFF816506FFBFF961EB33F975" author="Korshunova T & Martynov A" box="[1005,1364,1765,1790]" pageId="17" pageNumber="462" pagination="0242103" refId="ref25588" refString="Korshunova T, Martynov A. Consolidated data on the phylogeny and evolution of the family Tritoniidae (Gastropoda: Nudibranchia) contribute to genera reassessment and clarify the taxonomic status of the neuroscience models Tritonia and Tochuina. PLoS One 2020; 15: e 0242103. hups: // doi. org / 10.1371 / journal. pone. 0242103" type="journal article" year="2020">Korshunova and Martynov (2020)</bibRefCitation>
|
||
included in
|
||
<taxonomicName id="4C0F4D19FF816506FF14F881EDF1F897" authorityName="Cuvier" authorityYear="1798" box="[838,918,1797,1820]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF816506FF14F881EDF1F897" box="[838,918,1797,1820]" italics="true" pageId="17" pageNumber="462">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
mainly large tritoniids with numerous processes on the bilobed oral veil and a large number of lateral teeth, which also included the species assigned to
|
||
<taxonomicName id="4C0F4D19FF816506F8FCF8C7EA9EF8D0" authorityName="Odhner" authorityYear="1963" box="[1198,1273,1859,1883]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="462" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF816506F8FCF8C7EA9EF8D0" box="[1198,1273,1859,1883]" italics="true" pageId="17" pageNumber="462">Myrella</emphasis>
|
||
</taxonomicName>
|
||
in this study. The two genera are externally similar, with numerous oral processes and arborescent gills that intersperse in size, but can be distinguished by a unique combination of morphological characters, particularly the penial morphology of
|
||
<taxonomicName id="4C0F4D19FF826505F896FF15EB73FF23" authorityName="Cuvier" authorityYear="1798" box="[1220,1300,145,168]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF826505F896FF15EB73FF23" box="[1220,1300,145,168]" italics="true" pageId="18" pageNumber="463">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
, which is significantly larger than that of
|
||
<taxonomicName id="4C0F4D19FF826505F86EFF2BEABBFF4C" box="[1084,1244,175,199]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B97BEA88FF826505F86EFF2BEAE0FF4C" box="[1084,1159,175,199]" italics="true" pageId="18" pageNumber="463">Myrella</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Thus, we have characterized
|
||
<taxonomicName id="4C0F4D19FF826505FFD6FF4BEDB3FF6D" authorityName="Cuvier" authorityYear="1798" box="[900,980,207,230]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF826505FFD6FF4BEDB3FF6D" box="[900,980,207,230]" italics="true" pageId="18" pageNumber="463">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
by a unique combination of features: a broad and bilobed oral veil with several simple or branched velar processes; fully arborescent gills with at least seven large pairs and smaller ones in between; and a large, elongate or conical penis. Due to the high morphological variance, the radula of
|
||
<taxonomicName id="4C0F4D19FF826505F94DFEC9EB08FEEF" authorityName="Cuvier" authorityYear="1798" box="[1311,1391,333,356]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF826505F94DFEC9EB08FEEF" box="[1311,1391,333,356]" italics="true" pageId="18" pageNumber="463">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
is not a good diagnostic character. The masticatory border of the jaw is also the most morphologically variable in
|
||
<taxonomicName id="4C0F4D19FF826505F8B4FE0FEB3DFE28" baseAuthorityName="sensu Odhner" baseAuthorityYear="1963" box="[1254,1370,395,419]" class="Gastropoda" family="Tritoniidae" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="family">Tritoniidae</taxonomicName>
|
||
, with species having medium-sized denticles with conical bases and sharp cusps, e.g.
|
||
<taxonomicName id="4C0F4D19FF826505FFF0FE4EEBF0FE6A" authority="Korshunova and Martynov 2020" authorityName="Korshunova and Martynov" authorityYear="2020" box="[930,1431,457,482]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="exsulans">
|
||
<emphasis id="B97BEA88FF826505FFF0FE4EEA2EFE6A" box="[930,1097,457,481]" italics="true" pageId="18" pageNumber="463">Tritonia exsulans</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF826505F81EFE4DEBF0FE6A" author="Korshunova T & Martynov A" box="[1100,1431,457,482]" pageId="18" pageNumber="463" pagination="0242103" refId="ref25588" refString="Korshunova T, Martynov A. Consolidated data on the phylogeny and evolution of the family Tritoniidae (Gastropoda: Nudibranchia) contribute to genera reassessment and clarify the taxonomic status of the neuroscience models Tritonia and Tochuina. PLoS One 2020; 15: e 0242103. hups: // doi. org / 10.1371 / journal. pone. 0242103" type="journal article" year="2020">Korshunova and Martynov 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF826505FF6BFE6DEA39FD8A" authority="Stearns 1873" authorityName="Stearns" authorityYear="1873" box="[825,1118,489,513]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="festiva">
|
||
<emphasis id="B97BEA88FF826505FF6BFE6DEDAAFD8A" box="[825,973,489,513]" italics="true" pageId="18" pageNumber="463">Tritonia festiva</emphasis>
|
||
Stearns 1873
|
||
</taxonomicName>
|
||
; large denticles with conical bases and blunt cusps, e.g.
|
||
<taxonomicName id="4C0F4D19FF826505F85BFD8DEB4DFDAB" authority="Bouchet 1977" authorityName="Bouchet" authorityYear="1977" box="[1033,1322,520,544]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="griegi">
|
||
<emphasis id="B97BEA88FF826505F85BFD8DEAF4FDAB" box="[1033,1171,521,544]" italics="true" pageId="18" pageNumber="463">Tritonia griegi</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF826505F8CAFD8CEB4DFDAB" author="Bouchet P" box="[1176,1322,520,544]" pageId="18" pageNumber="463" pagination="28 - 60" refId="ref24570" refString="Bouchet P. Opisthobranches de profondeur de l'Ocean Atlantique, II Notaspidea et Nudibranchiata. Journal of Molluscan Studies 1977; 43: 28 - 60." type="journal article" year="1977">Bouchet 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
; or rod-shaped denticles, e.g.
|
||
<taxonomicName id="4C0F4D19FF826505FF82FDACEAECFDB4" authorityName="Cuvier" authorityYear="1803" box="[976,1163,551,575]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="hombergii">
|
||
<emphasis id="B97BEA88FF826505FF82FDACEAECFDB4" box="[976,1163,551,575]" italics="true" pageId="18" pageNumber="463">Tritonia hombergii</emphasis>
|
||
</taxonomicName>
|
||
(present study) and
|
||
<taxonomicName id="4C0F4D19FF826505F920FDACEA1EFDD4" authority="Zhang and Zhang 2021" authorityName="Zhang and Zhang" authorityYear="2021" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="iocasica">
|
||
<emphasis id="B97BEA88FF826505F920FDACEDE3FDD5" italics="true" pageId="18" pageNumber="463">Tritonia iocasica</emphasis>
|
||
<bibRefCitation id="EF9E4B6BFF826505FFD8FDC2EA1EFDD4" author="Zhang S & Zhang S" box="[906,1145,582,607]" pageId="18" pageNumber="463" pagination="1817 - 1829" refId="ref28172" refString="Zhang S, Zhang S. Tritonia iocasica sp. nov., a new tritoniid species from a seamount in the tropical western Pacific (Heterobranchia: Nudibranchia). Journal of Oceanology and Limnology 2021; 39: 1817 - 1829." type="journal article" year="2021">Zhang and Zhang 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF826505FD42FF5DECCCF83F" pageId="18" pageNumber="463">
|
||
<table id="F90FC43AFF829AE8FD42FF17ECCCF83E" box="[272,683,147,1973]" gridcols="2" gridrows="14" pageId="18" pageNumber="463">
|
||
<tr id="353F34D8FF829AE8FD42FF17ECCCFE91" box="[272,683,147,282]" gridrow="0" pageId="18" pageNumber="463" rowspan-1="1">
