435 lines
51 KiB
XML
435 lines
51 KiB
XML
<document id="0D6CB702E3837A24E6D1F5E15C642FEC" ID-CLB-Dataset="55782" ID-DOI="10.11646/zootaxa.5129.4.2" ID-GBIF-Dataset="ff4f0237-882f-45d1-a04a-898b7c0ea9f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="6504144" ID-ZooBank="E9ACC0E3-A5EE-4867-8CBA-56FBBB728EFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651218754646" checkinUser="plazi" docAuthor="Dantas, Galileu P. S., Amat, Eduardo, Hamada, Neusa & Giłka, Wojciech" docDate="2022" docId="03A487C8FFC1FFA8FF59FF03EF75ABBE" docLanguage="en" docName="zootaxa.5129.4.2.pdf" docOrigin="Zootaxa 5129 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus gnomon Dantas & Amat & Hamada & Giłka 2022, sp. nov." docType="treatment" docUuid="8FF8DF78-6BA1-4661-B38C-560195B62C38" docUuidSource="ZooBank" docVersion="8" lastPageNumber="515" masterDocId="FF9DFFB0FFC9FFA2FFCEFF94EB45A81B" masterDocTitle="Towards the systematics and diversity of Neotropical Tanytarsus van der Wulp (Diptera: Chironomidae): news from Colombia" masterLastPageNumber="529" masterPageNumber="505" pageNumber="513" updateTime="1699329456537" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="B5296327A6F73CB3ED1B7614396661C7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3E8855ABFC2F93A0FA536D079F0D2807">
|
||
<mods:title id="66FE88D6F4EC7AE0C2869CEA11AC2092">Towards the systematics and diversity of Neotropical Tanytarsus van der Wulp (Diptera: Chironomidae): news from Colombia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E11C6A32790198F18AB2F4AAEDE6556A" type="personal">
|
||
<mods:role id="D1FF974D4D859459AA0066960BB050FD">
|
||
<mods:roleTerm id="6836EA781A7446C7BD364BD13DA5AEDF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F01B3862878F0B53EE6DA9034095930B">Dantas, Galileu P. S.</mods:namePart>
|
||
<mods:nameIdentifier id="87EBA0AC118BE86E1581FDFF7B90E8EA" type="ORCID">0000-0002-9155-533X</mods:nameIdentifier>
|
||
<mods:affiliation id="DC13BCC888EE0187DFD0076504D06AB5">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt); Av. André Araújo, 2936, 69067 - 375, Manaus, Amazonas, Brazil. & galileu. psd @ gmail. com; https: // orcid. org / 0000 - 0002 - 9155 - 533 X</mods:affiliation>
|
||
<mods:nameIdentifier id="227949C4441A8F79A034FE6CBC4338F3" type="email">galileu.psd@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="E1C922D5EE699A3AE24C7056F096391C" type="personal">
|
||
<mods:role id="EFFE83C5865ED16A5E85BC4AF00389B5">
|
||
<mods:roleTerm id="79AF93948D452CB87680A3861A4A1966">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D1DF28EBA2D4703EF3AE1D29858BD5D3">Amat, Eduardo</mods:namePart>
|
||
<mods:affiliation id="40589339C79C4C405C52CC0A0BF2F651">Grupo de investigación BIOFORENSE, Facultad de Derecho y Ciencias Forenses, Tecnológico deAntioquia - Institución Universitaria; calle 78 B no. T 2 A- 22 MI MedellínI Colombia.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="02C9F49A495A92FDD5440A63A4649DA7" type="personal">
|
||
<mods:role id="3B3461D7EC80EA6F31137C333CAB6D80">
|
||
<mods:roleTerm id="94618F253E8873377F757EA7E7BDAD44">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FB6B49761CD8190E2A1197FD172A5572">Hamada, Neusa</mods:namePart>
|
||
<mods:nameIdentifier id="EB8E5D70645AA4716E24408FC2CAA7B1" type="ORCID">0000-0002-3526-5426</mods:nameIdentifier>
|
||
<mods:affiliation id="6FDBD67A6E0A3065CC82FB17D637FB38">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt); Av. André Araújo, 2936, 69067 - 375, Manaus, Amazonas, Brazil. & nhamada @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 3526 - 5426</mods:affiliation>
|
||
<mods:nameIdentifier id="2C6CB15B75175A8FDBEEB74EF7A60C4A" type="email">nhamada@inpa.gov.br</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="773F375E43665EF6C3DEE777C0326CD2" type="personal">
|
||
<mods:role id="C6E1380A8980542A88118676C9B4AB69">
|
||
<mods:roleTerm id="837640C41753E7B0379789898A9E6550">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="94DBC300C33B4DE00434DDC7D491BF2F">Giłka, Wojciech</mods:namePart>
|
||
<mods:affiliation id="DEEDD2D42D42939C41341EECEEA4AF95">University of Gdańsk, Faculty of Biology, Department of Invertebrate Zoology and Parasitology, Laboratory of Systematic Zoology; Wita Stwosza 59, 80 - 308 Gdańsk, Poland.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3B29C907B48D23934B1BAA20B0791D91">text</mods:typeOfResource>
|
||
<mods:relatedItem id="65339CF6E5641BF063D38CC6B7AC14E1" type="host">
|
||
<mods:titleInfo id="BE013BF25E4F04A0A1361477CD3D388F">
|
||
<mods:title id="BFAA7CDDAB2891A3261272AFD4EB2BFD">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="7A239036DD2E6006001AFBED2CB8E1A4">
|
||
<mods:date id="775FF5E2B77207732F51072A36FD9E27">2022</mods:date>
|
||
<mods:detail id="F3A5134D1F276D888DDC57F7E5CA0565" type="pubDate">
|
||
<mods:number id="6D0A7DBD1FA8035689644AB4827EB6AF">2022-04-29</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2B62EDF73A5078B8E151AF9298D0D45B" type="volume">
|
||
<mods:number id="88279B86EA6D7C69E7A35E6AE0AD5246">5129</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="6C096F483DDE5301142C12933EAE0E0D" type="issue">
|
||
<mods:number id="AE41161323376A4E145E0B40770C8304">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BFD57D8F967491A0E40C420DFB4C633C" unit="page">
|
||
<mods:start id="5357AEA525942B03E5B9C6CD4F0A7E1C">505</mods:start>
|
||
<mods:end id="7FE74A0F5EAD7C8D6363F4BFDCA62457">529</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="346824EF994344235AE4C2ACB6AFA84E">journal article</mods:classification>
|
||
<mods:identifier id="AE44A48734FCD363FB4764A6AF56B122" type="CLB-Dataset">55782</mods:identifier>
|
||
<mods:identifier id="0D238165D80644A6C2C1D36B6349D603" type="DOI">10.11646/zootaxa.5129.4.2</mods:identifier>
|
||
<mods:identifier id="0B768B9B6FB8CD30BF22929451A95E8C" type="GBIF-Dataset">ff4f0237-882f-45d1-a04a-898b7c0ea9f9</mods:identifier>
|
||
<mods:identifier id="59BCE2EEB641CC0E0CE63C19DF201424" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="18D5D701C882F83F534CECE52E2EC7C5" type="Zenodo-Dep">6504144</mods:identifier>
|
||
<mods:identifier id="75E24CE2E755FC0EA609AB600B328EA0" type="ZooBank">E9ACC0E3-A5EE-4867-8CBA-56FBBB728EFE</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A487C8FFC1FFA8FF59FF03EF75ABBE" ID-DOI="http://doi.org/10.5281/zenodo.6506256" ID-GBIF-Taxon="194877471" ID-Zenodo-Dep="6506256" LSID="urn:lsid:zoobank.org:act:8FF8DF78-6BA1-4661-B38C-560195B62C38" httpUri="http://treatment.plazi.org/id/03A487C8FFC1FFA8FF59FF03EF75ABBE" lastPageId="10" lastPageNumber="515" pageId="8" pageNumber="513">
|
||
<subSubSection id="C3176555FFC1FFAAFF59FF03EF4FA8CD" pageId="8" pageNumber="513" type="nomenclature">
|
||
<paragraph id="8BB236DEFFC1FFAAFF59FF03EAAEA8AA" blockId="8.[151,1034,151,249]" box="[151,491,151,177]" pageId="8" pageNumber="513">
|
||
<heading id="D0FA81B2FFC1FFAAFF59FF03EAAEA8AA" bold="true" box="[151,491,151,177]" fontSize="11" level="1" pageId="8" pageNumber="513" reason="1">
|
||
<emphasis id="B979EACCFFC1FFAAFF59FF03EAAEA8AA" bold="true" box="[151,491,151,177]" pageId="8" pageNumber="513">
|
||
<taxonomicName id="4C0D4D5DFFC1FFAAFF59FF03EAC3A8AA" ID-CoL="9J2NY" authority="Dantas & Amat & Hamada & Giłka, 2022" authorityName="Dantas & Amat & Hamada & Giłka" authorityYear="2022" box="[151,390,151,177]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="gnomon" status="sp. nov.">
|
||
<emphasis id="B979EACCFFC1FFAAFF59FF03EAC3A8AA" bold="true" box="[151,390,151,177]" italics="true" pageId="8" pageNumber="513">Tanytarsus gnomon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A57B7FFC1FFAAFE40FF03EAAEA8AA" box="[398,491,151,177]" pageId="8" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF59FF2FEF4FA8CD" blockId="8.[151,1034,151,249]" box="[151,1034,187,214]" pageId="8" pageNumber="513">
|
||
LSID:
|
||
<uri id="FF9C3ADCFFC1FFAAFF2AFF2FEF4FA8CD" box="[228,1034,187,214]" pageId="8" pageNumber="513">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFAB0C0BFFC1FFAAFE34FF2FEF4FA8CD" box="[506,1034,187,214]" pageId="8" pageNumber="513">8FF8DF78-6BA1-4661-B38C-560195B62C38</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176555FFC1FFAAFF59FF4BEA58A8E2" box="[151,285,223,249]" pageId="8" pageNumber="513" type="description">
|
||
<paragraph id="8BB236DEFFC1FFAAFF59FF4BEA58A8E2" blockId="8.[151,1034,151,249]" box="[151,285,223,249]" pageId="8" pageNumber="513">
|
||
(
|
||
<figureCitation id="13362A5BFFC1FFAAFF6EFF4BEA50A8E2" box="[160,277,223,249]" captionStart="FIGURE 4" captionStartId="9.[151,250,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-20@9.[151,1435,181,1920]" captionTargetPageId="9" captionText="FIGURE 4. Tanytarsus gnomon sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: arrangement of anal point, superior volsella and digitus; D: superior volsella and digitus; E, F: median volsella drawn (E) and photographed (F); (C–F magnified ca. 2–3 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504152" httpUri="https://zenodo.org/record/6504152/files/figure.png" pageId="8" pageNumber="513">Fig. 4A–F</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176555FFC1FFAAFF59FEB5EA1FA998" pageId="8" pageNumber="513" type="materials_examined">
|
||
<paragraph id="8BB236DEFFC1FFAAFF59FEB5EA1FA998" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF59FEB5EA0FA920" bold="true" box="[151,330,289,315]" pageId="8" pageNumber="513">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B653C83FFC1FFAAFEAAFEB5EA1FA998" ID-GBIF-Occurrence="3757950305" collectingDate="2018-03-06" collectionCode="R" collectorName="G. P. S. Dantas & Hernandez, E. C. G" country="Colombia" county="Departament" elevation="570" latitude="2.66135" location="Condominio Santa Helena" longLatPrecision="1" longitude="-75.51668" municipality="Yaguara" pageId="8" pageNumber="513" specimenCount="1" stateProvince="Huila" typeStatus="holotype">
|
||
<typeStatus id="54B6887CFFC1FFAAFEAAFEB5EA94A920" box="[356,465,289,315]" pageId="8" pageNumber="513" type="holotype">Holotype</typeStatus>
|
||
: male,
|
||
<collectingCountry id="F31A764EFFC1FFAAFDF9FEB5E98AA920" box="[567,719,289,315]" name="Colombia" pageId="8" pageNumber="513">COLOMBIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C9F83CFFC1FFAAFD28FEB5E861A920" box="[742,804,289,315]" country="Colombia" name="Huila" pageId="8" pageNumber="513">Huila</collectingRegion>
|
||
<collectingCounty id="62D34E52FFC1FFAAFCF9FEB6E88EA927" box="[823,971,290,316]" pageId="8" pageNumber="513">Departament</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BD6ACA4FFC1FFAAFC2EFEB5EF7AA920" box="[992,1087,289,315]" pageId="8" pageNumber="513">Yaguará</collectingMunicipality>
|
||
,
|
||
<location id="8ED26005FFC1FFAAFB98FEB5EEDDA920" LSID="urn:lsid:plazi:treatment:03A487C8FFC1FFA8FF59FF03EF75ABBE:8ED26005FFC1FFAAFB98FEB5EEDDA920" box="[1110,1432,289,315]" country="Colombia" county="Departament" latitude="2.66135" longLatPrecision="1" longitude="-75.51668" municipality="Yaguara" name="Condominio Santa Helena" pageId="8" pageNumber="513" stateProvince="Huila">Condominio Santa Helena</location>
|
||
,
|
||
<geoCoordinate id="EE395019FFC1FFAAFF59FED1EA70A944" box="[151,309,325,351]" degrees="2" direction="north" minutes="39" orientation="latitude" pageId="8" pageNumber="513" precision="1" seconds="40.86" value="2.66135">2°39’40.86”N</geoCoordinate>
|
||
<geoCoordinate id="EE395019FFC1FFAAFEF0FED0EAA7A944" box="[318,482,324,351]" degrees="75" direction="west" minutes="31" orientation="longitude" pageId="8" pageNumber="513" precision="1" seconds="0.03" value="-75.51668">75°31’0.03”W</geoCoordinate>
|
||
,
|
||
<elevation id="0020D1EDFFC1FFAAFE21FED0E933A944" box="[495,630,324,351]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="513" unit="m" value="570.0">
|
||
<quantity id="4CF59B3BFFC1FFAAFE21FED0E97DA944" box="[495,568,324,351]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="513" unit="m" value="570.0">570 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFB3101EFFC1FFAAFD4AFED1E870A944" box="[644,821,325,351]" pageId="8" pageNumber="513" value="2018-03-06">
|
||
<collectingDate id="EFF7E9F6FFC1FFAAFD4AFED1E870A944" box="[644,821,325,351]" pageId="8" pageNumber="513" value="2018-03-06">06 March 2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="524C4EC9FFC1FFAAFC8CFED1E8F0A944" box="[834,949,325,351]" pageId="8" pageNumber="513">sweep net</collectingMethod>
|
||
,
|
||
<collectorName id="26F85308FFC1FFAAFC0CFED1EF24A944" box="[962,1121,325,351]" pageId="8" pageNumber="513">G.P.S. Dantas</collectorName>
|
||
, S.M.
|
||
<collectionCode id="ED1CAE1BFFC1FFAAFB6DFED1EFFFA944" box="[1187,1210,325,351]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="513">R</collectionCode>
|
||
.
|
||
<collectorName id="26F85308FFC1FFAAFB08FED1EEDCA944" box="[1222,1433,325,351]" pageId="8" pageNumber="513">Hernández, E.C.G</collectorName>
|
||
. Amat (CETdeA).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176555FFC1FFAAFF09FE1FE924A9D2" pageId="8" pageNumber="513" type="etymology">
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FE1FE924A9D2" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FE1FEAD8A9BE" bold="true" box="[199,413,395,421]" italics="true" pageId="8" pageNumber="513">Derivatio nominis.</emphasis>
|
||
In reference to the shape of the anal point with its posterior bar, resembling a sundial clock hand (Lat.
|
||
<taxonomicName id="4C0D4D5DFFC1FFAAFEDFFE24EA2EA9D2" authorityName="Dantas & Amat & Hamada & Giłka" authorityYear="2022" box="[273,363,432,457]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="gnomon">
|
||
<emphasis id="B979EACCFFC1FFAAFEDFFE24EA2EA9D2" box="[273,363,432,457]" italics="true" pageId="8" pageNumber="513">gnomon</emphasis>
|
||
</taxonomicName>
|
||
). Noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176555FFC1FFAAFF09FE46E9E4AA85" pageId="8" pageNumber="513" type="diagnosis">
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FE46E9E4AA85" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FE46EA07A9F7" bold="true" box="[199,322,466,492]" pageId="8" pageNumber="513">Diagnosis.</emphasis>
|
||
Frontal tubercles small. Tergite IX covered with dense short microtrichia except for two bare fields lateral to anal point base. Anal point with well-developed anal crests extending towards tergite IX and flanking a small cone-shaped anterior bar, posterior bar slender, evenly curved upwards, with apex slightly enlarged and split. Superior volsella with posteromedian corner strongly narrowed, in shape of finger-like projection, with apex swollen and medially directed. Digitus subtriangular, pointed, reaching half length of superior volsella. Median volsella with 1 foliate and 3 setiform lamellae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176555FFC1FFAAFF09FD33E94CAFE1" pageId="8" pageNumber="513" type="description">
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FD33E973AADA" blockId="8.[151,1437,289,1274]" box="[199,566,679,706]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FD33EA1CAADA" bold="true" box="[199,345,679,705]" pageId="8" pageNumber="513">Description.</emphasis>
|
||
Adult male (n = 1).
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FD5FE933AB13" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FD5FEAB4AAFE" box="[199,497,715,741]" italics="true" pageId="8" pageNumber="513">Body size and proportions</emphasis>
|
||
. Total length
|
||
<quantity id="4CF59B3BFFC1FFAAFD5EFD5FE9BDAAFE" box="[656,760,715,741]" metricMagnitude="-3" metricUnit="m" metricValue="2.39" pageId="8" pageNumber="513" unit="mm" value="2.39">2.39 mm</quantity>
|
||
. Wing length
|
||
<quantity id="4CF59B3BFFC1FFAAFC59FD5FE8BAAAFE" box="[919,1023,715,742]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="513" unit="mm" value="1.25">1.25 mm</quantity>
|
||
. Total length/wing length ratio 1.90. Wing length/length of profemur ratio 1.71.
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FC86E8E7AB54" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FC86EA07AB37" box="[199,322,786,812]" italics="true" pageId="8" pageNumber="513">Coloration</emphasis>
|
||
. Eyes black. Antenna light brown. Head capsule and thorax yellow to light brown. Legs yellowish to light brown. Wing veins yellow, membrane pale. Abdomen yellow.
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FCCDE934ABC6" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FCCDEA41AB68" box="[199,260,857,883]" italics="true" pageId="8" pageNumber="513">Head</emphasis>
|
||
. Eyes bare, with well-developed dorsomedian extensions. Antenna with 13 flagellomeres; ultimate flagellomere 446 μm long; AR 1.01. Frontal tubercles small, 4 μm long. Tentorium 105 μm long. Temporal setae 9 on each side. Clypeus with 11 setae. Lengths of palpomeres 1–5 (in μm): 24, 28, 78, 90, 160; third palpomere with 2 sensilla clavata subapically, 12 μm long.
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FC72E8EEAC3E" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FC72EA5DAC1B" box="[199,280,998,1024]" italics="true" pageId="8" pageNumber="513">Thorax</emphasis>
|
||
. Ac 16, restricted to anterior region of scutum; Dc 5 on each side, uniserial; Pa 1 on each side; Scts 4. Scutum projected and rounded anteriorly, overreaching antepronotum.
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FBBAEE30AC70" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FBBAEA44AC5C" box="[199,257,1070,1095]" italics="true" pageId="8" pageNumber="513">Wing</emphasis>
|
||
. Typical of the genus. Almost all veins (except subcosta) and entire membrane posterior to radial veins area (except base of m and an cells) covered with macrotrichia. Brachiolum with 1 seta. VRCu 1.15. WW 0.28.
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF09FBE1EA69ACE2" blockId="8.[151,1437,289,1274]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09FBE1EBB9AC95" box="[199,252,1141,1166]" italics="true" pageId="8" pageNumber="513">Legs</emphasis>
|
||
. Foreleg tibia with straight lanceolate spur 15 μm long. Tibial combs of mid and hind legs separated; spurs of mid leg unequal: one bent, 24 μm long, second straight, 12 μm long; spurs of hind leg unequal: one bent 28 μm long, second straight 27 μm long. Basitarsus of mid leg without sensilla chaetica. Lengths and proportions of legs as in
|
||
<tableCitation id="C68F0365FFC1FFAAFF1CFB4BEA62ACE2" box="[210,295,1247,1273]" captionStart="TABLE 3" captionStartId="8.[151,239,1316,1340]" captionTargetPageId="8" captionText="TABLE 3. Lengths (in μm) and proportions of legs of Tanytarsus gnomon sp. nov., male (n = 1)." pageId="8" pageNumber="513">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF726656FFC1FFAAFF59FAB0EFE6AD27" box="[151,1187,1316,1340]" pageId="8" pageNumber="513" startId="8.[151,239,1316,1340]" targetBox="[166,1378,1356,1507]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="8BB236DEFFC1FFAAFF59FAB0EFE6AD27" blockId="8.[151,1187,1316,1340]" box="[151,1187,1316,1340]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF59FAB0EA4FAD27" bold="true" box="[151,266,1316,1340]" pageId="8" pageNumber="513">TABLE 3.</emphasis>
|
||
Lengths (in μm) and proportions of legs of
|
||
<taxonomicName id="4C0D4D5DFFC1FFAAFD11FAB1E8EAAD27" authority="Dantas & Amat & Hamada & Giłka, 2022" authorityName="Dantas & Amat & Hamada & Giłka" authorityYear="2022" box="[735,943,1317,1340]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="gnomon" status="sp. nov.">
|
||
<emphasis id="B979EACCFFC1FFAAFD11FAB1E8EAAD27" box="[735,943,1317,1340]" italics="true" pageId="8" pageNumber="513">Tanytarsus gnomon</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979EACCFFC1FFAAFC78FAB0EF4EAD27" bold="true" box="[950,1035,1316,1340]" pageId="8" pageNumber="513">
|
||
<taxonomicNameLabel id="A24A57B7FFC1FFAAFC78FAB0EF4EAD27" box="[950,1035,1316,1340]" pageId="8" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (n = 1).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB236DEFFC1FFAAFED4FAD8EE27ADC6" pageId="8" pageNumber="513">
|
||
<table id="F90DC47EFFC1005DFF68FAD8EE27ADF8" box="[166,1378,1356,1507]" colsContinueFrom="6.[166,1416,1093,1243]" colsContinueIn="12.[166,1416,877,1027]" gridcols="11" gridrows="4" pageId="8" pageNumber="513">
|
||
<tr id="353D349CFFC1005DFF68FAD8EE27AD70" box="[166,1378,1356,1387]" gridrow="0" pageId="8" pageNumber="513" rowspan-0="1">
|
||
<th id="76EC5DE0FFC1005DFED4FAD8EA05AD70" box="[282,320,1356,1387]" gridcol="1" gridrow="0" pageId="8" pageNumber="513">fe</th>
|
||
<th id="76EC5DE0FFC1005DFE41FAD8EAF0AD70" box="[399,437,1356,1387]" gridcol="2" gridrow="0" pageId="8" pageNumber="513">ti</th>
|
||
<th id="76EC5DE0FFC1005DFDCAFAD8E96FAD70" box="[516,554,1356,1387]" gridcol="3" gridrow="0" pageId="8" pageNumber="513">ta1</th>
|
||
<th id="76EC5DE0FFC1005DFDB7FAD8E9DAAD70" box="[633,671,1356,1387]" gridcol="4" gridrow="0" pageId="8" pageNumber="513">ta2</th>
|
||
<th id="76EC5DE0FFC1005DFD20FAD8E851AD70" box="[750,788,1356,1387]" gridcol="5" gridrow="0" pageId="8" pageNumber="513">ta3</th>
|
||
<th id="76EC5DE0FFC1005DFCACFAD8E8CDAD70" box="[866,904,1356,1387]" gridcol="6" gridrow="0" pageId="8" pageNumber="513">ta4</th>
|
||
<th id="76EC5DE0FFC1005DFC19FAD8E8B4AD70" box="[983,1009,1356,1387]" gridcol="7" gridrow="0" pageId="8" pageNumber="513">ta 5</th>
|
||
<th id="76EC5DE0FFC1005DFB82FAD8EF3DAD70" box="[1100,1144,1356,1387]" gridcol="8" gridrow="0" pageId="8" pageNumber="513">LR</th>
|
||
<th id="76EC5DE0FFC1005DFB0FFAD8EFA8AD70" box="[1217,1261,1356,1387]" gridcol="9" gridrow="0" pageId="8" pageNumber="513">BV</th>
|
||
<th id="76EC5DE0FFC1005DFAF8FAD8EE27AD70" box="[1334,1378,1356,1387]" gridcol="10" gridrow="0" pageId="8" pageNumber="513">SV</th>
|
||
</tr>
|
||
<tr id="353D349CFFC1005DFF68FAE0EE27AD89" box="[166,1378,1396,1426]" gridrow="1" pageId="8" pageNumber="513">
|
||
<th id="76EC5DE0FFC1005DFF68FAE0EBFCAD89" box="[166,185,1396,1426]" gridcol="0" gridrow="1" pageId="8" pageNumber="513">p1</th>
|
||
<td id="76EC5DE0FFC1005DFED4FAE0EA05AD89" box="[282,320,1396,1426]" gridcol="1" gridrow="1" pageId="8" pageNumber="513">734</td>
|
||
<td id="76EC5DE0FFC1005DFE41FAE0EAF0AD89" box="[399,437,1396,1426]" gridcol="2" gridrow="1" pageId="8" pageNumber="513">299</td>
|
||
<td id="76EC5DE0FFC1005DFDCAFAE0E96FAD89" box="[516,554,1396,1426]" gridcol="3" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
<td id="76EC5DE0FFC1005DFDB7FAE0E9DAAD89" box="[633,671,1396,1426]" gridcol="4" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
<td id="76EC5DE0FFC1005DFD20FAE0E851AD89" box="[750,788,1396,1426]" gridcol="5" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
<td id="76EC5DE0FFC1005DFCACFAE0E8CDAD89" box="[866,904,1396,1426]" gridcol="6" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
<td id="76EC5DE0FFC1005DFC19FAE0E8B4AD89" box="[983,1009,1396,1426]" gridcol="7" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
<td id="76EC5DE0FFC1005DFB82FAE0EF3DAD89" box="[1100,1144,1396,1426]" gridcol="8" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
<td id="76EC5DE0FFC1005DFB0FFAE0EFA8AD89" box="[1217,1261,1396,1426]" gridcol="9" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
<td id="76EC5DE0FFC1005DFAF8FAE0EE27AD89" box="[1334,1378,1396,1426]" gridcol="10" gridrow="1" pageId="8" pageNumber="513">–</td>
|
||
</tr>
|
||
<tr id="353D349CFFC1005DFF68FA08EE27ADA1" box="[166,1378,1436,1466]" gridrow="2" pageId="8" pageNumber="513">
|
||
<th id="76EC5DE0FFC1005DFF68FA08EBFCADA1" box="[166,185,1436,1466]" gridcol="0" gridrow="2" pageId="8" pageNumber="513">p2</th>
|
||
<td id="76EC5DE0FFC1005DFED4FA08EA05ADA1" box="[282,320,1436,1466]" gridcol="1" gridrow="2" pageId="8" pageNumber="513">668</td>
|
||
<td id="76EC5DE0FFC1005DFE41FA08EAF0ADA1" box="[399,437,1436,1466]" gridcol="2" gridrow="2" pageId="8" pageNumber="513">525</td>
|
||
<td id="76EC5DE0FFC1005DFDCAFA08E96FADA1" box="[516,554,1436,1466]" gridcol="3" gridrow="2" pageId="8" pageNumber="513">308</td>
|
||
<td id="76EC5DE0FFC1005DFDB7FA08E9DAADA1" box="[633,671,1436,1466]" gridcol="4" gridrow="2" pageId="8" pageNumber="513">136</td>
|
||
<td id="76EC5DE0FFC1005DFD20FA08E851ADA1" box="[750,788,1436,1466]" gridcol="5" gridrow="2" pageId="8" pageNumber="513">100</td>
|
||
<td id="76EC5DE0FFC1005DFCACFA08E8CDADA1" box="[866,904,1436,1466]" gridcol="6" gridrow="2" pageId="8" pageNumber="513">63</td>
|
||
<td id="76EC5DE0FFC1005DFC19FA08E8B4ADA1" box="[983,1009,1436,1466]" gridcol="7" gridrow="2" pageId="8" pageNumber="513">55</td>
|
||
<td id="76EC5DE0FFC1005DFB82FA08EF3DADA1" box="[1100,1144,1436,1466]" gridcol="8" gridrow="2" pageId="8" pageNumber="513">0.59</td>
|
||
<td id="76EC5DE0FFC1005DFB0FFA08EFA8ADA1" box="[1217,1261,1436,1466]" gridcol="9" gridrow="2" pageId="8" pageNumber="513">4.24</td>
|
||
<td id="76EC5DE0FFC1005DFAF8FA08EE27ADA1" box="[1334,1378,1436,1466]" gridcol="10" gridrow="2" pageId="8" pageNumber="513">3.86</td>
|
||
</tr>
|
||
<tr id="353D349CFFC1005DFF68FA51EE27ADF8" box="[166,1378,1477,1507]" gridrow="3" pageId="8" pageNumber="513">
|
||
<th id="76EC5DE0FFC1005DFF68FA51EBFCADF8" box="[166,185,1477,1507]" gridcol="0" gridrow="3" pageId="8" pageNumber="513">p 3</th>
|
||
<td id="76EC5DE0FFC1005DFED4FA51EA05ADF8" box="[282,320,1477,1507]" gridcol="1" gridrow="3" pageId="8" pageNumber="513">727</td>
|
||
<td id="76EC5DE0FFC1005DFE41FA51EAF0ADF8" box="[399,437,1477,1507]" gridcol="2" gridrow="3" pageId="8" pageNumber="513">610</td>
|
||
<td id="76EC5DE0FFC1005DFDCAFA51E96FADF8" box="[516,554,1477,1507]" gridcol="3" gridrow="3" pageId="8" pageNumber="513">449</td>
|
||
<td id="76EC5DE0FFC1005DFDB7FA51E9DAADF8" box="[633,671,1477,1507]" gridcol="4" gridrow="3" pageId="8" pageNumber="513">242</td>
|
||
<td id="76EC5DE0FFC1005DFD20FA51E851ADF8" box="[750,788,1477,1507]" gridcol="5" gridrow="3" pageId="8" pageNumber="513">215</td>
|
||
<td id="76EC5DE0FFC1005DFCACFA51E8CDADF8" box="[866,904,1477,1507]" gridcol="6" gridrow="3" pageId="8" pageNumber="513">125</td>
|
||
<td id="76EC5DE0FFC1005DFC19FA51E8B4ADF8" box="[983,1009,1477,1507]" gridcol="7" gridrow="3" pageId="8" pageNumber="513">86</td>
|
||
<td id="76EC5DE0FFC1005DFB82FA51EF3DADF8" box="[1100,1144,1477,1507]" gridcol="8" gridrow="3" pageId="8" pageNumber="513">0.74</td>
|
||
<td id="76EC5DE0FFC1005DFB0FFA51EFA8ADF8" box="[1217,1261,1477,1507]" gridcol="9" gridrow="3" pageId="8" pageNumber="513">2.67</td>
|
||
<td id="76EC5DE0FFC1005DFAF8FA51EE27ADF8" box="[1334,1378,1477,1507]" gridcol="10" gridrow="3" pageId="8" pageNumber="513">2.98</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BB236DEFFC1FFAAFF09F999E94CAFE1" blockId="8.[151,1437,1548,2043]" pageId="8" pageNumber="513">
|
||
<emphasis id="B979EACCFFC1FFAAFF09F999EA13AE3D" box="[199,342,1549,1574]" italics="true" pageId="8" pageNumber="513">Hypopygium</emphasis>
|
||
. Tergite IX covered with dense short microtrichia except for two bare fields lateral to anal point base, with 5 median setae placed between anal tergite bands and 4 setae on each side of anal point; lateral teeth present, minute; anal tergite bands V-shaped, widely separated, ending anterior to anal point base (
|
||
<figureCitation id="13362A5BFFC1FFAAFB39F9C0EE16AE75" box="[1271,1363,1620,1646]" captionStart="FIGURE 4" captionStartId="9.[151,250,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-20@9.[151,1435,181,1920]" captionTargetPageId="9" captionText="FIGURE 4. Tanytarsus gnomon sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: arrangement of anal point, superior volsella and digitus; D: superior volsella and digitus; E, F: median volsella drawn (E) and photographed (F); (C–F magnified ca. 2–3 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504152" httpUri="https://zenodo.org/record/6504152/files/figure.png" pageId="8" pageNumber="513">Fig. 4A</figureCitation>
|
||
). Anal point 55 μm long, somewhat lanceolate, with elongated and rounded tip, bearing well-developed, 22 µm long anal crests extending towards tergite IX and flanking a small, 8 μm long, cone-shaped anterior bar, posterior bar slender, evenly curved upwards, with apex slightly enlarged and split (
|
||
<figureCitation id="13362A5BFFC1FFAAFCA3F954E8ABAEC1" box="[877,1006,1728,1754]" captionStart="FIGURE 4" captionStartId="9.[151,250,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-20@9.[151,1435,181,1920]" captionTargetPageId="9" captionText="FIGURE 4. Tanytarsus gnomon sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: arrangement of anal point, superior volsella and digitus; D: superior volsella and digitus; E, F: median volsella drawn (E) and photographed (F); (C–F magnified ca. 2–3 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504152" httpUri="https://zenodo.org/record/6504152/files/figure.png" pageId="8" pageNumber="513">Fig. 4A, C</figureCitation>
|
||
). Superior volsella 44 μm long, with posteromedian corner strongly narrowed, in shape of finger-like projection swollen apically and directed medially; with 5 setae on dorsal surface and 3 ventral setae close to median margin (at least 2 setae placed on prominent tubercles), field of microtrichia on dorsal surface absent; digitus 11 μm long, subtriangular, pointed, reaching half length of superior volsella (
|
||
<figureCitation id="13362A5BFFC1FFAAFE04F8C4E902AF71" box="[458,583,1872,1898]" captionStart="FIGURE 4" captionStartId="9.[151,250,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-20@9.[151,1435,181,1920]" captionTargetPageId="9" captionText="FIGURE 4. Tanytarsus gnomon sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: arrangement of anal point, superior volsella and digitus; D: superior volsella and digitus; E, F: median volsella drawn (E) and photographed (F); (C–F magnified ca. 2–3 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504152" httpUri="https://zenodo.org/record/6504152/files/figure.png" pageId="8" pageNumber="513">Fig. 4A–D</figureCitation>
|
||
). Stem of median volsella 11 μm long, with 1 foliate and 3 setiform lamellae (
|
||
<figureCitation id="13362A5BFFC1FFAAFF6EF8E0EA7DAF94" box="[160,312,1908,1935]" captionStart="FIGURE 4" captionStartId="9.[151,250,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-20@9.[151,1435,181,1920]" captionTargetPageId="9" captionText="FIGURE 4. Tanytarsus gnomon sp. nov., male. A, B: hypopygium in dorsal (A) and ventral aspect (B); C: arrangement of anal point, superior volsella and digitus; D: superior volsella and digitus; E, F: median volsella drawn (E) and photographed (F); (C–F magnified ca. 2–3 times relative to A and B)." figureDoi="http://doi.org/10.5281/zenodo.6504152" httpUri="https://zenodo.org/record/6504152/files/figure.png" pageId="8" pageNumber="513">Fig. 4B, E, F</figureCitation>
|
||
). Inferior volsella ~60 μm long, covered with microtrichia, with distal half slightly curved, swollen and posteromedially directed. Phallapodeme sinuous, ~65 μm long; transverse sternapodeme ~50 μm long, with strong oral projections. Gonocoxite 95 μm long. Gonostylus 85 μm long, almost straight, evenly tapering towards pointed apex. HR 1.14. HV 2.81.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF726656FFC0FFABFF59F80CE9DFAFE3" ID-DOI="http://doi.org/10.5281/zenodo.6504152" ID-Zenodo-Dep="6504152" httpUri="https://zenodo.org/record/6504152/files/figure.png" pageId="9" pageNumber="514" startId="9.[151,250,1944,1968]" targetBox="[151,1435,181,1920]" targetPageId="9">
|
||
<paragraph id="8BB236DEFFC0FFABFF59F80CE9DFAFE3" blockId="9.[151,1436,1944,2040]" pageId="9" pageNumber="514">
|
||
<emphasis id="B979EACCFFC0FFABFF59F80CEA48AFAB" bold="true" box="[151,269,1944,1968]" pageId="9" pageNumber="514">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C0D4D5DFFC0FFABFED7F80DEA9AAFAB" authority="Dantas & Amat & Hamada & Giłka, 2022" authorityName="Dantas & Amat & Hamada & Giłka" authorityYear="2022" box="[281,479,1945,1968]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="gnomon" status="sp. nov.">
|
||
<emphasis id="B979EACCFFC0FFABFED7F80DEA9AAFAB" box="[281,479,1945,1968]" italics="true" pageId="9" pageNumber="514">Tanytarsus gnomon</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979EACCFFC0FFABFE2BF80CE970AFAB" bold="true" box="[485,565,1944,1968]" pageId="9" pageNumber="514">
|
||
<taxonomicNameLabel id="A24A57B7FFC0FFABFE2BF80CE970AFAB" box="[485,565,1944,1968]" pageId="9" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male.
|
||
<emphasis id="B979EACCFFC0FFABFDB1F80CE9D4AFAB" bold="true" box="[639,657,1944,1968]" pageId="9" pageNumber="514">A</emphasis>
|
||
,
|
||
<emphasis id="B979EACCFFC0FFABFD50F80CE9EAAFAB" bold="true" box="[670,687,1944,1968]" pageId="9" pageNumber="514">B</emphasis>
|
||
: hypopygium in dorsal (
|
||
<emphasis id="B979EACCFFC0FFABFC69F80CE8FCAFAB" bold="true" box="[935,953,1944,1968]" pageId="9" pageNumber="514">A</emphasis>
|
||
) and ventral aspect (
|
||
<emphasis id="B979EACCFFC0FFABFB43F80CEFDBAFAB" bold="true" box="[1165,1182,1944,1968]" pageId="9" pageNumber="514">B</emphasis>
|
||
);
|
||
<emphasis id="B979EACCFFC0FFABFB7DF80CEF80AFAB" bold="true" box="[1203,1221,1944,1968]" pageId="9" pageNumber="514">C</emphasis>
|
||
: arrangement of anal point, superior volsella and digitus;
|
||
<emphasis id="B979EACCFFC0FFABFDC5F828E958AFCF" bold="true" box="[523,541,1980,2004]" pageId="9" pageNumber="514">D</emphasis>
|
||
: superior volsella and digitus;
|
||
<emphasis id="B979EACCFFC0FFABFC90F828E82AAFCF" bold="true" box="[862,879,1980,2004]" pageId="9" pageNumber="514">E</emphasis>
|
||
,
|
||
<emphasis id="B979EACCFFC0FFABFCB3F828E8C9AFCF" bold="true" box="[893,908,1980,2004]" pageId="9" pageNumber="514">F</emphasis>
|
||
: median volsella drawn (
|
||
<emphasis id="B979EACCFFC0FFABFB5AF828EFE0AFCF" bold="true" box="[1172,1189,1980,2004]" pageId="9" pageNumber="514">E</emphasis>
|
||
) and photographed (
|
||
<emphasis id="B979EACCFFC0FFABFAB3F828EEC9AFCF" bold="true" box="[1405,1420,1980,2004]" pageId="9" pageNumber="514">F</emphasis>
|
||
); (C–F magnified
|
||
<emphasis id="B979EACCFFC0FFABFEF1F875EA18AFE3" box="[319,349,2017,2040]" italics="true" pageId="9" pageNumber="514">ca.</emphasis>
|
||
2–3 times relative to A and B).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3176555FFC3FFA8FF09FF02E985A906" pageId="10" pageNumber="515" type="distribution">
|
||
<paragraph id="8BB236DEFFC3FFA8FF09FF02E985A906" blockId="10.[151,1437,150,933]" pageId="10" pageNumber="515">
|
||
<emphasis id="B979EACCFFC3FFA8FF09FF02E913A8AA" bold="true" box="[199,598,150,177]" pageId="10" pageNumber="515">Distribution and ecological notes.</emphasis>
|
||
This species is known only from the
|
||
<typeStatus id="54B6887CFFC3FFA8FC34FF03EF6FA8AA" box="[1018,1066,151,177]" pageId="10" pageNumber="515">type</typeStatus>
|
||
locality in the
|
||
<collectingRegion id="49C9F83CFFC3FFA8FB1CFF03EEDDA8AA" box="[1234,1432,151,177]" country="Colombia" name="Huila" pageId="10" pageNumber="515">Huila department</collectingRegion>
|
||
, at the inter-Andean valley of
|
||
<collectingRegion id="49C9F83CFFC3FFA8FE26FF2FE921A8CE" box="[488,612,187,213]" country="Colombia" name="Magdalena" pageId="10" pageNumber="515">Magdalena</collectingRegion>
|
||
River basin, reaching the
|
||
<quantity id="4CF59B3BFFC3FFA8FC5AFF2EE899A8CE" box="[916,988,186,213]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="10" pageNumber="515" unit="m" value="570.0">570 m</quantity>
|
||
elevation. Next to the
|
||
<typeStatus id="54B6887CFFC3FFA8FB2AFF2FEE51A8CE" box="[1252,1300,187,213]" pageId="10" pageNumber="515">type</typeStatus>
|
||
locality are some small artificial lakes and a first-order stream. However, attempts to collect the immature stages of the new species in these environments were unsuccessful.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176555FFC3FFA8FF09FEB2EF75ABBE" pageId="10" pageNumber="515" type="discussion">
|
||
<paragraph id="8BB236DEFFC3FFA8FF09FEB2EF75ABBE" blockId="10.[151,1437,150,933]" pageId="10" pageNumber="515">
|
||
<emphasis id="B979EACCFFC3FFA8FF09FEB2EA09A95B" bold="true" box="[199,332,294,320]" pageId="10" pageNumber="515">Discussion.</emphasis>
|
||
Sanseverino (2006) discussed relationships between
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FC65FEB3EF82A95B" authorityName="Contreras-Lichtenberg" authorityYear="1988" box="[939,1223,295,320]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="curvicristatus">
|
||
<emphasis id="B979EACCFFC3FFA8FC65FEB3EF82A95B" box="[939,1223,295,320]" italics="true" pageId="10" pageNumber="515">Tanytarsus curvicristatus</emphasis>
|
||
</taxonomicName>
|
||
and species of the
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FF59FEDFEA32A97F" box="[151,375,331,356]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="signatus">
|
||
<emphasis id="B979EACCFFC3FFA8FF59FEDFEA32A97F" box="[151,375,331,356]" italics="true" pageId="10" pageNumber="515">Tanytarsus signatus</emphasis>
|
||
</taxonomicName>
|
||
group, as well as those of former
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FCCFFEDFE8CFA97E" authorityName="Sawedal" authorityYear="1981" box="[769,906,331,357]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FCCFFEDFE8CFA97E" box="[769,906,331,357]" italics="true" pageId="10" pageNumber="515">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FC0DFEDFEF31A97F" authorityName="Pinder" authorityYear="1982" box="[963,1140,331,356]" class="Insecta" family="Chironomidae" genus="Virgatanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FC0DFEDFEF31A97F" box="[963,1140,331,356]" italics="true" pageId="10" pageNumber="515">Virgatanytarsus</emphasis>
|
||
</taxonomicName>
|
||
—the names evidenced as synonyms of
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FEFFFEFBE934A992" authority="(Lin et al. 2018)" baseAuthorityName="Lin" baseAuthorityYear="2018" box="[305,625,367,393]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FEFFFEFBEAEEA993" box="[305,427,367,392]" italics="true" pageId="10" pageNumber="515">Tanytarsus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9C4B2FFFC3FFA8FE75FEFBE92CA992" author="Lin, X. - L. & Stur, E. & Ekrem, T." box="[443,617,367,393]" pageId="10" pageNumber="515" pagination="659 - 677" refId="ref13096" refString="Lin, X. - L., Stur, E. & Ekrem, T. (2018) Molecular phylogeny and temporal diversification of Tanytarsus van der Wulp (Diptera: Chironomidae) support generic synonymies, a new classification and centre of origin. Systematic Entomology, 43 (4), 659 - 677. https: // doi. org / 10.1111 / syen. 12292" type="journal article" year="2018">
|
||
Lin
|
||
<emphasis id="B979EACCFFC3FFA8FE22FEFBE960A992" box="[492,549,367,393]" italics="true" pageId="10" pageNumber="515">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, thus validating a previously postulated concept of
|
||
<bibRefCitation id="EF9C4B2FFFC3FFA8FB75FEFBEE36A992" author="Cranston, P. S." box="[1211,1395,367,393]" pageId="10" pageNumber="515" pagination="138 - 159" refId="ref12531" refString="Cranston, P. S. (2000) Monsoonal tropical Tanytarsus van der Wulp (Diptera: Chironomidae) reviewed: new species, life histories and significance as aquatic environmental indicators. Australian Journal of Entomology, 39, 138 - 159. https: // doi. org / 10.1046 / J. 1440 - 6055.2000.00170. X" type="journal article" year="2000">Cranston (2000)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FF59FE07EA0DA9B7" authorityName="Pinder" authorityYear="1982" box="[151,328,403,428]" class="Insecta" family="Chironomidae" genus="Virgatanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FF59FE07EA0DA9B7" box="[151,328,403,428]" italics="true" pageId="10" pageNumber="515">Virgatanytarsus</emphasis>
|
||
</taxonomicName>
|
||
. The Sanseverino’s analysis was based on morphology of male hypopygium, mainly the anal point and its bars—the structures present in all the taxa analysed. In the dissertation, relations between
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FB09FE23EE3EA9CB" authorityName="Contreras-Lichtenberg" authorityYear="1988" box="[1223,1403,439,464]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="curvicristatus">
|
||
<emphasis id="B979EACCFFC3FFA8FB09FE23EE3EA9CB" box="[1223,1403,439,464]" italics="true" pageId="10" pageNumber="515">T. curvicristatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FA46FE23EA33A9ED" authority="(Holarctic)" baseAuthorityName="Holarctic" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="signatus">
|
||
<emphasis id="B979EACCFFC3FFA8FA46FE23EBB1A9EF" italics="true" pageId="10" pageNumber="515">T. signatus</emphasis>
|
||
(Holarctic)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FE65FE4FE9CEA9ED" authority="(Australia)" authorityName="Glover" authorityYear="1973" baseAuthorityName="Australia" box="[427,651,475,502]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="liepae">
|
||
<emphasis id="B979EACCFFC3FFA8FE65FE4FE94FA9EE" box="[427,522,475,501]" italics="true" pageId="10" pageNumber="515">T. liepae</emphasis>
|
||
(
|
||
<collectingCountry id="F31A764EFFC3FFA8FDD7FE4FE9C6A9ED" box="[537,643,475,502]" name="Australia" pageId="10" pageNumber="515">Australia</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
were defined as close and possibly closer to each other than to species of the former
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FEF6FE6BEA84AA02" authorityName="Sawedal" authorityYear="1981" box="[312,449,511,537]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FEF6FE6BEA84AA02" box="[312,449,511,537]" italics="true" pageId="10" pageNumber="515">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FE32FE6BE9E8AA03" authorityName="Pinder" authorityYear="1982" box="[508,685,511,536]" class="Insecta" family="Chironomidae" genus="Virgatanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FE32FE6BE9E8AA03" box="[508,685,511,536]" italics="true" pageId="10" pageNumber="515">Virgatanytarsus</emphasis>
|
||
</taxonomicName>
|
||
. Due to a peculiar structure of the anal point in
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FB10FE6BEED0AA03" authorityName="Contreras-Lichtenberg" authorityYear="1988" box="[1246,1429,511,536]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="curvicristatus">
|
||
<emphasis id="B979EACCFFC3FFA8FB10FE6BEED0AA03" box="[1246,1429,511,536]" italics="true" pageId="10" pageNumber="515">T. curvicristatus</emphasis>
|
||
</taxonomicName>
|
||
, a new species group for this species was also suggested. A close species described later,
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FB41FDB7EED0AA26" authorityName="Trivinho-Strixino, Wiedenbrug et da Silva" authorityYear="2015" box="[1167,1429,547,573]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="pseudocurvicristatus">
|
||
<emphasis id="B979EACCFFC3FFA8FB41FDB7EED0AA26" box="[1167,1429,547,573]" italics="true" pageId="10" pageNumber="515">T. pseudocurvicristatus</emphasis>
|
||
</taxonomicName>
|
||
, supported this concept (
|
||
<bibRefCitation id="EF9C4B2FFFC3FFA8FE51FDD3E9A3AA7A" author="Trivinho-Strixino, S. & Wiedenbrug, S. & da Silva, F. L." box="[415,742,582,609]" pageId="10" pageNumber="515" pagination="92 - 100" refId="ref14804" refString="Trivinho-Strixino, S., Wiedenbrug, S. & da Silva, F. L. (2015) New species of Tanytarsus van der Wulp (Diptera: Chironomidae: Tanytarsini) from Brazil. European Journal of Environmental Sciences, 5 (1), 92 - 100. https: // doi. org / 10.14712 / 23361964.2015.82" type="journal article" year="2015">
|
||
Trivinho-Strixino
|
||
<emphasis id="B979EACCFFC3FFA8FDA5FDD3E9E6AA7A" box="[619,675,583,609]" italics="true" pageId="10" pageNumber="515">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). Further, molecular analyses indicated
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FB53FDD3EE50AA7B" box="[1181,1301,583,608]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="signatus">
|
||
<emphasis id="B979EACCFFC3FFA8FB53FDD3EE50AA7B" box="[1181,1301,583,608]" italics="true" pageId="10" pageNumber="515">T. signatus</emphasis>
|
||
</taxonomicName>
|
||
as a lineage genetically divergent from that of
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FDD9FDFFE98AAA9F" authorityName="Contreras-Lichtenberg" authorityYear="1988" box="[535,719,619,645]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="curvicristatus">
|
||
<emphasis id="B979EACCFFC3FFA8FDD9FDFFE962AA9F" box="[535,551,619,644]" italics="true" pageId="10" pageNumber="515">T</emphasis>
|
||
.
|
||
<emphasis id="B979EACCFFC3FFA8FDFBFDFFE98AAA9F" box="[565,719,619,644]" italics="true" pageId="10" pageNumber="515">curvicristatus</emphasis>
|
||
</taxonomicName>
|
||
, while the latter species and its morphologically close relatives, including
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FEC6FD1BE955AAB2" authorityName="Trivinho-Strixino, Wiedenbrug et da Silva" authorityYear="2015" box="[264,528,655,681]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="pseudocurvicristatus">
|
||
<emphasis id="B979EACCFFC3FFA8FEC6FD1BEA5DAAB3" box="[264,280,655,680]" italics="true" pageId="10" pageNumber="515">T</emphasis>
|
||
.
|
||
<emphasis id="B979EACCFFC3FFA8FEE9FD1BE955AAB2" box="[295,528,655,681]" italics="true" pageId="10" pageNumber="515">pseudocurvicristatus</emphasis>
|
||
</taxonomicName>
|
||
, again were suggested to be placed in a separate species group (
|
||
<bibRefCitation id="EF9C4B2FFFC3FFA8FB2AFD1BEED4AAB2" author="Lin, X. - L. & Stur, E. & Ekrem, T." box="[1252,1425,655,681]" pageId="10" pageNumber="515" pagination="659 - 677" refId="ref13096" refString="Lin, X. - L., Stur, E. & Ekrem, T. (2018) Molecular phylogeny and temporal diversification of Tanytarsus van der Wulp (Diptera: Chironomidae) support generic synonymies, a new classification and centre of origin. Systematic Entomology, 43 (4), 659 - 677. https: // doi. org / 10.1111 / syen. 12292" type="journal article" year="2018">
|
||
Lin
|
||
<emphasis id="B979EACCFFC3FFA8FADAFD1BEE08AAB2" box="[1300,1357,655,681]" italics="true" pageId="10" pageNumber="515">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). Another two species, presented here as new, made it possible to formulate a diagnosis based on characters found in all four species, and to establish the previously expected
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FCD9FD43EF77AAEB" authorityName="Contreras-Lichtenberg" authorityYear="1988" box="[791,1074,727,752]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="curvicristatus">
|
||
<emphasis id="B979EACCFFC3FFA8FCD9FD43EF77AAEB" box="[791,1074,727,752]" italics="true" pageId="10" pageNumber="515">Tanytarsus curvicristatus</emphasis>
|
||
</taxonomicName>
|
||
species group. These characters are the shape and arrangement of the anal point crests, bars, the lack of the spinulae typical of most
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FAD5FD6FEED0AB0F" box="[1307,1429,763,788]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FAD5FD6FEED0AB0F" box="[1307,1429,763,788]" italics="true" pageId="10" pageNumber="515">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
, as well as the shape of the superior volsella and digitus (see the group diagnosis). The anal point bar(s) shape/ arrangement seems to be crucial in phylogeny analyses based on morphology of
|
||
<taxonomicName id="4C0D4D5DFFC3FFA8FBECFCD7EFD9AB47" box="[1058,1180,835,860]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACCFFC3FFA8FBECFCD7EFD9AB47" box="[1058,1180,835,860]" italics="true" pageId="10" pageNumber="515">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
. These structures have recently been studied in both extant and fossil species (
|
||
<bibRefCitation id="EF9C4B2FFFC3FFA8FCCCFCF3EF4EAB9A" author="Trivinho-Strixino, S." box="[770,1035,871,897]" pageId="10" pageNumber="515" pagination="1 - 41" refId="ref14337" refString="Trivinho-Strixino, S. (2012) A systematic review of Neotropical Caladomyia Sawedal (Diptera: Chironomidae). Zootaxa, 3495 (1), 1 - 41. https: // doi. org / 10.11646 / zootaxa. 3495.1.1" type="journal article" year="2012">Trivinho-Strixino 2012</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9C4B2FFFC3FFA8FBD9FCF3EE06AB9A" author="Zakrzewska, M. & Gilka, W." box="[1047,1347,871,897]" pageId="10" pageNumber="515" pagination="397 - 403" refId="ref14923" refString="Zakrzewska, M. & Gilka, W. (2013) In the Eocene, the extant genus Caladomyia occurred in the Palaearctic (Diptera: Chironomidae: Tanytarsini). Polish Journal of Entomology, 82, 397 - 403. https: // doi. org / 10.2478 / pjen- 2013 - 0003" type="journal article" year="2013">Zakrzewska & Giłka 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9C4B2FFFC3FFA8FA80FCF3EA70ABBE" author="Dantas, G. P. S. & Gilka, W." pageId="10" pageNumber="515" pagination="281 - 291" refId="ref12727" refString="Dantas, G. P. S. & Gilka, W. (2017) New Tanytarsus van der Wulp from the Brazilian Amazonia indicate clues to intrageneric relations (Diptera: Chironomidae). Zootaxa, 4294 (2), 281 - 291. https: // doi. org / 10.11646 / zootaxa. 4294.2.10" type="journal article" year="2017">Dantas & Giłka 2017</bibRefCitation>
|
||
), although they are found in only a few species groups of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |