treatments-xml/data/03/A4/87/03A487F4FF03467DD9AE2FC22366F82B.xml

<document id="58C790B93D911D11A7E14A39E895C4E5" ID-CLB-Dataset="276813" ID-DOI="10.11646/zootaxa.5134.1.5" ID-GBIF-Dataset="dd45000d-7fe4-4aa0-9f55-15c7a9de107b" ID-ISSN="1175-5326" ID-Zenodo-Dep="6531370" ID-ZooBank="85F3C1AE-CD82-4A3E-817C-CBEC9FCA6C8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652087510310" checkinUser="plazi" docAuthor="Zhou, Mingwei, He, Anyou, Wang, Fangxin, Li, Yusen &amp; Li, Chenhong" docDate="2022" docId="03A487F4FF03467DD9AE2FC22366F82B" docLanguage="en" docName="zootaxa.5134.1.5.pdf" docOrigin="Zootaxa 5134 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neodontobutis lani , Zhou and Li 2022, sp. nov." docType="treatment" docVersion="10" lastPageNumber="122" masterDocId="FF9DFF8CFF014674D9392B30203AFFFE" masterDocTitle="Neodontobutis lani, a new sleeper fish of the family Odontobutidae (Teleostei: Gobiiformes) from Guangxi, southern China" masterLastPageNumber="124" masterPageNumber="113" pageNumber="115" updateTime="1699333007954" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CE3E1CF660969D90A235C147507B262B">Neodontobutis lani, a new sleeper fish of the family Odontobutidae (Teleostei: Gobiiformes) from Guangxi, southern China</mods:title>
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<mods:namePart id="E954B79CB3850249B794B299F05C4138">Zhou, Mingwei</mods:namePart>
<mods:affiliation id="DF6B663B295BBEF7B7E452980A41DECE">Guangxi Key Laboratory of Aquatic Genetic Breeding and Healthy Aquaculture, Guangxi Academy of Fishery Sciences, Nanning 530021, China &amp; Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, China &amp; Shanghai Collaborative Innovation for Aquatic Animal Genetics and Breeding, Shanghai Ocean University, Shanghai 201306, China authors contributed equally to this work</mods:affiliation>
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<mods:namePart id="A0E2B8A5C67F2686097525BCDCB4B716">He, Anyou</mods:namePart>
<mods:affiliation id="E68A58FE8C56420BBEB3B9FF045D308C">Guangxi Key Laboratory of Aquatic Genetic Breeding and Healthy Aquaculture, Guangxi Academy of Fishery Sciences, Nanning 530021, China</mods:affiliation>
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<mods:namePart id="48A2F331BDA286C004694684565453D7">Wang, Fangxin</mods:namePart>
<mods:affiliation id="3D591428891A38D5D75A1B4E9F2CD4CE">Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, China &amp; Shanghai Collaborative Innovation for Aquatic Animal Genetics and Breeding, Shanghai Ocean University, Shanghai 201306, China authors contributed equally to this work</mods:affiliation>
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<mods:namePart id="8C0FBE6C4F4474DC577BF97C2DDDF8D7">Li, Yusen</mods:namePart>
<mods:affiliation id="46B07C3F5DB4D2C4950861B9DF9D5C49">Guangxi Key Laboratory of Aquatic Genetic Breeding and Healthy Aquaculture, Guangxi Academy of Fishery Sciences, Nanning 530021, China</mods:affiliation>
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<mods:namePart id="9F98067754D690971817EE1CA5BC1167">Li, Chenhong</mods:namePart>
<mods:affiliation id="FCF06FF697EC97563C1E76D4518917F8">Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, China &amp; Shanghai Collaborative Innovation for Aquatic Animal Genetics and Breeding, Shanghai Ocean University, Shanghai 201306, China authors contributed equally to this work</mods:affiliation>
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<taxonomicName id="4C0D4D61FF034676D9AE2FC22210FAF2" authority="Zhou and Li" authorityName=", Zhou and Li" authorityYear="2022" box="[151,554,1266,1293]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="115" phylum="Chordata" rank="species" species="lani" status="sp. nov.">Neodontobutis lani, Zhou and Li</taxonomicName>
, 
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<materialsCitation id="3B653CBFFF034676D9AE2E0B241BFA63" ID-GBIF-Occurrence="4435882301" collectingDate="2019-10" collectorName="J. H. Lan" country="China" county="Longzhou Town" location="Zuojiang River" municipality="Pearl River" pageId="2" pageNumber="115" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B979EAF0FF034676D9AE2E0B2131FAAB" bold="true" box="[151,267,1339,1365]" pageId="2" pageNumber="115">
<typeStatus id="54B68840FF034676D9AE2E0B213CFAAB" box="[151,262,1339,1365]" pageId="2" pageNumber="115" type="holotype">Holotype</typeStatus>
.
</emphasis>
SOU1801007-1 (25911), female, 
<quantity id="4CF59B07FF034676DBB52E0B22CBFAAB" box="[652,753,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="6.24" pageId="2" pageNumber="115" unit="mm" value="62.4">62.4 mm</quantity>
standard length, obtained from a local market and presumed to be collected from the 
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, a tributary of the 
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of the 
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basin, at 
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, Chongzuo City, 
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, 
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, collected by 
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, 
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</date>
(
<figureCitation id="13362A67FF034676DA882EB32429FA63" box="[945,1043,1411,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1866,1890]" captionTargetBox="[359,1301,1022,1842]" captionTargetId="figure-105@5.[285,1301,1004,1842]" captionTargetPageId="5" captionText="FIGURE 3. Neodontobutis lani sp. nov., SOU1801007-1, holotype, 62.4 mm standard length, Longzhou Town, Guangxi. A. dorsal, B. lateral, C. ventral views." figureDoi="http://doi.org/10.5281/zenodo.6531376" httpUri="https://zenodo.org/record/6531376/files/figure.png" pageId="2" pageNumber="115">Figure 3</figureCitation>
).
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<emphasis id="B979EAF0FF034676D9AE2C8B2128F82D" bold="true" box="[151,274,1979,2003]" pageId="2" pageNumber="115">FIGURE 1.</emphasis>
The sampling location of 
<taxonomicName id="4C0D4D61FF034676DB2E2C8B22EDF82D" authority="Zhou &amp; He &amp; Wang &amp; Li &amp; Li, 2022" authorityName="Zhou &amp; He &amp; Wang &amp; Li &amp; Li" authorityYear="2022" box="[535,727,1979,2003]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="115" phylum="Chordata" rank="species" species="lani" status="sp. nov.">
<emphasis id="B979EAF0FF034676DB2E2C8B22EDF82D" box="[535,727,1979,2003]" italics="true" pageId="2" pageNumber="115">Neodontobutis lani</emphasis>
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<taxonomicNameLabel id="A24A578BFF034676DBE52C8B231DF82D" box="[732,807,1979,2003]" pageId="2" pageNumber="115" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
, indicated by the triangle. The inset shows the sampling region within the map of China.
</paragraph>
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<emphasis id="B979EAF0FF024677D9AE2BA72136FF51" bold="true" box="[151,268,151,175]" pageId="3" pageNumber="116">TABLE 2.</emphasis>
Details of comparative materials. Abbreviation of “Material type”: S = Specimen, G = Genome DNA, N = Raw reads of nuclear target gene capture dataset. M = Mitochondrial genome, for extracting 
<emphasis id="B979EAF0FF024677DD4D2B8C249DFF2A" box="[1140,1191,188,212]" italics="true" pageId="3" pageNumber="116">cox1</emphasis>
sequence.
</paragraph>
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<paragraph id="8BB236E2FF024677D99F2BD4235BFAFB" pageId="3" pageNumber="116">
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<th id="76EC5DDCFF02B98BD99F2BD421D8FEDE" box="[166,482,228,288]" gridcol="0" gridrow="0" pageId="3" pageNumber="116">Collection number or sequence ID</th>
<th id="76EC5DDCFF02B98BDB322BD4225BFEDE" box="[523,609,228,288]" gridcol="1" gridrow="0" pageId="3" pageNumber="116">Material type</th>
<th id="76EC5DDCFF02B98BDBA22BD42394FEDE" box="[667,942,228,288]" gridcol="2" gridrow="0" pageId="3" pageNumber="116">Species</th>
<th id="76EC5DDCFF02B98BDD1C2BD42549FEDE" box="[1061,1395,228,288]" gridcol="3" gridrow="0" pageId="3" pageNumber="116">Location</th>
</tr>
<tr id="353D34A0FF02B98BD99F2A1F2549FE95" box="[166,1395,303,363]" gridrow="1" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F2A1F21D8FE95" box="[166,482,303,363]" gridcol="0" gridrow="1" pageId="3" pageNumber="116">25902, 25904, 25905, 25461, 25462, 25464, 25465</th>
<td id="76EC5DDCFF02B98BDB322A1F225BFE95" box="[523,609,303,363]" gridcol="1" gridrow="1" pageId="3" pageNumber="116">S, G</td>
<td id="76EC5DDCFF02B98BDBA22A1F2394FE95" box="[667,942,303,363]" gridcol="2" gridrow="1" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22A1F2394FEB9" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[667,942,303,327]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF024677DBA22A1F2394FEB9" box="[667,942,303,327]" italics="true" pageId="3" pageNumber="116">Neodontobutis hainanensis</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C2A1F2549FE95" box="[1061,1395,303,363]" gridcol="3" gridrow="1" pageId="3" pageNumber="116">Chongzuo, Guangxi, China</td>
</tr>
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<th id="76EC5DDCFF02B98BD99F2A4B21D8FE6A" box="[166,482,379,404]" gridcol="0" gridrow="2" pageId="3" pageNumber="116">25901, 25903, 25906</th>
<td id="76EC5DDCFF02B98BDB322A4B225BFE6A" box="[523,609,379,404]" gridcol="1" gridrow="2" pageId="3" pageNumber="116">S</td>
<td id="76EC5DDCFF02B98BDBA22A4B2394FE6A" box="[667,942,379,404]" gridcol="2" gridrow="2" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22A4C230BFE6A" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[667,817,380,404]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF024677DBA22A4C230BFE6A" box="[667,817,380,404]" italics="true" pageId="3" pageNumber="116">N. hainanensis</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C2A4B2549FE6A" box="[1061,1395,379,404]" gridcol="3" gridrow="2" pageId="3" pageNumber="116">Chongzuo, Guangxi, China</td>
</tr>
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<th id="76EC5DDCFF02B98BD99F2A9421D8FE1E" box="[166,482,420,480]" gridcol="0" gridrow="3" pageId="3" pageNumber="116">20272, 20276, 20278, 202710, 202712</th>
<td id="76EC5DDCFF02B98BDB322A94225BFE1E" box="[523,609,420,480]" gridcol="1" gridrow="3" pageId="3" pageNumber="116">G</td>
<td id="76EC5DDCFF02B98BDBA22A942394FE1E" box="[667,942,420,480]" gridcol="2" gridrow="3" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22A95230BFE42" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[667,817,420,444]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF024677DBA22A95230BFE42" box="[667,817,420,444]" italics="true" pageId="3" pageNumber="116">N. hainanensis</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C2A942549FE1E" box="[1061,1395,420,480]" gridcol="3" gridrow="3" pageId="3" pageNumber="116">Hainan, China</td>
</tr>
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<th id="76EC5DDCFF02B98BD99F2AC021D8FDF6" box="[166,482,496,520]" gridcol="0" gridrow="4" pageId="3" pageNumber="116">25012</th>
<td id="76EC5DDCFF02B98BDB322AC0225BFDF6" box="[523,609,496,520]" gridcol="1" gridrow="4" pageId="3" pageNumber="116">G</td>
<td id="76EC5DDCFF02B98BDBA22AC02394FDF6" box="[667,942,496,520]" gridcol="2" gridrow="4" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22AC0234FFDF6" authorityName="Herre" authorityYear="1940" box="[667,885,496,520]" class="Actinopterygii" family="Odontobutidae" genus="Sineleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="saccharae">
<emphasis id="B979EAF0FF024677DBA22AC0234FFDF6" box="[667,885,496,520]" italics="true" pageId="3" pageNumber="116">Sineleotris saccharae</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C2AC02549FDF6" box="[1061,1395,496,520]" gridcol="3" gridrow="4" pageId="3" pageNumber="116">Fangchenggang, Guangxi, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F29282549FDCE" box="[166,1395,536,560]" gridrow="5" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F292821D8FDCE" box="[166,482,536,560]" gridcol="0" gridrow="5" pageId="3" pageNumber="116">CL1279_4</th>
<td id="76EC5DDCFF02B98BDB322928225BFDCE" box="[523,609,536,560]" gridcol="1" gridrow="5" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA229282394FDCE" box="[667,942,536,560]" gridcol="2" gridrow="5" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22929230BFDCE" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[667,817,536,560]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF024677DBA22929230BFDCE" box="[667,817,536,560]" italics="true" pageId="3" pageNumber="116">N. hainanensis</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C29282549FDCE" box="[1061,1395,536,560]" gridcol="3" gridrow="5" pageId="3" pageNumber="116">Yangjiang, Guangdong, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F29712549FDA7" box="[166,1395,577,601]" gridrow="6" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F297121D8FDA7" box="[166,482,577,601]" gridcol="0" gridrow="6" pageId="3" pageNumber="116">CL632_1</th>
<td id="76EC5DDCFF02B98BDB322971225BFDA7" box="[523,609,577,601]" gridcol="1" gridrow="6" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA229712394FDA7" box="[667,942,577,601]" gridcol="2" gridrow="6" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22971237DFDA7" authorityName="Dybowski" authorityYear="1877" box="[667,839,577,601]" class="Actinopterygii" family="Odontobutidae" genus="Perccottus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="glenii">
<emphasis id="B979EAF0FF024677DBA22971237DFDA7" box="[667,839,577,601]" italics="true" pageId="3" pageNumber="116">Perccottus glenii</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C29712549FDA7" box="[1061,1395,577,601]" gridcol="3" gridrow="6" pageId="3" pageNumber="116">Harbin, Heilongjiang, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F29592549FD7F" box="[166,1395,617,641]" gridrow="7" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F295921D8FD7F" box="[166,482,617,641]" gridcol="0" gridrow="7" pageId="3" pageNumber="116">CL808_11</th>
<td id="76EC5DDCFF02B98BDB322959225BFD7F" box="[523,609,617,641]" gridcol="1" gridrow="7" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA229592394FD7F" box="[667,942,617,641]" gridcol="2" gridrow="7" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA229592354FD7F" authorityName="Wu, Chen &amp; Chong" authorityYear="2002" box="[667,878,617,641]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="B979EAF0FF024677DBA229592354FD7F" box="[667,878,617,641]" italics="true" pageId="3" pageNumber="116">Odontobutis sinensis</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C29592549FD7F" box="[1061,1395,617,641]" gridcol="3" gridrow="7" pageId="3" pageNumber="116">Guilin, Guangxi, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F29A12549FD57" box="[166,1395,657,681]" gridrow="8" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F29A121D8FD57" box="[166,482,657,681]" gridcol="0" gridrow="8" pageId="3" pageNumber="116">CL1221_2</th>
<td id="76EC5DDCFF02B98BDB3229A1225BFD57" box="[523,609,657,681]" gridcol="1" gridrow="8" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA229A12394FD57" box="[667,942,657,681]" gridcol="2" gridrow="8" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA229A22300FD57" baseAuthorityName="Guenther" baseAuthorityYear="1861" box="[667,826,657,681]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="potamophila">
<emphasis id="B979EAF0FF024677DBA229A22300FD57" box="[667,826,657,681]" italics="true" pageId="3" pageNumber="116">O. potamophila</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C29A12549FD57" box="[1061,1395,657,681]" gridcol="3" gridrow="8" pageId="3" pageNumber="116">Meicheng, Zhejiang, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F29892549FD2F" box="[166,1395,697,721]" gridrow="9" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F298921D8FD2F" box="[166,482,697,721]" gridcol="0" gridrow="9" pageId="3" pageNumber="116">CL1275_4</th>
<td id="76EC5DDCFF02B98BDB322989225BFD2F" box="[523,609,697,721]" gridcol="1" gridrow="9" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA229892394FD2F" box="[667,942,697,721]" gridcol="2" gridrow="9" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA2298A2322FD2F" authorityName="Wu, Wu &amp; Xie" authorityYear="1993" box="[667,792,697,721]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="yaluensis">
<emphasis id="B979EAF0FF024677DBA2298A2322FD2F" box="[667,792,697,721]" italics="true" pageId="3" pageNumber="116">O. yaluensis</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C29892549FD2F" box="[1061,1395,697,721]" gridcol="3" gridrow="9" pageId="3" pageNumber="116">Dandong, Liaoning, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F29D22549FD04" box="[166,1395,738,762]" gridrow="10" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F29D221D8FD04" box="[166,482,738,762]" gridcol="0" gridrow="10" pageId="3" pageNumber="116">CL1224_3</th>
<td id="76EC5DDCFF02B98BDB3229D2225BFD04" box="[523,609,738,762]" gridcol="1" gridrow="10" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA229D22394FD04" box="[667,942,738,762]" gridcol="2" gridrow="10" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA229D2234FFD04" authorityName="Herre" authorityYear="1940" box="[667,885,738,762]" class="Actinopterygii" family="Odontobutidae" genus="Sineleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="saccharae">
<emphasis id="B979EAF0FF024677DBA229D2234FFD04" box="[667,885,738,762]" italics="true" pageId="3" pageNumber="116">Sineleotris saccharae</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C29D22549FD04" box="[1061,1395,738,762]" gridcol="3" gridrow="10" pageId="3" pageNumber="116">Huizhou, Guangdong, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F283A2549FCDC" box="[166,1395,778,802]" gridrow="11" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F283A21D8FCDC" box="[166,482,778,802]" gridcol="0" gridrow="11" pageId="3" pageNumber="116">CL8_3</th>
<td id="76EC5DDCFF02B98BDB32283A225BFCDC" box="[523,609,778,802]" gridcol="1" gridrow="11" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA2283A2394FCDC" box="[667,942,778,802]" gridcol="2" gridrow="11" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA2283B23AEFCDC" baseAuthorityName="Gunther" baseAuthorityYear="1873" box="[667,916,778,802]" class="Actinopterygii" family="Odontobutidae" genus="Micropercops" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="swinhonis">
<emphasis id="B979EAF0FF024677DBA2283B23AEFCDC" box="[667,916,778,802]" italics="true" pageId="3" pageNumber="116">Micropercops swinhonis</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C283A2549FCDC" box="[1061,1395,778,802]" gridcol="3" gridrow="11" pageId="3" pageNumber="116">Langxi, Anhui, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F28022549FCB4" box="[166,1395,818,842]" gridrow="12" pageId="3" pageNumber="116">
<th id="76EC5DDCFF02B98BD99F280221D8FCB4" box="[166,482,818,842]" gridcol="0" gridrow="12" pageId="3" pageNumber="116">CL_929</th>
<td id="76EC5DDCFF02B98BDB322802225BFCB4" box="[523,609,818,842]" gridcol="1" gridrow="12" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA228022394FCB4" box="[667,942,818,842]" gridcol="2" gridrow="12" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22802235CFCB4" box="[667,870,818,842]" class="Bryopsida" family="Dicranaceae" genus="Microdus" kingdom="Plantae" order="Dicranales" pageId="3" pageNumber="116" phylum="Bryophyta" rank="species" species="chalmersi">
<emphasis id="B979EAF0FF024677DBA22802235CFCB4" box="[667,870,818,842]" italics="true" pageId="3" pageNumber="116">Microdus chalmersi</emphasis>
</taxonomicName>
</td>
<td id="76EC5DDCFF02B98BDD1C28022549FCB4" box="[1061,1395,818,842]" gridcol="3" gridrow="12" pageId="3" pageNumber="116">Wuzhishan, Hainan, China</td>
</tr>
<tr id="353D34A0FF02B98BD99F286A2549FC8D" box="[166,1395,858,883]" gridrow="13" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F286A21D8FC8D" box="[166,482,858,883]" gridcol="0" gridrow="13" pageId="3" pageNumber="116">CL303_1</th>
<td id="76EC5DDCFF02B98BDB32286A225BFC8D" box="[523,609,858,883]" gridcol="1" gridrow="13" pageId="3" pageNumber="116">N</td>
<td id="76EC5DDCFF02B98BDBA2286A2394FC8D" box="[667,942,858,883]" gridcol="2" gridrow="13" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA2286A235BFC8C" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[667,865,858,882]" class="Actinopterygii" family="Rhyacichthyidae" genus="Rhyacichthys" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="aspro">
<emphasis id="B979EAF0FF024677DBA2286A235BFC8C" box="[667,865,858,882]" italics="true" pageId="3" pageNumber="116">Rhyacichthys aspro</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F28B32549FC65" box="[166,1395,899,923]" gridrow="14" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F28B321D8FC65" box="[166,482,899,923]" gridcol="0" gridrow="14" pageId="3" pageNumber="116">NC_021763.1</th>
<td id="76EC5DDCFF02B98BDB3228B3225BFC65" box="[523,609,899,923]" gridcol="1" gridrow="14" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA228B32394FC65" box="[667,942,899,923]" gridcol="2" gridrow="14" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA228B323AEFC65" baseAuthorityName="Gunther" baseAuthorityYear="1873" box="[667,916,899,923]" class="Actinopterygii" family="Odontobutidae" genus="Micropercops" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="swinhonis">
<emphasis id="B979EAF0FF024677DBA228B323AEFC65" box="[667,916,899,923]" italics="true" pageId="3" pageNumber="116">Micropercops swinhonis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F289B2549FC3D" box="[166,1395,939,963]" gridrow="15" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F289B21D8FC3D" box="[166,482,939,963]" gridcol="0" gridrow="15" pageId="3" pageNumber="116">NC_022818.1</th>
<td id="76EC5DDCFF02B98BDB32289B225BFC3D" box="[523,609,939,963]" gridcol="1" gridrow="15" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA2289B2394FC3D" box="[667,942,939,963]" gridcol="2" gridrow="15" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA2289B2354FC3D" authorityName="Wu, Chen &amp; Chong" authorityYear="2002" box="[667,878,939,963]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="B979EAF0FF024677DBA2289B2354FC3D" box="[667,878,939,963]" italics="true" pageId="3" pageNumber="116">Odontobutis sinensis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F28E32549FC15" box="[166,1395,979,1003]" gridrow="16" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F28E321D8FC15" box="[166,482,979,1003]" gridcol="0" gridrow="16" pageId="3" pageNumber="116">NC_036056.1</th>
<td id="76EC5DDCFF02B98BDB3228E3225BFC15" box="[523,609,979,1003]" gridcol="1" gridrow="16" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA228E32394FC15" box="[667,942,979,1003]" gridcol="2" gridrow="16" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA228E42303FC15" authorityName="Chen" authorityYear="1985" box="[667,825,979,1003]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="haifengensis">
<emphasis id="B979EAF0FF024677DBA228E42303FC15" box="[667,825,979,1003]" italics="true" pageId="3" pageNumber="116">O. haifengensis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F28CB2549FBED" box="[166,1395,1019,1043]" gridrow="17" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F28CB21D8FBED" box="[166,482,1019,1043]" gridcol="0" gridrow="17" pageId="3" pageNumber="116">NC_027583.1</th>
<td id="76EC5DDCFF02B98BDB3228CB225BFBED" box="[523,609,1019,1043]" gridcol="1" gridrow="17" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA228CB2394FBED" box="[667,942,1019,1043]" gridcol="2" gridrow="17" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA228CC231BFBED" authorityName="Iwata &amp; Jeon" authorityYear="1985" box="[667,801,1020,1043]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="interrupta">
<emphasis id="B979EAF0FF024677DBA228CC231BFBED" box="[667,801,1020,1043]" italics="true" pageId="3" pageNumber="116">O. interrupta</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F2F142549FBC2" box="[166,1395,1060,1084]" gridrow="18" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F2F1421D8FBC2" box="[166,482,1060,1084]" gridcol="0" gridrow="18" pageId="3" pageNumber="116">NC_022706.1</th>
<td id="76EC5DDCFF02B98BDB322F14225BFBC2" box="[523,609,1060,1084]" gridcol="1" gridrow="18" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA22F142394FBC2" box="[667,942,1060,1084]" gridcol="2" gridrow="18" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22F142300FBC2" baseAuthorityName="Guenther" baseAuthorityYear="1861" box="[667,826,1060,1084]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="potamophila">
<emphasis id="B979EAF0FF024677DBA22F142300FBC2" box="[667,826,1060,1084]" italics="true" pageId="3" pageNumber="116">O. potamophila</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F2F7C2549FB9A" box="[166,1395,1100,1124]" gridrow="19" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F2F7C21D8FB9A" box="[166,482,1100,1124]" gridcol="0" gridrow="19" pageId="3" pageNumber="116">NC_010199.1</th>
<td id="76EC5DDCFF02B98BDB322F7C225BFB9A" box="[523,609,1100,1124]" gridcol="1" gridrow="19" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA22F7C2394FB9A" box="[667,942,1100,1124]" gridcol="2" gridrow="19" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22F7D2307FB9A" authorityName="Iwata &amp; Jeon" authorityYear="1985" box="[667,829,1100,1124]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="platycephala">
<emphasis id="B979EAF0FF024677DBA22F7D2307FB9A" box="[667,829,1100,1124]" italics="true" pageId="3" pageNumber="116">O. platycephala</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F2F442549FB72" box="[166,1395,1140,1164]" gridrow="20" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F2F4421D8FB72" box="[166,482,1140,1164]" gridcol="0" gridrow="20" pageId="3" pageNumber="116">NC_027160.1</th>
<td id="76EC5DDCFF02B98BDB322F44225BFB72" box="[523,609,1140,1164]" gridcol="1" gridrow="20" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA22F442394FB72" box="[667,942,1140,1164]" gridcol="2" gridrow="20" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22F452322FB72" authorityName="Wu, Wu &amp; Xie" authorityYear="1993" box="[667,792,1140,1164]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="yaluensis">
<emphasis id="B979EAF0FF024677DBA22F452322FB72" box="[667,792,1140,1164]" italics="true" pageId="3" pageNumber="116">O. yaluensis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F2FAC2549FB4B" box="[166,1395,1180,1205]" gridrow="21" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F2FAC21D8FB4B" box="[166,482,1180,1205]" gridcol="0" gridrow="21" pageId="3" pageNumber="116">NC_045932.1</th>
<td id="76EC5DDCFF02B98BDB322FAC225BFB4B" box="[523,609,1180,1205]" gridcol="1" gridrow="21" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA22FAC2394FB4B" box="[667,942,1180,1205]" gridcol="2" gridrow="21" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22FAD2349FB4B" baseAuthorityName="Nichols &amp; Pope" baseAuthorityYear="1927" box="[667,883,1181,1205]" class="Actinopteri" family="Odontobutidae" genus="Microdous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiiformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="chalmersi">
<emphasis id="B979EAF0FF024677DBA22FAD2349FB4B" box="[667,883,1181,1205]" italics="true" pageId="3" pageNumber="116">Microdous chalmersi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F2FF52549FB23" box="[166,1395,1221,1245]" gridrow="22" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F2FF521D8FB23" box="[166,482,1221,1245]" gridcol="0" gridrow="22" pageId="3" pageNumber="116">NC_020350.1</th>
<td id="76EC5DDCFF02B98BDB322FF5225BFB23" box="[523,609,1221,1245]" gridcol="1" gridrow="22" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA22FF52394FB23" box="[667,942,1221,1245]" gridcol="2" gridrow="22" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22FF6237DFB23" authorityName="Dybowski" authorityYear="1877" box="[667,839,1221,1245]" class="Actinopterygii" family="Odontobutidae" genus="Perccottus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="glenii">
<emphasis id="B979EAF0FF024677DBA22FF6237DFB23" box="[667,839,1221,1245]" italics="true" pageId="3" pageNumber="116">Perccottus glenii</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="353D34A0FF02B98BD99F2FDD2549FAFB" box="[166,1395,1261,1285]" gridrow="23" pageId="3" pageNumber="116" rowspan-3="1">
<th id="76EC5DDCFF02B98BD99F2FDD21D8FAFB" box="[166,482,1261,1285]" gridcol="0" gridrow="23" pageId="3" pageNumber="116">NC_004414.1</th>
<td id="76EC5DDCFF02B98BDB322FDD225BFAFB" box="[523,609,1261,1285]" gridcol="1" gridrow="23" pageId="3" pageNumber="116">M</td>
<td id="76EC5DDCFF02B98BDBA22FDD2394FAFB" box="[667,942,1261,1285]" gridcol="2" gridrow="23" pageId="3" pageNumber="116">
<taxonomicName id="4C0D4D61FF024677DBA22FDD235BFAFB" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[667,865,1261,1285]" class="Actinopterygii" family="Rhyacichthyidae" genus="Rhyacichthys" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="aspro">Rhyacichthys aspro</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB236E2FF024677D9FE2E05238BFA8D" blockId="3.[151,1437,1333,2043]" pageId="3" pageNumber="116">
<emphasis id="B979EAF0FF024677D9FE2E05217CFAB1" bold="true" box="[199,326,1333,1359]" pageId="3" pageNumber="116">
<typeStatus id="54B68840FF024677D9FE2E05217BFAB1" box="[199,321,1333,1359]" pageId="3" pageNumber="116" type="paratype">Paratypes</typeStatus>
.
</emphasis>
SOU1801007-3 (25913), female, 
<quantity id="4CF59B07FF024677DBE92E05230DFAB1" box="[720,823,1333,1360]" metricMagnitude="-2" metricUnit="m" metricValue="5.88" pageId="3" pageNumber="116" unit="mm" value="58.8">58.8 mm</quantity>
standard length; SOU1801007-5 (25915), male, 60.0 mm standard length. Collection locality and date same as the 
<typeStatus id="54B68840FF024677DA7E2E692397FA8D" box="[839,941,1369,1395]" pageId="3" pageNumber="116" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176569FF024677D9FE2E4C2288F8FE" pageId="3" pageNumber="116" type="diagnosis">
<paragraph id="8BB236E2FF024677D9FE2E4C2288F8FE" blockId="3.[151,1437,1333,2043]" pageId="3" pageNumber="116">
<emphasis id="B979EAF0FF024677D9FE2E4C2178FA68" bold="true" box="[199,322,1404,1430]" pageId="3" pageNumber="116">Diagnosis.</emphasis>
<taxonomicName id="4C0D4D61FF024677D8762E4D2216FA69" authorityName="Zhou &amp; He &amp; Wang &amp; Li &amp; Li" authorityYear="2022" box="[335,556,1405,1431]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF024677D8762E4D2216FA69" box="[335,556,1405,1431]" italics="true" pageId="3" pageNumber="116">Neodontobutis lani</emphasis>
</taxonomicName>
can be distinguished from other 
<taxonomicName id="4C0D4D61FF024677DAFF2E4D24F2FA69" box="[966,1224,1405,1431]" pageId="3" pageNumber="116">
<emphasis id="B979EAF0FF024677DAFF2E4D2453FA69" box="[966,1129,1405,1431]" italics="true" pageId="3" pageNumber="116">Neodontobutis</emphasis>
species
</taxonomicName>
by the following characteristics: anterior part of head relatively flat, with interorbital width / eye diameter =1.4–1.9 (mean=1.6), while the other 
<taxonomicName id="4C0D4D61FF024677D8772EF52276FA21" box="[334,588,1477,1503]" pageId="3" pageNumber="116">
<emphasis id="B979EAF0FF024677D8772EF521CBFA21" box="[334,497,1477,1503]" italics="true" pageId="3" pageNumber="116">Neodontobutis</emphasis>
species
</taxonomicName>
possess rather compressed head and body, with mean interorbital width / eye diameter 1.0 in 
<taxonomicName id="4C0D4D61FF024677D8462EDA2212F9FD" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[383,552,1513,1539]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF024677D8462EDA2212F9FD" box="[383,552,1513,1539]" italics="true" pageId="3" pageNumber="116">N. hainanensis</emphasis>
</taxonomicName>
, less than 
<quantity id="4CF59B07FF024677DB9C2ED922E9F9FD" box="[677,723,1513,1539]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="116" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C0D4D61FF024677DBE52EDA235BF9FD" box="[732,865,1514,1539]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="auarmus">
<emphasis id="B979EAF0FF024677DBE52EDA235BF9FD" box="[732,865,1514,1539]" italics="true" pageId="3" pageNumber="116">N. auarmus</emphasis>
</taxonomicName>
, 
<quantity id="4CF59B07FF024677DA4B2ED9238EF9FD" box="[882,948,1513,1539]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" pageId="3" pageNumber="116" unit="in" value="1.3">1.3 in</quantity>
<taxonomicName id="4C0D4D61FF024677DA842EDA24ADF9FD" authorityName="Mai" authorityYear="1978" box="[957,1175,1513,1539]" class="Actinopterygii" family="Odontobutidae" genus="Micropercops" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="macropectoralis">
<emphasis id="B979EAF0FF024677DA842EDA24ADF9FD" box="[957,1175,1513,1539]" italics="true" pageId="3" pageNumber="116">N. macropectoralis</emphasis>
</taxonomicName>
, 
<quantity id="4CF59B07FF024677DD912ED924D0F9FD" box="[1192,1258,1513,1539]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="3" pageNumber="116" unit="in" value="1.2">1.2 in</quantity>
<taxonomicName id="4C0D4D61FF024677DDCA2EDA25AEF9FD" baseAuthorityName="Mai" baseAuthorityYear="1978" box="[1267,1428,1513,1539]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="species" species="tonkinensis">
<emphasis id="B979EAF0FF024677DDCA2EDA25AEF9FD" box="[1267,1428,1513,1539]" italics="true" pageId="3" pageNumber="116">N. tonkinensis</emphasis>
</taxonomicName>
; sensory papilla on ventral tip of lower jaw form two oblong clusters (
<figureCitation id="13362A67FF024677DA962D3D2425F9D9" box="[943,1055,1549,1575]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1267]" captionTargetBox="[163,1420,191,1210]" captionTargetId="figure-28@6.[151,1436,181,1220]" captionTargetPageId="6" captionText="FIGURE 4. Patterns of main cephalic sensory papilla lines (row of spots) of Neodontobutis lani sp. nov. (SOU1801007-1, holotype) from lateral (a), dorsal (b) and ventral (c) view." figureDoi="http://doi.org/10.5281/zenodo.6531378" httpUri="https://zenodo.org/record/6531378/files/figure.png" pageId="3" pageNumber="116">Figure 4c</figureCitation>
), whereas in other 
<taxonomicName id="4C0D4D61FF024677DDC02D3D25A6F9D9" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1273,1436,1549,1575]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF024677DDC02D3D25A6F9D9" box="[1273,1436,1549,1575]" italics="true" pageId="3" pageNumber="116">Neodontobutis</emphasis>
</taxonomicName>
papillae form two single lines; several rows of transforming ctenii on posterior edge of body scales (
<figureCitation id="13362A67FF024677DC242D0125B4F9B2" box="[1309,1422,1585,1612]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[180,1408,1328,1919]" captionTargetId="figure-59@6.[179,1408,1328,1919]" captionTargetPageId="6" captionText="FIGURE 5. Posterior edge of scale on dorsal part of Neodontobutis lani SOU1801007-5, paratype (a) and N. hainanensis 25901 (b), ct, ctenii; tr, transforming ctenii. Sensory papilla on lower jaw of N. lani (c), p, sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.6531380" httpUri="https://zenodo.org/record/6531380/files/figure.png" pageId="3" pageNumber="116">Figure 5a</figureCitation>
), whereas single row of transforming ctenii on scale of the other 
<taxonomicName id="4C0D4D61FF024677DA5A2D65243CF991" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[867,1030,1621,1647]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF024677DA5A2D65243CF991" box="[867,1030,1621,1647]" italics="true" pageId="3" pageNumber="116">Neodontobutis</emphasis>
</taxonomicName>
(
<figureCitation id="13362A67FF024677DD2C2D6524BEF98E" box="[1045,1156,1621,1648]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[180,1408,1328,1919]" captionTargetId="figure-59@6.[179,1408,1328,1919]" captionTargetPageId="6" captionText="FIGURE 5. Posterior edge of scale on dorsal part of Neodontobutis lani SOU1801007-5, paratype (a) and N. hainanensis 25901 (b), ct, ctenii; tr, transforming ctenii. Sensory papilla on lower jaw of N. lani (c), p, sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.6531380" httpUri="https://zenodo.org/record/6531380/files/figure.png" pageId="3" pageNumber="116">Figure 5b</figureCitation>
) (Chen, Kottelat, &amp; Wu, 2002; 
<bibRefCitation id="EF9C4B13FF024677D9E62D49215DF96D" author="Iwata, A." box="[223,359,1657,1683]" pageId="3" pageNumber="116" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF024677D84D2D492212F96D" author="Kottelat, M." box="[372,552,1657,1683]" pageId="3" pageNumber="116" refId="ref5721" refString="Kottelat, M. (2001 b) Freshwater Fishes of Northern Vietnam. World Bank, Washington, 11 pp." type="book" year="2001">Kottelat, 2001b</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF024677DB0D2D492334F96A" author="Vidthayanon, C." box="[564,782,1657,1684]" pageId="3" pageNumber="116" pagination="235 - 242" refId="ref6186" refString="Vidthayanon, C. (1995) Odontobutis aurarmus, a new species of odontobutid goby from wetlands of the Mekong basin, Thailand. Ichthyological Exploration of Freshwaters, 6, 235 - 242." type="journal article" year="1995">Vidthayanon, 1995</bibRefCitation>
). This species can be distinguished from 
<taxonomicName id="4C0D4D61FF024677DDDC2D492555F96D" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1253,1391,1657,1683]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF024677DDDC2D492555F96D" box="[1253,1391,1657,1683]" italics="true" pageId="3" pageNumber="116">Odontobutis</emphasis>
</taxonomicName>
by: presence of barbel-like projection on sensory papilla (
<figureCitation id="13362A67FF024677DBC52DAD2350F946" box="[764,874,1693,1720]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[180,1408,1328,1919]" captionTargetId="figure-59@6.[179,1408,1328,1919]" captionTargetPageId="6" captionText="FIGURE 5. Posterior edge of scale on dorsal part of Neodontobutis lani SOU1801007-5, paratype (a) and N. hainanensis 25901 (b), ct, ctenii; tr, transforming ctenii. Sensory papilla on lower jaw of N. lani (c), p, sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.6531380" httpUri="https://zenodo.org/record/6531380/files/figure.png" pageId="3" pageNumber="116">Figure 5c</figureCitation>
) (absent in 
<taxonomicName id="4C0D4D61FF024677DACA2DAD2447F949" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1011,1149,1693,1719]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF024677DACA2DAD2447F949" box="[1011,1149,1693,1719]" italics="true" pageId="3" pageNumber="116">Odontobutis</emphasis>
</taxonomicName>
), separated right and left gill membranes (joined in 
<taxonomicName id="4C0D4D61FF024677D8F22DF1226FF925" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[459,597,1729,1755]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF024677D8F22DF1226FF925" box="[459,597,1729,1755]" italics="true" pageId="3" pageNumber="116">Odontobutis</emphasis>
</taxonomicName>
) and cycloid scales on predorsal region and abdomen (ctenoid scales in 
<taxonomicName id="4C0D4D61FF024677D9AE2DD5211BF901" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[151,289,1765,1791]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="116" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF024677D9AE2DD5211BF901" box="[151,289,1765,1791]" italics="true" pageId="3" pageNumber="116">Odontobutis</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EF9C4B13FF024677D8032DD521FBF901" author="Iwata, A." box="[314,449,1765,1791]" pageId="3" pageNumber="116" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF024677D8F42DD5229CF8FE" author="Vidthayanon, C." box="[461,678,1765,1792]" pageId="3" pageNumber="116" pagination="235 - 242" refId="ref6186" refString="Vidthayanon, C. (1995) Odontobutis aurarmus, a new species of odontobutid goby from wetlands of the Mekong basin, Thailand. Ichthyological Exploration of Freshwaters, 6, 235 - 242." type="journal article" year="1995">Vidthayanon, 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176569FF024673D9FE2C382178FD7B" lastPageId="7" lastPageNumber="120" pageId="3" pageNumber="116" type="description">
<paragraph id="8BB236E2FF024677D9FE2C38253BF892" blockId="3.[151,1437,1333,2043]" pageId="3" pageNumber="116">
<emphasis id="B979EAF0FF024677D9FE2C382163F8DC" bold="true" box="[199,345,1800,1826]" pageId="3" pageNumber="116">Description.</emphasis>
Morphometric and meristic data for the 
<typeStatus id="54B68840FF024677DA322C392357F8DD" box="[779,877,1801,1827]" pageId="3" pageNumber="116" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54B68840FF024677DAA72C392433F8DD" box="[926,1033,1801,1827]" pageId="3" pageNumber="116" type="paratype">paratypes</typeStatus>
are presented in 
<tableCitation id="C68F0359FF024677DD852C392536F8DD" box="[1212,1292,1801,1827]" captionStart="TABLE 3" captionStartId="4.[151,239,296,320]" captionTargetPageId="4" captionText="TABLE 3. Morphological characters for the type specimens of Neodontobutis lani." httpUri="http://table.plazi.org/id/DF72666AFF054670D9AE2A18242AFEBE" pageId="3" pageNumber="116" tableUuid="DF72666AFF054670D9AE2A18242AFEBE">Table 3</tableCitation>
respectively. Dorsal fin rays VII; I, 9; anal fin rays I, 6–7; pectoral fin rays 15, pelvic fin rays I, 5; caudal fin rays 14–15; lateral scales 34–36; transverse scales 12–15; predorsal scales 18–19; gill rakers 3+6; vertebrae 28 (13+15).
</paragraph>
<paragraph id="8BB236E2FF024677D9FE2C452381F829" blockId="3.[151,1437,1333,2043]" pageId="3" pageNumber="116">
Body stout, slightly compressed posteriorly. Head large, slightly flat, interorbital distance larger than eye diameter. Lower jaw slightly longer than upper. Gill membrane separated, attached to the isthmus. Cephalic sensory canals absent. Cephalic sensory papilla pattern is illustrated in 
<figureCitation id="13362A67FF024677DA6F2C8D238DF829" box="[854,951,1981,2007]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1267]" captionTargetBox="[163,1420,191,1210]" captionTargetId="figure-28@6.[151,1436,181,1220]" captionTargetPageId="6" captionText="FIGURE 4. Patterns of main cephalic sensory papilla lines (row of spots) of Neodontobutis lani sp. nov. (SOU1801007-1, holotype) from lateral (a), dorsal (b) and ventral (c) view." figureDoi="http://doi.org/10.5281/zenodo.6531378" httpUri="https://zenodo.org/record/6531378/files/figure.png" pageId="3" pageNumber="116">Figure 4</figureCitation>
.
</paragraph>
<paragraph id="8BB236E2FF024670D9FE2CD1239BFF07" blockId="3.[151,1437,1333,2043]" lastBlockId="4.[151,1437,151,249]" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116">Dorsal spines nearly equal length except for the last shorter one, not reaching origin of the second dorsal fin. Origin of the anal fin slightly posterior to the origin of the second dorsal fin. Pelvic fin short, close but separated. Pectoral fins and caudal fin wide, rounded. Most body scales ctenoid with several rows of transforming ctenii on the posterior edge. Predorsal scales and part of abdominal scales cycloid.</paragraph>
<caption id="DF72666AFF054670D9AE2A18242AFEBE" ID-Table-UUID="DF72666AFF054670D9AE2A18242AFEBE" box="[151,1040,296,321]" httpUri="http://table.plazi.org/id/DF72666AFF054670D9AE2A18242AFEBE" pageId="4" pageNumber="117" startId="4.[151,239,296,320]" targetBox="[166,1368,336,1966]" targetIsTable="true" targetPageId="4">
<paragraph id="8BB236E2FF054670D9AE2A18242AFEBE" blockId="4.[151,1040,296,321]" box="[151,1040,296,321]" pageId="4" pageNumber="117">
<emphasis id="B979EAF0FF054670D9AE2A182130FEBE" bold="true" box="[151,266,296,320]" pageId="4" pageNumber="117">TABLE 3.</emphasis>
Morphological characters for the type specimens of 
<taxonomicName id="4C0D4D61FF054670DA042A182432FEBE" authorityName="Zhou &amp; He &amp; Wang &amp; Li &amp; Li" authorityYear="2022" box="[829,1032,296,320]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="117" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF054670DA042A182432FEBE" box="[829,1032,296,320]" italics="true" pageId="4" pageNumber="117">Neodontobutis lani</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BB236E2FF054670D99F2A6124DEF850" pageId="4" pageNumber="117">
<table id="F90DC442FF05B98BD99F2A602562F850" box="[166,1368,336,1966]" gridcols="4" gridrows="40" pageId="4" pageNumber="117">
<tr id="353D34A0FF05B98BD99F2A602562FE97" box="[166,1368,336,361]" gridrow="0" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2A602275FE97" box="[166,591,336,361]" gridcol="0" gridrow="0" pageId="4" pageNumber="117">Characters</th>
<th id="76EC5DDCFF05B98BDBF52A602357FE97" box="[716,877,336,361]" gridcol="1" gridrow="0" pageId="4" pageNumber="117">SOU1801007-1</th>
<th id="76EC5DDCFF05B98BDAFB2A602457FE97" box="[962,1133,336,361]" gridcol="2" gridrow="0" pageId="4" pageNumber="117">SOU1801007-3</th>
<th id="76EC5DDCFF05B98BDD812A602562FE97" box="[1208,1368,336,361]" gridcol="3" gridrow="0" pageId="4" pageNumber="117">SOU1801007-5</th>
</tr>
<tr id="353D34A0FF05B98BD99F2A492562FE6F" box="[166,1368,377,401]" gridrow="1" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2A492562FE6F" box="[166,1368,377,401]" colspan="4" colspanRight="3" gridcol="0" gridrow="1" pageId="4" pageNumber="117">Counts</th>
</tr>
<tr id="353D34A0FF05B98BD99F2A912562FE47" box="[166,1368,417,441]" gridrow="2" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2A912275FE47" box="[166,591,417,441]" gridcol="0" gridrow="2" pageId="4" pageNumber="117">Dorsal fins</th>
<td id="76EC5DDCFF05B98BDBF52A912357FE47" box="[716,877,417,441]" gridcol="1" gridrow="2" pageId="4" pageNumber="117">VII;I,9</td>
<td id="76EC5DDCFF05B98BDAFB2A912457FE47" box="[962,1133,417,441]" gridcol="2" gridrow="2" pageId="4" pageNumber="117">VII;I,9</td>
<td id="76EC5DDCFF05B98BDD812A912562FE47" box="[1208,1368,417,441]" gridcol="3" gridrow="2" pageId="4" pageNumber="117">VII;I,9</td>
</tr>
<tr id="353D34A0FF05B98BD99F2AF92562FE1F" box="[166,1368,457,481]" gridrow="3" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2AF92275FE1F" box="[166,591,457,481]" gridcol="0" gridrow="3" pageId="4" pageNumber="117">Pectoral fins</th>
<td id="76EC5DDCFF05B98BDBF52AF92357FE1F" box="[716,877,457,481]" gridcol="1" gridrow="3" pageId="4" pageNumber="117">15</td>
<td id="76EC5DDCFF05B98BDAFB2AF92457FE1F" box="[962,1133,457,481]" gridcol="2" gridrow="3" pageId="4" pageNumber="117">15</td>
<td id="76EC5DDCFF05B98BDD812AF92562FE1F" box="[1208,1368,457,481]" gridcol="3" gridrow="3" pageId="4" pageNumber="117">15</td>
</tr>
<tr id="353D34A0FF05B98BD99F2AC12562FDF4" box="[166,1368,497,522]" gridrow="4" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2AC12275FDF4" box="[166,591,497,522]" gridcol="0" gridrow="4" pageId="4" pageNumber="117">Pelvic fins</th>
<td id="76EC5DDCFF05B98BDBF52AC12357FDF4" box="[716,877,497,522]" gridcol="1" gridrow="4" pageId="4" pageNumber="117">I,5</td>
<td id="76EC5DDCFF05B98BDAFB2AC12457FDF4" box="[962,1133,497,522]" gridcol="2" gridrow="4" pageId="4" pageNumber="117">I,5</td>
<td id="76EC5DDCFF05B98BDD812AC12562FDF4" box="[1208,1368,497,522]" gridcol="3" gridrow="4" pageId="4" pageNumber="117">I,5</td>
</tr>
<tr id="353D34A0FF05B98BD99F292A2562FDCC" box="[166,1368,538,562]" gridrow="5" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F292A2275FDCC" box="[166,591,538,562]" gridcol="0" gridrow="5" pageId="4" pageNumber="117">Anal fin</th>
<td id="76EC5DDCFF05B98BDBF5292A2357FDCC" box="[716,877,538,562]" gridcol="1" gridrow="5" pageId="4" pageNumber="117">I,7</td>
<td id="76EC5DDCFF05B98BDAFB292A2457FDCC" box="[962,1133,538,562]" gridcol="2" gridrow="5" pageId="4" pageNumber="117">I,7</td>
<td id="76EC5DDCFF05B98BDD81292A2562FDCC" box="[1208,1368,538,562]" gridcol="3" gridrow="5" pageId="4" pageNumber="117">I,6</td>
</tr>
<tr id="353D34A0FF05B98BD99F29722562FDA4" box="[166,1368,578,602]" gridrow="6" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F29722275FDA4" box="[166,591,578,602]" gridcol="0" gridrow="6" pageId="4" pageNumber="117">Caudal fin</th>
<td id="76EC5DDCFF05B98BDBF529722357FDA4" box="[716,877,578,602]" gridcol="1" gridrow="6" pageId="4" pageNumber="117">15</td>
<td id="76EC5DDCFF05B98BDAFB29722457FDA4" box="[962,1133,578,602]" gridcol="2" gridrow="6" pageId="4" pageNumber="117">15</td>
<td id="76EC5DDCFF05B98BDD8129722562FDA4" box="[1208,1368,578,602]" gridcol="3" gridrow="6" pageId="4" pageNumber="117">14</td>
</tr>
<tr id="353D34A0FF05B98BD99F295A2562FD7C" box="[166,1368,618,642]" gridrow="7" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F295A2275FD7C" box="[166,591,618,642]" gridcol="0" gridrow="7" pageId="4" pageNumber="117">Lateral scales</th>
<td id="76EC5DDCFF05B98BDBF5295A2357FD7C" box="[716,877,618,642]" gridcol="1" gridrow="7" pageId="4" pageNumber="117">34</td>
<td id="76EC5DDCFF05B98BDAFB295A2457FD7C" box="[962,1133,618,642]" gridcol="2" gridrow="7" pageId="4" pageNumber="117">36</td>
<td id="76EC5DDCFF05B98BDD81295A2562FD7C" box="[1208,1368,618,642]" gridcol="3" gridrow="7" pageId="4" pageNumber="117">36</td>
</tr>
<tr id="353D34A0FF05B98BD99F29A22562FD55" box="[166,1368,658,683]" gridrow="8" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F29A22275FD55" box="[166,591,658,683]" gridcol="0" gridrow="8" pageId="4" pageNumber="117">Transverse scales</th>
<td id="76EC5DDCFF05B98BDBF529A22357FD55" box="[716,877,658,683]" gridcol="1" gridrow="8" pageId="4" pageNumber="117">12</td>
<td id="76EC5DDCFF05B98BDAFB29A22457FD55" box="[962,1133,658,683]" gridcol="2" gridrow="8" pageId="4" pageNumber="117">15</td>
<td id="76EC5DDCFF05B98BDD8129A22562FD55" box="[1208,1368,658,683]" gridcol="3" gridrow="8" pageId="4" pageNumber="117">13</td>
</tr>
<tr id="353D34A0FF05B98BD99F298B2562FD2D" box="[166,1368,699,723]" gridrow="9" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F298B2275FD2D" box="[166,591,699,723]" gridcol="0" gridrow="9" pageId="4" pageNumber="117">Predorsal scales</th>
<td id="76EC5DDCFF05B98BDBF5298B2357FD2D" box="[716,877,699,723]" gridcol="1" gridrow="9" pageId="4" pageNumber="117">18</td>
<td id="76EC5DDCFF05B98BDAFB298B2457FD2D" box="[962,1133,699,723]" gridcol="2" gridrow="9" pageId="4" pageNumber="117">19</td>
<td id="76EC5DDCFF05B98BDD81298B2562FD2D" box="[1208,1368,699,723]" gridcol="3" gridrow="9" pageId="4" pageNumber="117">18</td>
</tr>
<tr id="353D34A0FF05B98BD99F29D32562FD02" box="[166,1368,739,764]" gridrow="10" pageId="4" pageNumber="117" rowspan-3="1">
<th id="76EC5DDCFF05B98BD99F29D32275FD02" box="[166,591,739,764]" gridcol="0" gridrow="10" pageId="4" pageNumber="117">Gill rakers</th>
<td id="76EC5DDCFF05B98BDBF529D32457FD02" box="[716,1133,739,764]" colspan="2" colspanRight="1" gridcol="1" gridrow="10" pageId="4" pageNumber="117">3+6 (counted on a non-type specimen)</td>
</tr>
<tr id="353D34A0FF05B98BD99F283B2562FCDA" box="[166,1368,779,804]" gridrow="11" pageId="4" pageNumber="117" rowspan-3="1">
<th id="76EC5DDCFF05B98BD99F283B2275FCDA" box="[166,591,779,804]" gridcol="0" gridrow="11" pageId="4" pageNumber="117">Vertebrae</th>
<td id="76EC5DDCFF05B98BDBF5283B2457FCDA" box="[716,1133,779,804]" colspan="2" colspanRight="1" gridcol="1" gridrow="11" pageId="4" pageNumber="117">13+15 (counted on a non-type specimen)</td>
</tr>
<tr id="353D34A0FF05B98BD99F28032562FCB2" box="[166,1368,819,844]" gridrow="12" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F28032275FCB2" box="[166,591,819,844]" gridcol="0" gridrow="12" pageId="4" pageNumber="117">Total Length</th>
<td id="76EC5DDCFF05B98BDBF528032357FCB2" box="[716,877,819,844]" gridcol="1" gridrow="12" pageId="4" pageNumber="117">70.6</td>
<td id="76EC5DDCFF05B98BDAFB28032457FCB2" box="[962,1133,819,844]" gridcol="2" gridrow="12" pageId="4" pageNumber="117">69.8</td>
<td id="76EC5DDCFF05B98BDD8128032562FCB2" box="[1208,1368,819,844]" gridcol="3" gridrow="12" pageId="4" pageNumber="117">68.8</td>
</tr>
<tr id="353D34A0FF05B98BD99F286C2562FC8A" box="[166,1368,860,884]" gridrow="13" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F286C2275FC8A" box="[166,591,860,884]" gridcol="0" gridrow="13" pageId="4" pageNumber="117">Standard Length</th>
<td id="76EC5DDCFF05B98BDBF5286C2357FC8A" box="[716,877,860,884]" gridcol="1" gridrow="13" pageId="4" pageNumber="117">60.5</td>
<td id="76EC5DDCFF05B98BDAFB286C2457FC8A" box="[962,1133,860,884]" gridcol="2" gridrow="13" pageId="4" pageNumber="117">54.9</td>
<td id="76EC5DDCFF05B98BDD81286C2562FC8A" box="[1208,1368,860,884]" gridcol="3" gridrow="13" pageId="4" pageNumber="117">57.2</td>
</tr>
<tr id="353D34A0FF05B98BD99F28B42562FC63" box="[166,1368,900,925]" gridrow="14" pageId="4" pageNumber="117" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76EC5DDCFF05B98BD99F28B42275FC63" box="[166,591,900,925]" gridcol="0" gridrow="14" pageId="4" pageNumber="117">Percentages of standard length (%)</th>
</tr>
<tr id="353D34A0FF05B98BD99F289C2562FC3A" box="[166,1368,940,964]" gridrow="15" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F289C2275FC3A" box="[166,591,940,964]" gridcol="0" gridrow="15" pageId="4" pageNumber="117">Head length</th>
<td id="76EC5DDCFF05B98BDBF5289C2357FC3A" box="[716,877,940,964]" gridcol="1" gridrow="15" pageId="4" pageNumber="117">36.7</td>
<td id="76EC5DDCFF05B98BDAFB289C2457FC3A" box="[962,1133,940,964]" gridcol="2" gridrow="15" pageId="4" pageNumber="117">37.7</td>
<td id="76EC5DDCFF05B98BDD81289C2562FC3A" box="[1208,1368,940,964]" gridcol="3" gridrow="15" pageId="4" pageNumber="117">38.1</td>
</tr>
<tr id="353D34A0FF05B98BD99F28E42562FC13" box="[166,1368,980,1005]" gridrow="16" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F28E42275FC13" box="[166,591,980,1005]" gridcol="0" gridrow="16" pageId="4" pageNumber="117">Predorsal length</th>
<td id="76EC5DDCFF05B98BDBF528E42357FC13" box="[716,877,980,1005]" gridcol="1" gridrow="16" pageId="4" pageNumber="117">43.5</td>
<td id="76EC5DDCFF05B98BDAFB28E42457FC13" box="[962,1133,980,1005]" gridcol="2" gridrow="16" pageId="4" pageNumber="117">43.7</td>
<td id="76EC5DDCFF05B98BDD8128E42562FC13" box="[1208,1368,980,1005]" gridcol="3" gridrow="16" pageId="4" pageNumber="117">44.8</td>
</tr>
<tr id="353D34A0FF05B98BD99F28CD2562FBEB" box="[166,1368,1021,1045]" gridrow="17" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F28CD2275FBEB" box="[166,591,1021,1045]" gridcol="0" gridrow="17" pageId="4" pageNumber="117">Snout – second Dorsal fin</th>
<td id="76EC5DDCFF05B98BDBF528CD2357FBEB" box="[716,877,1021,1045]" gridcol="1" gridrow="17" pageId="4" pageNumber="117">63.3</td>
<td id="76EC5DDCFF05B98BDAFB28CD2457FBEB" box="[962,1133,1021,1045]" gridcol="2" gridrow="17" pageId="4" pageNumber="117">63.8</td>
<td id="76EC5DDCFF05B98BDD8128CD2562FBEB" box="[1208,1368,1021,1045]" gridcol="3" gridrow="17" pageId="4" pageNumber="117">64.0</td>
</tr>
<tr id="353D34A0FF05B98BD99F2F152562FBC3" box="[166,1368,1061,1085]" gridrow="18" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2F152275FBC3" box="[166,591,1061,1085]" gridcol="0" gridrow="18" pageId="4" pageNumber="117">Snout – anus</th>
<td id="76EC5DDCFF05B98BDBF52F152357FBC3" box="[716,877,1061,1085]" gridcol="1" gridrow="18" pageId="4" pageNumber="117">57.9</td>
<td id="76EC5DDCFF05B98BDAFB2F152457FBC3" box="[962,1133,1061,1085]" gridcol="2" gridrow="18" pageId="4" pageNumber="117">59.0</td>
<td id="76EC5DDCFF05B98BDD812F152562FBC3" box="[1208,1368,1061,1085]" gridcol="3" gridrow="18" pageId="4" pageNumber="117">60.0</td>
</tr>
<tr id="353D34A0FF05B98BD99F2F7D2562FB9B" box="[166,1368,1101,1125]" gridrow="19" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2F7D2275FB9B" box="[166,591,1101,1125]" gridcol="0" gridrow="19" pageId="4" pageNumber="117">Snout – anal fin</th>
<td id="76EC5DDCFF05B98BDBF52F7D2357FB9B" box="[716,877,1101,1125]" gridcol="1" gridrow="19" pageId="4" pageNumber="117">63.6</td>
<td id="76EC5DDCFF05B98BDAFB2F7D2457FB9B" box="[962,1133,1101,1125]" gridcol="2" gridrow="19" pageId="4" pageNumber="117">66.3</td>
<td id="76EC5DDCFF05B98BDD812F7D2562FB9B" box="[1208,1368,1101,1125]" gridcol="3" gridrow="19" pageId="4" pageNumber="117">64.3</td>
</tr>
<tr id="353D34A0FF05B98BD99F2F452562FB70" box="[166,1368,1141,1166]" gridrow="20" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2F452275FB70" box="[166,591,1141,1166]" gridcol="0" gridrow="20" pageId="4" pageNumber="117">Snout – Pelvic fins</th>
<td id="76EC5DDCFF05B98BDBF52F452357FB70" box="[716,877,1141,1166]" gridcol="1" gridrow="20" pageId="4" pageNumber="117">35.5</td>
<td id="76EC5DDCFF05B98BDAFB2F452457FB70" box="[962,1133,1141,1166]" gridcol="2" gridrow="20" pageId="4" pageNumber="117">37.3</td>
<td id="76EC5DDCFF05B98BDD812F452562FB70" box="[1208,1368,1141,1166]" gridcol="3" gridrow="20" pageId="4" pageNumber="117">36.2</td>
</tr>
<tr id="353D34A0FF05B98BD99F2FAE2562FB48" box="[166,1368,1182,1206]" gridrow="21" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2FAE2275FB48" box="[166,591,1182,1206]" gridcol="0" gridrow="21" pageId="4" pageNumber="117">Length of caudal peduncle</th>
<td id="76EC5DDCFF05B98BDBF52FAE2357FB48" box="[716,877,1182,1206]" gridcol="1" gridrow="21" pageId="4" pageNumber="117">28.4</td>
<td id="76EC5DDCFF05B98BDAFB2FAE2457FB48" box="[962,1133,1182,1206]" gridcol="2" gridrow="21" pageId="4" pageNumber="117">28.4</td>
<td id="76EC5DDCFF05B98BDD812FAE2562FB48" box="[1208,1368,1182,1206]" gridcol="3" gridrow="21" pageId="4" pageNumber="117">27.1</td>
</tr>
<tr id="353D34A0FF05B98BD99F2FF62562FB20" box="[166,1368,1222,1246]" gridrow="22" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2FF62275FB20" box="[166,591,1222,1246]" gridcol="0" gridrow="22" pageId="4" pageNumber="117">Depth of caudal peduncle</th>
<td id="76EC5DDCFF05B98BDBF52FF62357FB20" box="[716,877,1222,1246]" gridcol="1" gridrow="22" pageId="4" pageNumber="117">12.2</td>
<td id="76EC5DDCFF05B98BDAFB2FF62457FB20" box="[962,1133,1222,1246]" gridcol="2" gridrow="22" pageId="4" pageNumber="117">12.4</td>
<td id="76EC5DDCFF05B98BDD812FF62562FB20" box="[1208,1368,1222,1246]" gridcol="3" gridrow="22" pageId="4" pageNumber="117">12.1</td>
</tr>
<tr id="353D34A0FF05B98BD99F2FDE2562FAF8" box="[166,1368,1262,1286]" gridrow="23" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2FDE2275FAF8" box="[166,591,1262,1286]" gridcol="0" gridrow="23" pageId="4" pageNumber="117">Body depth at first dorsal fin origin</th>
<td id="76EC5DDCFF05B98BDBF52FDE2357FAF8" box="[716,877,1262,1286]" gridcol="1" gridrow="23" pageId="4" pageNumber="117">20.8</td>
<td id="76EC5DDCFF05B98BDAFB2FDE2457FAF8" box="[962,1133,1262,1286]" gridcol="2" gridrow="23" pageId="4" pageNumber="117">22.4</td>
<td id="76EC5DDCFF05B98BDD812FDE2562FAF8" box="[1208,1368,1262,1286]" gridcol="3" gridrow="23" pageId="4" pageNumber="117">21.9</td>
</tr>
<tr id="353D34A0FF05B98BD99F2E262562FAD1" box="[166,1368,1302,1327]" gridrow="24" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2E262275FAD1" box="[166,591,1302,1327]" gridcol="0" gridrow="24" pageId="4" pageNumber="117">Body width at anal fin origin</th>
<td id="76EC5DDCFF05B98BDBF52E262357FAD1" box="[716,877,1302,1327]" gridcol="1" gridrow="24" pageId="4" pageNumber="117">12.1</td>
<td id="76EC5DDCFF05B98BDAFB2E262457FAD1" box="[962,1133,1302,1327]" gridcol="2" gridrow="24" pageId="4" pageNumber="117">11.3</td>
<td id="76EC5DDCFF05B98BDD812E262562FAD1" box="[1208,1368,1302,1327]" gridcol="3" gridrow="24" pageId="4" pageNumber="117">11.9</td>
</tr>
<tr id="353D34A0FF05B98BD99F2E0F2562FAA9" box="[166,1368,1343,1367]" gridrow="25" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2E0F2275FAA9" box="[166,591,1343,1367]" gridcol="0" gridrow="25" pageId="4" pageNumber="117">Length of first dorsal fin base</th>
<td id="76EC5DDCFF05B98BDBF52E0F2357FAA9" box="[716,877,1343,1367]" gridcol="1" gridrow="25" pageId="4" pageNumber="117">16.7</td>
<td id="76EC5DDCFF05B98BDAFB2E0F2457FAA9" box="[962,1133,1343,1367]" gridcol="2" gridrow="25" pageId="4" pageNumber="117">13.3</td>
<td id="76EC5DDCFF05B98BDD812E0F2562FAA9" box="[1208,1368,1343,1367]" gridcol="3" gridrow="25" pageId="4" pageNumber="117">17.5</td>
</tr>
<tr id="353D34A0FF05B98BD99F2E572562FA81" box="[166,1368,1383,1407]" gridrow="26" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2E572275FA81" box="[166,591,1383,1407]" gridcol="0" gridrow="26" pageId="4" pageNumber="117">Length of second dorsal fin base</th>
<td id="76EC5DDCFF05B98BDBF52E572357FA81" box="[716,877,1383,1407]" gridcol="1" gridrow="26" pageId="4" pageNumber="117">17.7</td>
<td id="76EC5DDCFF05B98BDAFB2E572457FA81" box="[962,1133,1383,1407]" gridcol="2" gridrow="26" pageId="4" pageNumber="117">21.7</td>
<td id="76EC5DDCFF05B98BDD812E572562FA81" box="[1208,1368,1383,1407]" gridcol="3" gridrow="26" pageId="4" pageNumber="117">17.1</td>
</tr>
<tr id="353D34A0FF05B98BD99F2EBF2562FA59" box="[166,1368,1423,1447]" gridrow="27" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2EBF2275FA59" box="[166,591,1423,1447]" gridcol="0" gridrow="27" pageId="4" pageNumber="117">Length of anal fin base</th>
<td id="76EC5DDCFF05B98BDBF52EBF2357FA59" box="[716,877,1423,1447]" gridcol="1" gridrow="27" pageId="4" pageNumber="117">14.9</td>
<td id="76EC5DDCFF05B98BDAFB2EBF2457FA59" box="[962,1133,1423,1447]" gridcol="2" gridrow="27" pageId="4" pageNumber="117">16.6</td>
<td id="76EC5DDCFF05B98BDD812EBF2562FA59" box="[1208,1368,1423,1447]" gridcol="3" gridrow="27" pageId="4" pageNumber="117">12.1</td>
</tr>
<tr id="353D34A0FF05B98BD99F2E872562FA2E" box="[166,1368,1463,1488]" gridrow="28" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2E872275FA2E" box="[166,591,1463,1488]" gridcol="0" gridrow="28" pageId="4" pageNumber="117">Length of caudal fin base</th>
<td id="76EC5DDCFF05B98BDBF52E872357FA2E" box="[716,877,1463,1488]" gridcol="1" gridrow="28" pageId="4" pageNumber="117">13.4</td>
<td id="76EC5DDCFF05B98BDAFB2E872457FA2E" box="[962,1133,1463,1488]" gridcol="2" gridrow="28" pageId="4" pageNumber="117">14.0</td>
<td id="76EC5DDCFF05B98BDD812E872562FA2E" box="[1208,1368,1463,1488]" gridcol="3" gridrow="28" pageId="4" pageNumber="117">12.8</td>
</tr>
<tr id="353D34A0FF05B98BD99F2ED02562FA06" box="[166,1368,1504,1528]" gridrow="29" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2ED02275FA06" box="[166,591,1504,1528]" gridcol="0" gridrow="29" pageId="4" pageNumber="117">Length of pectoral fin</th>
<td id="76EC5DDCFF05B98BDBF52ED02357FA06" box="[716,877,1504,1528]" gridcol="1" gridrow="29" pageId="4" pageNumber="117">18.3</td>
<td id="76EC5DDCFF05B98BDAFB2ED02457FA06" box="[962,1133,1504,1528]" gridcol="2" gridrow="29" pageId="4" pageNumber="117">17.9</td>
<td id="76EC5DDCFF05B98BDD812ED02562FA06" box="[1208,1368,1504,1528]" gridcol="3" gridrow="29" pageId="4" pageNumber="117">19.4</td>
</tr>
<tr id="353D34A0FF05B98BD99F2D382562F9DE" box="[166,1368,1544,1568]" gridrow="30" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2D382275F9DE" box="[166,591,1544,1568]" gridcol="0" gridrow="30" pageId="4" pageNumber="117">Length of pelvic fin</th>
<td id="76EC5DDCFF05B98BDBF52D382357F9DE" box="[716,877,1544,1568]" gridcol="1" gridrow="30" pageId="4" pageNumber="117">14.5</td>
<td id="76EC5DDCFF05B98BDAFB2D382457F9DE" box="[962,1133,1544,1568]" gridcol="2" gridrow="30" pageId="4" pageNumber="117">16.8</td>
<td id="76EC5DDCFF05B98BDD812D382562F9DE" box="[1208,1368,1544,1568]" gridcol="3" gridrow="30" pageId="4" pageNumber="117">15.9</td>
</tr>
<tr id="353D34A0FF05B98BD99F2D002562F9B7" box="[166,1368,1584,1609]" gridrow="31" pageId="4" pageNumber="117" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76EC5DDCFF05B98BD99F2D002275F9B7" box="[166,591,1584,1609]" gridcol="0" gridrow="31" pageId="4" pageNumber="117">Percentage of head length (%)</th>
</tr>
<tr id="353D34A0FF05B98BD99F2D682562F98F" box="[166,1368,1624,1649]" gridrow="32" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2D682275F98F" box="[166,591,1624,1649]" gridcol="0" gridrow="32" pageId="4" pageNumber="117">Snout length</th>
<td id="76EC5DDCFF05B98BDBF52D682357F98F" box="[716,877,1624,1649]" gridcol="1" gridrow="32" pageId="4" pageNumber="117">22.5</td>
<td id="76EC5DDCFF05B98BDAFB2D682457F98F" box="[962,1133,1624,1649]" gridcol="2" gridrow="32" pageId="4" pageNumber="117">24.2</td>
<td id="76EC5DDCFF05B98BDD812D682562F98F" box="[1208,1368,1624,1649]" gridcol="3" gridrow="32" pageId="4" pageNumber="117">20.2</td>
</tr>
<tr id="353D34A0FF05B98BD99F2DB02562F967" box="[166,1368,1664,1689]" gridrow="33" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2DB02275F967" box="[166,591,1664,1689]" gridcol="0" gridrow="33" pageId="4" pageNumber="117">Postorbital length</th>
<td id="76EC5DDCFF05B98BDBF52DB02357F967" box="[716,877,1664,1689]" gridcol="1" gridrow="33" pageId="4" pageNumber="117">57.2</td>
<td id="76EC5DDCFF05B98BDAFB2DB02457F967" box="[962,1133,1664,1689]" gridcol="2" gridrow="33" pageId="4" pageNumber="117">59.9</td>
<td id="76EC5DDCFF05B98BDD812DB02562F967" box="[1208,1368,1664,1689]" gridcol="3" gridrow="33" pageId="4" pageNumber="117">59.6</td>
</tr>
<tr id="353D34A0FF05B98BD99F2D992562F91B" box="[166,1368,1705,1765]" gridrow="34" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2D992275F91B" box="[166,591,1705,1765]" gridcol="0" gridrow="34" pageId="4" pageNumber="117">Head depth on cheek (at the middle of the preopercular)</th>
<td id="76EC5DDCFF05B98BDBF52D992357F91B" box="[716,877,1705,1765]" gridcol="1" gridrow="34" pageId="4" pageNumber="117">46.4</td>
<td id="76EC5DDCFF05B98BDAFB2D992457F91B" box="[962,1133,1705,1765]" gridcol="2" gridrow="34" pageId="4" pageNumber="117">49.8</td>
<td id="76EC5DDCFF05B98BDD812D992562F91B" box="[1208,1368,1705,1765]" gridcol="3" gridrow="34" pageId="4" pageNumber="117">45.4</td>
</tr>
<tr id="353D34A0FF05B98BD99F2DC52562F8F3" box="[166,1368,1781,1805]" gridrow="35" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2DC52275F8F3" box="[166,591,1781,1805]" gridcol="0" gridrow="35" pageId="4" pageNumber="117">Maximum head depth</th>
<td id="76EC5DDCFF05B98BDBF52DC52357F8F3" box="[716,877,1781,1805]" gridcol="1" gridrow="35" pageId="4" pageNumber="117">56.3</td>
<td id="76EC5DDCFF05B98BDAFB2DC52457F8F3" box="[962,1133,1781,1805]" gridcol="2" gridrow="35" pageId="4" pageNumber="117">54.1</td>
<td id="76EC5DDCFF05B98BDD812DC52562F8F3" box="[1208,1368,1781,1805]" gridcol="3" gridrow="35" pageId="4" pageNumber="117">53.7</td>
</tr>
<tr id="353D34A0FF05B98BD99F2C2D2562F8CB" box="[166,1368,1821,1845]" gridrow="36" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2C2D2275F8CB" box="[166,591,1821,1845]" gridcol="0" gridrow="36" pageId="4" pageNumber="117">Maximum head width</th>
<td id="76EC5DDCFF05B98BDBF52C2D2357F8CB" box="[716,877,1821,1845]" gridcol="1" gridrow="36" pageId="4" pageNumber="117">55.0</td>
<td id="76EC5DDCFF05B98BDAFB2C2D2457F8CB" box="[962,1133,1821,1845]" gridcol="2" gridrow="36" pageId="4" pageNumber="117">56.5</td>
<td id="76EC5DDCFF05B98BDD812C2D2562F8CB" box="[1208,1368,1821,1845]" gridcol="3" gridrow="36" pageId="4" pageNumber="117">54.6</td>
</tr>
<tr id="353D34A0FF05B98BD99F2C752562F8A0" box="[166,1368,1861,1886]" gridrow="37" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2C752275F8A0" box="[166,591,1861,1886]" gridcol="0" gridrow="37" pageId="4" pageNumber="117">Interorbital width</th>
<td id="76EC5DDCFF05B98BDBF52C752357F8A0" box="[716,877,1861,1886]" gridcol="1" gridrow="37" pageId="4" pageNumber="117">28.8</td>
<td id="76EC5DDCFF05B98BDAFB2C752457F8A0" box="[962,1133,1861,1886]" gridcol="2" gridrow="37" pageId="4" pageNumber="117">29.5</td>
<td id="76EC5DDCFF05B98BDD812C752562F8A0" box="[1208,1368,1861,1886]" gridcol="3" gridrow="37" pageId="4" pageNumber="117">25.2</td>
</tr>
<tr id="353D34A0FF05B98BD99F2C5E2562F878" box="[166,1368,1902,1926]" gridrow="38" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2C5E2275F878" box="[166,591,1902,1926]" gridcol="0" gridrow="38" pageId="4" pageNumber="117">Eye diameter</th>
<td id="76EC5DDCFF05B98BDBF52C5E2357F878" box="[716,877,1902,1926]" gridcol="1" gridrow="38" pageId="4" pageNumber="117">18.5</td>
<td id="76EC5DDCFF05B98BDAFB2C5E2457F878" box="[962,1133,1902,1926]" gridcol="2" gridrow="38" pageId="4" pageNumber="117">17.9</td>
<td id="76EC5DDCFF05B98BDD812C5E2562F878" box="[1208,1368,1902,1926]" gridcol="3" gridrow="38" pageId="4" pageNumber="117">17.0</td>
</tr>
<tr id="353D34A0FF05B98BD99F2CA62562F850" box="[166,1368,1942,1966]" gridrow="39" pageId="4" pageNumber="117">
<th id="76EC5DDCFF05B98BD99F2CA62275F850" box="[166,591,1942,1966]" gridcol="0" gridrow="39" pageId="4" pageNumber="117">Lower jaw length</th>
<td id="76EC5DDCFF05B98BDBF52CA62357F850" box="[716,877,1942,1966]" gridcol="1" gridrow="39" pageId="4" pageNumber="117">29.7</td>
<td id="76EC5DDCFF05B98BDAFB2CA62457F850" box="[962,1133,1942,1966]" gridcol="2" gridrow="39" pageId="4" pageNumber="117">36.7</td>
<td id="76EC5DDCFF05B98BDD812CA62562F850" box="[1208,1368,1942,1966]" gridcol="3" gridrow="39" pageId="4" pageNumber="117">34.4</td>
</tr>
</table>
</paragraph>
<caption id="DF72666AFF044671D9AE28382342FCDE" ID-DOI="http://doi.org/10.5281/zenodo.6531374" ID-Zenodo-Dep="6531374" box="[151,888,776,800]" httpUri="https://zenodo.org/record/6531374/files/figure.png" pageId="5" pageNumber="118" startId="5.[151,250,776,800]" targetBox="[151,1437,181,752]" targetPageId="5">
<paragraph id="8BB236E2FF044671D9AE28382342FCDE" blockId="5.[151,888,776,800]" box="[151,888,776,800]" pageId="5" pageNumber="118">
<emphasis id="B979EAF0FF044671D9AE28382129FCDE" bold="true" box="[151,275,776,800]" pageId="5" pageNumber="118">FIGURE 2.</emphasis>
<taxonomicName id="4C0D4D61FF044671D820283821E1FCDE" authority="Zhou &amp; He &amp; Wang &amp; Li &amp; Li, 2022" authorityName="Zhou &amp; He &amp; Wang &amp; Li &amp; Li" authorityYear="2022" box="[281,475,776,800]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="118" phylum="Chordata" rank="species" species="lani" status="sp. nov.">
<emphasis id="B979EAF0FF044671D820283821E1FCDE" box="[281,475,776,800]" italics="true" pageId="5" pageNumber="118">Neodontobutis lani</emphasis>
</taxonomicName>
<emphasis id="B979EAF0FF044671D8D828382208FCDE" bold="true" box="[481,562,776,800]" pageId="5" pageNumber="118">
<taxonomicNameLabel id="A24A578BFF044671D8D828382208FCDE" box="[481,562,776,800]" pageId="5" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, a live uncatalogued individual.
</paragraph>
</caption>
<paragraph id="8BB236E2FF044671D9FE287E24E1FC2A" blockId="5.[151,1437,846,981]" pageId="5" pageNumber="118">
Coloration: Head and body with dark brown mottling in live individuals. Several dark bars radiate from orbitals. Dark stripes on each side of body. A shiny, greenish blotch on upper side of operculum. A large dark blotch on base of caudal fin. Dorsal fins, anal fin and caudal fin mottled (
<figureCitation id="13362A67FF044671DA1B28A623BFFC4E" box="[802,901,918,944]" captionStart="FIGURE 2" captionStartId="5.[151,250,776,800]" captionTargetBox="[151,1437,181,752]" captionTargetId="figure-18@5.[151,1437,181,752]" captionTargetPageId="5" captionText="FIGURE 2. Neodontobutis lani sp. nov., a live uncatalogued individual." figureDoi="http://doi.org/10.5281/zenodo.6531374" httpUri="https://zenodo.org/record/6531374/files/figure.png" pageId="5" pageNumber="118">Figure 2</figureCitation>
). Preserved specimens light brown, with stripes and irregular darker blotches on each side of body. Abdominal pale, slightly yellowish (
<figureCitation id="13362A67FF044671DD52288A24F7FC2A" box="[1131,1229,954,980]" captionStart="FIGURE 3" captionStartId="5.[151,250,1866,1890]" captionTargetBox="[359,1301,1022,1842]" captionTargetId="figure-105@5.[285,1301,1004,1842]" captionTargetPageId="5" captionText="FIGURE 3. Neodontobutis lani sp. nov., SOU1801007-1, holotype, 62.4 mm standard length, Longzhou Town, Guangxi. A. dorsal, B. lateral, C. ventral views." figureDoi="http://doi.org/10.5281/zenodo.6531376" httpUri="https://zenodo.org/record/6531376/files/figure.png" pageId="5" pageNumber="118">Figure 3</figureCitation>
).
</paragraph>
<caption id="DF72666AFF044671D9AE2C7A21C2F878" ID-DOI="http://doi.org/10.5281/zenodo.6531376" ID-Zenodo-Dep="6531376" httpUri="https://zenodo.org/record/6531376/files/figure.png" pageId="5" pageNumber="118" startId="5.[151,250,1866,1890]" targetBox="[359,1301,1022,1842]" targetPageId="5">
<paragraph id="8BB236E2FF044671D9AE2C7A21C2F878" blockId="5.[151,1437,1866,1927]" pageId="5" pageNumber="118">
<emphasis id="B979EAF0FF044671D9AE2C7A212EF89C" bold="true" box="[151,276,1866,1890]" pageId="5" pageNumber="118">FIGURE 3.</emphasis>
<taxonomicName id="4C0D4D61FF044671D8252C7A21E5F89C" authority="Zhou &amp; He &amp; Wang &amp; Li &amp; Li, 2022" authorityName="Zhou &amp; He &amp; Wang &amp; Li &amp; Li" authorityYear="2022" box="[284,479,1866,1890]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="118" phylum="Chordata" rank="species" species="lani" status="sp. nov.">
<emphasis id="B979EAF0FF044671D8252C7A21E5F89C" box="[284,479,1866,1890]" italics="true" pageId="5" pageNumber="118">Neodontobutis lani</emphasis>
</taxonomicName>
<emphasis id="B979EAF0FF044671D8DE2C7B2203F89D" bold="true" box="[487,569,1867,1891]" pageId="5" pageNumber="118">
<taxonomicNameLabel id="A24A578BFF044671D8DE2C7B2203F89D" box="[487,569,1867,1891]" pageId="5" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, SOU1801007-1, holotype, 62.4 mm standard length, Longzhou Town, Guangxi. A. dorsal, B. lateral, C. ventral views.
</paragraph>
</caption>
<caption id="DF72666AFF074672D9AE2FEB22E4FAE6" ID-DOI="http://doi.org/10.5281/zenodo.6531378" ID-Zenodo-Dep="6531378" httpUri="https://zenodo.org/record/6531378/files/figure.png" pageId="6" pageNumber="119" startId="6.[151,250,1243,1267]" targetBox="[163,1420,191,1210]" targetPageId="6">
<paragraph id="8BB236E2FF074672D9AE2FEB22E4FAE6" blockId="6.[151,1436,1243,1304]" pageId="6" pageNumber="119">
<emphasis id="B979EAF0FF074672D9AE2FEB212FFB0D" bold="true" box="[151,277,1243,1267]" pageId="6" pageNumber="119">FIGURE 4.</emphasis>
Patterns of main cephalic sensory papilla lines (row of spots) of 
<taxonomicName id="4C0D4D61FF074672DAFE2FEC24B0FB0A" authority="Zhou &amp; He &amp; Wang &amp; Li &amp; Li, 2022" authorityName="Zhou &amp; He &amp; Wang &amp; Li &amp; Li" authorityYear="2022" box="[967,1162,1244,1268]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="lani" status="sp. nov.">
<emphasis id="B979EAF0FF074672DAFE2FEC24B0FB0A" box="[967,1162,1244,1268]" italics="true" pageId="6" pageNumber="119">Neodontobutis lani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A578BFF074672DDAD2FEC24DFFB0D" box="[1172,1253,1244,1267]" pageId="6" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
(SOU1801007-1, holotype) from lateral (a), dorsal (b) and ventral (c) view.
</paragraph>
</caption>
<caption id="DF72666AFF074672D9AE2CA724ABF82A" ID-DOI="http://doi.org/10.5281/zenodo.6531380" ID-Zenodo-Dep="6531380" httpUri="https://zenodo.org/record/6531380/files/figure.png" pageId="6" pageNumber="119" startId="6.[151,250,1943,1967]" targetBox="[180,1408,1328,1919]" targetPageId="6">
<paragraph id="8BB236E2FF074672D9AE2CA724ABF82A" blockId="6.[151,1436,1943,2004]" pageId="6" pageNumber="119">
<emphasis id="B979EAF0FF074672D9AE2CA72128F84E" bold="true" box="[151,274,1943,1968]" pageId="6" pageNumber="119">FIGURE 5.</emphasis>
Posterior edge of scale on dorsal part of 
<taxonomicName id="4C0D4D61FF074672DB912CA82399F851" authority="SOU" authorityName="SOU" box="[680,931,1943,1968]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF074672DB912CA82352F84E" box="[680,872,1944,1968]" italics="true" pageId="6" pageNumber="119">Neodontobutis lani</emphasis>
SOU
</taxonomicName>
1801007-5, paratype (a) and 
<taxonomicName id="4C0D4D61FF074672DDFB2CA82562F84E" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[1218,1368,1944,1968]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF074672DDFB2CA82562F84E" box="[1218,1368,1944,1968]" italics="true" pageId="6" pageNumber="119">N. hainanensis</emphasis>
</taxonomicName>
25901 (b), ct, ctenii; tr, transforming ctenii. Sensory papilla on lower jaw of 
<taxonomicName id="4C0D4D61FF074672DA612C8C23A7F82A" box="[856,925,1980,2004]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF074672DA612C8C23A7F82A" box="[856,925,1980,2004]" italics="true" pageId="6" pageNumber="119">N. lani</emphasis>
</taxonomicName>
(c), p, sensory papillae.
</paragraph>
</caption>
<paragraph id="8BB236E2FF064673D9FE2BA62178FD7B" blockId="7.[151,1437,150,969]" pageId="7" pageNumber="120">
<emphasis id="B979EAF0FF064673D9FE2BA621F0FF4E" bold="true" box="[199,458,150,176]" pageId="7" pageNumber="120">Phylogenetic analysis.</emphasis>
Four individuals of 
<taxonomicName id="4C0D4D61FF064673DB9F2BA72341FF4F" box="[678,891,151,177]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DB9F2BA72341FF4F" box="[678,891,151,177]" italics="true" pageId="7" pageNumber="120">Neodontobutis lani</emphasis>
</taxonomicName>
(25912, 25913, 25914, 25916) and representative species of all genera in family 
<taxonomicName id="4C0D4D61FF064673D8C62B8B2292FF2B" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[511,680,187,213]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
, except for the genus 
<emphasis id="B979EAF0FF064673DA932B8B240AFF2B" box="[938,1072,187,213]" italics="true" pageId="7" pageNumber="120">Terateleoris</emphasis>
, were used in the concatenated nuclear gene tree reconstruction. 
<taxonomicName id="4C0D4D61FF064673DB362BEF23F8FF04" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[527,962,223,250]" class="Actinopterygii" family="Rhyacichthyidae" genus="Rhyacichthys" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="aspro">
<emphasis id="B979EAF0FF064673DB362BEF22D6FF06" box="[527,748,223,249]" italics="true" pageId="7" pageNumber="120">Rhyacichthys aspro</emphasis>
(Rhyacichthyidae)
</taxonomicName>
was used as the outgroup according to Li (H. 
<bibRefCitation id="EF9C4B13FF064673D9FB2A332158FEE3" author="Li, H. &amp; He, Y. &amp; Jiang, J. &amp; Liu, Z. &amp; Li, C." box="[194,354,259,285]" pageId="7" pageNumber="120" pagination="1 - 11" refId="ref5860" refString="Li, H., He, Y., Jiang, J., Liu, Z. &amp; Li, C. (2018) Molecular systematics and phylogenetic analysis of the Asian endemic freshwater sleepers (Gobiiformes: Odontobutidae). Molecular Phylogenetics and Evolution, 121, 1 - 11. https: // doi. org / 10.1016 / j. ympev. 2017.12.026" type="journal article" year="2018">
Li 
<emphasis id="B979EAF0FF064673D9D82A332122FEE3" box="[225,280,259,285]" italics="true" pageId="7" pageNumber="120">et al.</emphasis>
, 2018
</bibRefCitation>
). For each sample, 
<geoCoordinate id="EE395025FF064673DB0F2A33224FFEE3" box="[566,629,259,285]" degrees="1,027" direction="north" orientation="latitude" pageId="7" pageNumber="120" precision="55" value="1.027">1,027</geoCoordinate>
<geoCoordinate id="EE395025FF064673DB4D2A3322FBFEE3" box="[628,705,259,285]" degrees="1,688" direction="west" orientation="longitude" pageId="7" pageNumber="120" precision="55" value="-1.688">–1,688</geoCoordinate>
loci from the 4,434 targeted ones were obtained after assembling. After excluding loci with more than 50% missing data, 2,021 loci were kept for phylogenetic analysis, the length of concatenated alignment was 336,090 bp (electronic appendix). The reconstructed maximum likelihood tree is shown in 
<figureCitation id="13362A67FF064673D83A2A5F2159FE77" box="[259,355,367,393]" captionStart="FIGURE 6" captionStartId="8.[151,250,1804,1828]" captionTargetBox="[159,1434,682,1769]" captionTargetId="figure-255@8.[151,1436,669,1781]" captionTargetPageId="8" captionText="FIGURE 6. Concatenated maximum likelihood tree of Neodontobutis lani, and species representing all genus of the Odontobutidae, except for Terateleotris, based on 2,076 single-copy nuclear loci. *SOU1801007-3 is a paratype." figureDoi="http://doi.org/10.5281/zenodo.6531382" httpUri="https://zenodo.org/record/6531382/files/figure.png" pageId="7" pageNumber="120">Figure 6</figureCitation>
. All genera of 
<taxonomicName id="4C0D4D61FF064673DB3D2A5F2290FE77" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[516,682,367,393]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
involved in this research formed a monophyletic group. The genera 
<taxonomicName id="4C0D4D61FF064673D9AE2AA32100FE53" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[151,314,403,429]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673D9AE2AA32100FE53" box="[151,314,403,429]" italics="true" pageId="7" pageNumber="120">Neodontobutis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0D4D61FF064673D8732AA321FBFE52" authorityName="Dybowski" authorityYear="1877" box="[330,449,403,428]" class="Actinopterygii" family="Odontobutidae" genus="Perccottus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673D8732AA321FBFE52" box="[330,449,403,428]" italics="true" pageId="7" pageNumber="120">Perccottus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D61FF064673D8C02AA322B9FE53" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[505,643,403,429]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673D8C02AA322B9FE53" box="[505,643,403,429]" italics="true" pageId="7" pageNumber="120">Odontobutis</emphasis>
</taxonomicName>
were all monophyletic. 
<taxonomicName id="4C0D4D61FF064673DAA12AA3244AFE53" box="[920,1136,403,429]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DAA12AA3244AFE53" box="[920,1136,403,429]" italics="true" pageId="7" pageNumber="120">Neodontobutis lani</emphasis>
</taxonomicName>
was the sister group of 
<taxonomicName id="4C0D4D61FF064673DCBA2AA32127FE2F" baseAuthorityName="Chen" baseAuthorityYear="1985" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF064673DCBA2AA32127FE2F" italics="true" pageId="7" pageNumber="120">N. hainanensis</emphasis>
</taxonomicName>
and then formed a clade with 
<taxonomicName id="4C0D4D61FF064673DBBC2A87237FFE2F" authorityName="Dybowski" authorityYear="1877" box="[645,837,439,465]" class="Actinopterygii" family="Odontobutidae" genus="Perccottus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="glenii">
<emphasis id="B979EAF0FF064673DBBC2A87237FFE2F" box="[645,837,439,465]" italics="true" pageId="7" pageNumber="120">Perccottus glenii</emphasis>
</taxonomicName>
. Aligned 
<emphasis id="B979EAF0FF064673DA802A8723D3FE2F" box="[953,1001,439,465]" italics="true" pageId="7" pageNumber="120">COI</emphasis>
sequences can be found in electronic appendix. The 
<emphasis id="B979EAF0FF064673D87D2AEB214EFE0B" box="[324,372,475,501]" italics="true" pageId="7" pageNumber="120">COI</emphasis>
gene tree which involved more individuals of 
<taxonomicName id="4C0D4D61FF064673DAAC2AEB23DFFE0B" box="[917,997,475,501]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DAAC2AEB23DFFE0B" box="[917,997,475,501]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
and more 
<taxonomicName id="4C0D4D61FF064673DD5D2AEB2537FE0B" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[1124,1293,475,501]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF064673DD5D2AEB2537FE0B" box="[1124,1293,475,501]" italics="true" pageId="7" pageNumber="120">N. hainanensis</emphasis>
</taxonomicName>
is shown in 
<figureCitation id="13362A67FF064673D9AE2ACF20C7FDE7" box="[151,253,511,537]" captionStart="FIGURE 7" captionStartId="9.[151,250,1368,1392]" captionTargetBox="[158,1425,191,1335]" captionTargetId="figure-18@9.[151,1436,181,1344]" captionTargetPageId="9" captionText="FIGURE 7. Maximum likelihood tree of Neodontobutis lani, and comparative species of the Odontobutidae, based on partial sequence (~ 1500 bp) of cytochrome c oxidase subunit I (COI) gene. *SOU1801007-1 is the Holotype, SOU1801007-3 and SOU1801007-5 are the paratypes." figureDoi="http://doi.org/10.5281/zenodo.6531384" httpUri="https://zenodo.org/record/6531384/files/figure.png" pageId="7" pageNumber="120">Figure 7</figureCitation>
. Its topological structure is same as the concatenated nuclear gene tree at genus level except for the position of genus 
<taxonomicName id="4C0D4D61FF064673D85C291321E7FDC3" box="[357,477,547,573]" class="Actinopteri" family="Odontobutidae" genus="Microdous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiiformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673D85C291321E7FDC3" box="[357,477,547,573]" italics="true" pageId="7" pageNumber="120">Microdous</emphasis>
</taxonomicName>
. All 
<taxonomicName id="4C0D4D61FF064673DB212913225CFDC3" box="[536,614,547,573]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DB212913225CFDC3" box="[536,614,547,573]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
individuals formed a distinct clade apart from 
<taxonomicName id="4C0D4D61FF064673DDBA29132511FDC3" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[1155,1323,547,573]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF064673DDBA29132511FDC3" box="[1155,1323,547,573]" italics="true" pageId="7" pageNumber="120">N. hainanensis</emphasis>
</taxonomicName>
. Both the nuclear and mitochondrial trees suggest that 
<taxonomicName id="4C0D4D61FF064673DBB4297722E3FD9F" box="[653,729,583,609]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DBB4297722E3FD9F" box="[653,729,583,609]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
is an independent species and should be placed in the genus of 
<taxonomicName id="4C0D4D61FF064673D9AE295B2100FD7B" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[151,314,619,645]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673D9AE295B2100FD7B" box="[151,314,619,645]" italics="true" pageId="7" pageNumber="120">Neodontobutis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176569FF064673D9FE29BE2328FD0F" pageId="7" pageNumber="120" type="distribution">
<paragraph id="8BB236E2FF064673D9FE29BE2328FD0F" blockId="7.[151,1437,150,969]" pageId="7" pageNumber="120">
<emphasis id="B979EAF0FF064673D9FE29BE215BFD56" bold="true" box="[199,353,654,680]" pageId="7" pageNumber="120">Distribution.</emphasis>
Specimens of 
<taxonomicName id="4C0D4D61FF064673DB2C29BF22CBFD57" box="[533,753,655,681]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DB2C29BF22CBFD57" box="[533,753,655,681]" italics="true" pageId="7" pageNumber="120">Neodontobutis lani</emphasis>
</taxonomicName>
were bought from a farmers’ market in Longzhou Town, 
<collectingRegion id="49C9F800FF064673D9AE298320C0FD33" box="[151,250,691,717]" country="China" name="Guangxi" pageId="7" pageNumber="120">Guangxi</collectingRegion>
, which were collected by a local fisherman from tributaries of the Zuojiang River (
<figureCitation id="13362A67FF064673DDA2298324C3FD33" box="[1179,1273,691,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2003]" captionTargetBox="[157,1436,1458,1945]" captionTargetId="figure-517@2.[151,1436,1458,1952]" captionTargetPageId="2" captionText="FIGURE 1. The sampling location of Neodontobutis lani sp.nov., indicated by the triangle.The inset shows the sampling region within the map of China." figureDoi="http://doi.org/10.5281/zenodo.6531372" httpUri="https://zenodo.org/record/6531372/files/figure.png" pageId="7" pageNumber="120">Figure 1</figureCitation>
). The detailed distribution range of 
<taxonomicName id="4C0D4D61FF064673D8BA29E721F5FD0F" box="[387,463,727,753]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673D8BA29E721F5FD0F" box="[387,463,727,753]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
requires further exploration.
</paragraph>
</subSubSection>
<subSubSection id="C3176569FF064673D9FE29CA2541FC7F" pageId="7" pageNumber="120" type="biology_ecology">
<paragraph id="8BB236E2FF064673D9FE29CA2541FC7F" blockId="7.[151,1437,150,969]" pageId="7" pageNumber="120">
<emphasis id="B979EAF0FF064673D9FE29CA211DFCEA" bold="true" box="[199,295,762,788]" pageId="7" pageNumber="120">Biology.</emphasis>
According to the measurement of specimens collected in this research, maximum size of 
<taxonomicName id="4C0D4D61FF064673DDC029CB20F9FCC7" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DDC029CB20F9FCC7" italics="true" pageId="7" pageNumber="120">Neodontobutis lani</emphasis>
</taxonomicName>
is approximately 
<quantity id="4CF59B07FF064673D8B3282F21D7FCC7" box="[394,493,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="6.24" pageId="7" pageNumber="120" unit="mm" value="62.4">62.4 mm</quantity>
standard length. An unidentified small fish (about 
<quantity id="4CF59B07FF064673DD18282E2452FCC7" box="[1057,1128,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="120" unit="mm" value="25.0">25mm</quantity>
standard length) was found in the mouth of a non-type specimen, indicating that 
<taxonomicName id="4C0D4D61FF064673DBDB28732314FCA3" box="[738,814,835,861]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DBDB28732314FCA3" box="[738,814,835,861]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
is carnivorous. Ovaries with mature eggs were found in a non-type specimen (
<quantity id="4CF59B07FF064673D8B4285721D6FC7F" box="[397,492,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="4.82" pageId="7" pageNumber="120" unit="mm" value="48.2">48.2mm</quantity>
standard length), holds approximately 
<specimenCount id="9D0BFD6BFF064673DA9B28572433FC7F" box="[930,1033,871,897]" pageId="7" pageNumber="120" type="egg">414 eggs</specimenCount>
, with average diameter of 
<quantity id="4CF59B07FF064673DC0A2857254DFC7F" box="[1331,1399,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="120" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176569FF064673D9FE28BB2134FC37" pageId="7" pageNumber="120" type="etymology">
<paragraph id="8BB236E2FF064673D9FE28BB2134FC37" blockId="7.[151,1437,150,969]" pageId="7" pageNumber="120">
<emphasis id="B979EAF0FF064673D9FE28BB2174FC5B" bold="true" box="[199,334,907,933]" pageId="7" pageNumber="120">Etymology.</emphasis>
The new species is named after Jiahu Lan, who found the specimens. The Chinese name is 
<emphasis id="B979EAF0FF064673DC5D28BB213DFC38" bold="true" pageId="7" pageNumber="120">Ầ氏 NJ沙dzffi</emphasis>
.
</paragraph>
</subSubSection>
<caption id="DF72666AFF064673D9AE28C821DFFBCB" ID-Table-UUID="DF72666AFF064673D9AE28C821DFFBCB" httpUri="http://table.plazi.org/id/DF72666AFF064673D9AE28C821DFFBCB" pageId="7" pageNumber="120" startId="7.[151,239,1016,1040]" targetBox="[166,1410,1093,1583]" targetIsTable="true" targetPageId="7">
<paragraph id="8BB236E2FF064673D9AE28C821DFFBCB" blockId="7.[151,1436,1016,1077]" pageId="7" pageNumber="120">
<emphasis id="B979EAF0FF064673D9AE28C82138FBEE" bold="true" box="[151,258,1016,1040]" pageId="7" pageNumber="120">TABLE 4</emphasis>
. Comparison on external characters of the new species 
<taxonomicName id="4C0D4D61FF064673DA7428C923AFFBEE" box="[845,917,1016,1040]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DA7428C923AFFBEE" box="[845,917,1016,1040]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
to those of the genera 
<taxonomicName id="4C0D4D61FF064673DDBA28C82525FBEE" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1155,1311,1016,1040]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673DDBA28C82525FBEE" box="[1155,1311,1016,1040]" italics="true" pageId="7" pageNumber="120">Neodontobutis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0D4D61FF064673DC1228C925A6FBEE" authorityName="Dybowski" authorityYear="1877" box="[1323,1436,1017,1040]" class="Actinopterygii" family="Odontobutidae" genus="Perccottus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673DC1228C925A6FBEE" box="[1323,1436,1017,1040]" italics="true" pageId="7" pageNumber="120">Perccottus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D61FF064673D9FD2F2C21DBFBCB" authority="(Iwata, 2011)" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[196,481,1052,1077]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673D9FD2F2C2172FBCA" box="[196,328,1052,1076]" italics="true" pageId="7" pageNumber="120">Odontobutis</emphasis>
(
<bibRefCitation id="EF9C4B13FF064673D86F2F2D21E3FBCB" author="Iwata, A." box="[342,473,1053,1077]" pageId="7" pageNumber="120" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BB236E2FF064673D99F2F75252BF9D1" pageId="7" pageNumber="120">
<table id="F90DC442FF06B98BD99F2F7525B8F9D1" box="[166,1410,1093,1583]" gridcols="5" gridrows="9" pageId="7" pageNumber="120">
<tr id="353D34A0FF06B98BD99F2F7525B8FBA3" box="[166,1410,1093,1117]" gridrow="0" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2F752238FBA3" box="[166,514,1093,1117]" gridcol="0" gridrow="0" pageId="7" pageNumber="120">Trait</th>
<th id="76EC5DDCFF06B98BDB152F7522C9FBA3" box="[556,755,1093,1117]" gridcol="1" gridrow="0" pageId="7" pageNumber="120">
<emphasis id="B979EAF0FF064673DB152F7522C9FBA3" box="[556,755,1093,1117]" italics="true" pageId="7" pageNumber="120">
<taxonomicName id="4C0D4D61FF064673DB152F7522F9FBA3" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[556,707,1093,1117]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">Neodontobutis</taxonomicName>
. 
<taxonomicName id="4C0D4D61FF064673DBF52F7522C9FBA3" box="[716,755,1093,1117]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">lani</taxonomicName>
</emphasis>
</th>
<th id="76EC5DDCFF06B98BDA252F7523DEFBA3" box="[796,996,1093,1117]" gridcol="2" gridrow="0" pageId="7" pageNumber="120">
<taxonomicName id="4C0D4D61FF064673DA252F75238AFBA3" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[796,944,1093,1117]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673DA252F75238AFBA3" box="[796,944,1093,1117]" italics="true" pageId="7" pageNumber="120">Neodontobutis</emphasis>
</taxonomicName>
</th>
<th id="76EC5DDCFF06B98BDD3C2F752489FBA3" box="[1029,1203,1093,1117]" gridcol="3" gridrow="0" pageId="7" pageNumber="120">
<taxonomicName id="4C0D4D61FF064673DD3C2F75244BFBA2" authorityName="Dybowski" authorityYear="1877" box="[1029,1137,1093,1116]" class="Actinopterygii" family="Odontobutidae" genus="Perccottus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673DD3C2F75244BFBA2" box="[1029,1137,1093,1116]" italics="true" pageId="7" pageNumber="120">Perccottus</emphasis>
</taxonomicName>
</th>
<th id="76EC5DDCFF06B98BDDE92F7525B8FBA3" box="[1232,1410,1093,1117]" gridcol="4" gridrow="0" pageId="7" pageNumber="120">
<taxonomicName id="4C0D4D61FF064673DDE92F752577FBA3" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1232,1357,1093,1117]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673DDE92F752577FBA3" box="[1232,1357,1093,1117]" italics="true" pageId="7" pageNumber="120">Odontobutis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353D34A0FF06B98BD99F2F5D25B8FB57" box="[166,1410,1133,1193]" gridrow="1" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2F5D2238FB57" box="[166,514,1133,1193]" gridcol="0" gridrow="1" pageId="7" pageNumber="120">Barbel-like projection on sensory papilla</th>
<td id="76EC5DDCFF06B98BDB152F5D22C9FB57" box="[556,755,1133,1193]" gridcol="1" gridrow="1" pageId="7" pageNumber="120">present</td>
<td id="76EC5DDCFF06B98BDA252F5D23DEFB57" box="[796,996,1133,1193]" gridcol="2" gridrow="1" pageId="7" pageNumber="120">present</td>
<td id="76EC5DDCFF06B98BDD3C2F5D2489FB57" box="[1029,1203,1133,1193]" gridcol="3" gridrow="1" pageId="7" pageNumber="120">absent</td>
<td id="76EC5DDCFF06B98BDDE92F5D25B8FB57" box="[1232,1410,1133,1193]" gridcol="4" gridrow="1" pageId="7" pageNumber="120">absent</td>
</tr>
<tr id="353D34A0FF06B98BD99F2F8925B8FB0B" box="[166,1410,1209,1269]" gridrow="2" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2F892238FB0B" box="[166,514,1209,1269]" gridcol="0" gridrow="2" pageId="7" pageNumber="120">Sensory papilla distribution on lower jaw</th>
<td id="76EC5DDCFF06B98BDB152F8922C9FB0B" box="[556,755,1209,1269]" gridcol="1" gridrow="2" pageId="7" pageNumber="120">two oblong patches</td>
<td id="76EC5DDCFF06B98BDA252F8923DEFB0B" box="[796,996,1209,1269]" gridcol="2" gridrow="2" pageId="7" pageNumber="120">two single lines</td>
<td id="76EC5DDCFF06B98BDD3C2F892489FB0B" box="[1029,1203,1209,1269]" gridcol="3" gridrow="2" pageId="7" pageNumber="120">two single lines</td>
<td id="76EC5DDCFF06B98BDDE92F8925B8FB0B" box="[1232,1410,1209,1269]" gridcol="4" gridrow="2" pageId="7" pageNumber="120">two patches</td>
</tr>
<tr id="353D34A0FF06B98BD99F2E3525B8FABF" box="[166,1410,1285,1345]" gridrow="3" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2E352238FABF" box="[166,514,1285,1345]" gridcol="0" gridrow="3" pageId="7" pageNumber="120">Interorbital width versus eye diameter</th>
<td id="76EC5DDCFF06B98BDB152E3522C9FABF" box="[556,755,1285,1345]" gridcol="1" gridrow="3" pageId="7" pageNumber="120">wider</td>
<td id="76EC5DDCFF06B98BDA252E3523DEFABF" box="[796,996,1285,1345]" gridcol="2" gridrow="3" pageId="7" pageNumber="120">narrower to slightly wider</td>
<td id="76EC5DDCFF06B98BDD3C2E352489FABF" box="[1029,1203,1285,1345]" gridcol="3" gridrow="3" pageId="7" pageNumber="120">nearly equal or slightly wider</td>
<td id="76EC5DDCFF06B98BDDE92E3525B8FABF" box="[1232,1410,1285,1345]" gridcol="4" gridrow="3" pageId="7" pageNumber="120">wider</td>
</tr>
<tr id="353D34A0FF06B98BD99F2E6225B8FA94" box="[166,1410,1362,1386]" gridrow="4" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2E622238FA94" box="[166,514,1362,1386]" gridcol="0" gridrow="4" pageId="7" pageNumber="120">Right and left gill membrane</th>
<td id="76EC5DDCFF06B98BDB152E6222C9FA94" box="[556,755,1362,1386]" gridcol="1" gridrow="4" pageId="7" pageNumber="120">separated</td>
<td id="76EC5DDCFF06B98BDA252E6223DEFA94" box="[796,996,1362,1386]" gridcol="2" gridrow="4" pageId="7" pageNumber="120">separated</td>
<td id="76EC5DDCFF06B98BDD3C2E622489FA94" box="[1029,1203,1362,1386]" gridcol="3" gridrow="4" pageId="7" pageNumber="120">separated</td>
<td id="76EC5DDCFF06B98BDDE92E6225B8FA94" box="[1232,1410,1362,1386]" gridcol="4" gridrow="4" pageId="7" pageNumber="120">united</td>
</tr>
<tr id="353D34A0FF06B98BD99F2E4A25B8FA6C" box="[166,1410,1402,1426]" gridrow="5" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2E4A2238FA6C" box="[166,514,1402,1426]" gridcol="0" gridrow="5" pageId="7" pageNumber="120">Predorsal scales</th>
<td id="76EC5DDCFF06B98BDB152E4A22C9FA6C" box="[556,755,1402,1426]" gridcol="1" gridrow="5" pageId="7" pageNumber="120">cycloid</td>
<td id="76EC5DDCFF06B98BDA252E4A23DEFA6C" box="[796,996,1402,1426]" gridcol="2" gridrow="5" pageId="7" pageNumber="120">cycloid</td>
<td id="76EC5DDCFF06B98BDD3C2E4A2489FA6C" box="[1029,1203,1402,1426]" gridcol="3" gridrow="5" pageId="7" pageNumber="120">cycloid</td>
<td id="76EC5DDCFF06B98BDDE92E4A25B8FA6C" box="[1232,1410,1402,1426]" gridcol="4" gridrow="5" pageId="7" pageNumber="120">ctenoid</td>
</tr>
<tr id="353D34A0FF06B98BD99F2E9225B8FA20" box="[166,1410,1442,1502]" gridrow="6" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2E922238FA20" box="[166,514,1442,1502]" gridcol="0" gridrow="6" pageId="7" pageNumber="120">Predorsal scales extension to</th>
<td id="76EC5DDCFF06B98BDB152E9222C9FA20" box="[556,755,1442,1502]" gridcol="1" gridrow="6" pageId="7" pageNumber="120">posterior part of interorbital space</td>
<td id="76EC5DDCFF06B98BDA252E9223DEFA20" box="[796,996,1442,1502]" gridcol="2" gridrow="6" pageId="7" pageNumber="120">posterior part of interorbital space</td>
<td id="76EC5DDCFF06B98BDD3C2E922489FA20" box="[1029,1203,1442,1502]" gridcol="3" gridrow="6" pageId="7" pageNumber="120">posterior part of interorbital space</td>
<td id="76EC5DDCFF06B98BDDE92E9225B8FA20" box="[1232,1410,1442,1502]" gridcol="4" gridrow="6" pageId="7" pageNumber="120">to anterior part of interorbital space</td>
</tr>
<tr id="353D34A0FF06B98BD99F2EDE25B8F9F8" box="[166,1410,1518,1542]" gridrow="7" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2EDE2238F9F8" box="[166,514,1518,1542]" gridcol="0" gridrow="7" pageId="7" pageNumber="120">Transforming ctenii on body scale</th>
<td id="76EC5DDCFF06B98BDB152EDE22C9F9F8" box="[556,755,1518,1542]" gridcol="1" gridrow="7" pageId="7" pageNumber="120">several rows</td>
<td id="76EC5DDCFF06B98BDA252EDE23DEF9F8" box="[796,996,1518,1542]" gridcol="2" gridrow="7" pageId="7" pageNumber="120">single row</td>
<td id="76EC5DDCFF06B98BDD3C2EDE2489F9F8" box="[1029,1203,1518,1542]" gridcol="3" gridrow="7" pageId="7" pageNumber="120">absent</td>
<td id="76EC5DDCFF06B98BDDE92EDE25B8F9F8" box="[1232,1410,1518,1542]" gridcol="4" gridrow="7" pageId="7" pageNumber="120">several rows</td>
</tr>
<tr id="353D34A0FF06B98BD99F2D2725B8F9D1" box="[166,1410,1559,1583]" gridrow="8" pageId="7" pageNumber="120">
<th id="76EC5DDCFF06B98BD99F2D272238F9D1" box="[166,514,1559,1583]" gridcol="0" gridrow="8" pageId="7" pageNumber="120">Pre-vomerine teeth</th>
<td id="76EC5DDCFF06B98BDB152D2722C9F9D1" box="[556,755,1559,1583]" gridcol="1" gridrow="8" pageId="7" pageNumber="120">absent</td>
<td id="76EC5DDCFF06B98BDA252D2723DEF9D1" box="[796,996,1559,1583]" gridcol="2" gridrow="8" pageId="7" pageNumber="120">absent</td>
<td id="76EC5DDCFF06B98BDD3C2D272489F9D1" box="[1029,1203,1559,1583]" gridcol="3" gridrow="8" pageId="7" pageNumber="120">present</td>
<td id="76EC5DDCFF06B98BDDE92D2725B8F9D1" box="[1232,1410,1559,1583]" gridcol="4" gridrow="8" pageId="7" pageNumber="120">absent</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3176569FF06467DD9FE2D6E2366F82B" lastPageId="9" lastPageNumber="122" pageId="7" pageNumber="120" type="discussion">
<paragraph id="8BB236E2FF06467CD9FE2D6E23FEFE9B" blockId="7.[151,1437,1630,2017]" lastBlockId="8.[151,1437,151,609]" lastPageId="8" lastPageNumber="121" pageId="7" pageNumber="120">
<emphasis id="B979EAF0FF064673D9FE2D6E2176F986" bold="true" box="[199,332,1630,1656]" pageId="7" pageNumber="120">Discussion.</emphasis>
The genus 
<taxonomicName id="4C0D4D61FF064673D8F42D6E224AF986" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[461,624,1630,1656]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF064673D8F42D6E224AF986" box="[461,624,1630,1656]" italics="true" pageId="7" pageNumber="120">Neodontobutis</emphasis>
</taxonomicName>
was established by 
<bibRefCitation id="EF9C4B13FF064673DA762D6E2426F986" author="Chen, I. S. &amp; Kottelat, M. &amp; Wu, H." box="[847,1052,1630,1656]" pageId="7" pageNumber="120" pagination="229 - 235" refId="ref5442" refString="Chen, I. S., Kottelat, M. &amp; Wu, H. (2002) A new genus of freshwater sleeper (Teleostei: Odontobutididae) from southern China and mainland Southeast Asia. Journal of the Fisheries Society of Taiwan, 29, 229 - 235." type="journal article" year="2002">
Chen 
<emphasis id="B979EAF0FF064673DAA82D6F23F3F986" box="[913,969,1630,1656]" italics="true" pageId="7" pageNumber="120">et al.</emphasis>
(2002)
</bibRefCitation>
, based on 
<taxonomicName id="4C0D4D61FF064673DDA92D6F250DF986" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[1168,1335,1630,1656]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF064673DDA92D6F250DF986" box="[1168,1335,1630,1656]" italics="true" pageId="7" pageNumber="120">N. hainanensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9C4B13FF064673DC7C2D6E20C4F962" author="Chen, I. S. &amp; Kottelat, M. &amp; Wu, H." pageId="7" pageNumber="120" pagination="229 - 235" refId="ref5442" refString="Chen, I. S., Kottelat, M. &amp; Wu, H. (2002) A new genus of freshwater sleeper (Teleostei: Odontobutididae) from southern China and mainland Southeast Asia. Journal of the Fisheries Society of Taiwan, 29, 229 - 235." type="journal article" year="2002">
Chen 
<emphasis id="B979EAF0FF064673DCB12D6F208EF962" italics="true" pageId="7" pageNumber="120">et al.</emphasis>
, 2002
</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF064673D8302DB321AAF962" author="Iwata, A." box="[265,400,1666,1693]" pageId="7" pageNumber="120" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF064673D8A22DB32234F962" author="Wu, H. L. &amp; Zhong, J. S. &amp; Chen, I. - S. &amp; Chong, D. H. &amp; Ni, Y. &amp; Shen, GY &amp; Zhao, S. L. &amp; Shao, K. T." box="[411,526,1666,1693]" pageId="7" pageNumber="120" refId="ref6288" refString="Wu, H. L., Zhong, J. S., Chen, I. - S., Chong, D. H., Ni, Y., Shen, GY., Zhao, S. L. &amp; Shao, K. T. (2008) Fauna Sinica, Ostichthyes, Perciformes (V), Gobioidei. Beijing: Science Press." type="book" year="2008">Wu, 2008</bibRefCitation>
). This genus can be distinguished from other genera of 
<taxonomicName id="4C0D4D61FF064673DD442DB22519F962" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[1149,1315,1666,1692]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
mainly by the following characters: (1) sensory papilla forming a barbel-like projection, which is the most distinctive character; (2) right and left gill membranes separate; (3) rather compressed head and body, interorbital width narrower than eye diameter; (4) predorsal scales cycloid, reaching posterior part of interorbital space; (5) sensory papilla on lower jaw arranged in two single lines (
<bibRefCitation id="EF9C4B13FF064673D8E72C222252F8D2" author="Iwata, A." box="[478,616,1810,1836]" pageId="7" pageNumber="120" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
). However, some relevant characters of 
<taxonomicName id="4C0D4D61FF064673DD122C232445F8D2" box="[1067,1151,1810,1836]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lanii">
<emphasis id="B979EAF0FF064673DD122C232445F8D2" box="[1067,1151,1810,1836]" italics="true" pageId="7" pageNumber="120">N. lanii</emphasis>
</taxonomicName>
do not fit the description above: 
<taxonomicName id="4C0D4D61FF064673D9DE2C072108F8AE" box="[231,306,1846,1872]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673D9DE2C072108F8AE" box="[231,306,1846,1872]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
possess relatively flat head, with mean interorbital width / eye diameter = 1.6 (range = 1.4–1.9), higher than that of 
<taxonomicName id="4C0D4D61FF064673D8182C6B21F2F88A" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[289,456,1882,1908]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF064673D8182C6B21F2F88A" box="[289,456,1882,1908]" italics="true" pageId="7" pageNumber="120">N. hainanensis</emphasis>
</taxonomicName>
(mean = 1.0; range = 0.8–1.1), 
<taxonomicName id="4C0D4D61FF064673DA0A2C6B238DF88A" box="[819,951,1883,1908]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="auarmus">
<emphasis id="B979EAF0FF064673DA0A2C6B238DF88A" box="[819,951,1883,1908]" italics="true" pageId="7" pageNumber="120">N. auarmus</emphasis>
</taxonomicName>
(1.0 in male, 
<quantity id="4CF59B07FF064673DD6E2C6A24A2F88A" box="[1111,1176,1882,1908]" metricMagnitude="-2" metricUnit="m" metricValue="2.286" pageId="7" pageNumber="120" unit="in" value="0.9">0.9 in</quantity>
female) (
<bibRefCitation id="EF9C4B13FF064673DC3D2C6A20E8F867" author="Vidthayanon, C." pageId="7" pageNumber="120" pagination="235 - 242" refId="ref6186" refString="Vidthayanon, C. (1995) Odontobutis aurarmus, a new species of odontobutid goby from wetlands of the Mekong basin, Thailand. Ichthyological Exploration of Freshwaters, 6, 235 - 242." type="journal article" year="1995">Vidthayanon, 1995</bibRefCitation>
), 
<taxonomicName id="4C0D4D61FF064673D9D12C4F21F8F866" authorityName="Mai" authorityYear="1978" box="[232,450,1918,1944]" class="Actinopterygii" family="Odontobutidae" genus="Micropercops" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="macropectoralis">
<emphasis id="B979EAF0FF064673D9D12C4F21F8F866" box="[232,450,1918,1944]" italics="true" pageId="7" pageNumber="120">N. macropectoralis</emphasis>
</taxonomicName>
(1.3) (
<bibRefCitation id="EF9C4B13FF064673DB292C4E22F8F867" author="Kottelat, M." box="[528,706,1918,1945]" pageId="7" pageNumber="120" refId="ref5721" refString="Kottelat, M. (2001 b) Freshwater Fishes of Northern Vietnam. World Bank, Washington, 11 pp." type="book" year="2001">Kottelat, 2001b</bibRefCitation>
) and 
<taxonomicName id="4C0D4D61FF064673DA3D2C4F239FF866" baseAuthorityName="Mai" baseAuthorityYear="1978" box="[772,933,1918,1944]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="tonkinensis">
<emphasis id="B979EAF0FF064673DA3D2C4F239FF866" box="[772,933,1918,1944]" italics="true" pageId="7" pageNumber="120">N. tonkinensis</emphasis>
</taxonomicName>
(1.2) (
<bibRefCitation id="EF9C4B13FF064673DACA2C4E249CF867" author="Kottelat, M." box="[1011,1190,1918,1945]" pageId="7" pageNumber="120" refId="ref5721" refString="Kottelat, M. (2001 b) Freshwater Fishes of Northern Vietnam. World Bank, Washington, 11 pp." type="book" year="2001">Kottelat, 2001b</bibRefCitation>
). The mean value of interorbital width / eye diameter of 
<taxonomicName id="4C0D4D61FF064673DB122C932243F842" box="[555,633,1954,1980]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF064673DB122C932243F842" box="[555,633,1954,1980]" italics="true" pageId="7" pageNumber="120">N. lani</emphasis>
</taxonomicName>
was significantly higher than that of 
<taxonomicName id="4C0D4D61FF064673DD182C9324F2F842" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[1057,1224,1954,1980]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF064673DD182C9324F2F842" box="[1057,1224,1954,1980]" italics="true" pageId="7" pageNumber="120">N. hainanensis</emphasis>
</taxonomicName>
(P &lt;0.01) (
<figureCitation id="13362A67FF064673DC682C92209CF81E" captionStart="FIGURE 8" captionStartId="10.[151,250,887,911]" captionTargetBox="[153,1436,189,859]" captionTargetId="figure-28@10.[151,1436,181,863]" captionTargetPageId="10" captionText="FIGURE 8. Comparison of the head region between Neodontobutis lani and N. hainanensis. (a) boxplot illustrates interorbital width / Eye diameter, including 12 individuals of N. lani (39.0–60.5 mm standard length) and 10 individuals of N. hainanensis (32.7–41.3 mm standard length), t value=11.171** (raw data can be found in electronic appendix). (b) Comparison of head shape of N. hainanensis (left, 41.3 mm standard length, 25901) and N. lani (right, 39.1 mm standard length, 259122)." figureDoi="http://doi.org/10.5281/zenodo.6531386" httpUri="https://zenodo.org/record/6531386/files/figure.png" pageId="7" pageNumber="120">Figure 8</figureCitation>
), with the raw measurements can be found in electronic appendix. Moreover, several rows of transforming ctenii present on posterior edge of body scale of 
<taxonomicName id="4C0D4D61FF09467CDB4B2BA72285FF4F" box="[626,703,151,177]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF09467CDB4B2BA72285FF4F" box="[626,703,151,177]" italics="true" pageId="8" pageNumber="121">N. lani</emphasis>
</taxonomicName>
, while only one row of transforming ctenii present on that part of the other 
<taxonomicName id="4C0D4D61FF09467CD8392B8B21C0FF2B" box="[256,506,187,213]" pageId="8" pageNumber="121">
<emphasis id="B979EAF0FF09467CD8392B8B2199FF2B" box="[256,419,187,213]" italics="true" pageId="8" pageNumber="121">Neodontobutis</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EF9C4B13FF09467CDB312B8B22B5FF2B" author="Iwata, A." box="[520,655,187,213]" pageId="8" pageNumber="121" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF09467CDBA32B8B2348FF2B" author="Vidthayanon, C." box="[666,882,186,213]" pageId="8" pageNumber="121" pagination="235 - 242" refId="ref6186" refString="Vidthayanon, C. (1995) Odontobutis aurarmus, a new species of odontobutid goby from wetlands of the Mekong basin, Thailand. Ichthyological Exploration of Freshwaters, 6, 235 - 242." type="journal article" year="1995">Vidthayanon, 1995</bibRefCitation>
). In addition, sensory papilla on the lower jaw are arranged in two oblong clusters in 
<taxonomicName id="4C0D4D61FF09467CDB262BEF2256FF07" box="[543,620,223,249]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF09467CDB262BEF2256FF07" box="[543,620,223,249]" italics="true" pageId="8" pageNumber="121">N. lani</emphasis>
</taxonomicName>
, which differs from other 
<taxonomicName id="4C0D4D61FF09467CDAAC2BEF24AEFF07" box="[917,1172,223,249]" pageId="8" pageNumber="121">
<emphasis id="B979EAF0FF09467CDAAC2BEF2402FF07" box="[917,1080,223,249]" italics="true" pageId="8" pageNumber="121">Neodontobutis</emphasis>
species
</taxonomicName>
, which have papioae in two single lines on the underside of the jaw (
<bibRefCitation id="EF9C4B13FF09467CDBBF2A332335FEE3" author="Iwata, A." box="[646,783,259,285]" pageId="8" pageNumber="121" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
). Interestingly, these characteristics of 
<taxonomicName id="4C0D4D61FF09467CDD842A332533FEE3" box="[1213,1289,259,285]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF09467CDD842A332533FEE3" box="[1213,1289,259,285]" italics="true" pageId="8" pageNumber="121">N. lani</emphasis>
</taxonomicName>
closely align with character states described for 
<taxonomicName id="4C0D4D61FF09467CDB142A17228DFEBF" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[557,695,295,321]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDB142A17228DFEBF" box="[557,695,295,321]" italics="true" pageId="8" pageNumber="121">Odontobutis</emphasis>
</taxonomicName>
. Some critical external characters of genus 
<taxonomicName id="4C0D4D61FF09467CDD812A172561FEBF" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1208,1371,295,321]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDD812A172561FEBF" box="[1208,1371,295,321]" italics="true" pageId="8" pageNumber="121">Neodontobutis</emphasis>
</taxonomicName>
were compared to that of 
<taxonomicName id="4C0D4D61FF09467CD8402A7B21CAFE9A" authorityName="Dybowski" authorityYear="1877" box="[377,496,331,356]" class="Actinopterygii" family="Odontobutidae" genus="Perccottus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CD8402A7B21CAFE9A" box="[377,496,331,356]" italics="true" pageId="8" pageNumber="121">Perccottus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D61FF09467CDB1F2A7B228AFE9B" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[550,688,331,357]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDB1F2A7B228AFE9B" box="[550,688,331,357]" italics="true" pageId="8" pageNumber="121">Odontobutis</emphasis>
</taxonomicName>
in table 4 (
<bibRefCitation id="EF9C4B13FF09467CDA172A7B2382FE9B" author="Iwata, A." box="[814,952,331,357]" pageId="8" pageNumber="121" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
).
</paragraph>
<paragraph id="8BB236E2FF09467CD9FE2A5F2543FD9F" blockId="8.[151,1437,151,609]" pageId="8" pageNumber="121">
Despite some similarities to 
<taxonomicName id="4C0D4D61FF09467CDB332A5F22AEFE77" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[522,660,367,393]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDB332A5F22AEFE77" box="[522,660,367,393]" italics="true" pageId="8" pageNumber="121">Odontobutis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0D4D61FF09467CDB9D2A5F2341FE77" box="[676,891,367,393]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF09467CDB9D2A5F2341FE77" box="[676,891,367,393]" italics="true" pageId="8" pageNumber="121">Neodontobutis lani</emphasis>
</taxonomicName>
was assigned to the genus 
<taxonomicName id="4C0D4D61FF09467CDD882A5F256EFE77" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1201,1364,367,393]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDD882A5F256EFE77" box="[1201,1364,367,393]" italics="true" pageId="8" pageNumber="121">Neodontobutis</emphasis>
</taxonomicName>
based on following reasons: (1) the most critical external characters, such as shape of sensory papilla (presence of barbellike projection), connection of gill membrane (not connected) and predorsal scales (cycloid, ctenoid in 
<taxonomicName id="4C0D4D61FF09467CDC312A8725A8FE2F" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[1288,1426,439,465]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDC312A8725A8FE2F" box="[1288,1426,439,465]" italics="true" pageId="8" pageNumber="121">Odontobutis</emphasis>
</taxonomicName>
) consistent with that of the genus 
<taxonomicName id="4C0D4D61FF09467CDB232AEB2287FE0B" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[538,701,475,501]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDB232AEB2287FE0B" box="[538,701,475,501]" italics="true" pageId="8" pageNumber="121">Neodontobutis</emphasis>
</taxonomicName>
; (2) molecular phylogenetic analysis shows that 
<taxonomicName id="4C0D4D61FF09467CDDCE2AEB257CFE0B" box="[1271,1350,475,501]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF09467CDDCE2AEB257CFE0B" box="[1271,1350,475,501]" italics="true" pageId="8" pageNumber="121">N. lani</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D61FF09467CDCBA2AEB2127FDE7" baseAuthorityName="Chen" baseAuthorityYear="1985" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF09467CDCBA2AEB2127FDE7" italics="true" pageId="8" pageNumber="121">N. hainanensis</emphasis>
</taxonomicName>
, the 
<typeStatus id="54B68840FF09467CD8652ACF21B6FDE7" box="[348,396,511,537]" pageId="8" pageNumber="121">type</typeStatus>
species of the genus 
<taxonomicName id="4C0D4D61FF09467CDBA92ACF2309FDE7" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[656,819,511,537]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CDBA92ACF2309FDE7" box="[656,819,511,537]" italics="true" pageId="8" pageNumber="121">Neodontobutis</emphasis>
</taxonomicName>
, formed a monophyletic group with a 100 bootstrap support (
<figureCitation id="13362A67FF09467CD9C329132161FDC3" box="[250,347,547,573]" captionStart="FIGURE 6" captionStartId="8.[151,250,1804,1828]" captionTargetBox="[159,1434,682,1769]" captionTargetId="figure-255@8.[151,1436,669,1781]" captionTargetPageId="8" captionText="FIGURE 6. Concatenated maximum likelihood tree of Neodontobutis lani, and species representing all genus of the Odontobutidae, except for Terateleotris, based on 2,076 single-copy nuclear loci. *SOU1801007-3 is a paratype." figureDoi="http://doi.org/10.5281/zenodo.6531382" httpUri="https://zenodo.org/record/6531382/files/figure.png" pageId="8" pageNumber="121">Figure 6</figureCitation>
&amp; 
<figureCitation id="13362A67FF09467CD846291321B5FDC3" box="[383,399,547,573]" captionStart="FIGURE 7" captionStartId="9.[151,250,1368,1392]" captionTargetBox="[158,1425,191,1335]" captionTargetId="figure-18@9.[151,1436,181,1344]" captionTargetPageId="9" captionText="FIGURE 7. Maximum likelihood tree of Neodontobutis lani, and comparative species of the Odontobutidae, based on partial sequence (~ 1500 bp) of cytochrome c oxidase subunit I (COI) gene. *SOU1801007-1 is the Holotype, SOU1801007-3 and SOU1801007-5 are the paratypes." figureDoi="http://doi.org/10.5281/zenodo.6531384" httpUri="https://zenodo.org/record/6531384/files/figure.png" pageId="8" pageNumber="121">7</figureCitation>
). Due to inadequate taxon sampling, specimens and tissue samples of 
<taxonomicName id="4C0D4D61FF09467CDD9129132511FDC2" box="[1192,1323,547,572]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="auarmus">
<emphasis id="B979EAF0FF09467CDD9129132511FDC2" box="[1192,1323,547,572]" italics="true" pageId="8" pageNumber="121">N. auarmus</emphasis>
</taxonomicName>
and other relevant species are not available for this study. The relationship of species of this genus needs further research.
</paragraph>
<caption id="DF72666AFF09467CD9AE2C3C2534F8B7" ID-DOI="http://doi.org/10.5281/zenodo.6531382" ID-Zenodo-Dep="6531382" httpUri="https://zenodo.org/record/6531382/files/figure.png" pageId="8" pageNumber="121" startId="8.[151,250,1804,1828]" targetBox="[159,1434,682,1769]" targetPageId="8">
<paragraph id="8BB236E2FF09467CD9AE2C3C2534F8B7" blockId="8.[151,1436,1804,1865]" pageId="8" pageNumber="121">
<emphasis id="B979EAF0FF09467CD9AE2C3C2120F8DA" bold="true" box="[151,282,1804,1828]" pageId="8" pageNumber="121">FIGURE 6.</emphasis>
Concatenated maximum likelihood tree of 
<taxonomicName id="4C0D4D61FF09467CDBC42C3C23FFF8DA" box="[765,965,1804,1828]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF09467CDBC42C3C23FFF8DA" box="[765,965,1804,1828]" italics="true" pageId="8" pageNumber="121">Neodontobutis lani</emphasis>
</taxonomicName>
, and species representing all genus of the 
<taxonomicName id="4C0D4D61FF09467CD9AE2C00210AF8B6" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[151,304,1840,1864]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
, except for 
<taxonomicName id="4C0D4D61FF09467CD89E2C002212F8B6" authorityName="Shibukawa, Iwata &amp; Viravong" authorityYear="2001" box="[423,552,1840,1864]" class="Actinopterygii" family="Odontobutidae" genus="Terateleotris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF09467CD89E2C002212F8B6" box="[423,552,1840,1864]" italics="true" pageId="8" pageNumber="121">Terateleotris</emphasis>
</taxonomicName>
, based on 2,076 single-copy nuclear loci. *SOU1801007-3 is a paratype.
</paragraph>
</caption>
<caption id="DF72666AFF08467DD9AE2E6821CBFA47" ID-DOI="http://doi.org/10.5281/zenodo.6531384" ID-Zenodo-Dep="6531384" httpUri="https://zenodo.org/record/6531384/files/figure.png" pageId="9" pageNumber="122" startId="9.[151,250,1368,1392]" targetBox="[158,1425,191,1335]" targetPageId="9">
<paragraph id="8BB236E2FF08467DD9AE2E6821CBFA47" blockId="9.[151,1437,1368,1465]" pageId="9" pageNumber="122">
<emphasis id="B979EAF0FF08467DD9AE2E68212EFA8F" bold="true" box="[151,276,1368,1393]" pageId="9" pageNumber="122">FIGURE 7.</emphasis>
Maximum likelihood tree of 
<taxonomicName id="4C0D4D61FF08467DDB7A2E68233FFA8E" box="[579,773,1368,1392]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF08467DDB7A2E68233FFA8E" box="[579,773,1368,1392]" italics="true" pageId="9" pageNumber="122">Neodontobutis lani</emphasis>
</taxonomicName>
, and comparative species of the 
<taxonomicName id="4C0D4D61FF08467DDD6F2E6824D5FA8E" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[1110,1263,1368,1392]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
, based on partial sequence (~ 1500 bp) of 
<emphasis id="B979EAF0FF08467DD8A72E4C22DAFA6A" box="[414,736,1404,1428]" italics="true" pageId="9" pageNumber="122">cytochrome c oxidase subunit I</emphasis>
(
<emphasis id="B979EAF0FF08467DDBC82E4D2327FA6A" box="[753,797,1405,1428]" italics="true" pageId="9" pageNumber="122">COI</emphasis>
) gene. *SOU1801007-1 is the Holotype, SOU1801007-3 and SOU1801007-5 are the paratypes.
</paragraph>
</caption>
<paragraph id="8BB236E2FF08467DD9FE2ED723BBF927" blockId="9.[151,1437,1511,2005]" pageId="9" pageNumber="122">
<emphasis id="B979EAF0FF08467DD9FE2ED7217AF9FF" box="[199,320,1511,1537]" pageId="9" pageNumber="122">Specimens</emphasis>
of 
<taxonomicName id="4C0D4D61FF08467DD8502ED7227BF9FF" box="[361,577,1511,1537]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF08467DD8502ED7227BF9FF" box="[361,577,1511,1537]" italics="true" pageId="9" pageNumber="122">Neodontobutis lani</emphasis>
</taxonomicName>
were collected from Longzhou Town, the Zuojiang River basin, which is a tributary of the Pearl River. 
<taxonomicName id="4C0D4D61FF08467DD8E92D3B233AF9DB" baseAuthorityName="Chen" baseAuthorityYear="1985" box="[464,768,1547,1573]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B979EAF0FF08467DD8E92D3B233AF9DB" box="[464,768,1547,1573]" italics="true" pageId="9" pageNumber="122">Neodontobutis hainanensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D61FF08467DDA0C2D3B238AF9DA" authorityName="Wu, Chen &amp; Chong" authorityYear="2002" box="[821,944,1547,1573]" class="Actinopterygii" family="Odontobutidae" genus="Odontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="B979EAF0FF08467DDA0C2D3B238AF9DA" box="[821,944,1547,1573]" italics="true" pageId="9" pageNumber="122">O. sinensis</emphasis>
</taxonomicName>
are other 
<taxonomicName id="4C0D4D61FF08467DDD192D3B24FCF9DB" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[1056,1222,1547,1573]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
species distributed in the Zuojiang Basin, so the differentiation of their niches would be an interesting question. Moreover, other 
<taxonomicName id="4C0D4D61FF08467DD9AE2D6321A2F993" box="[151,408,1619,1645]" pageId="9" pageNumber="122">
<emphasis id="B979EAF0FF08467DD9AE2D632100F993" box="[151,314,1619,1645]" italics="true" pageId="9" pageNumber="122">Neodontobutis</emphasis>
species
</taxonomicName>
, 
<taxonomicName id="4C0D4D61FF08467DD89F2D632216F992" box="[422,556,1619,1644]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="auarmus">
<emphasis id="B979EAF0FF08467DD89F2D632216F992" box="[422,556,1619,1644]" italics="true" pageId="9" pageNumber="122">N. auarmus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0D4D61FF08467DDB562D63232BF993" baseAuthorityName="Mai" baseAuthorityYear="1978" box="[623,785,1619,1645]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="tonkinensis">
<emphasis id="B979EAF0FF08467DDB562D63232BF993" box="[623,785,1619,1645]" italics="true" pageId="9" pageNumber="122">N. tonkinensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D61FF08467DDA742D6323C1F993" authorityName="X. K. Nguyen et D. H. Nguyen" authorityYear="2011" box="[845,1019,1619,1645]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="ngheanensis">
<emphasis id="B979EAF0FF08467DDA742D6323C1F993" box="[845,1019,1619,1645]" italics="true" pageId="9" pageNumber="122">N. ngheanensis</emphasis>
</taxonomicName>
, which are tentatively placed in the genus 
<taxonomicName id="4C0D4D61FF08467DD9DA2D472214F96F" authority="(Iwata, 2011)" baseAuthorityName="Iwata" baseAuthorityYear="2011" box="[227,558,1655,1681]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="genus">
<emphasis id="B979EAF0FF08467DD9DA2D4721BCF96F" box="[227,390,1655,1681]" italics="true" pageId="9" pageNumber="122">Neodontobutis</emphasis>
(
<bibRefCitation id="EF9C4B13FF08467DD8A32D47221CF96F" author="Iwata, A." box="[410,550,1655,1681]" pageId="9" pageNumber="122" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
)
</taxonomicName>
, are distributed in the Indo-China Peninsula (
<bibRefCitation id="EF9C4B13FF08467DDD7C2D472516F96F" author="Khoa, N. &amp; Duc, N." box="[1093,1324,1655,1681]" pageId="9" pageNumber="122" refId="ref5645" refString="Khoa, N. &amp; Duc, N. (2012) New species of genus Neodontobutis (Gobioidei: Odontobutidae) from Khe Khang stream in Pu Mat national park. TAP CHI SINH HOC, 33. https: // doi. org / 10.15625 / 0866 - 7160 / v 33 n 1.728" type="book" year="2012">Khoa &amp; Duc, 2012</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF08467DDC022D4720E5F94B" author="Kottelat, M." pageId="9" pageNumber="122" refId="ref5701" refString="Kottelat, M. (2001 a) Fishes of Laos. WHT Publications, Colombo, 198 pp." type="book" year="2001">Kottelat, 2001a</bibRefCitation>
, 
<bibRefCitation id="EF9C4B13FF08467DD9D12DAB2108F94B" author="Kottelat, M." box="[232,306,1691,1717]" pageId="9" pageNumber="122" refId="ref5721" refString="Kottelat, M. (2001 b) Freshwater Fishes of Northern Vietnam. World Bank, Washington, 11 pp." type="book" year="2001">2001b</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF08467DD8022DAB2229F94B" author="Vidthayanon, C." box="[315,531,1690,1717]" pageId="9" pageNumber="122" pagination="235 - 242" refId="ref6186" refString="Vidthayanon, C. (1995) Odontobutis aurarmus, a new species of odontobutid goby from wetlands of the Mekong basin, Thailand. Ichthyological Exploration of Freshwaters, 6, 235 - 242." type="journal article" year="1995">Vidthayanon, 1995</bibRefCitation>
). Investigation on the relationship of 
<taxonomicName id="4C0D4D61FF08467DDA972DAB23C3F94B" box="[942,1017,1691,1717]" class="Actinopterygii" family="Odontobutidae" genus="Neodontobutis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="species" species="lani">
<emphasis id="B979EAF0FF08467DDA972DAB23C3F94B" box="[942,1017,1691,1717]" italics="true" pageId="9" pageNumber="122">N. lani</emphasis>
</taxonomicName>
and other 
<taxonomicName id="4C0D4D61FF08467DDD552DAB255CF94B" box="[1132,1382,1691,1717]" pageId="9" pageNumber="122">
<emphasis id="B979EAF0FF08467DDD552DAB2535F94B" box="[1132,1295,1691,1717]" italics="true" pageId="9" pageNumber="122">Neodontobutis</emphasis>
species
</taxonomicName>
may shed light on the origin, diversification and dispersal of these taxa.
</paragraph>
<paragraph id="8BB236E2FF08467DD9FE2DD32366F82B" blockId="9.[151,1437,1511,2005]" pageId="9" pageNumber="122">
According to 
<bibRefCitation id="EF9C4B13FF08467DD85D2DD3222EF903" author="Li, H. &amp; He, Y. &amp; Jiang, J. &amp; Liu, Z. &amp; Li, C." box="[356,532,1763,1789]" pageId="9" pageNumber="122" pagination="1 - 11" refId="ref5860" refString="Li, H., He, Y., Jiang, J., Liu, Z. &amp; Li, C. (2018) Molecular systematics and phylogenetic analysis of the Asian endemic freshwater sleepers (Gobiiformes: Odontobutidae). Molecular Phylogenetics and Evolution, 121, 1 - 11. https: // doi. org / 10.1016 / j. ympev. 2017.12.026" type="journal article" year="2018">
Li 
<emphasis id="B979EAF0FF08467DD8BC2DD32185F903" box="[389,447,1763,1789]" italics="true" pageId="9" pageNumber="122">et al.</emphasis>
(2018)
</bibRefCitation>
, southern 
<collectingCountry id="F31A7672FF08467DDBB32DD322F7F903" box="[650,717,1763,1789]" name="China" pageId="9" pageNumber="122">China</collectingCountry>
may be the region where the family 
<taxonomicName id="4C0D4D61FF08467DDD4C2DD32521F903" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[1141,1307,1763,1789]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
originated. Indeed, southern 
<collectingCountry id="F31A7672FF08467DD86F2C3721A3F8DF" box="[342,409,1799,1825]" name="China" pageId="9" pageNumber="122">China</collectingCountry>
(
<collectingRegion id="49C9F800FF08467DD89F2C372217F8DF" box="[422,557,1799,1825]" country="China" name="Guangdong" pageId="9" pageNumber="122">Guangdong</collectingRegion>
, 
<collectingRegion id="49C9F800FF08467DDB0E2C3722ADF8DF" box="[567,663,1799,1825]" country="China" name="Guangxi" pageId="9" pageNumber="122">Guangxi</collectingRegion>
and 
<collectingRegion id="49C9F800FF08467DDBF52C37231BF8DC" box="[716,801,1799,1826]" country="China" name="Hainan" pageId="9" pageNumber="122">Hainan</collectingRegion>
), and the adjacent Indo-China Peninsula, are the diversity centers of the 
<taxonomicName id="4C0D4D61FF08467DD8052C1B21DFF8BB" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[316,485,1835,1861]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
, which is home to 7 species (belonging to 5 genera) and 6 species (belonging to 3 genera) respectively, with 2 shared genera (
<bibRefCitation id="EF9C4B13FF08467DDBB32C7F236CF897" author="Chen, I. S. &amp; Kottelat, M. &amp; Wu, H." box="[650,854,1871,1897]" pageId="9" pageNumber="122" pagination="229 - 235" refId="ref5442" refString="Chen, I. S., Kottelat, M. &amp; Wu, H. (2002) A new genus of freshwater sleeper (Teleostei: Odontobutididae) from southern China and mainland Southeast Asia. Journal of the Fisheries Society of Taiwan, 29, 229 - 235." type="journal article" year="2002">
Chen 
<emphasis id="B979EAF0FF08467DDBE92C7F2330F897" box="[720,778,1871,1897]" italics="true" pageId="9" pageNumber="122">et al.</emphasis>
, 2002
</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF08467DDA5A2C7F23D6F897" author="Iwata, A." box="[867,1004,1871,1897]" pageId="9" pageNumber="122" pagination="61 - 77" refId="ref5568" refString="Iwata, A. (2011) Systematics of odontobutidae. The Biology of Gobies. Science Publishers, Enfield, 61 - 77." type="book chapter" year="2011">Iwata, 2011</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF08467DDAC32C7F24E3F897" author="Khoa, N. &amp; Duc, N." box="[1018,1241,1871,1897]" pageId="9" pageNumber="122" refId="ref5645" refString="Khoa, N. &amp; Duc, N. (2012) New species of genus Neodontobutis (Gobioidei: Odontobutidae) from Khe Khang stream in Pu Mat national park. TAP CHI SINH HOC, 33. https: // doi. org / 10.15625 / 0866 - 7160 / v 33 n 1.728" type="book" year="2012">Khoa &amp; Duc, 2012</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF08467DDDDF2C7F25A2F897" author="Kottelat, M." box="[1254,1432,1871,1897]" pageId="9" pageNumber="122" refId="ref5701" refString="Kottelat, M. (2001 a) Fishes of Laos. WHT Publications, Colombo, 198 pp." type="book" year="2001">Kottelat, 2001a</bibRefCitation>
, 
<bibRefCitation id="EF9C4B13FF08467DD9AE2C4320DBF873" author="Kottelat, M." box="[151,225,1907,1933]" pageId="9" pageNumber="122" refId="ref5721" refString="Kottelat, M. (2001 b) Freshwater Fishes of Northern Vietnam. World Bank, Washington, 11 pp." type="book" year="2001">2001b</bibRefCitation>
; 
<bibRefCitation id="EF9C4B13FF08467DD9D72C432159F873" author="Wu, H. L. &amp; Zhong, J. S. &amp; Chen, I. - S. &amp; Chong, D. H. &amp; Ni, Y. &amp; Shen, GY &amp; Zhao, S. L. &amp; Shao, K. T." box="[238,355,1907,1933]" pageId="9" pageNumber="122" refId="ref6288" refString="Wu, H. L., Zhong, J. S., Chen, I. - S., Chong, D. H., Ni, Y., Shen, GY., Zhao, S. L. &amp; Shao, K. T. (2008) Fauna Sinica, Ostichthyes, Perciformes (V), Gobioidei. Beijing: Science Press." type="book" year="2008">Wu, 2008</bibRefCitation>
). Due to their 
<emphasis id="B979EAF0FF08467DDB352C432273F873" box="[524,585,1907,1933]" pageId="9" pageNumber="122">small</emphasis>
size, benthic habit and low economic importance, as well as their remote distribution, diversity of 
<taxonomicName id="4C0D4D61FF08467DD8852CA72258F84F" authorityName="Hoese &amp; A.C.Gill" authorityYear="1993" box="[444,610,1943,1969]" class="Actinopterygii" family="Odontobutidae" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="122" phylum="Chordata" rank="family">Odontobutidae</taxonomicName>
in Southern 
<collectingCountry id="F31A7672FF08467DDA382CA7237EF84F" box="[769,836,1943,1969]" name="China" pageId="9" pageNumber="122">China</collectingCountry>
and Indo-China Peninsula is understudied. Further exploration may uncover more new species from these regions.
</paragraph>
</subSubSection>
</treatment>
</document>