|
||
<th id="76EE5DA4FF829AE8FD42FF17EF4EFE91" box="[272,297,147,282]" gridcol="0" gridrow="0" pageId="18" pageNumber="463">within genera</th>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FE9FECCCFEFC" box="[272,683,283,375]" gridrow="1" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FE9FECCCFEFC" box="[272,683,283,375]" colspan="2" colspanRight="1" gridcol="0" gridrow="1" pageId="18" pageNumber="463">Distance 0 –17.2 0.3– 19.5 0.5 –16.5 0.2 0 – 14 0 0 16 0 – 13.6 0.6 – 19 3.8 –22.5</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FE1AECCCFE67" box="[272,683,414,492]" gridrow="2" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FE1AEF4EFE67" box="[272,297,414,492]" gridcol="0" gridrow="2" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FE57EF4EFE67" bold="true" box="[272,297,467,492]" pageId="18" pageNumber="463">10</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FE1AECCCFE67" box="[354,683,414,492]" gridcol="1" gridrow="2" pageId="18" pageNumber="463">14.5 – 22</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FD98ECCCFDE1" box="[272,683,540,618]" gridrow="3" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FD98EF4EFDE1" box="[272,297,540,618]" gridcol="0" gridrow="3" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FDD9EF4EFDE1" bold="true" box="[272,297,605,618]" pageId="18" pageNumber="463">9</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FD98ECCCFDE1" box="[354,683,540,618]" gridcol="1" gridrow="3" pageId="18" pageNumber="463">19 – 22.5 19 – 24</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FD1DECCCFD72" box="[272,683,665,761]" gridrow="4" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FD1DEF4EFD72" box="[272,297,665,761]" gridcol="0" gridrow="4" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FD68EF4EFD72" bold="true" box="[272,297,748,761]" pageId="18" pageNumber="463">8</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FD1DECCCFD72" box="[354,683,665,761]" gridcol="1" gridrow="4" pageId="18" pageNumber="463">17.7 – 19.8 18 – 22.7 19 – 21.3</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FCACECCCFCFC" box="[272,683,808,887]" gridrow="5" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FCACEF4EFCFC" box="[272,297,808,887]" gridcol="0" gridrow="5" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FCEEEF4EFCFC" bold="true" box="[272,297,874,887]" pageId="18" pageNumber="463">7</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FCACECCCFCFC" box="[354,683,808,887]" gridcol="1" gridrow="5" pageId="18" pageNumber="463">17 – 18 17 – 19 17 – 22.5 17 – 22.7</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FC22ECCCFC7E" box="[272,683,934,1013]" gridrow="6" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FC22EF4EFC7E" box="[272,297,934,1013]" gridcol="0" gridrow="6" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FC6CEF4EFC7E" bold="true" box="[272,297,1000,1013]" pageId="18" pageNumber="463">6</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FC22ECCCFC7E" box="[354,683,934,1013]" gridcol="1" gridrow="6" pageId="18" pageNumber="463">17.2 19.6 – 21 20 – 21 18.7 – 20 17 – 21</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FBA0ECCCFB0F" box="[272,683,1060,1156]" gridrow="7" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FBA0EF4EFB0F" box="[272,297,1060,1156]" gridcol="0" gridrow="7" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FBF3EF4EFB0F" bold="true" box="[272,297,1143,1156]" pageId="18" pageNumber="463">5</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FBA0ECCCFB0F" box="[354,683,1060,1156]" gridcol="1" gridrow="7" pageId="18" pageNumber="463">15 – 18 16– 19 17 – 20.5 17.8 – 21 17 – 22.5 16.3 – 21.5</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FB37ECCCFA98" box="[272,683,1203,1299]" gridrow="8" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FB37EF4EFA98" box="[272,297,1203,1299]" gridcol="0" gridrow="8" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FA82EF4EFA98" bold="true" box="[272,297,1286,1299]" pageId="18" pageNumber="463">4</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FB37ECCCFA98" box="[354,683,1203,1299]" gridcol="1" gridrow="8" pageId="18" pageNumber="463">18 – 19.6 19.8 – 20 19.7 – 20 19 – 22.5 17.5 – 19.5 18.5 – 21.4 19 – 22</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FAC6ECCCFA29" box="[272,683,1346,1442]" gridrow="9" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FAC6EF4EFA29" box="[272,297,1346,1442]" gridcol="0" gridrow="9" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42FA11EF4EFA29" bold="true" box="[272,297,1429,1442]" pageId="18" pageNumber="463">3</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FAC6ECCCFA29" box="[354,683,1346,1442]" gridcol="1" gridrow="9" pageId="18" pageNumber="463">16 – 19 16.5 – 18.5 18.5 – 20 18.4 – 19 18 – 21 16.3 – 19 16.6 – 21.3 17 – 22</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42FA52ECCCF9BE" box="[272,683,1494,1589]" gridrow="10" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42FA52EF4EF9BE" box="[272,297,1494,1589]" gridcol="0" gridrow="10" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42F9A0EF4EF9BA" bold="true" box="[272,297,1572,1585]" pageId="18" pageNumber="463">2</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30FA52ECCCF9BE" box="[354,683,1494,1589]" gridcol="1" gridrow="10" pageId="18" pageNumber="463">15 – 21.5 18 – 21.5 17 – 21 18 – 22 16.6 – 20 19 – 22.5 17 – 22 16.5 – 22 16.3 – 22.5</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42F9D9ECCCF94B" box="[272,683,1629,1728]" gridrow="11" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42F9D9EF4EF94B" box="[272,297,1629,1728]" gridcol="0" gridrow="11" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42F937EF4EF94B" bold="true" box="[272,297,1715,1728]" pageId="18" pageNumber="463">1</emphasis>
|
||
</td>
|
||
<td id="76EE5DA4FF829AE8FD30F9D9ECCCF94B" box="[354,683,1629,1728]" gridcol="1" gridrow="11" pageId="18" pageNumber="463">16.5 – 21 15 – 20 15.5 – 17.5 16 – 19.6 17 – 21 16.5 – 19 17 – 20.5 16 – 18.2 16.4 – 21.7 15.8 – 21</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42F96BECCCF8EE" box="[272,683,1775,1893]" gridrow="12" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42F96BECCCF8EE" box="[272,683,1775,1893]" colspan="2" colspanRight="1" gridcol="0" gridrow="12" pageId="18" pageNumber="463">
|
||
<emphasis id="B97BEA88FF826505FD42F898EF4EF8EE" bold="true" box="[272,297,1820,1893]" pageId="18" pageNumber="463">Genera</emphasis>
|
||
<emphasis id="B97BEA88FF826505FD12F89FECCCF8EF" box="[320,683,1775,1893]" italics="true" pageId="18" pageNumber="463">
|
||
<taxonomicName id="4C0F4D19FF826505FD12F89FEF3EF8EE" authorityName="Cuvier" authorityYear="1798" box="[320,345,1819,1893]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Tritonia</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FD30F88AEF1CF8EE" authorityName="Gray" authorityYear="1850" box="[354,379,1806,1893]" class="Gastropoda" family="Tritoniidae" genus="Candiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Candiella</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FDD6F885EFFAF8EE" authorityName="Korshunova and Martynov" authorityYear="2020" box="[388,413,1793,1893]" class="Gastropoda" family="Tritoniidae" genus="Tritonicula" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Tritonicula</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FDF7F97DEFD9F8EE" authorityName="Bergh" authorityYear="1907" box="[421,446,1785,1893]" class="Gastropoda" family="Tritoniidae" genus="Tritonidoxa" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Tritonidoxa</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FD95F97FEF87F8EE" authorityName="Eliot" authorityYear="1905" box="[455,480,1787,1893]" class="Gastropoda" family="Tritoniidae" genus="Tritoniopsis" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Tritoniopsis</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FDBBF887EC65F8EE" baseAuthorityName="Odhner" baseAuthorityYear="1963" box="[489,514,1795,1893]" class="Gastropoda" family="Tritoniidae" genus="Marianina" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Marianina</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FE59F883EC43F8EE" baseAuthorityName="Wagele" baseAuthorityYear="1989" box="[523,548,1799,1893]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Tritoniella</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FE7FF895EC21F8EE" baseAuthorityName="MacFarland" baseAuthorityYear="1966" box="[557,582,1809,1893]" class="Gastropoda" family="Tritoniidae" genus="Tochuina" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Tochuina</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FE1DF89BEC0FF8EE" authorityName="Odhner" authorityYear="1963" box="[591,616,1823,1893]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Myrella</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FE23F88BECEDF8EF" authorityName="Vayssiere" authorityYear="1877" box="[625,650,1807,1892]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Marionia</taxonomicName>
|
||
<taxonomicName id="4C0F4D19FF826505FEC0F96BECCCF8EF" authorityName="Odhner" authorityYear="1934" box="[658,683,1775,1892]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">Marioniopsis</taxonomicName>
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="353F34D8FF829AE8FD42F818ECCCF83E" box="[272,683,1948,1973]" gridrow="13" pageId="18" pageNumber="463">
|
||
<td id="76EE5DA4FF829AE8FD42F818ECCCF83E" box="[272,683,1948,1973]" colspan="2" colspanRight="1" gridcol="0" gridrow="13" pageId="18" pageNumber="463">1 2 3 4 5 6 7 8 9 10 11</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF826505FF07FDE2EAA5FB39" blockId="18.[825,1474,144,1202]" pageId="18" pageNumber="463">
|
||
The recently described
|
||
<taxonomicName id="4C0F4D19FF826505F812FDE3EA86FDF5" authorityName="Zhang and Zhang" authorityYear="2021" box="[1088,1249,615,638]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="iocasica">
|
||
<emphasis id="B97BEA88FF826505F812FDE3EA86FDF5" box="[1088,1249,615,638]" italics="true" pageId="18" pageNumber="463">Tritonia iocasica</emphasis>
|
||
</taxonomicName>
|
||
clearly belongs to this genus based on its molecular and morphological data (
|
||
<bibRefCitation id="EF9E4B6BFF826505F9D2FD01ED8AFD36" author="Zhang S & Zhang S" pageId="18" pageNumber="463" pagination="1817 - 1829" refId="ref28172" refString="Zhang S, Zhang S. Tritonia iocasica sp. nov., a new tritoniid species from a seamount in the tropical western Pacific (Heterobranchia: Nudibranchia). Journal of Oceanology and Limnology 2021; 39: 1817 - 1829." type="journal article" year="2021">Zhang and Zhang 2021</bibRefCitation>
|
||
). The species
|
||
<taxonomicName id="4C0F4D19FF826505F8C0FD21EB06FD37" box="[1170,1377,676,700]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="callogorgiae">
|
||
<emphasis id="B97BEA88FF826505F8C0FD21EB06FD37" box="[1170,1377,676,700]" italics="true" pageId="18" pageNumber="463">Tritonia callogorgiae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF826505F920FD21ED09FD57" authorityName="Bouchet" authorityYear="1977" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="griegi">
|
||
<emphasis id="B97BEA88FF826505F920FD21ED09FD57" italics="true" pageId="18" pageNumber="463">Tritonia griegi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF826505FF2EFD41EA54FD57" authorityName="Ortea and Moro" authorityYear="2020" box="[892,1075,708,732]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="hirondelle">
|
||
<emphasis id="B97BEA88FF826505FF2EFD41EA54FD57" box="[892,1075,708,732]" italics="true" pageId="18" pageNumber="463">Tritonia hirondelle</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF826505F813FD41EABBFD57" authorityName="Bergh" authorityYear="1904" box="[1089,1244,709,732]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="incerta">
|
||
<emphasis id="B97BEA88FF826505F813FD41EABBFD57" box="[1089,1244,709,732]" italics="true" pageId="18" pageNumber="463">Tritonia incerta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF826505F8B8FD41EBF3FD57" authorityName="Bergh" authorityYear="1907" box="[1258,1428,708,732]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="indecora">
|
||
<emphasis id="B97BEA88FF826505F8B8FD41EBF3FD57" box="[1258,1428,708,732]" italics="true" pageId="18" pageNumber="463">Tritonia indecora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF826505FF6BFD60EA6FFD70" authorityName="Minichev" authorityYear="1971" box="[825,1032,740,763]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="primorjensis">
|
||
<emphasis id="B97BEA88FF826505FF6BFD60EA6FFD70" box="[825,1032,740,763]" italics="true" pageId="18" pageNumber="463">Tritonia primorjensis</emphasis>
|
||
</taxonomicName>
|
||
for which no molecular data are available, are included based on their original description and recent morphological evidence, which fit our proposed diagnosis (
|
||
<bibRefCitation id="EF9E4B6BFF826505F9D7FCA6ED0AFCD2" author="Bergh LSR" pageId="18" pageNumber="463" pagination="1 - 56" refId="ref24457" refString="Bergh LSR. Malacologische Untersuchungen. Reisen im Archipel der Philippinen von Dr. Carl Gottflied Semper. Zweiter Theil. Wissenschaflliche Resultate. Band 9, Theil 6, Lief. 1, 1904, pp 1 - 56." type="book chapter" year="1904">Bergh 1904</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9E4B6BFF826505FF29FCC5EDC8FCD2" author="Bergh R" box="[891,943,833,857]" pageId="18" pageNumber="463" pagination="1 - 144" refId="ref24498" refString="Bergh R. The Opisthobranchiata of South Africa. Transactions of the South Aflican Philosophical Society 1907; 17: 1 - 144." type="journal article" year="1907">1907</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9E4B6BFF826505FFECFCC5EA2BFCD2" author="Odhner N" box="[958,1100,833,857]" pageId="18" pageNumber="463" pagination="1 - 47" refId="ref26522" refString="Odhner N. Norwegian opisthobranchiate Mollusca in the collection of the zoological Museum of Kristiania. Nyt Magazin for Naturvidenskaberne 1922; 60: 1 - 47." type="journal article" year="1922">Odhner 1922</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9E4B6BFF826505F80BFCC5EB50FCD2" author="Chimienti G & Angeleui L & Furfaro G" box="[1113,1335,833,857]" pageId="18" pageNumber="463" pagination="103364" refId="ref24637" refString="Chimienti G, Angeleui L, Furfaro G, et al. Habitat, morphology and trophism of Tritonia callogorgiae sp. nov., a large nudibranch inhabiting Callogorgia verticillata forests in the Mediterranean Sea. Deep Sea Research Part I: Oceanographic Research Papers 2020; 165: 103364." type="journal article" year="2020">
|
||
Chimienti
|
||
<emphasis id="B97BEA88FF826505F898FCC6EA92FCD2" box="[1226,1269,833,857]" italics="true" pageId="18" pageNumber="463">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9E4B6BFF826505F917FCC5EA68FCF2" author="Korshunova T & Martynov A" pageId="18" pageNumber="463" pagination="0242103" refId="ref25588" refString="Korshunova T, Martynov A. Consolidated data on the phylogeny and evolution of the family Tritoniidae (Gastropoda: Nudibranchia) contribute to genera reassessment and clarify the taxonomic status of the neuroscience models Tritonia and Tochuina. PLoS One 2020; 15: e 0242103. hups: // doi. org / 10.1371 / journal. pone. 0242103" type="journal article" year="2020">Korshunova and Martynov 2020</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9E4B6BFF826505F84DFCE5EB69FCF2" author="Ortea J & Moro L" box="[1055,1294,864,889]" pageId="18" pageNumber="463" pagination="29 - 34" refId="ref26633" refString="Ortea J, Moro L. El FS Poseidon rinde homenaje al hirondelle: descripcion de una nueva especie de Tritonia Cuvier, 1798 (Mollusca: Nudibranchia) de las aguas profundas de las islas de Cabo Verde para celebrar las singladuras de un yate carismatico. Avicennia 2020; 27: 29 - 34." type="journal article" year="2020">Ortea and Moro 2020</bibRefCitation>
|
||
). The remaining genetically unexplored species currently assigned to
|
||
<taxonomicName id="4C0F4D19FF826505F91CFC04EBF9FC1C" authorityName="Cuvier" authorityYear="1798" box="[1358,1438,896,919]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF826505F91CFC04EBF9FC1C" box="[1358,1438,896,919]" italics="true" pageId="18" pageNumber="463">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
are not included in the genus in this study due to inconsistencies with our proposed diagnosis. For example,
|
||
<taxonomicName id="4C0F4D19FF826505F951FC3BEBA5FC5D" authorityName="Bouchet" authorityYear="1977" box="[1283,1474,958,982]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="episcopalis">
|
||
<emphasis id="B97BEA88FF826505F951FC3BEBA5FC5D" box="[1283,1474,958,982]" italics="true" pageId="18" pageNumber="463">Tritonia episcopalis</emphasis>
|
||
</taxonomicName>
|
||
has an entire oral veil and accessory denticles on the first lateral tooth (
|
||
<bibRefCitation id="EF9E4B6BFF826505FFD7FC79EA7EFB9E" author="Bouchet P" box="[901,1049,1021,1045]" pageId="18" pageNumber="463" pagination="28 - 60" refId="ref24570" refString="Bouchet P. Opisthobranches de profondeur de l'Ocean Atlantique, II Notaspidea et Nudibranchiata. Journal of Molluscan Studies 1977; 43: 28 - 60." type="journal article" year="1977">Bouchet 1977</bibRefCitation>
|
||
), which are characteristic of
|
||
<taxonomicName id="4C0F4D19FF826505F902FC79EBDBFB9E" authorityName="Korshunova and Martynov" authorityYear="2020" box="[1360,1468,1021,1045]" class="Gastropoda" family="Tritoniidae" genus="Tritonicula" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF826505F902FC79EBDBFB9E" box="[1360,1468,1021,1045]" italics="true" pageId="18" pageNumber="463">Tritonicula</emphasis>
|
||
</taxonomicName>
|
||
. The transfer of Antarctic and sub-Antarctic
|
||
<taxonomicName id="4C0F4D19FF826505F952FB99EBC1FBBF" box="[1280,1446,1052,1076]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B97BEA88FF826505F952FB99EB37FBBF" box="[1280,1360,1053,1076]" italics="true" pageId="18" pageNumber="463">Tritonia</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C0F4D19FF826505FF6BFBB8EDE3FBDF" authorityName="Odhner" authorityYear="1963" box="[825,900,1084,1108]" class="Gastropoda" family="Tritoniidae" genus="Myrella" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF826505FF6BFBB8EDE3FBDF" box="[825,900,1084,1108]" italics="true" pageId="18" pageNumber="463">Myrella</emphasis>
|
||
</taxonomicName>
|
||
proposed in this study helps to alleviate the taxonomic issue of
|
||
<taxonomicName id="4C0F4D19FF826505FFDFFBD8EDBAFBF8" authorityName="Cuvier" authorityYear="1798" box="[909,989,1116,1139]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="463" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF826505FFDFFBD8EDBAFBF8" box="[909,989,1116,1139]" italics="true" pageId="18" pageNumber="463">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
. However, new collection efforts are needed to investigate potentially hidden generic lineages within the genus and to define clear generic boundaries.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |