treatments-xml/data/03/A4/87/03A487C8FFC3FFAFFF59FA83EF53AAB2.xml
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<document id="FEF5AE7E1A72BD963FCD63026060CD54" ID-CLB-Dataset="55782" ID-DOI="10.11646/zootaxa.5129.4.2" ID-GBIF-Dataset="ff4f0237-882f-45d1-a04a-898b7c0ea9f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="6504144" ID-ZooBank="E9ACC0E3-A5EE-4867-8CBA-56FBBB728EFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651218754646" checkinUser="plazi" docAuthor="Dantas, Galileu P. S., Amat, Eduardo, Hamada, Neusa &amp; Giłka, Wojciech" docDate="2022" docId="03A487C8FFC3FFAFFF59FA83EF53AAB2" docLanguage="en" docName="zootaxa.5129.4.2.pdf" docOrigin="Zootaxa 5129 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus lulu Dantas &amp; Amat &amp; Hamada &amp; Giłka 2022, sp. nov." docType="treatment" docUuid="0FFBC66B-AF75-4301-ACA1-96195E22440C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="517" masterDocId="FF9DFFB0FFC9FFA2FFCEFF94EB45A81B" masterDocTitle="Towards the systematics and diversity of Neotropical Tanytarsus van der Wulp (Diptera: Chironomidae): news from Colombia" masterLastPageNumber="529" masterPageNumber="505" pageNumber="515" updateTime="1699329456537" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B9828E2DE36391A9A39AC264E262728C">Towards the systematics and diversity of Neotropical Tanytarsus van der Wulp (Diptera: Chironomidae): news from Colombia</mods:title>
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<mods:namePart id="A22D0800C978D21C4E8F23588CD08EDC">Dantas, Galileu P. S.</mods:namePart>
<mods:nameIdentifier id="DE473A2E3B96FBE226D4A0295B813A79" type="ORCID">0000-0002-9155-533X</mods:nameIdentifier>
<mods:affiliation id="48B58D92A7F0E510F38B5969456150CF">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt); Av. André Araújo, 2936, 69067 - 375, Manaus, Amazonas, Brazil. &amp; galileu. psd @ gmail. com; https: // orcid. org / 0000 - 0002 - 9155 - 533 X</mods:affiliation>
<mods:nameIdentifier id="2EA7607AE804D135DF6D7196FAE190E3" type="email">galileu.psd@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BD625512F4BC4842F16600AAC51AB482">Amat, Eduardo</mods:namePart>
<mods:affiliation id="6DBFB456D85EF59305ABEB809CA171CB">Grupo de investigación BIOFORENSE, Facultad de Derecho y Ciencias Forenses, Tecnológico deAntioquia - Institución Universitaria; calle 78 B no. T 2 A- 22 MI MedellínI Colombia.</mods:affiliation>
</mods:name>
<mods:name id="58234C0BD8E5A6BBD89F70F945D5B70A" type="personal">
<mods:role id="934E92DBE41F724C15CC13B90B014689">
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<mods:namePart id="EE16F13E1C9A4A521BAF28CE359420CE">Hamada, Neusa</mods:namePart>
<mods:nameIdentifier id="A5811E9E9558F48D1FF45D3ACD1A7E0E" type="ORCID">0000-0002-3526-5426</mods:nameIdentifier>
<mods:affiliation id="176EA653051998D3B91755B135B65F8C">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt); Av. André Araújo, 2936, 69067 - 375, Manaus, Amazonas, Brazil. &amp; nhamada @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 3526 - 5426</mods:affiliation>
<mods:nameIdentifier id="7A89D6616C6EDDB3493A74AAB6FE24EB" type="email">nhamada@inpa.gov.br</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="26E2B0872A9D75D8A345EA07CD7B75D6">Giłka, Wojciech</mods:namePart>
<mods:affiliation id="AD2FBBA827EB0B8F83009C2C0CAC938F">University of Gdańsk, Faculty of Biology, Department of Invertebrate Zoology and Parasitology, Laboratory of Systematic Zoology; Wita Stwosza 59, 80 - 308 Gdańsk, Poland.</mods:affiliation>
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<mods:title id="320A9BDFC07FB7794AF0F235CB8F4C3D">Zootaxa</mods:title>
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<mods:date id="A51D63FCF1C19DF5C4C3C0D5F8FD3133">2022</mods:date>
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<treatment id="03A487C8FFC3FFAFFF59FA83EF53AAB2" ID-DOI="http://doi.org/10.5281/zenodo.6506262" ID-GBIF-Taxon="194877469" ID-Zenodo-Dep="6506262" LSID="urn:lsid:zoobank.org:act:0FFBC66B-AF75-4301-ACA1-96195E22440C" httpUri="http://treatment.plazi.org/id/03A487C8FFC3FFAFFF59FA83EF53AAB2" lastPageId="13" lastPageNumber="517" pageId="10" pageNumber="515">
<subSubSection id="C3176555FFC3FFA8FF59FA83EF56AD4E" pageId="10" pageNumber="515" type="nomenclature">
<paragraph id="8BB236DEFFC3FFA8FF59FA83EAFCAD2A" blockId="10.[151,1043,1302,1401]" box="[151,441,1302,1329]" pageId="10" pageNumber="515">
<heading id="D0FA81B2FFC3FFA8FF59FA83EAFCAD2A" bold="true" box="[151,441,1302,1329]" fontSize="11" level="1" pageId="10" pageNumber="515" reason="1">
<emphasis id="B979EACCFFC3FFA8FF59FA83EAFCAD2A" bold="true" box="[151,441,1302,1329]" pageId="10" pageNumber="515">
<taxonomicName id="4C0D4D5DFFC3FFA8FF59FA83EA11AD2A" ID-CoL="9J2NV" authority="Dantas &amp; Amat &amp; Hamada &amp; Giłka, 2022" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[151,340,1302,1329]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="lulu" status="sp. nov.">
<emphasis id="B979EACCFFC3FFA8FF59FA83EA11AD2A" bold="true" box="[151,340,1302,1329]" italics="true" pageId="10" pageNumber="515">Tanytarsus lulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A57B7FFC3FFA8FE95FA83EAFCAD2A" box="[347,441,1303,1329]" pageId="10" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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LSID:
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urn:lsid:zoobank.org:act:
<uuid id="FFAB0C0BFFC3FFA8FE3AFAAEEF56AD4E" box="[500,1043,1338,1365]" pageId="10" pageNumber="515">0FFBC66B-AF75-4301-ACA1-96195E22440C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3176555FFC3FFA8FF59FACBEA5AAD62" box="[151,287,1374,1401]" pageId="10" pageNumber="515" type="description">
<paragraph id="8BB236DEFFC3FFA8FF59FACBEA5AAD62" blockId="10.[151,1043,1302,1401]" box="[151,287,1374,1401]" pageId="10" pageNumber="515">
(
<figureCitation id="13362A5BFFC3FFA8FF6EFACBEA52AD62" box="[160,279,1374,1401]" captionStart="FIGURE 5" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[151,1435,181,1871]" captionTargetId="figure-20@11.[151,1435,181,1878]" captionTargetPageId="11" captionText="FIGURE 5. Tanytarsus lulu sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: anal point; D: superior volsella and digitus; E: median volsella; (CE magnified ca. 34 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504154" httpUri="https://zenodo.org/record/6504154/files/figure.png" pageId="10" pageNumber="515">Fig. 5AE</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3176555FFC3FFA8FF59FA33EAF8AE12" pageId="10" pageNumber="515" type="materials_examined">
<paragraph id="8BB236DEFFC3FFA8FF59FA33EAF8AE12" blockId="10.[151,1437,1446,2013]" pageId="10" pageNumber="515">
<emphasis id="B979EACCFFC3FFA8FF59FA33EA02ADDB" bold="true" box="[151,327,1446,1473]" pageId="10" pageNumber="515">Type material.</emphasis>
<materialsCitation id="3B653C83FFC3FFA8FE9EFA33EFFAADFD" ID-GBIF-Occurrence="3757950307" collectingDate="2021-01-01" collectingDateMax="2021-01-03" collectingDateMin="2021-01-01" collectionCode="R" collectorName="G. P. S. Dantas &amp; Hernandez" country="Colombia" elevation="206" latitude="4.136389" location="Puerto Lopez" longLatPrecision="21" longitude="-72.88139" pageId="10" pageNumber="515" specimenCount="1" stateProvince="Meta" typeStatus="holotype">
<typeStatus id="54B6887CFFC3FFA8FE9EFA33EAFEADDA" box="[336,443,1447,1473]" pageId="10" pageNumber="515" type="holotype">Holotype</typeStatus>
, adult male:
<collectingCountry id="F31A764EFFC3FFA8FD9EFA33E9ADADDA" box="[592,744,1447,1473]" name="Colombia" pageId="10" pageNumber="515">COLOMBIA</collectingCountry>
,
<collectingRegion id="49C9F83CFFC3FFA8FD3BFA33E8FAADDA" box="[757,959,1447,1473]" country="Colombia" name="Meta" pageId="10" pageNumber="515">Meta Department</collectingRegion>
,
<location id="8ED26005FFC3FFA8FC02FA33EF22ADDA" LSID="urn:lsid:plazi:treatment:03A487C8FFC3FFAFFF59FA83EF53AAB2:8ED26005FFC3FFA8FC02FA33EF22ADDA" box="[972,1127,1447,1473]" country="Colombia" latitude="4.136389" longLatPrecision="21" longitude="-72.88139" name="Puerto Lopez" pageId="10" pageNumber="515" stateProvince="Meta">Puerto Lopez</location>
,
<geoCoordinate id="EE395019FFC3FFA8FBBAFA33EFBBADDA" box="[1140,1278,1447,1473]" degrees="04" direction="north" minutes="08" orientation="latitude" pageId="10" pageNumber="515" precision="15" seconds="11" value="4.136389">04º0811N</geoCoordinate>
<geoCoordinate id="EE395019FFC3FFA8FAC9FA32EEDDADDA" box="[1287,1432,1446,1473]" degrees="72" direction="west" minutes="52" orientation="longitude" pageId="10" pageNumber="515" precision="15" seconds="53" value="-72.88139">72º5253W</geoCoordinate>
,
<elevation id="0020D1EDFFC3FFA8FF59FA5FEA5BADFE" box="[151,286,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="10" pageNumber="515" unit="m" value="206.0">
<quantity id="4CF59B3BFFC3FFA8FF59FA5FEBA5ADFE" box="[151,224,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="10" pageNumber="515" unit="m" value="206.0">206 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB3101EFFC3FFA8FEE2FA5FE953ADFE" box="[300,534,1483,1509]" pageId="10" pageNumber="515" value="2021-01-01" valueMax="2021-01-03" valueMin="2021-01-01">
<collectingDate id="EFF7E9F6FFC3FFA8FEE2FA5FE953ADFE" box="[300,534,1483,1509]" pageId="10" pageNumber="515" value="2021-01-01" valueMax="2021-01-03" valueMin="2021-01-01">0103 January 2021</collectingDate>
</date>
,
<collectingMethod id="524C4EC9FFC3FFA8FDECFA5FE9F7ADFE" box="[546,690,1483,1509]" pageId="10" pageNumber="515">Malaise trap</collectingMethod>
,
<collectorName id="26F85308FFC3FFA8FD71FA5FE81BADFE" box="[703,862,1483,1509]" pageId="10" pageNumber="515">G.P.S. Dantas</collectorName>
, S.M.
<collectionCode id="ED1CAE1BFFC3FFA8FC6EFA5FE8F2ADFE" box="[928,951,1483,1509]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="515">R</collectionCode>
.
<collectorName id="26F85308FFC3FFA8FC0DFA5FEF79ADFE" box="[963,1084,1483,1509]" pageId="10" pageNumber="515">Hernández</collectorName>
(CETdeA)
</materialsCitation>
.
<materialsCitation id="3B653C83FFC3FFA8FB05FA5FEAFCAE12" ID-GBIF-Occurrence="3757950306" collectingDate="2021-01-01" collectingDateMax="2021-01-03" collectingDateMin="2021-01-01" collectionCode="R" collectorName="G. P. S. Dantas &amp; Hernandez" country="Colombia" elevation="206" latitude="4.136389" location="Puerto Lopez" longLatPrecision="21" longitude="-72.88139" pageId="10" pageNumber="515" specimenCount="1" specimenCount-male="1" stateProvince="Meta" typeStatus="paratype">
<typeStatus id="54B6887CFFC3FFA8FB05FA5FEE79ADFE" box="[1227,1340,1483,1509]" pageId="10" pageNumber="515" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0BFD57FFC3FFA8FA86FA5FEEDDADFE" box="[1352,1432,1483,1509]" pageId="10" pageNumber="515" type="male">1 male</specimenCount>
, same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176555FFC3FFA8FF09F987E810AE82" pageId="10" pageNumber="515" type="etymology">
<paragraph id="8BB236DEFFC3FFA8FF09F987E810AE82" blockId="10.[151,1437,1446,2013]" pageId="10" pageNumber="515">
<emphasis id="B979EACCFFC3FFA8FF09F987EADAAE36" bold="true" box="[199,415,1555,1581]" italics="true" pageId="10" pageNumber="515">Derivatio nominis.</emphasis>
The name is a tribute to the couple, Luz Helena Rangel and Luiz M. Hernández, for their invaluable help during fieldwork in Puerto Lopez, and especially for all the support and warm welcome given during the first authors stay in
<collectingCountry id="F31A764EFFC3FFA8FE36F9CFE92FAE6E" box="[504,618,1627,1653]" name="Colombia" pageId="10" pageNumber="515">Colombia</collectingCountry>
. The specific epithet is formed by the junction of the first syllable of Luz and Luiz (
<taxonomicName id="4C0D4D5DFFC3FFA8FEC4F9EBEA73AE82" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[266,310,1663,1689]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC3FFA8FEC4F9EBEA73AE82" box="[266,310,1663,1689]" italics="true" pageId="10" pageNumber="515">lulu</emphasis>
</taxonomicName>
) and should be considered a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3176555FFC3FFA8FF09F936EAFDAF6A" pageId="10" pageNumber="515" type="diagnosis">
<paragraph id="8BB236DEFFC3FFA8FF09F936EAFDAF6A" blockId="10.[151,1437,1446,2013]" pageId="10" pageNumber="515">
<emphasis id="B979EACCFFC3FFA8FF09F936EA7FAEA7" bold="true" box="[199,314,1698,1724]" pageId="10" pageNumber="515">Diagnosis</emphasis>
. AR&gt; 1. Frontal tubercles well-developed, over two times as long as wide. Mid leg with apical row of 47 sensilla chaetica. Tergite IX with lateral teeth. Orolateral spines of laterosternite IX in shape of small processes. Anal point with apex spoon-shaped, smooth, lacking serrations. Superior volsella oval, with posteromedian corner slightly projecting rounded, median margin slightly concave. Digitus pointed, extending slightly beyond median margin of superior volsella. Median volsella with 23 setiform and 3 pectinate lamellae. Inferior volsella with angular dorsal protrusion.
</paragraph>
</subSubSection>
<subSubSection id="C3176555FFC3FFAEFF09F8EEE94CADEC" lastPageId="12" lastPageNumber="517" pageId="10" pageNumber="515" type="description">
<paragraph id="8BB236DEFFC3FFA8FF09F8EEE973AF8E" blockId="10.[151,1437,1446,2013]" box="[199,566,1914,1941]" pageId="10" pageNumber="515">
<emphasis id="B979EACCFFC3FFA8FF09F8EEEA1CAF8F" bold="true" box="[199,345,1914,1940]" pageId="10" pageNumber="515">Description.</emphasis>
Adult male (n = 2).
</paragraph>
<paragraph id="8BB236DEFFC3FFA8FF09F80BE828AFC6" blockId="10.[151,1437,1446,2013]" pageId="10" pageNumber="515">
<emphasis id="B979EACCFFC3FFA8FF09F80BEAB5AFA3" box="[199,496,1951,1977]" italics="true" pageId="10" pageNumber="515">Body size and proportions</emphasis>
. Total length
<quantity id="4CF59B3BFFC3FFA8FD41F80BE873AFA2" box="[655,822,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.86" metricValueMin="2.34" pageId="10" pageNumber="515" unit="mm" value="2.5999999999999996" valueMax="2.86" valueMin="2.34">2.342.86 mm</quantity>
. Wing length
<quantity id="4CF59B3BFFC3FFA8FC1BF80BEF39AFA2" box="[981,1148,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="1.3399999999999999" metricValueMax="1.46" metricValueMin="1.22" pageId="10" pageNumber="515" unit="mm" value="1.3399999999999999" valueMax="1.46" valueMin="1.22">1.221.46 mm</quantity>
. Total length/wing length ratio 1.921.96. Wing length/length of profemur ratio 2.142.26.
</paragraph>
<caption id="DF726656FFC2FFA9FF59F8F9EF7CAFB1" ID-DOI="http://doi.org/10.5281/zenodo.6504154" ID-Zenodo-Dep="6504154" httpUri="https://zenodo.org/record/6504154/files/figure.png" pageId="11" pageNumber="516" startId="11.[151,250,1901,1925]" targetBox="[151,1435,181,1871]" targetPageId="11">
<paragraph id="8BB236DEFFC2FFA9FF59F8F9EF7CAFB1" blockId="11.[151,1436,1901,1962]" pageId="11" pageNumber="516">
<emphasis id="B979EACCFFC2FFA9FF59F8F9EA48AF9D" bold="true" box="[151,269,1901,1926]" pageId="11" pageNumber="516">FIGURE 5</emphasis>
.
<taxonomicName id="4C0D4D5DFFC2FFA9FED4F8FAEAF3AF9D" authority="Dantas &amp; Amat &amp; Hamada &amp; Giłka, 2022" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[282,438,1902,1926]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="516" phylum="Arthropoda" rank="species" species="lulu" status="sp. nov.">
<emphasis id="B979EACCFFC2FFA9FED4F8FAEAF3AF9D" box="[282,438,1902,1926]" italics="true" pageId="11" pageNumber="516">Tanytarsus lulu</emphasis>
</taxonomicName>
<emphasis id="B979EACCFFC2FFA9FE73F8FAE94BAF9D" bold="true" box="[445,526,1902,1926]" pageId="11" pageNumber="516">
<taxonomicNameLabel id="A24A57B7FFC2FFA9FE73F8FAE94BAF9D" box="[445,526,1902,1926]" pageId="11" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B979EACCFFC2FFA9FD94F8F9E929AF9E" bold="true" box="[602,620,1901,1925]" pageId="11" pageNumber="516">A</emphasis>
,
<emphasis id="B979EACCFFC2FFA9FDB4F8FAE9CEAF9D" bold="true" box="[634,651,1902,1926]" pageId="11" pageNumber="516">B</emphasis>
: hypopygium in dorsal (
<emphasis id="B979EACCFFC2FFA9FC49F8F9E8DCAF9E" bold="true" box="[903,921,1901,1925]" pageId="11" pageNumber="516">A</emphasis>
) and ventral aspect (
<emphasis id="B979EACCFFC2FFA9FBA1F8FAEFC5AF9D" bold="true" box="[1135,1152,1902,1926]" pageId="11" pageNumber="516">B</emphasis>
);
<emphasis id="B979EACCFFC2FFA9FB58F8F9EFEDAF9E" bold="true" box="[1174,1192,1901,1925]" pageId="11" pageNumber="516">C</emphasis>
: anal point;
<emphasis id="B979EACCFFC2FFA9FAE7F8FAEE7EAF9D" bold="true" box="[1321,1339,1902,1926]" pageId="11" pageNumber="516">D</emphasis>
: superior volsella and digitus;
<emphasis id="B979EACCFFC2FFA9FEA4F806EA3EAFB1" bold="true" box="[362,379,1938,1962]" pageId="11" pageNumber="516">E</emphasis>
: median volsella; (CE magnified
<emphasis id="B979EACCFFC2FFA9FD10F806E9B8AFB2" box="[734,765,1938,1961]" italics="true" pageId="11" pageNumber="516">ca.</emphasis>
34 times relative to A and B).
</paragraph>
</caption>
<paragraph id="8BB236DEFFC5FFAEFF09FF03EAC4A8E2" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09FF03EA15A8AA" box="[199,336,151,177]" italics="true" pageId="12" pageNumber="517">Colouration</emphasis>
. Eyes black. Antenna brown. Head capsule and thorax yellow to light brown. Legs: proximal half of femora pale, distal half of femora plus tibiae and tarsi light brown. Wing veins yellowish brown, membrane pale. Abdomen yellowish.
</paragraph>
<paragraph id="8BB236DEFFC5FFAEFF09FE97EAC8A9B6" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09FE97EA41A906" box="[199,260,259,285]" italics="true" pageId="12" pageNumber="517">Head</emphasis>
. Eyes bare, with moderately developed dorsomedian extensions. Antenna with 13 flagellomeres; ultimate flagellomere 425 μm long; AR 1.04. Frontal tubercles well-developed, 2729 μm long, 1215 μm wide, ca. 2.2 times as long as wide. Tentorium 140 μm long. Temporal setae 89 on each side. Clypeus with 1718 setae. Lengths of palpomeres 15 (in μm): 2627, 2627, 6780, 84106, 153169; third palpomere with 1 subapical sensillum clavatum 14 μm long.
</paragraph>
<paragraph id="8BB236DEFFC5FFAEFF09FE23EFBDA9EE" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09FE23EA5DA9CA" box="[199,280,439,465]" italics="true" pageId="12" pageNumber="517">Thorax</emphasis>
. Ac 12, restricted to anterior region of scutum; Dc 67 on each side, uniserial; Pa 1 on each side; Scts 4. Scutum projected and rounded anteriorly, overreaching antepronotum. Halteres with at least 4 setae.
</paragraph>
<paragraph id="8BB236DEFFC5FFAEFF09FE6BEE69AA9E" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09FE6BEA44AA03" box="[199,257,511,536]" italics="true" pageId="12" pageNumber="517">Wing</emphasis>
. Somewhat oblong, with anal lobe reduced. Brachiolum with 1 seta, Sc bare, R with 1618 setae, R
<subScript id="1789349BFFC5FFAEFA92FD85EE21AA04" attach="left" box="[1372,1380,529,543]" fontSize="6" pageId="12" pageNumber="517">1</subScript>
with 1320 setae, R
<subScript id="1789349BFFC5FFAEFEF1FDA1EA1CAA58" attach="left" box="[319,345,565,579]" fontSize="6" pageId="12" pageNumber="517">4+5</subScript>
with 1118 setae, M bare, M
<subScript id="1789349BFFC5FFAEFD66FDA1E987AA58" attach="left" box="[680,706,565,579]" fontSize="6" pageId="12" pageNumber="517">1+2</subScript>
with 3540 setae, M
<subScript id="1789349BFFC5FFAEFC7CFDA1E889AA58" attach="left" box="[946,972,565,579]" fontSize="6" pageId="12" pageNumber="517">3+4</subScript>
with 30 setae, Cu with 4 setae, Cu
<subScript id="1789349BFFC5FFAEFA94FDA1EE27AA58" attach="left" box="[1370,1378,565,579]" fontSize="6" pageId="12" pageNumber="517">1</subScript>
with 712 setae, postcubitus with 34 setae, An with 1320 setae. Cell m bare (false vein bare), r
<subScript id="1789349BFFC5FFAEFB7DFDCDEF88AA7C" attach="left" box="[1203,1229,601,615]" fontSize="6" pageId="12" pageNumber="517">4+5</subScript>
with 7580 setae, m
<subScript id="1789349BFFC5FFAEFF63FDE9EB83AA90" attach="left" box="[173,198,637,651]" fontSize="6" pageId="12" pageNumber="517">1+2</subScript>
with 7080 setae (+48 setae on false vein), m
<subScript id="1789349BFFC5FFAEFD07FDE9E9A6AA90" attach="left" box="[713,739,637,651]" fontSize="6" pageId="12" pageNumber="517">3+4</subScript>
with 2028 setae, cu and an bare. VRCu 1.201.23.
</paragraph>
<paragraph id="8BB236DEFFC5FFAEFF09FD1BE845AB0E" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09FD1BEBB9AAB3" box="[199,252,655,680]" italics="true" pageId="12" pageNumber="517">Legs</emphasis>
. Foreleg tibia with short lanceolate spur 1520 μm long. Lengths of combs of mid tibia 89 μm (with 1112 μm long spur) and 1317 μm (with 1521 μm long spur); lengths of combs of hind tibia 1415 μm (with 1718 μm spur) and 1719 μm (with 2225 μm long spur). Basitarsus of mid leg with apical row of 47 sensilla chaetica. Lengths and proportions of legs as in
<tableCitation id="C68F0365FFC5FFAEFD69FD6FE9B9AB0E" box="[679,764,763,789]" captionStart="TABLE 4" captionStartId="12.[151,239,836,860]" captionTargetPageId="12" captionText="TABLE 4. Lengths (in μm) and proportions of legs of Tanytarsus lulu sp. nov., male (n = 2)." pageId="12" pageNumber="517">Table 4</tableCitation>
.
</paragraph>
<caption id="DF726656FFC5FFAEFF59FCD0EF33AB46" box="[151,1142,836,861]" pageId="12" pageNumber="517" startId="12.[151,239,836,860]" targetBox="[166,1416,877,1027]" targetIsTable="true" targetPageId="12">
<paragraph id="8BB236DEFFC5FFAEFF59FCD0EF33AB46" blockId="12.[151,1142,836,861]" box="[151,1142,836,861]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF59FCD0EA4FAB47" bold="true" box="[151,266,836,860]" pageId="12" pageNumber="517">TABLE 4.</emphasis>
Lengths (in μm) and proportions of legs of
<taxonomicName id="4C0D4D5DFFC5FFAEFD11FCD1E8C7AB47" authority="Dantas &amp; Amat &amp; Hamada &amp; Giłka, 2022" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[735,898,836,860]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="lulu" status="sp. nov.">
<emphasis id="B979EACCFFC5FFAEFD11FCD1E8C7AB47" box="[735,898,836,860]" italics="true" pageId="12" pageNumber="517">Tanytarsus lulu</emphasis>
</taxonomicName>
<emphasis id="B979EACCFFC5FFAEFC47FCD1E89AAB46" bold="true" box="[905,991,837,861]" pageId="12" pageNumber="517">
<taxonomicNameLabel id="A24A57B7FFC5FFAEFC47FCD1E89AAB46" box="[905,991,837,861]" pageId="12" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (n = 2).
</paragraph>
</caption>
<paragraph id="8BB236DEFFC5FFAEFF29FCF9EECDABE6" pageId="12" pageNumber="517">
<table id="F90DC47EFFC5005DFF68FCF9EECDAC18" box="[166,1416,877,1027]" colsContinueFrom="8.[166,1378,1356,1507]" gridcols="11" gridrows="4" pageId="12" pageNumber="517">
<tr id="353D349CFFC5005DFF68FCF9EECDAB97" box="[166,1416,877,908]" gridrow="0" pageId="12" pageNumber="517" rowspan-0="1">
<th id="76EC5DE0FFC5005DFF29FCF9EA05AB97" box="[231,320,877,908]" gridcol="1" gridrow="0" pageId="12" pageNumber="517">fe</th>
<th id="76EC5DE0FFC5005DFE90FCF9EAF2AB97" box="[350,439,877,908]" gridcol="2" gridrow="0" pageId="12" pageNumber="517">ti</th>
<th id="76EC5DE0FFC5005DFE18FCF9E96AAB97" box="[470,559,877,908]" gridcol="3" gridrow="0" pageId="12" pageNumber="517">ta1</th>
<th id="76EC5DE0FFC5005DFD9FFCF9E9EFAB97" box="[593,682,877,908]" gridcol="4" gridrow="0" pageId="12" pageNumber="517">ta2</th>
<th id="76EC5DE0FFC5005DFD04FCF9E866AB97" box="[714,803,877,908]" gridcol="5" gridrow="0" pageId="12" pageNumber="517">ta3</th>
<th id="76EC5DE0FFC5005DFC8EFCF9E8DCAB97" box="[832,921,877,908]" gridcol="6" gridrow="0" pageId="12" pageNumber="517">ta4</th>
<th id="76EC5DE0FFC5005DFC78FCF9E8B3AB97" box="[950,1014,877,908]" gridcol="7" gridrow="0" pageId="12" pageNumber="517">ta 5</th>
<th id="76EC5DE0FFC5005DFBD5FCF9EFC4AB97" box="[1051,1153,877,908]" gridcol="8" gridrow="0" pageId="12" pageNumber="517">LR</th>
<th id="76EC5DE0FFC5005DFB51FCF9EE40AB97" box="[1183,1285,877,908]" gridcol="9" gridrow="0" pageId="12" pageNumber="517">BV</th>
<th id="76EC5DE0FFC5005DFAEDFCF9EECDAB97" box="[1315,1416,877,908]" gridcol="10" gridrow="0" pageId="12" pageNumber="517">SV</th>
</tr>
<tr id="353D349CFFC5005DFF68FC01EECDABA8" box="[166,1416,917,947]" gridrow="1" pageId="12" pageNumber="517">
<th id="76EC5DE0FFC5005DFF68FC01EBFCABA8" box="[166,185,917,947]" gridcol="0" gridrow="1" pageId="12" pageNumber="517">p 1</th>
<td id="76EC5DE0FFC5005DFF29FC01EA05ABA8" box="[231,320,917,947]" gridcol="1" gridrow="1" pageId="12" pageNumber="517">543681</td>
<td id="76EC5DE0FFC5005DFE90FC01EAF2ABA8" box="[350,439,917,947]" gridcol="2" gridrow="1" pageId="12" pageNumber="517">265330</td>
<td id="76EC5DE0FFC5005DFE18FC01E96AABA8" box="[470,559,917,947]" gridcol="3" gridrow="1" pageId="12" pageNumber="517">758</td>
<td id="76EC5DE0FFC5005DFD9FFC01E9EFABA8" box="[593,682,917,947]" gridcol="4" gridrow="1" pageId="12" pageNumber="517">392</td>
<td id="76EC5DE0FFC5005DFD04FC01E866ABA8" box="[714,803,917,947]" gridcol="5" gridrow="1" pageId="12" pageNumber="517">323</td>
<td id="76EC5DE0FFC5005DFC8EFC01E8DCABA8" box="[832,921,917,947]" gridcol="6" gridrow="1" pageId="12" pageNumber="517">230</td>
<td id="76EC5DE0FFC5005DFC78FC01E8B3ABA8" box="[950,1014,917,947]" gridcol="7" gridrow="1" pageId="12" pageNumber="517">81</td>
<td id="76EC5DE0FFC5005DFBD5FC01EFC4ABA8" box="[1051,1153,917,947]" gridcol="8" gridrow="1" pageId="12" pageNumber="517">2.86</td>
<td id="76EC5DE0FFC5005DFB51FC01EE40ABA8" box="[1183,1285,917,947]" gridcol="9" gridrow="1" pageId="12" pageNumber="517">1.53</td>
<td id="76EC5DE0FFC5005DFAEDFC01EECDABA8" box="[1315,1416,917,947]" gridcol="10" gridrow="1" pageId="12" pageNumber="517">1.07</td>
</tr>
<tr id="353D349CFFC5005DFF68FC29EECDABC0" box="[166,1416,957,987]" gridrow="2" pageId="12" pageNumber="517">
<th id="76EC5DE0FFC5005DFF68FC29EBFCABC0" box="[166,185,957,987]" gridcol="0" gridrow="2" pageId="12" pageNumber="517">p2</th>
<td id="76EC5DE0FFC5005DFF29FC29EA05ABC0" box="[231,320,957,987]" gridcol="1" gridrow="2" pageId="12" pageNumber="517">580721</td>
<td id="76EC5DE0FFC5005DFE90FC29EAF2ABC0" box="[350,439,957,987]" gridcol="2" gridrow="2" pageId="12" pageNumber="517">446558</td>
<td id="76EC5DE0FFC5005DFE18FC29E96AABC0" box="[470,559,957,987]" gridcol="3" gridrow="2" pageId="12" pageNumber="517">287347</td>
<td id="76EC5DE0FFC5005DFD9FFC29E9EFABC0" box="[593,682,957,987]" gridcol="4" gridrow="2" pageId="12" pageNumber="517">135166</td>
<td id="76EC5DE0FFC5005DFD04FC29E866ABC0" box="[714,803,957,987]" gridcol="5" gridrow="2" pageId="12" pageNumber="517">100121</td>
<td id="76EC5DE0FFC5005DFC8EFC29E8DCABC0" box="[832,921,957,987]" gridcol="6" gridrow="2" pageId="12" pageNumber="517">7077</td>
<td id="76EC5DE0FFC5005DFC78FC29E8B3ABC0" box="[950,1014,957,987]" gridcol="7" gridrow="2" pageId="12" pageNumber="517">5157</td>
<td id="76EC5DE0FFC5005DFBD5FC29EFC4ABC0" box="[1051,1153,957,987]" gridcol="8" gridrow="2" pageId="12" pageNumber="517">0.620.64</td>
<td id="76EC5DE0FFC5005DFB51FC29EE40ABC0" box="[1183,1285,957,987]" gridcol="9" gridrow="2" pageId="12" pageNumber="517">3.693.86</td>
<td id="76EC5DE0FFC5005DFAEDFC29EECDABC0" box="[1315,1416,957,987]" gridcol="10" gridrow="2" pageId="12" pageNumber="517">3.573.69</td>
</tr>
<tr id="353D349CFFC5005DFF68FC71EECDAC18" box="[166,1416,997,1027]" gridrow="3" pageId="12" pageNumber="517">
<th id="76EC5DE0FFC5005DFF68FC71EBFCAC18" box="[166,185,997,1027]" gridcol="0" gridrow="3" pageId="12" pageNumber="517">p3</th>
<td id="76EC5DE0FFC5005DFF29FC71EA05AC18" box="[231,320,997,1027]" gridcol="1" gridrow="3" pageId="12" pageNumber="517">600732</td>
<td id="76EC5DE0FFC5005DFE90FC71EAF2AC18" box="[350,439,997,1027]" gridcol="2" gridrow="3" pageId="12" pageNumber="517">597732</td>
<td id="76EC5DE0FFC5005DFE18FC71E96AAC18" box="[470,559,997,1027]" gridcol="3" gridrow="3" pageId="12" pageNumber="517">420500</td>
<td id="76EC5DE0FFC5005DFD9FFC71E9EFAC18" box="[593,682,997,1027]" gridcol="4" gridrow="3" pageId="12" pageNumber="517">255294</td>
<td id="76EC5DE0FFC5005DFD04FC71E866AC18" box="[714,803,997,1027]" gridcol="5" gridrow="3" pageId="12" pageNumber="517">223266</td>
<td id="76EC5DE0FFC5005DFC8EFC71E8DCAC18" box="[832,921,997,1027]" gridcol="6" gridrow="3" pageId="12" pageNumber="517">147169</td>
<td id="76EC5DE0FFC5005DFC78FC71E8B3AC18" box="[950,1014,997,1027]" gridcol="7" gridrow="3" pageId="12" pageNumber="517">7692</td>
<td id="76EC5DE0FFC5005DFBD5FC71EFC4AC18" box="[1051,1153,997,1027]" gridcol="8" gridrow="3" pageId="12" pageNumber="517">0.680.70</td>
<td id="76EC5DE0FFC5005DFB51FC71EE40AC18" box="[1183,1285,997,1027]" gridcol="9" gridrow="3" pageId="12" pageNumber="517">2.312.39</td>
<td id="76EC5DE0FFC5005DFAEDFC71EECDAC18" box="[1315,1416,997,1027]" gridcol="10" gridrow="3" pageId="12" pageNumber="517">2.852.93</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB236DEFFC5FFAEFF09FBBAE94CADEC" blockId="12.[151,1437,1069,2031]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09FBBAEA13AC5C" box="[199,342,1070,1095]" italics="true" pageId="12" pageNumber="517">Hypopygium</emphasis>
. Tergite IX covered with microtrichia except for small bare area at base of anal point; lateral teeth present; orolateral spines of laterosternite IX in shape of small processes; anal tergite bands T-type, fused on median part of tergite, ending close to anal point base (
<figureCitation id="13362A5BFFC5FFAEFCE1FBE1E8EEAC8B" box="[815,939,1141,1168]" captionStart="FIGURE 5" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[151,1435,181,1871]" captionTargetId="figure-20@11.[151,1435,181,1878]" captionTargetPageId="11" captionText="FIGURE 5. Tanytarsus lulu sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: anal point; D: superior volsella and digitus; E: median volsella; (CE magnified ca. 34 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504154" httpUri="https://zenodo.org/record/6504154/files/figure.png" pageId="12" pageNumber="517">Fig. 5A, B</figureCitation>
). Anal point 3540 μm long, elongate, with spoon-shaped apex with margin smooth (without serrations), pair of dorsal setae at base, two lateral setae on each side and pair of ventral setae subapically; microtrichia and spinulae absent (
<figureCitation id="13362A5BFFC5FFAEFC34FB29EF3DACCC" box="[1018,1144,1213,1240]" captionStart="FIGURE 5" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[151,1435,181,1871]" captionTargetId="figure-20@11.[151,1435,181,1878]" captionTargetPageId="11" captionText="FIGURE 5. Tanytarsus lulu sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: anal point; D: superior volsella and digitus; E: median volsella; (CE magnified ca. 34 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504154" httpUri="https://zenodo.org/record/6504154/files/figure.png" pageId="12" pageNumber="517">Fig. 5A, C</figureCitation>
). Superior volsella 3235 μm long, oval, posteromedian corner slightly projecting and rounded, median margin slightly concave; 67 setae on dorsal surface, 2 setae on median margin, and 1 ventral seta; field of microtrichia on dorsal surface absent; digitus 2123 μm long, pointed, slightly extending beyond median margin of superior volsella (
<figureCitation id="13362A5BFFC5FFAEFB44FABDEE68AD5F" box="[1162,1325,1321,1348]" captionStart="FIGURE 5" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[151,1435,181,1871]" captionTargetId="figure-20@11.[151,1435,181,1878]" captionTargetPageId="11" captionText="FIGURE 5. Tanytarsus lulu sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: anal point; D: superior volsella and digitus; E: median volsella; (CE magnified ca. 34 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504154" httpUri="https://zenodo.org/record/6504154/files/figure.png" pageId="12" pageNumber="517">Fig. 5A, B, D</figureCitation>
). Stem of median volsella 1315 μm long, with 23 setiform and 3 pectinate lamellae (
<figureCitation id="13362A5BFFC5FFAEFBC8FAD9EFC5AD73" box="[1030,1152,1357,1384]" captionStart="FIGURE 5" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[151,1435,181,1871]" captionTargetId="figure-20@11.[151,1435,181,1878]" captionTargetPageId="11" captionText="FIGURE 5. Tanytarsus lulu sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: anal point; D: superior volsella and digitus; E: median volsella; (CE magnified ca. 34 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504154" httpUri="https://zenodo.org/record/6504154/files/figure.png" pageId="12" pageNumber="517">Fig. 5B, E</figureCitation>
). Inferior volsella 75 μm long, covered with microtrichia, slightly curved and posteromedially directed, apex with angular dorsal protrusion. Phallapodeme ~100105 μm long, strongly sinuous; transverse sternapodeme ~3550 μm long, with well-developed oral projections. Gonocoxite 85100 μm long. Gonostylus 125 μm long, narrow, slightly curved, tapering towards pointed apex. HR 0.80. HV 2.29.
</paragraph>
</subSubSection>
<subSubSection id="C3176555FFC5FFAEFF09F995EA75AE7F" pageId="12" pageNumber="517" type="distribution">
<paragraph id="8BB236DEFFC5FFAEFF09F995EA75AE7F" blockId="12.[151,1437,1069,2031]" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09F995E913AE00" bold="true" box="[199,598,1537,1563]" pageId="12" pageNumber="517">Distribution and ecological notes.</emphasis>
The adult male specimens examined were collected together with those of three other species described in the present paper. For details on ecology and bionomics see notes under
<taxonomicName id="4C0D4D5DFFC5FFAEFAECF9B2EA6CAE78" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="colombiensis">
<emphasis id="B979EACCFFC5FFAEFAECF9B2EA6CAE78" italics="true" pageId="12" pageNumber="517">Tanytarsus colombiensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176555FFC5FFAFFF09F9F9EF53AAB2" lastPageId="13" lastPageNumber="518" pageId="12" pageNumber="517" type="discussion">
<paragraph id="8BB236DEFFC5FFAFFF09F9F9EF53AAB2" blockId="12.[151,1437,1069,2031]" lastBlockId="13.[151,1437,151,681]" lastPageId="13" lastPageNumber="518" pageId="12" pageNumber="517">
<emphasis id="B979EACCFFC5FFAEFF09F9F9EA00AE9C" bold="true" box="[199,325,1645,1671]" pageId="12" pageNumber="517">Discussion</emphasis>
.
<taxonomicName id="4C0D4D5DFFC5FFAEFE97F9FAE94EAE9C" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[345,523,1645,1671]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC5FFAEFE97F9FAE94EAE9C" box="[345,523,1645,1671]" italics="true" pageId="12" pageNumber="517">Tanytarsus lulu</emphasis>
</taxonomicName>
and several Neotropical species, i.e.
<taxonomicName id="4C0D4D5DFFC5FFAEFC04F9FAEF08AE9C" authorityName="Reiss" authorityYear="1972" box="[970,1101,1645,1671]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFC04F9FAEF08AE9C" box="[970,1101,1645,1671]" italics="true" pageId="12" pageNumber="517">T. ligulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0D4D5DFFC5FFAEFBAFF9FAEE5DAE9C" authorityName="Reiss" authorityYear="1972" box="[1121,1304,1645,1671]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="paraligulatus">
<emphasis id="B979EACCFFC5FFAEFBAFF9FAEE5DAE9C" box="[1121,1304,1645,1671]" italics="true" pageId="12" pageNumber="517">T. paraligulatus</emphasis>
</taxonomicName>
as well as “
<taxonomicName id="4C0D4D5DFFC5FFAEFF6AF906EAEDAEB0" authorityName="Reiss" authorityYear="1972" box="[164,424,1681,1707]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFF6AF906EA5BAEB0" box="[164,286,1682,1707]" italics="true" pageId="12" pageNumber="517">Tanytarsus</emphasis>
cf.
<emphasis id="B979EACCFFC5FFAEFE88F905EAEDAEB0" box="[326,424,1681,1707]" italics="true" pageId="12" pageNumber="517">ligulatus</emphasis>
</taxonomicName>
” [pre-named and described by Sanseverino (2006), but finally not delimited as new species nor yet validated] are distinct by the anal point crests forming an extraordinary structure—the round, spoon-shaped apex (cf.
<bibRefCitation id="EF9C4B2FFFC5FFAEFECCF94DEAC7AEE8" author="Reiss, F." box="[258,386,1753,1779]" pageId="12" pageNumber="517" pagination="49 - 94" refId="ref13403" refString="Reiss, F. (1972) Die Tanytarsini (Chironomidae, Diptera) Sudchiles und Westpatagoniens. Mit Hinweisen auf die Tanytarsini- Fauna der Neotropis. Studies on Neotropical Fauna, 7, 49 - 94. https: // doi. org / 10.1080 / 01650527209360434" type="journal article" year="1972">Reiss 1972</bibRefCitation>
, Sanseverino 2006). This character state prompted us to treat it as a main apomorphy for the
<taxonomicName id="4C0D4D5DFFC5FFAEFF59F969EBBCAF0C" authorityName="Reiss" authorityYear="1972" box="[151,249,1789,1815]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFF59F969EBBCAF0C" box="[151,249,1789,1815]" italics="true" pageId="12" pageNumber="517">ligulatus</emphasis>
</taxonomicName>
species group, proposed here. Several other diagnostic characters in common among the species mentioned above support this concept. All species analysed have well-developed, long frontal tubercles. They are 3.5 times as long as wide in
<taxonomicName id="4C0D4D5DFFC5FFAEFE89F8D2EA81AF44" authorityName="Reiss" authorityYear="1972" box="[327,452,1861,1887]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFE89F8D2EA81AF44" box="[327,452,1861,1887]" italics="true" pageId="12" pageNumber="517">T. ligulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D5DFFC5FFAEFE34F8D2E9DFAF44" authorityName="Reiss" authorityYear="1972" box="[506,666,1861,1887]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFE34F8D2E94AAF44" box="[506,527,1862,1887]" italics="true" pageId="12" pageNumber="517">T.</emphasis>
cf.
<emphasis id="B979EACCFFC5FFAEFDF6F8D1E9DFAF44" box="[568,666,1861,1887]" italics="true" pageId="12" pageNumber="517">ligulatus</emphasis>
</taxonomicName>
, the ratio is
<quantity id="4CF59B3BFFC5FFAEFCEAF8D1E80BAF44" box="[804,846,1861,1887]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="12" pageNumber="517" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C0D4D5DFFC5FFAEFC9BF8D2EF42AF44" authorityName="Reiss" authorityYear="1972" box="[853,1031,1861,1887]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="paraligulatus">
<emphasis id="B979EACCFFC5FFAEFC9BF8D2EF42AF44" box="[853,1031,1861,1887]" italics="true" pageId="12" pageNumber="517">T. paraligulatus</emphasis>
</taxonomicName>
, while
<quantity id="4CF59B3BFFC5FFAEFB97F8D1EFDCAF44" box="[1113,1177,1861,1887]" metricMagnitude="-2" metricUnit="m" metricValue="5.588" pageId="12" pageNumber="517" unit="in" value="2.2">2.2 in</quantity>
<taxonomicName id="4C0D4D5DFFC5FFAEFB51F8D2EFA3AF44" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[1183,1254,1861,1887]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC5FFAEFB51F8D2EFA3AF44" box="[1183,1254,1861,1887]" italics="true" pageId="12" pageNumber="517">T. lulu</emphasis>
</taxonomicName>
. In the
<taxonomicName id="4C0D4D5DFFC5FFAEFAF4F8D1EED9AF44" authorityName="Reiss" authorityYear="1972" box="[1338,1436,1861,1887]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFAF4F8D1EED9AF44" box="[1338,1436,1861,1887]" italics="true" pageId="12" pageNumber="517">ligulatus</emphasis>
</taxonomicName>
group, the superior volsella is more or less heart-shaped, with the median margin concave slightly at least (strongly concave in
<taxonomicName id="4C0D4D5DFFC5FFAEFED6F81AEAD3AFBC" authorityName="Reiss" authorityYear="1972" box="[280,406,1933,1959]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFED6F81AEAD3AFBC" box="[280,406,1933,1959]" italics="true" pageId="12" pageNumber="517">T. ligulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D5DFFC5FFAEFE03F81AE92AAFBC" authorityName="Reiss" authorityYear="1972" box="[461,623,1933,1959]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFE03F81AEAA7AFBC" box="[461,482,1934,1959]" italics="true" pageId="12" pageNumber="517">T.</emphasis>
cf.
<emphasis id="B979EACCFFC5FFAEFDC3F819E92AAFBC" box="[525,623,1933,1959]" italics="true" pageId="12" pageNumber="517">ligulatus</emphasis>
</taxonomicName>
, nearly straight in
<taxonomicName id="4C0D4D5DFFC5FFAEFC8AF81AE8B2AFBC" authorityName="Reiss" authorityYear="1972" box="[836,1015,1933,1959]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="paraligulatus">
<emphasis id="B979EACCFFC5FFAEFC8AF81AE8B2AFBC" box="[836,1015,1933,1959]" italics="true" pageId="12" pageNumber="517">T. paraligulatus</emphasis>
</taxonomicName>
, slightly concave in
<taxonomicName id="4C0D4D5DFFC5FFAEFB2CF81AEE6FAFBC" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[1250,1322,1933,1959]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC5FFAEFB2CF81AEE6FAFBC" box="[1250,1322,1933,1959]" italics="true" pageId="12" pageNumber="517">T. lulu</emphasis>
</taxonomicName>
). The key character within the
<taxonomicName id="4C0D4D5DFFC5FFAEFEB4F825EA99AFD0" authorityName="Reiss" authorityYear="1972" box="[378,476,1969,1995]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC5FFAEFEB4F825EA99AFD0" box="[378,476,1969,1995]" italics="true" pageId="12" pageNumber="517">ligulatus</emphasis>
</taxonomicName>
group is also the size of the digitus, well-developed in all the species. However,
<taxonomicName id="4C0D4D5DFFC5FFAEFA98F826EED8AFD0" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[1366,1437,1969,1995]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC5FFAEFA98F826EED8AFD0" box="[1366,1437,1969,1995]" italics="true" pageId="12" pageNumber="517">T. lulu</emphasis>
</taxonomicName>
has the digitus pointed, extending slightly beyond the median margin of the superior volsella, while the digitus is finger-like, rounded apically, extending well beyond the superior volsella in
<taxonomicName id="4C0D4D5DFFC4FFAFFC28FF03EF26A8AA" authorityName="Reiss" authorityYear="1972" box="[998,1123,151,177]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFC28FF03EF26A8AA" box="[998,1123,151,177]" italics="true" pageId="13" pageNumber="518">T. ligulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D5DFFC4FFAFFB56FF03EE0CA8AA" authorityName="Reiss" authorityYear="1972" box="[1176,1353,151,177]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="paraligulatus">
<emphasis id="B979EACCFFC4FFAFFB56FF03EE0CA8AA" box="[1176,1353,151,177]" italics="true" pageId="13" pageNumber="518">T. paraligulatus</emphasis>
</taxonomicName>
. In this character
<taxonomicName id="4C0D4D5DFFC4FFAFFECBFF2FEA08A8CE" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[261,333,187,213]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFECBFF2FEA08A8CE" box="[261,333,187,213]" italics="true" pageId="13" pageNumber="518">T. lulu</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C0D4D5DFFC4FFAFFE11FF2FE9C4A8CE" authorityName="Reiss" authorityYear="1972" box="[479,641,187,213]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFE11FF2FEAB1A8CF" box="[479,500,187,212]" italics="true" pageId="13" pageNumber="518">T.</emphasis>
cf.
<emphasis id="B979EACCFFC4FFAFFDD1FF2FE9C4A8CE" box="[543,641,187,213]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
</taxonomicName>
. These two, possibly the closest species share also some other features that differentiate them from both
<taxonomicName id="4C0D4D5DFFC4FFAFFDDCFF4BE9D4A8E2" authorityName="Reiss" authorityYear="1972" box="[530,657,223,249]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFDDCFF4BE9D4A8E2" box="[530,657,223,249]" italics="true" pageId="13" pageNumber="518">T. ligulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D5DFFC4FFAFFD04FF4BE83BA8E2" authorityName="Reiss" authorityYear="1972" box="[714,894,223,249]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="paraligulatus">
<emphasis id="B979EACCFFC4FFAFFD04FF4BE83BA8E2" box="[714,894,223,249]" italics="true" pageId="13" pageNumber="518">T. paraligulatus</emphasis>
</taxonomicName>
: the spoon-shaped anal point apex is round and have no serrations (teeth) on margin in
<taxonomicName id="4C0D4D5DFFC4FFAFFD87FE97E9D5A906" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[585,656,259,285]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFD87FE97E9D5A906" box="[585,656,259,285]" italics="true" pageId="13" pageNumber="518">T. lulu</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D5DFFC4FFAFFD0AFE97E826A906" authorityName="Reiss" authorityYear="1972" box="[708,867,259,285]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFD0AFE97E99CA907" box="[708,729,259,284]" italics="true" pageId="13" pageNumber="518">T.</emphasis>
cf.
<emphasis id="B979EACCFFC4FFAFFCCFFE97E826A906" box="[769,867,259,285]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
</taxonomicName>
, while the apex is flattened, with margin toothed in
<taxonomicName id="4C0D4D5DFFC4FFAFFF59FEB3EA51A95A" authorityName="Reiss" authorityYear="1972" box="[151,276,295,321]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFF59FEB3EA51A95A" box="[151,276,295,321]" italics="true" pageId="13" pageNumber="518">T. ligulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D5DFFC4FFAFFE85FEB3EAB8A95A" authorityName="Reiss" authorityYear="1972" box="[331,509,295,321]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="paraligulatus">
<emphasis id="B979EACCFFC4FFAFFE85FEB3EAB8A95A" box="[331,509,295,321]" italics="true" pageId="13" pageNumber="518">T. paraligulatus</emphasis>
</taxonomicName>
. Despite the aforementioned similarities,
<taxonomicName id="4C0D4D5DFFC4FFAFFC1FFEB3EF5DA95A" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[977,1048,295,321]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFC1FFEB3EF5DA95A" box="[977,1048,295,321]" italics="true" pageId="13" pageNumber="518">T. lulu</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D5DFFC4FFAFFB80FEB3EFABA95A" authorityName="Reiss" authorityYear="1972" box="[1102,1262,295,321]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFB80FEB3EF26A95B" box="[1102,1123,295,320]" italics="true" pageId="13" pageNumber="518">T.</emphasis>
cf.
<emphasis id="B979EACCFFC4FFAFFB42FEB3EFABA95A" box="[1164,1262,295,321]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
</taxonomicName>
display several differences: the AR is greater than 1 (1.04) in
<taxonomicName id="4C0D4D5DFFC4FFAFFD50FEDFE9A3A97E" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[670,742,331,357]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFD50FEDFE9A3A97E" box="[670,742,331,357]" italics="true" pageId="13" pageNumber="518">T. lulu</emphasis>
</taxonomicName>
, while less than 1 (0.84) in
<taxonomicName id="4C0D4D5DFFC4FFAFFBEFFEDFEF81A97E" authorityName="Reiss" authorityYear="1972" box="[1057,1220,331,357]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFBEFFEDFEF73A97F" box="[1057,1078,331,356]" italics="true" pageId="13" pageNumber="518">T.</emphasis>
cf.
<emphasis id="B979EACCFFC4FFAFFBACFEDFEF81A97E" box="[1122,1220,331,357]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
</taxonomicName>
; other differentiate characters pertain to the frontal tubercle length/width ratio, the shape of the superior volsella (see above) and the wing chaetotaxy and macrotrichia—numerous in
<taxonomicName id="4C0D4D5DFFC4FFAFFD1DFE07E85BA9B6" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[723,798,403,429]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFD1DFE07E85BA9B6" box="[723,798,403,429]" italics="true" pageId="13" pageNumber="518">T. lulu</emphasis>
</taxonomicName>
relative to
<taxonomicName id="4C0D4D5DFFC4FFAFFC69FE07EF15A9B6" authorityName="Reiss" authorityYear="1972" box="[935,1104,403,429]" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFC69FE07E8F8A9B7" box="[935,957,403,428]" italics="true" pageId="13" pageNumber="518">T.</emphasis>
cf.
<emphasis id="B979EACCFFC4FFAFFC20FE07EF15A9B6" box="[1006,1104,403,429]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
</taxonomicName>
(cf. Sanseverino 2006). The structure of an auxiliary importance in the male
<taxonomicName id="4C0D4D5DFFC4FFAFFD0AFE23E807A9CA" authorityName="Zavrel" authorityYear="1917" box="[708,834,439,465]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="tribe" tribe="Tanytarsini">Tanytarsini</taxonomicName>
diagnostics are the hypopygial anterolateral teeth or orolateral spines of laterosternite IX, however, they usually are minute thus weakly observable. The spines are in shape of small processes in
<taxonomicName id="4C0D4D5DFFC4FFAFFE38FE6BE9E1AA02" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[502,676,511,537]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFE38FE6BE9E1AA02" box="[502,676,511,537]" italics="true" pageId="13" pageNumber="518">Tanytarsus lulu</emphasis>
</taxonomicName>
but not confirmed in other species of the
<taxonomicName id="4C0D4D5DFFC4FFAFFB48FE6BEFADAA02" authorityName="Reiss" authorityYear="1972" box="[1158,1256,511,537]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFB48FE6BEFADAA02" box="[1158,1256,511,537]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
</taxonomicName>
group. Another character that needs re-examination is the presence of sensilla chaetica on the mid leg basitarsus which form a distinct row (comb) in
<taxonomicName id="4C0D4D5DFFC4FFAFFE5BFDD3EA99AA7A" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[405,476,583,609]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFE5BFDD3EA99AA7A" box="[405,476,583,609]" italics="true" pageId="13" pageNumber="518">T. lulu</emphasis>
</taxonomicName>
, but have not been mentioned in descriptions of
<taxonomicName id="4C0D4D5DFFC4FFAFFC39FDD3EF31AA7A" authorityName="Reiss" authorityYear="1972" box="[1015,1140,583,609]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFC39FDD3EF31AA7A" box="[1015,1140,583,609]" italics="true" pageId="13" pageNumber="518">T. ligulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0D4D5DFFC4FFAFFB4FFDD3EE76AA7A" authorityName="Reiss" authorityYear="1972" box="[1153,1331,583,609]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="paraligulatus">
<emphasis id="B979EACCFFC4FFAFFB4FFDD3EE76AA7A" box="[1153,1331,583,609]" italics="true" pageId="13" pageNumber="518">T. paraligulatus</emphasis>
</taxonomicName>
nor
<taxonomicName id="4C0D4D5DFFC4FFAFFAABFDD3EAA4AA9D" authority="(Sanseverino 2006)" baseAuthorityName="Sanseverino" baseAuthorityYear="2006" class="Insecta" family="Chironomidae" genus="Tanytarsus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFAABFDD3EE3FAA7B" box="[1381,1402,583,608]" italics="true" pageId="13" pageNumber="518">T.</emphasis>
cf.
<emphasis id="B979EACCFFC4FFAFFF59FDFFEBBCAA9E" box="[151,249,619,645]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
(Sanseverino 2006)
</taxonomicName>
. All species of the
<taxonomicName id="4C0D4D5DFFC4FFAFFD75FDFFE858AA9E" baseAuthorityName="Sanseverino" baseAuthorityYear="2006" box="[699,797,619,645]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B979EACCFFC4FFAFFD75FDFFE858AA9E" box="[699,797,619,645]" italics="true" pageId="13" pageNumber="518">ligulatus</emphasis>
</taxonomicName>
group are close morphologically, thus their delimitation has to be based on detailed descriptions (given in the extended form for
<taxonomicName id="4C0D4D5DFFC4FFAFFC71FD1BEF42AAB2" authorityName="Dantas &amp; Amat &amp; Hamada &amp; Giłka" authorityYear="2022" box="[959,1031,655,681]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="lulu">
<emphasis id="B979EACCFFC4FFAFFC71FD1BEF42AAB2" box="[959,1031,655,681]" italics="true" pageId="13" pageNumber="518">T. lulu</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>