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<document id="65C8D2CA3DA418039CFC170CDAF51D74" ID-CLB-Dataset="9458" ID-DOI="10.11646/zootaxa.4889.1.1" ID-GBIF-Dataset="11a122a9-8f3e-4a1c-a8d3-5ef3bea09b59" ID-ISSN="1175-5326" ID-Zenodo-Dep="4297816" ID-ZooBank="0B8630F6-2EF0-44E6-9D3A-7386BF949FD0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606731381730" checkinUser="plazi" docAuthor="Grzymala, Traci L. &amp; Leschen, Richard A. B." docDate="2020" docId="03A48794FFFBFFC76F854BF17543FD81" docLanguage="en" docName="zootaxa.4889.1.1.pdf" docOrigin="Zootaxa 4889 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Zenascus incensum Grzymala &amp; Leschen 2020, sp. n." docType="treatment" docUuid="620CE468-9C1D-415E-8749-7ED246EDF478" docUuidSource="ZooBank" docVersion="13" lastPageNumber="39" masterDocId="FF9DFFECFFDEFFE16F124B66712FFFBC" masterDocTitle="Sexual Dimorphism of New Zealand Puppet Beetles (Aderidae, Coleoptera, Tenebrionoidea): Systematic Revision, Description of Three New Genera, and Phylogeny for Zenascus, gen. n." masterLastPageNumber="59" masterPageNumber="1" pageNumber="38" updateTime="1698846900633" updateUser="ExternalLinkService">
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<mods:title id="F81E6B03CC95EC3B6C5BC1D03D3CA78C">Sexual Dimorphism of New Zealand Puppet Beetles (Aderidae, Coleoptera, Tenebrionoidea): Systematic Revision, Description of Three New Genera, and Phylogeny for Zenascus, gen. n.</mods:title>
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<mods:namePart id="9AB256F7A903FE026FCF4F06D169D7EF">Grzymala, Traci L.</mods:namePart>
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<mods:namePart id="C7B1D22B2ED1C4253E6CBC89DEF7D73E">Leschen, Richard A. B.</mods:namePart>
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<mods:date id="E77A2CA5A458495D192B7EF0533DBDF5">2020</mods:date>
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<mods:number id="64A95E58142BBBE15DD7A7250FFF23CC">2020-11-30</mods:number>
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<treatment id="03A48794FFFBFFC76F854BF17543FD81" ID-DOI="http://doi.org/10.5281/zenodo.4334368" ID-GBIF-Taxon="170606610" ID-Zenodo-Dep="4334368" LSID="urn:lsid:zoobank.org:act:620CE468-9C1D-415E-8749-7ED246EDF478" httpUri="http://treatment.plazi.org/id/03A48794FFFBFFC76F854BF17543FD81" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="C3176509FFFBFFC46F854BF170E0FF0D" box="[151,463,151,177]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BB23682FFFBFFC46F854BF170E0FF0D" blockId="37.[151,707,151,214]" box="[151,463,151,177]" pageId="37" pageNumber="38">
<heading id="D0FA81EEFFFBFFC46F854BF170E0FF0D" bold="true" box="[151,463,151,177]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
<emphasis id="B979EA90FFFBFFC46F854BF170E0FF0D" bold="true" box="[151,463,151,177]" pageId="37" pageNumber="38">
<taxonomicName id="4C0D4D01FFFBFFC46F854BF170A9FF0D" authority="Grzymala &amp; Leschen, 2020" authorityName="Grzymala &amp; Leschen" authorityYear="2020" box="[151,390,151,177]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="incensum" status="sp. nov.">
<emphasis id="B979EA90FFFBFFC46F854BF170A9FF0D" bold="true" box="[151,390,151,177]" italics="true" pageId="37" pageNumber="38">Zenascus incensum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A57EBFFFBFFC46E9F4BF170E0FF0D" box="[397,463,151,177]" pageId="37" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3176509FFFBFFC46F854BDD73ECFF69" box="[151,707,187,214]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BB23682FFFBFFC46F854BDD73ECFF69" blockId="37.[151,707,151,214]" box="[151,707,187,214]" pageId="37" pageNumber="38">
<figureCitation id="13362A07FFFBFFC46F854BDD71D1FF69" box="[151,254,187,213]" captionStart="FIGURES 12" captionStartId="31.[151,264,1912,1936]" captionTargetBox="[158,1428,787,1880]" captionTargetId="figure-291@31.[151,1436,777,1889]" captionTargetPageId="31" captionText="FIGURES 12AE. Dorsal habitus images. (A) Zenascus antennalis (Broun), male; (B) Zenascus xenarthrus (Broun), male; (C) Zenascus roberti sp. n., male; (D) Zenascus luniger (Champion), male; (E) Zenascus incensum sp. n., male. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4297842" httpUri="https://zenodo.org/record/4297842/files/figure.png" pageId="37" pageNumber="38">Figs 12E</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46E184BDD701CFF69" box="[266,307,187,213]" captionStart="FIGURES 14" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[268,1319,181,1901]" captionTargetId="figure-17@33.[268,1319,181,1919]" captionTargetPageId="33" captionText="FIGURES 14AJ. Antennal illustrations for Zenascus species, male on left, female on right.(A) Z. coloratus; (B) Z. obscurus; (C) Z. aurum, sp. n.; (D) Z. nitidus; (E) Z. elenae, sp. n.; (F) Z. antennalis; (G) Z. xenarthrus; (H) Z. roberti, sp. n.; (I) Z. luniger; (J) Z. incensum, sp. n. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4297849" httpUri="https://zenodo.org/record/4297849/files/figure.png" pageId="37" pageNumber="38">14J</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46E2C4BDD705FFF6A" box="[318,368,187,214]" captionStart="FIGURES 15" captionStartId="38.[151,264,1639,1663]" captionTargetBox="[152,1435,1095,1614]" captionTargetId="figure-366@38.[152,1436,1095,1615]" captionTargetPageId="38" captionText="FIGURES 15AC. Scanning electron microscope images of Zenascus species with close-ups of male antennomeres 79. (A) Z. roberti, sp. n.; (B) Z. luniger; (C) Z. incensum, sp. n.. Scale bars = 20μm." figureDoi="http://doi.org/10.5281/zenodo.4297851" httpUri="https://zenodo.org/record/4297851/files/figure.png" pageId="37" pageNumber="38">15C</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46E694BDD7085FF69" box="[379,426,187,213]" captionStart="FIGURES 19" captionStartId="44.[151,264,1421,1445]" captionTargetBox="[157,1433,185,1393]" captionTargetId="figure-28@44.[151,1436,181,1397]" captionTargetPageId="44" captionText="FIGURES 19AE. Scanning electron microscope images, Zenascus species, elytron, dorsal view. (A) Z. antennalis; (B) Z. xenarthrus; (C) Z. roberti, sp. n.; (D) Z. luniger; (E) Z. incensum, sp. n. Scale bars = 10μm." figureDoi="http://doi.org/10.5281/zenodo.4297859" httpUri="https://zenodo.org/record/4297859/files/figure.png" pageId="37" pageNumber="38">19E</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46EA44BDD70F0FF69" box="[438,479,187,213]" captionStart="FIGURES 20" captionStartId="45.[151,264,1411,1435]" captionTargetBox="[151,1436,184,1384]" captionTargetId="figure-17@45.[151,1436,181,1387]" captionTargetPageId="45" captionText="FIGURES 20AJ. Scanning electron microscope images, ventral view of metafemora for Zenascus species. (A) Z. coloratus, male; (B) Z. obscurus, male; (C) Z. aurum, sp. n., male; (D) Z. nitidus, female; (E) Z. elenae, sp. n., male; (F) Z. antennalis, female; (G) Z. xenarthrus, male; (H) Z. roberti, sp. n., male; (I) Z. luniger, male; (J) Z. incensum, sp. n., male. Scale bars = 60μm." figureDoi="http://doi.org/10.5281/zenodo.4297861" httpUri="https://zenodo.org/record/4297861/files/figure.png" pageId="37" pageNumber="38">20J</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46EF94BDD7334FF69" box="[491,539,187,213]" captionStart="FIGURES 22" captionStartId="47.[151,264,1163,1187]" captionTargetBox="[151,1436,186,1140]" captionTargetId="figure-17@47.[151,1436,181,1140]" captionTargetPageId="47" captionText="FIGURES 22AE. Abdominal ventrite illustrations for Zenascus species showing generalized punctation pattern. (A) Z. antennalis, male; (B) Z. xenarthrus, male; (C) Z. roberti, sp. n., male; (D) Z. luniger, male; (E) Z. incensum, sp. n., male." figureDoi="http://doi.org/10.5281/zenodo.4297873" httpUri="https://zenodo.org/record/4297873/files/figure.png" pageId="37" pageNumber="38">22E</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46D344BDD737FFF69" box="[550,592,187,213]" captionStart="FIGURES 23" captionStartId="49.[151,264,1943,1967]" captionTargetBox="[231,1359,181,1919]" captionTargetId="figure-17@49.[227,1359,181,1919]" captionTargetPageId="49" captionText="FIGURES 23AJ. Male genitalia illustrations for Zenascus species. (A) Z. coloratus; (B) Z. obscurus; (C) Z. aurum, sp. n.; (D) Z. nitidus; (E) Z. elenae, sp. n.; (F) Z. antennalis; (G) Z. xenarthrus; (H) Z. roberti, sp. n.; (I) Z. luniger; (J) Z. incensum, sp. n." figureDoi="http://doi.org/10.5281/zenodo.4297877" httpUri="https://zenodo.org/record/4297877/files/figure.png" pageId="37" pageNumber="38">23J</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46D494BDD73A8FF69" box="[603,647,187,213]" captionStart="FIGURES 24" captionStartId="50.[151,264,1642,1666]" captionTargetBox="[159,1431,183,1612]" captionTargetId="figure-28@50.[151,1436,181,1619]" captionTargetPageId="50" captionText="FIGURES 24AF. Scanning electron micrographs of gland pores on metafemur of Zenascus spp. (A) Z. aurum, sp. n.; (B) Z. antennalis; (C) Z. xenarthrus; (D) Z. roberti, sp. n.; (E) Z. luniger; (F) Z. incensum, sp. n. Scale bars = 2μm." figureDoi="http://doi.org/10.5281/zenodo.4297881" httpUri="https://zenodo.org/record/4297881/files/figure.png" pageId="37" pageNumber="38">24F</figureCitation>
,
<figureCitation id="13362A07FFFBFFC46D804BDD73ECFF69" box="[658,707,187,213]" captionStart="FIGURES 27" captionStartId="54.[151,264,1939,1963]" captionTargetBox="[161,1428,193,1906]" captionTargetId="figure-28@54.[151,1436,181,1916]" captionTargetPageId="54" captionText="FIGURES 27AD. Distribution maps. (A) Zenascus aurum, sp. n., Zenascus incensum, sp. n., and Zenascus xenarthrus, (B) Zenascus coloratus, (C) Zenascus elenae, sp. n. and (D) Zenascus luniger." figureDoi="http://doi.org/10.5281/zenodo.4297887" httpUri="https://zenodo.org/record/4297887/files/figure.png" pageId="37" pageNumber="38">27A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFFBFFC46F854A657295FEDA" pageId="37" pageNumber="38" type="etymology">
<paragraph id="8BB23682FFFBFFC46F854A657295FEDA" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">
<emphasis id="B979EA90FFFBFFC46F854A657031FEA1" bold="true" box="[151,286,259,285]" pageId="37" pageNumber="38">Etymology.</emphasis>
The specific epithet is in reference to frankincense, one of the gifts brought by the three kings on the Epiphany, which refers to the Three Kings Islands (Manawatâwhi) where this species appears to be biogeographically restricted. The name is formed from the Latin
<taxonomicName id="4C0D4D01FFFBFFC46DC94A2A726AFED9" authorityName="Grzymala &amp; Leschen" authorityYear="2020" box="[731,837,332,357]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="incensum">
<emphasis id="B979EA90FFFBFFC46DC94A2A726AFED9" box="[731,837,332,357]" italics="true" pageId="37" pageNumber="38">incensum</emphasis>
</taxonomicName>
(incense).
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFFBFFC46FD54A0973DDFDDD" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BB23682FFFBFFC46FD54A0973DDFDDD" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">
<emphasis id="B979EA90FFFBFFC46FD54A09706DFE35" bold="true" box="[199,322,367,393]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
This species is distinguished from congeners by the biseriate pubescence of the pronotum and elytra, the highly modified antennomeres 79 of males, and the slightly impressed basal pronotal fovea.
<taxonomicName id="4C0D4D01FFFBFFC46A264AF2702EFE6D" authorityName="Grzymala &amp; Leschen" authorityYear="2020" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="incensum">
<emphasis id="B979EA90FFFBFFC46A264AF2702EFE6D" italics="true" pageId="37" pageNumber="38">Zenascus incensum</emphasis>
</taxonomicName>
is most easily confused with
<taxonomicName id="4C0D4D01FFFBFFC46D484ADE7232FE6D" authorityName="Grzymala &amp; Leschen" authorityYear="2020" baseAuthorityName="Grzymala &amp; Leschen" baseAuthorityYear="2020" box="[602,797,439,465]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="luniger">
<emphasis id="B979EA90FFFBFFC46D484ADE7232FE6D" box="[602,797,439,465]" italics="true" pageId="37" pageNumber="38">Zenascus luniger</emphasis>
</taxonomicName>
, but males are distinguished between these two species based on the presence of a distinct vertexal ridge in
<taxonomicName id="4C0D4D01FFFBFFC46DC84ABA724EFE49" authorityName="Grzymala &amp; Leschen" authorityYear="2020" box="[730,865,476,501]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="incensum">
<emphasis id="B979EA90FFFBFFC46DC84ABA724EFE49" box="[730,865,476,501]" italics="true" pageId="37" pageNumber="38">Z. incensum</emphasis>
</taxonomicName>
and the modified shape of antennomeres 79 (
<figureCitation id="13362A07FFFBFFC46A784ABD7033FDA6" captionStart="FIGURES 15" captionStartId="38.[151,264,1639,1663]" captionTargetBox="[152,1435,1095,1614]" captionTargetId="figure-366@38.[152,1436,1095,1615]" captionTargetPageId="38" captionText="FIGURES 15AC. Scanning electron microscope images of Zenascus species with close-ups of male antennomeres 79. (A) Z. roberti, sp. n.; (B) Z. luniger; (C) Z. incensum, sp. n.. Scale bars = 20μm." figureDoi="http://doi.org/10.5281/zenodo.4297851" httpUri="https://zenodo.org/record/4297851/files/figure.png" pageId="37" pageNumber="38">Figs 15B &amp; 15C</figureCitation>
). Females are more difficult to separate, but currently
<taxonomicName id="4C0D4D01FFFBFFC46C9249667527FDA5" authorityName="Grzymala &amp; Leschen" authorityYear="2020" box="[896,1032,512,537]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="incensum">
<emphasis id="B979EA90FFFBFFC46C9249667527FDA5" box="[896,1032,512,537]" italics="true" pageId="37" pageNumber="38">Z. incensum</emphasis>
</taxonomicName>
has only been collected from Three Kings Island whereas
<taxonomicName id="4C0D4D01FFFBFFC46E9F494270D4FD81" authorityName="Grzymala &amp; Leschen" authorityYear="2020" baseAuthorityName="Grzymala &amp; Leschen" baseAuthorityYear="2020" box="[397,507,547,573]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="luniger">
<emphasis id="B979EA90FFFBFFC46E9F494270D4FD81" box="[397,507,547,573]" italics="true" pageId="37" pageNumber="38">Z. luniger</emphasis>
</taxonomicName>
has only been collected from the North Island of
<collectingCountry id="F31A7612FFFBFFC46B3049427594FD81" box="[1058,1211,547,574]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
. This biogeographic separation differentiates females of these two species.
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFFBFFC76FD5490D75D3FEFD" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="8BB23682FFFBFFC46FD5490D73A7F8CD" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">
<emphasis id="B979EA90FFFBFFC46FD5490D7076FD39" bold="true" box="[199,345,619,645]" pageId="37" pageNumber="38">Description.</emphasis>
Length
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. Head, scape, pedicel, antennomeres 711, and pronotum piceus to rufotestaceous; antennomeres 36, mouthparts, and elytra brunneous to golden. Dorsal integument matte, vestiture biseriate, one decumbent, thin, short seta arising anterad of each primary puncture, length about 3× puncture length, with 34 additional decumbent, thin, shorter setae between each pair of punctures, length about 2/3× primary seta. Ventral vestiture similarly biseriate. Head, HW
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, HL
<quantity id="4CF59B67FFFBFFC46C81499D7517FCAA" box="[915,1080,763,790]" metricMagnitude="-4" metricUnit="m" metricValue="1.65" metricValueMax="1.7999999999999998" metricValueMin="1.5" pageId="37" pageNumber="38" unit="mm" value="0.16499999999999998" valueMax="0.18" valueMin="0.15">0.150.18 mm</quantity>
, punctation consisting of a few, deep punctures, unevenly spaced on vertex, lacking from base to posterior edge of eyes, strongly impressed and concave between antennal ridge and vertex, occiput of head in male with distinct transverse ridge. Antennae with distance between antennal insertions small, approximately equal to diameter of two antennal insertions; length reaching past basal ½ of elytra when extended backward; scape broad and flattened, approximately 3× length of pedicel; pedicel subglobular; antennomere 3 shorter than pedicel and shorter than antennomere 4; antennomere 4 approximately 3× length of antennomere 3, longer than antennomere 5; antennomeres 5 and 6 subequal in length and width, slightly broader than antennomere 4; antennomere 7 greatly modified, flattened, broad, and branched just anterad of apex, with apex of outer branch terminating in broad triangle, with apex of inner branch connecting to remaining antennomeres, with apex acute, extending ¼ past antennomere 8; antennomere 8 longer than wide, with strong outwards curvature; antennomere 9 with modified outer apex, projecting anteriorly, apex rounded, with dense pilosity extending behind; antennomere 10 subquadrate, slightly expanded apically; antennomere 11 longer than 10; antennomeres 311 covered in moderately dense, suberect pubescence, antennomeres 79 with additional pilosity; pedicel and antennomeres 311 with apical ring of elongate setae, antennomere 11 with additional medial ring of setae. Pronotum, PW
<quantity id="4CF59B67FFFBFFC46ECB4F957352FAB2" box="[473,637,1267,1294]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="37" pageNumber="38" unit="mm" value="0.395" valueMax="0.41" valueMin="0.38">0.380.41 mm</quantity>
, PL
<quantity id="4CF59B67FFFBFFC46DBC4F95727DFAB2" box="[686,850,1267,1294]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="37" pageNumber="38" unit="mm" value="0.395" valueMax="0.41" valueMin="0.38">0.380.41 mm</quantity>
, subquadrate, width subequal to slightly greater than length, width 1.01.1× length; pronotal width 0.530.59× elytral width, pronotal width 0.810.87× head width; sides slightly rounded, posterior angles rounded; disc with two slightly impressed basal fovea, with slight transverse sulcus just anterad of center; punctaion consisting of deep, small elongate punctures, unevenly spaced, more sparse basally and laterally. Elytra 1.671.87× longer than wide and 3.153.22× longer than pronotal length, EW
<quantity id="4CF59B67FFFBFFC46F854EC17012FA7E" box="[151,317,1447,1474]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.199999999999999" metricValueMin="6.8999999999999995" pageId="37" pageNumber="38" unit="mm" value="0.705" valueMax="0.72" valueMin="0.69">0.690.72 mm</quantity>
, EL
<quantity id="4CF59B67FFFBFFC46E634EC17337FA7E" box="[369,536,1447,1474]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" metricValueMax="1.32" metricValueMin="1.2" pageId="37" pageNumber="38" unit="mm" value="1.26" valueMax="1.32" valueMin="1.2">1.201.32 mm</quantity>
; slightly impressed, oblique subscutellar depression present; punctation consisting of elongate punctures, separated by an average of two punctural lengths. Proleg with tarsomere 1 approximately 23× longer than tarsomere 2, without a ventral short, stout spine; tarsomere 3 with ventral adhesive setae. Midleg with tibia gently curved inward from base to apex. Hindleg with numerous deep, elongate punctures along posterior margin of coxa; femur with distinctly thickened setae on posterior face, overlying slight medial excavation along length of femur, apico-ventral impression absent; tibia gradually expanded in width from base to apex, apex inner face triangular and apex densely pubescent; tarsomere 1 elongate, length 10× width; tarsomere 2 expanded ventrally. Abdomen with lateral length of ventrite 2 longer than its medial length; deep punctures confined to abdominal process, basal margin of ventrite 1, basal margin and medially on ventrite 2, and a few scattered on ventrite 3; small shallow micropunctures present on ventrites 4 and 5; ventrite 5 without medial impression. Phallobase broadly rounded anteriorly, laterally delimited from apicale; apicale posteriorly narrowed; accessory lobes present, with four setae, one long and two short located at apex, one long located basal to apex on inner face; penis with anterior struts elongate, extending slightly past phallobase.
</paragraph>
<paragraph id="8BB23682FFFBFFC76FD54C1D750EFF45" blockId="37.[151,1437,259,2014]" lastBlockId="38.[151,1436,151,573]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<emphasis id="B979EA90FFFBFFC46FD54C1D701FF829" bold="true" box="[199,304,1915,1941]" pageId="37" pageNumber="38">Females.</emphasis>
Length
<quantity id="4CF59B67FFFBFFC46E834C1D70D6F82A" box="[401,505,1915,1942]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="37" pageNumber="38" unit="mm" value="1.84">1.84 mm</quantity>
, HW
<quantity id="4CF59B67FFFBFFC46D2F4C1D738AF82A" box="[573,677,1915,1942]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="37" pageNumber="38" unit="mm" value="0.48">0.48 mm</quantity>
, HL
<quantity id="4CF59B67FFFBFFC46DCD4C1D7268F82A" box="[735,839,1915,1942]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="37" pageNumber="38" unit="mm" value="0.18">0.18 mm</quantity>
, PW
<quantity id="4CF59B67FFFBFFC46C944C1D72C1F82A" box="[902,1006,1915,1942]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="37" pageNumber="38" unit="mm" value="0.45">0.45 mm</quantity>
, PL
<quantity id="4CF59B67FFFBFFC46B364C1D75A4F82A" box="[1060,1163,1915,1942]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="37" pageNumber="38" unit="mm" value="0.41">0.41 mm</quantity>
, EW
<quantity id="4CF59B67FFFBFFC46BDE4C1D741BF82A" box="[1228,1332,1915,1942]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="37" pageNumber="38" unit="mm" value="0.78">0.78 mm</quantity>
, EL
<quantity id="4CF59B67FFFBFFC46A794C1D71E9F806" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="37" pageNumber="38" unit="mm" value="1.43">1.43 mm</quantity>
. Head very slightly impressed, concave between antennal ridge and vertex. Antennae with scape laterally flattened, slightly longer than wide, pedicel subglobular; antennomere 3 slightly reduced in width compared to pedicel, slightly expanded apically; antennomere 4 slightly longer than 3; antennomeres 46 subequal in length and width; antennomeres 810 subsequently decreasing slightly in length and increasing slightly in width; antennomere 11 elongate; antennomeres 311 covered in moderately dense, suberect pubescence.
</paragraph>
<paragraph id="8BB23682FFF8FFC76FD54A6575D3FEFD" blockId="38.[151,1436,151,573]" pageId="38" pageNumber="39">
<emphasis id="B979EA90FFF8FFC76FD54A6570AAFEA0" bold="true" box="[199,389,258,285]" pageId="38" pageNumber="39">Natural history.</emphasis>
This Three Kings Islands endemic has primarily been collected by beating and sweeping vegetation, specifically from grasses and sedges. It has additionally been collected by sweeping kânuka.
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFF8FFC76FD54A2C7263FED9" box="[199,844,330,357]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BB23682FFF8FFC76FD54A2C7263FED9" blockId="38.[151,1436,151,573]" box="[199,844,330,357]" pageId="38" pageNumber="39">
<emphasis id="B979EA90FFF8FFC76FD54A2C704EFED8" bold="true" box="[199,353,330,356]" pageId="38" pageNumber="39">Distribution.</emphasis>
<materialsCitation id="3B653CDFFFF8FFC76E754A2C7263FED9" ID-GBIF-Occurrence="2989339322" box="[359,844,330,357]" country="New Zealand" location="Offshore island" pageId="38" pageNumber="39" specimenCount="1">
<emphasis id="B979EA90FFF8FFC76E754A2C7306FED8" bold="true" box="[359,553,330,356]" pageId="38" pageNumber="39">
<location id="8ED26059FFF8FFC76E754A2C730BFED8" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76E754A2C730BFED8" box="[359,548,330,356]" name="Offshore island" pageId="38" pageNumber="39">Offshore island</location>
:
</emphasis>
<location id="8ED26059FFF8FFC76D3D4A2D722AFED9" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76D3D4A2D722AFED9" box="[559,773,331,357]" name="Three Kings Island" pageId="38" pageNumber="39">Three Kings Island</location>
(TH).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFF8FFC76FD54A097543FD81" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BB23682FFF8FFC76FD54A097543FD81" blockId="38.[151,1436,151,573]" pageId="38" pageNumber="39">
<emphasis id="B979EA90FFF8FFC76FD54A0970DFFE34" bold="true" box="[199,496,366,393]" pageId="38" pageNumber="39">Type material examined.</emphasis>
<materialsCitation id="3B653CDFFFF8FFC76EE54A0971F0FE6D" ID-GBIF-Occurrence="2988225369" collectingDate="1983-11-27" collectionCode="NZAC" collectorName="C. F. Butcher" country="New Zealand" determinerName="S. E. Thorpe" location="THREE KINGS IS" pageId="38" pageNumber="39" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B68820FFF8FFC76EE54A09734CFE35" box="[503,611,367,393]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
.
<specimenCount id="9D0BFD0BFFF8FFC76D7C4A097387FE35" box="[622,680,367,393]" pageId="38" pageNumber="39" type="male">Male</specimenCount>
(
<collectionCode id="ED1CAE47FFF8FFC76DA54A097226FE35" box="[695,777,367,393]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" pageId="38" pageNumber="39">NZAC</collectionCode>
), labeled: “
<location id="8ED26059FFF8FFC76C944A09754FFE35" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76C944A09754FFE35" box="[902,1120,367,393]" country="New Zealand" name="THREE KINGS IS" pageId="38" pageNumber="39">THREE KINGS IS</location>
<collectingCountry id="F31A7612FFF8FFC76B7A4A0975A2FE35" box="[1128,1165,367,393]" name="New Zealand" pageId="38" pageNumber="39">NZ</collectingCountry>
/ Great
<date id="FFB31042FFF8FFC76BFB4A09740AFE35" box="[1257,1317,367,393]" bridgedPair="/" pageId="38" pageNumber="39" value="1927-01">
I /
<collectingDate id="EFF7E9AAFFF8FFC76A1B4A09740AFE35" box="[1289,1317,367,393]" pageId="38" pageNumber="39" value="1983-11-27">27</collectingDate>
</date>
Nov 1983 /
<collectorName id="26F85354FFF8FFC76FB44AF5701BFE11" box="[166,308,403,429]" pageId="38" pageNumber="39">C.F. Butcher</collectorName>
//ex. Grasses and / sedges // Xylophilus luniger /
<bibRefCitation id="EF9C4B73FFF8FFC76C7C4AF5750EFE11" author="Champion, G. C." box="[878,1057,403,429]" pageId="38" pageNumber="39" pagination="1 - 64" refId="ref39507" refString="Champion, G. C. (1916) I. On new or little known Xylophilidae. Transactions of the Royal Entomological Society of London, 64 (1), 1 - 64. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03119. x" type="journal article" year="1916">Champion 1916</bibRefCitation>
[male symbol] / det.
<determinerName id="7E6EDA46FFF8FFC76A1D4AF574B7FE11" box="[1295,1432,403,429]" pageId="38" pageNumber="39">S.E. Thorpe</determinerName>
, 2002”
</materialsCitation>
.
<materialsCitation id="3B653CDFFFF8FFC76FF84AD170D0FE49" ID-GBIF-Occurrence="2988225380" collectingDate="1983-11-27" collectingMethod="sweeping vegetation mainly" collectionCode="NZAC" collectorName="C. F. Butcher" country="New Zealand" location="Offshore island" municipality="Great Is" pageId="38" pageNumber="39" specimenCount="3" typeStatus="paratype">
<typeStatus id="54B68820FFF8FFC76FF84AD17078FE6D" box="[234,343,439,465]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
(17).
<emphasis id="B979EA90FFF8FFC76E894AD073BDFE6D" bold="true" box="[411,658,438,465]" pageId="38" pageNumber="39">
<location id="8ED26059FFF8FFC76E894AD07379FE6C" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76E894AD07379FE6C" box="[411,598,438,464]" country="New Zealand" municipality="Great Is" name="Offshore island" pageId="38" pageNumber="39">Offshore island</location>
. TH:
</emphasis>
<location id="8ED26059FFF8FFC76D884AD173D9FE6D" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76D884AD173D9FE6D" box="[666,758,439,465]" country="New Zealand" municipality="Great Is" name="Great Is" pageId="38" pageNumber="39">Great Is</location>
,
<date id="FFB31042FFF8FFC76C104AD172ACFE6D" box="[770,899,439,465]" pageId="38" pageNumber="39" value="1983-11-27">
<collectingDate id="EFF7E9AAFFF8FFC76C104AD172ACFE6D" box="[770,899,439,465]" pageId="38" pageNumber="39" value="1983-11-27">27/11/1983</collectingDate>
</date>
,
<collectingMethod id="524C4E95FFF8FFC76C9D4AD175E5FE6D" box="[911,1226,439,465]" pageId="38" pageNumber="39">sweeping vegetation mainly</collectingMethod>
<emphasis id="B979EA90FFF8FFC76BC34AD17442FE6C" box="[1233,1389,439,464]" italics="true" pageId="38" pageNumber="39">Leptopermum</emphasis>
<taxonomicName id="4C0D4D01FFF8FFC76A664AD17055FE49" authority=", C. F. Butcher" authorityName="C. F. Butcher" class="Magnoliopsida" family="Myrtaceae" genus="Kunzea" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="ericoides">
<emphasis id="B979EA90FFF8FFC76A664AD171F0FE49" italics="true" pageId="38" pageNumber="39">ericoides</emphasis>
,
<collectorName id="26F85354FFF8FFC76FFF4ABD7055FE49" box="[237,378,475,501]" pageId="38" pageNumber="39">C.F. Butcher</collectorName>
</taxonomicName>
(
<specimenCount id="9D0BFD0BFFF8FFC76E994ABD70B5FE49" box="[395,410,475,501]" pageId="38" pageNumber="39">3</specimenCount>
,
<collectionCode id="ED1CAE47FFF8FFC76EB44ABD70D8FE49" box="[422,503,475,501]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" country="New Zealand" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="38" pageNumber="39" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B653CDFFFF8FFC76D184ABD72D4FE49" ID-GBIF-Occurrence="2989339313" box="[522,1019,475,501]" collectingDate="1983-11-27" collectingMethod="sweeping vegetation mainly" collectionCode="NZAC" collectorName="C. F. Butcher" country="New Zealand" location="Offshore island" municipality="Great Is" pageId="38" pageNumber="39" specimenCount="3">
same, but, ex grasses and sedges (
<specimenCount id="9D0BFD0BFFF8FFC76C954ABD72B9FE49" box="[903,918,475,501]" pageId="38" pageNumber="39">3</specimenCount>
,
<collectionCode id="ED1CAE47FFF8FFC76CB04ABD72DCFE49" box="[930,1011,475,501]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" country="New Zealand" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="38" pageNumber="39" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B653CDFFFF8FFC76B144ABD7037FDA5" ID-GBIF-Occurrence="2988225328" collectingDate="11 / 1970" collectionCode="NZAC" collectorName="J. C. Watt" country="New Zealand" location="Great Is" pageId="38" pageNumber="39" specimenCount="1" typeStatus="paratype">
<location id="8ED26059FFF8FFC76B144ABD754EFE49" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76B144ABD754EFE49" box="[1030,1121,475,501]" country="New Zealand" name="Great Is" pageId="38" pageNumber="39">Great Is</location>
,
<collectingDate id="EFF7E9AAFFF8FFC76B7F4ABD75E5FE49" box="[1133,1226,475,501]" pageId="38" pageNumber="39">11/1970</collectingDate>
, beating,
<collectorName id="26F85354FFF8FFC76A244ABD74B3FE49" box="[1334,1436,475,501]" pageId="38" pageNumber="39">J.C. Watt</collectorName>
(
<specimenCount id="9D0BFD0BFFF8FFC76FB14A99719FFDA5" box="[163,176,511,537]" pageId="38" pageNumber="39">1</specimenCount>
,
<collectionCode id="ED1CAE47FFF8FFC76FAD4A99703FFDA5" box="[191,272,511,537]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" country="New Zealand" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="38" pageNumber="39" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B653CDFFFF8FFC76E344A99727DFDA5" ID-GBIF-Occurrence="2989339301" box="[294,850,511,537]" collectingDate="11 / 1970" collectionCode="NZAC" collectorName="J. C. Watt" country="New Zealand" location="Great Is" pageId="38" pageNumber="39" specimenCount="1">
same, but without collecting method (
<specimenCount id="9D0BFD0BFFF8FFC76DCE4A9973C6FDA5" box="[732,745,511,537]" pageId="38" pageNumber="39">1</specimenCount>
,
<collectionCode id="ED1CAE47FFF8FFC76DEB4A997265FDA5" box="[761,842,511,537]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" country="New Zealand" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="38" pageNumber="39" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B653CDFFFF8FFC76C4D4A9971DEFD81" ID-GBIF-Occurrence="2988225373" collectingDate="11 / 1970" collectionCode="NZAC" collectorName="G. Kushel" country="New Zealand" location="Great Is" municipality="Castaway Camp" pageId="38" pageNumber="39" specimenCount="5" typeStatus="paratype">
<location id="8ED26059FFF8FFC76C4D4A997292FDA5" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76C4D4A997292FDA5" box="[863,957,511,537]" country="New Zealand" municipality="Castaway Camp" name="Great Is" pageId="38" pageNumber="39">Great Is</location>
,
<location id="8ED26059FFF8FFC76CDE4A9975A7FDA5" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76CDE4A9975A7FDA5" box="[972,1160,511,537]" country="New Zealand" municipality="Castaway Camp" name="Castaway Camp" pageId="38" pageNumber="39">Castaway Camp</location>
,
<collectingDate id="EFF7E9AAFFF8FFC76B874A9975DDFDA5" box="[1173,1266,511,537]" pageId="38" pageNumber="39">11/1970</collectingDate>
,
<collectorName id="26F85354FFF8FFC76A124A99745BFDA5" box="[1280,1396,511,537]" pageId="38" pageNumber="39">G. Kushel</collectorName>
(
<specimenCount id="9D0BFD0BFFF8FFC76A9A4A9874B8FDA5" box="[1416,1431,510,537]" pageId="38" pageNumber="39">5</specimenCount>
,
<collectionCode id="ED1CAE47FFF8FFC76F85494571C7FD81" box="[151,232,547,573]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" country="New Zealand" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="38" pageNumber="39" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B653CDFFFF8FFC76FEE4945735AFD81" ID-GBIF-Occurrence="2988225366" box="[252,629,547,573]" collectingDate="11 / 1970" collectionCode="NZAC" collectorName="G. Ramsay" country="New Zealand" location="Great Is" municipality="Castaway Camp" pageId="38" pageNumber="39" specimenCount="4" typeStatus="paratype">
same, but
<collectorName id="26F85354FFF8FFC76E7C494570C4FD81" box="[366,491,547,573]" pageId="38" pageNumber="39">G. Ramsay</collectorName>
, (
<specimenCount id="9D0BFD0BFFF8FFC76EED49457320FD81" box="[511,527,547,573]" pageId="38" pageNumber="39">4</specimenCount>
,
<collectionCode id="ED1CAE47FFF8FFC76D0E49457342FD81" box="[540,621,547,573]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" country="New Zealand" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="38" pageNumber="39" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B653CDFFFF8FFC76D9249457547FD81" ID-GBIF-Occurrence="2988225315" box="[640,1128,547,573]" collectingDate="1983-12-01" collectorName="J. C. Watt" country="New Zealand" location="Great Is" municipality="North East" pageId="38" pageNumber="39" specimenCount="1" typeStatus="paratype">
<location id="8ED26059FFF8FFC76D92494573F4FD81" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76D92494573F4FD81" box="[640,731,547,573]" country="New Zealand" municipality="North East" name="Great Is" pageId="38" pageNumber="39">Great Is</location>
,
<location id="8ED26059FFF8FFC76DF54945724FFD81" LSID="urn:lsid:plazi:treatment:03A48794FFFBFFC76F854BF17543FD81:8ED26059FFF8FFC76DF54945724FFD81" box="[743,864,547,573]" country="New Zealand" municipality="North East" name="North East" pageId="38" pageNumber="39">North East</location>
I,
<date id="FFB31042FFF8FFC76C6C494572DDFD81" box="[894,1010,547,573]" pageId="38" pageNumber="39" value="1983-12-01">
<collectingDate id="EFF7E9AAFFF8FFC76C6C494572DDFD81" box="[894,1010,547,573]" pageId="38" pageNumber="39" value="1983-12-01">1/12/1983</collectingDate>
</date>
,
<collectorName id="26F85354FFF8FFC76CEC49457547FD81" box="[1022,1128,547,573]" pageId="38" pageNumber="39">J.C. Watt</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>