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<mods:title id="47EAEAE613B49D1DCE438D99842F8E34">Sexual Dimorphism of New Zealand Puppet Beetles (Aderidae, Coleoptera, Tenebrionoidea): Systematic Revision, Description of Three New Genera, and Phylogeny for Zenascus, gen. n.</mods:title>
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<taxonomicName id="4C0D4D01FFC2FFFD6F854DF9736BF904" authority="(Broun, 1893)" authorityName="Grzymala &amp; Leschen" authorityYear="2020" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[151,580,1694,1721]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="antennalis" status="comb. nov.">
<emphasis id="B979EA90FFC2FFFD6F854DF970BFF905" bold="true" box="[151,400,1694,1721]" italics="true" pageId="28" pageNumber="29">Zenascus antennalis</emphasis>
(
<bibRefCitation id="EF9C4B73FFC2FFFD6E8D4DF87313F904" author="Broun, T." box="[415,572,1694,1720]" pageId="28" pageNumber="29" pagination="975 - 1504" refId="ref39241" refString="Broun, T. (1893) Manual of the New Zealand Coleoptera. Parts V-VII. Government Printer, Wellington, pp. XVII + 975 - 1504." type="book chapter" year="1893">Broun, 1893</bibRefCitation>
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<taxonomicNameLabel id="A24A57EBFFC2FFFD6D404DF87392F904" box="[594,701,1694,1720]" pageId="28" pageNumber="29" rank="species">comb. n.</taxonomicNameLabel>
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<subSubSection id="C3176509FFC2FFFD6F854DA473B3F960" box="[151,668,1730,1756]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BB23682FFC2FFFD6F854DA473B3F960" blockId="28.[151,701,1694,1756]" box="[151,668,1730,1756]" pageId="28" pageNumber="29">
<figureCitation id="13362A07FFC2FFFD6F854DA4702DF960" box="[151,258,1730,1756]" captionStart="FIGURES 12" captionStartId="31.[151,264,1912,1936]" captionTargetBox="[158,1428,787,1880]" captionTargetId="figure-291@31.[151,1436,777,1889]" captionTargetPageId="31" captionText="FIGURES 12A–E. Dorsal habitus images. (A) Zenascus antennalis (Broun), male; (B) Zenascus xenarthrus (Broun), male; (C) Zenascus roberti sp. n., male; (D) Zenascus luniger (Champion), male; (E) Zenascus incensum sp. n., male. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4297842" httpUri="https://zenodo.org/record/4297842/files/figure.png" pageId="28" pageNumber="29">Figs 12A</figureCitation>
, 
<figureCitation id="13362A07FFC2FFFD6E1F4DA47016F960" box="[269,313,1730,1756]" captionStart="FIGURES 14" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[268,1319,181,1901]" captionTargetId="figure-17@33.[268,1319,181,1919]" captionTargetPageId="33" captionText="FIGURES 14A–J. Antennal illustrations for Zenascus species, male on left, female on right.(A) Z. coloratus; (B) Z. obscurus; (C) Z. aurum, sp. n.; (D) Z. nitidus; (E) Z. elenae, sp. n.; (F) Z. antennalis; (G) Z. xenarthrus; (H) Z. roberti, sp. n.; (I) Z. luniger; (J) Z. incensum, sp. n. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4297849" httpUri="https://zenodo.org/record/4297849/files/figure.png" pageId="28" pageNumber="29">14F</figureCitation>
, 
<figureCitation id="13362A07FFC2FFFD6E564DA47058F960" box="[324,375,1730,1756]" captionStart="FIGURES 19" captionStartId="44.[151,264,1421,1445]" captionTargetBox="[157,1433,185,1393]" captionTargetId="figure-28@44.[151,1436,181,1397]" captionTargetPageId="44" captionText="FIGURES 19A–E. Scanning electron microscope images, Zenascus species, elytron, dorsal view. (A) Z. antennalis; (B) Z. xenarthrus; (C) Z. roberti, sp. n.; (D) Z. luniger; (E) Z. incensum, sp. n. Scale bars = 10μm." figureDoi="http://doi.org/10.5281/zenodo.4297859" httpUri="https://zenodo.org/record/4297859/files/figure.png" pageId="28" pageNumber="29">19A</figureCitation>
, 
<figureCitation id="13362A07FFC2FFFD6E904DA47081F960" box="[386,430,1730,1756]" captionStart="FIGURES 20" captionStartId="45.[151,264,1411,1435]" captionTargetBox="[151,1436,184,1384]" captionTargetId="figure-17@45.[151,1436,181,1387]" captionTargetPageId="45" captionText="FIGURES 20A–J. Scanning electron microscope images, ventral view of metafemora for Zenascus species. (A) Z. coloratus, male; (B) Z. obscurus, male; (C) Z. aurum, sp. n., male; (D) Z. nitidus, female; (E) Z. elenae, sp. n., male; (F) Z. antennalis, female; (G) Z. xenarthrus, male; (H) Z. roberti, sp. n., male; (I) Z. luniger, male; (J) Z. incensum, sp. n., male. Scale bars = 60μm." figureDoi="http://doi.org/10.5281/zenodo.4297861" httpUri="https://zenodo.org/record/4297861/files/figure.png" pageId="28" pageNumber="29">20F</figureCitation>
, 
<figureCitation id="13362A07FFC2FFFD6EA84DA470C3F960" box="[442,492,1730,1756]" captionStart="FIGURES 22" captionStartId="47.[151,264,1163,1187]" captionTargetBox="[151,1436,186,1140]" captionTargetId="figure-17@47.[151,1436,181,1140]" captionTargetPageId="47" captionText="FIGURES 22A–E. Abdominal ventrite illustrations for Zenascus species showing generalized punctation pattern. (A) Z. antennalis, male; (B) Z. xenarthrus, male; (C) Z. roberti, sp. n., male; (D) Z. luniger, male; (E) Z. incensum, sp. n., male." figureDoi="http://doi.org/10.5281/zenodo.4297873" httpUri="https://zenodo.org/record/4297873/files/figure.png" pageId="28" pageNumber="29">22A</figureCitation>
, 
<figureCitation id="13362A07FFC2FFFD6EEA4DA4730CF960" box="[504,547,1730,1756]" captionStart="FIGURES 23" captionStartId="49.[151,264,1943,1967]" captionTargetBox="[231,1359,181,1919]" captionTargetId="figure-17@49.[227,1359,181,1919]" captionTargetPageId="49" captionText="FIGURES 23A–J. Male genitalia illustrations for Zenascus species. (A) Z. coloratus; (B) Z. obscurus; (C) Z. aurum, sp. n.; (D) Z. nitidus; (E) Z. elenae, sp. n.; (F) Z. antennalis; (G) Z. xenarthrus; (H) Z. roberti, sp. n.; (I) Z. luniger; (J) Z. incensum, sp. n." figureDoi="http://doi.org/10.5281/zenodo.4297877" httpUri="https://zenodo.org/record/4297877/files/figure.png" pageId="28" pageNumber="29">23F</figureCitation>
, 
<figureCitation id="13362A07FFC2FFFD6D3D4DA4734EF960" box="[559,609,1730,1756]" captionStart="FIGURES 24" captionStartId="50.[151,264,1642,1666]" captionTargetBox="[159,1431,183,1612]" captionTargetId="figure-28@50.[151,1436,181,1619]" captionTargetPageId="50" captionText="FIGURES 24A–F. Scanning electron micrographs of gland pores on metafemur of Zenascus spp. (A) Z. aurum, sp. n.; (B) Z. antennalis; (C) Z. xenarthrus; (D) Z. roberti, sp. n.; (E) Z. luniger; (F) Z. incensum, sp. n. Scale bars = 2μm." figureDoi="http://doi.org/10.5281/zenodo.4297881" httpUri="https://zenodo.org/record/4297881/files/figure.png" pageId="28" pageNumber="29">24B</figureCitation>
, 
<figureCitation id="13362A07FFC2FFFD6D7E4DA473B3F960" box="[620,668,1730,1756]" captionStart="FIGURES 26" captionStartId="53.[151,264,1952,1976]" captionTargetBox="[163,1421,191,1918]" captionTargetId="figure-17@53.[155,1433,181,1928]" captionTargetPageId="53" captionText="FIGURES 26A–D. Distribution maps. (A) Scraptogetus anthracinus, (B) Scraptogetus arboreus, (C) Transrenus thulater, sp. n. and Pseudozena denticulata, sp. n., and (D) Zenascus antennalis." figureDoi="http://doi.org/10.5281/zenodo.4297885" httpUri="https://zenodo.org/record/4297885/files/figure.png" pageId="28" pageNumber="29">26D</figureCitation>
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<taxonomicName id="4C0D4D01FFC2FFFD6F854C617356F8A3" authority="Broun, 1893: 1163 - 1164" authorityName="Broun" authorityPageNumber="1163 - 1164" authorityYear="1893" box="[151,633,1799,1823]" class="Insecta" family="Aderidae" genus="Xylophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="antennalis">
<emphasis id="B979EA90FFC2FFFD6F854C61705BF8A3" box="[151,372,1799,1823]" italics="true" pageId="28" pageNumber="29">Xylophilus antennalis</emphasis>
<bibRefCitation id="EF9C4B73FFC2FFFD6E684C617356F8A3" author="Broun, T." box="[378,633,1799,1823]" pageId="28" pageNumber="29" pagination="975 - 1504" refId="ref39241" refString="Broun, T. (1893) Manual of the New Zealand Coleoptera. Parts V-VII. Government Printer, Wellington, pp. XVII + 975 - 1504." type="book chapter" year="1893">Broun, 1893: 1163–1164</bibRefCitation>
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<bibRefCitation id="EF9C4B73FFC2FFFD6D914C617236F8A3" author="Pic, M." box="[643,793,1799,1823]" pageId="28" pageNumber="29" pagination="427 - 436" refId="ref40981" refString="Pic, M. (1894) Liste complete des Xylophilides decrits jusqu'en 1894, avec descriptions d'especes nouvelles. Memoires de la Societe Zoologique de France, 7, 427 - 436." type="journal article" year="1894">Pic, 1894: 428</bibRefCitation>
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<bibRefCitation id="EF9C4B73FFC2FFFD6CB94C617540F8A3" author="Hudson, G. V." box="[939,1135,1799,1823]" pageId="28" pageNumber="29" refId="ref40063" refString="Hudson, G. V. (1934) New Zealand beetles and their larvae: an elementary introduction to the study of our native Coleoptera, with seventeeen coloured plates. Ferguson &amp; Osborn, Wellington, 236 pp., 17 pls." type="book" year="1934">Hudson, 1934: 204</bibRefCitation>
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<bibRefCitation id="EF9C4B73FFC2FFFD6B684C61747AF8A3" author="Maddison, P. M." box="[1146,1365,1799,1823]" pageId="28" pageNumber="29" refId="ref40799" refString="Maddison, P. M. (2010) Checklist of New Zealand Hexapoda. Order Coleoptera. [Chapter] NINE: Phylum Arthropoda: Subphylum Hexapoda: Protura, springtails, Diplura, and insects. In: Gordon, D. P. (Ed.), New Zealand Inventory of Biodiversity. Vol. 2. Kingdom Animalia. Chaetognatha, Ecdysozoa, ichnofossils. Canterbury University Press, Christchurch, pp. 233 - 467." type="book" year="2010">Maddison, 2010: 428</bibRefCitation>
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<taxonomicName id="4C0D4D01FFC2FFFD6F854C437058F881" authority="Pic, 1902" box="[151,375,1829,1853]" class="Aves" family="Vireonidae" genus="Hylophilus" kingdom="Animalia" order="Passeriformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="antennalis">
<emphasis id="B979EA90FFC2FFFD6F854C437058F881" box="[151,375,1829,1853]" italics="true" pageId="28" pageNumber="29">Hylophilus antennalis</emphasis>
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<bibRefCitation id="EF9C4B73FFC2FFFD6E964C43732FF881" author="Pic, M." box="[388,512,1829,1853]" pageId="28" pageNumber="29" pagination="1 - 14" refId="ref41041" refString="Pic, M. (1902) Fascicule 8. Coleoptera Heteromera Fam. Hylophilidae. In: Wytsman, P. (Ed.), Genera Insectorum. V. Verteneuil &amp; L. Desmet, Brussels, pp. 1 - 14." type="book chapter" year="1902">Pic, 1902: 6</bibRefCitation>
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<bibRefCitation id="EF9C4B73FFC2FFFD6D194C43738FF881" author="Pic, M." box="[523,672,1829,1853]" pageId="28" pageNumber="29" pagination="181 - 286" refId="ref41115" refString="Pic, M. (1905) Contribution a l'etude generale des Hylophilidae. Annales de la Societe Entomologique de France, 74, 181 - 286. https: // doi. org / 10.5962 / bhl. part. 26096" type="journal article" year="1905">Pic, 1905: 216</bibRefCitation>
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, 230; 
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<bibRefCitation id="EF9C4B73FFC2FFFD6DCC4C437275F881" author="Pic, M." box="[734,858,1829,1853]" pageId="28" pageNumber="29" pagination="1 - 25" refId="ref41157" refString="Pic, M. (1910) Pars 14: Hylophilidae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 25. https: // doi. org / 10.1002 / mmnd. 4801910103" type="book chapter" year="1910">Pic, 1910: 5</bibRefCitation>
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<paragraph id="8BB23682FFC2FFFC6F854C0D72A7FF69" blockId="28.[151,1436,1899,2034]" lastBlockId="29.[151,1437,151,1797]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B979EA90FFC2FFFD6F854C0D703DF839" bold="true" box="[151,274,1899,1925]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
This species is distinguished from congeners by the shining, black integument without secondary pubescence on the elytra, brunneous femora and tibiae, the black antennal scape, antennomeres 8–11 darkened in coloration, and the modified antennae of males (
<figureCitation id="13362A07FFC2FFFD6D794CD573E1F872" box="[619,718,1971,1998]" captionStart="FIGURES 14" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[268,1319,181,1901]" captionTargetId="figure-17@33.[268,1319,181,1919]" captionTargetPageId="33" captionText="FIGURES 14A–J. Antennal illustrations for Zenascus species, male on left, female on right.(A) Z. coloratus; (B) Z. obscurus; (C) Z. aurum, sp. n.; (D) Z. nitidus; (E) Z. elenae, sp. n.; (F) Z. antennalis; (G) Z. xenarthrus; (H) Z. roberti, sp. n.; (I) Z. luniger; (J) Z. incensum, sp. n. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4297849" httpUri="https://zenodo.org/record/4297849/files/figure.png" pageId="28" pageNumber="29">Fig. 14F</figureCitation>
). 
<taxonomicName id="4C0D4D01FFC2FFFD6DF34CD272EBF871" authorityName="Grzymala &amp; Leschen" authorityYear="2020" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[737,964,1971,1997]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="antennalis">
<emphasis id="B979EA90FFC2FFFD6DF34CD272EBF871" box="[737,964,1971,1997]" italics="true" pageId="28" pageNumber="29">Zenascus antennalis</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C0D4D01FFC2FFFD6B834CD275D5F871" authorityName="Grzymala &amp; Leschen" authorityYear="2020" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[1169,1274,1971,1997]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B979EA90FFC2FFFD6B834CD275D5F871" box="[1169,1274,1971,1997]" italics="true" pageId="28" pageNumber="29">Z. nitidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D01FFC2FFFD6A3D4CD274BAF871" authorityName="Grzymala &amp; Leschen" authorityYear="2020" box="[1327,1429,1971,1997]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="B979EA90FFC2FFFD6A3D4CD274BAF871" box="[1327,1429,1971,1997]" italics="true" pageId="28" pageNumber="29">Z. elenae</emphasis>
</taxonomicName>
, which all have the shining black integument. 
<taxonomicName id="4C0D4D01FFC2FFFD6D864CBE7256F84D" authorityName="Grzymala &amp; Leschen" authorityYear="2020" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[660,889,2007,2033]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="antennalis">
<emphasis id="B979EA90FFC2FFFD6D864CBE7256F84D" box="[660,889,2007,2033]" italics="true" pageId="28" pageNumber="29">Zenascus antennalis</emphasis>
</taxonomicName>
can be separated by the brunneus femora, tibiae, and the dark coloration of antennomeres 8–11 present in both males and females. Both 
<taxonomicName id="4C0D4D01FFC3FFFC6B4D4BF175E7FF0D" authorityName="Grzymala &amp; Leschen" authorityYear="2020" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[1119,1224,151,177]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B979EA90FFC3FFFC6B4D4BF175E7FF0D" box="[1119,1224,151,177]" italics="true" pageId="29" pageNumber="30">Z. nitidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D01FFC3FFFC6BEC4BF1744BFF0D" authorityName="Grzymala &amp; Leschen" authorityYear="2020" box="[1278,1380,151,177]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="B979EA90FFC3FFFC6BEC4BF1744BFF0D" box="[1278,1380,151,177]" italics="true" pageId="29" pageNumber="30">Z. elenae</emphasis>
</taxonomicName>
conversely have the femora, tibiae, and antennomeres 8–11 testaceous.
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFC3FFFC6FD54BB875E7FA59" pageId="29" pageNumber="30" type="description">
<paragraph id="8BB23682FFC3FFFC6FD54BB872C7FAB1" blockId="29.[151,1437,151,1797]" pageId="29" pageNumber="30">
<emphasis id="B979EA90FFC3FFFC6FD54BB8707EFF44" bold="true" box="[199,337,222,248]" pageId="29" pageNumber="30">Description</emphasis>
. Length 
<quantity id="4CF59B67FFC3FFFC6EA54BB97371FF45" box="[439,606,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" metricValueMax="1.82" metricValueMin="1.6" pageId="29" pageNumber="30" unit="mm" value="1.71" valueMax="1.82" valueMin="1.6">1.60–1.82 mm</quantity>
. Head, antennal scape, antennomeres 8–11, femora, tibiae, pronotum, and elytra piceous to brunneous; antennal pedicel, antennomeres 3–7, mouthparts, and tarsi testaceous. Dorsal integument shining, vestiture uniseriate, one decumbent, thin, long seta arising anterad of each primary puncture, length about 3× puncture length, without additional setae between each pair of punctures. Ventral vestiture biseriate. Head, HW 
<quantity id="4CF59B67FFC3FFFC6FDD4A087058FE35" box="[207,375,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="29" pageNumber="30" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
, HL 
<quantity id="4CF59B67FFC3FFFC6EA74A097372FE35" box="[437,605,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.7999999999999998" metricValueMin="1.7" pageId="29" pageNumber="30" unit="mm" value="0.175" valueMax="0.18" valueMin="0.17">0.17–0.18 mm</quantity>
, with numerous deep punctures, approximately evenly spaced, covering frons and vertex, absent from base to just anterad of head constriction; impression between antennal ridge and vertex absent. Antennae with distance between antennal insertions wide, approximately equal to the diameter of five antennal insertions; alength reaching past basal ¼ of elytra when extended backward; scape longer than broad, approximately 2× length of pedicel; pedicel subglobular; antennomere 3 longer than pedicel and longer than antennomere 4; antennomeres 4–8 subequal in length; antennomere 7 with apico-anterior projection, apex of projection rounded; antennomere 8 with additional, larger apico-anterior projection, apex of projection pointed forward; antennomere 9 with small, truncate apico-anterior projection; antennomeres 9 and 10 subequal in length; antennomere 11 elongate, greater in length than any preceding antennal segment; antennomeres 3–11 covered in moderately dense suberect pubescence. Pronotum, PW 
<quantity id="4CF59B67FFC3FFFC6D5149D573C4FD71" box="[579,747,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="29" pageNumber="30" unit="mm" value="0.4" valueMax="0.42" valueMin="0.38">0.38–0.42 mm</quantity>
, PL 
<quantity id="4CF59B67FFC3FFFC6C3149D472E4FD71" box="[803,971,690,717]" metricMagnitude="-4" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.5" metricValueMin="3.2" pageId="29" pageNumber="30" unit="mm" value="0.33499999999999996" valueMax="0.35" valueMin="0.32">0.32–0.35 mm</quantity>
, subquadrate, width subequal to slightly greater than length, width 1.19–1.20× length; pronotal width 0.48–0.51× elytral width, pronotal width 0.84–0.88× head width; sides slightly sinuate; disc with two faint basal fovea, with slight transverse suture just anterad of center; disc with deep elongate punctures, approximately evenly spaced from one another, separated by an average of two punctural lengths. Elytra 1.69–1.71× longer than wide and 4.00–4.20× longer than pronotal length, EW 
<quantity id="4CF59B67FFC3FFFC6A3E482471EAFC3D" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.7" metricValueMin="7.5" pageId="29" pageNumber="30" unit="mm" value="0.81" valueMax="0.87" valueMin="0.75">0.75–0.87 mm</quantity>
, EL 
<quantity id="4CF59B67FFC3FFFC6FEA480170B2FC3D" box="[248,413,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.47" metricValueMin="1.28" pageId="29" pageNumber="30" unit="mm" value="1.375" valueMax="1.47" valueMin="1.28">1.28–1.47 mm</quantity>
; subscutellar oblique depression slightly impressed; disc with elongate punctures, separated by an average of two punctural lengths. Proleg with tarsomere 1 approximately 2–3× longer than tarsomere 2, without a ventral short, stout spine; ventral surface of tarsomere 3 with adhesive setae. Midleg with tibia straight from base to apex. Hindleg with numerous distinct, deep punctures along posterior margin of coxa; femur with distinct lines of thickened setae on posterior face, apico-ventral impression absent; tibia gradually expanded in width from base to apex, apex inner face triangular and densely pubescent. Abdomen with lateral and medial lengths of ventrite 2 subequal; deep punctures confined to abdominal process and basal margin of ventrite 1 and present centrally on ventrites 2–4; ventrite 5 with few deep punctures scattered basally and lacking medial impression. Phallobase broadly rounded anteriorly and without clear delimitation between phallobase and apicale, fused medially and laterally; apicale posteriorly narrowly acute, accessory lobes present, short and thickened, width about ½× length, and curved inward towards apex, with four short setae, two on inner lateral surface, one on outer lateral surface, one at apex; penis with anterior struts elongate, extending slightly past phallobase.
</paragraph>
<paragraph id="8BB23682FFC3FFFC6FD54E7175E7FA59" blockId="29.[151,1437,151,1797]" pageId="29" pageNumber="30">
<emphasis id="B979EA90FFC3FFFC6FD54E71701FFA8D" bold="true" box="[199,304,1303,1329]" pageId="29" pageNumber="30">Females.</emphasis>
Distance between antennal insertions wide, approximately equal to diameter of three antennal insertions; scape slightly longer than wide; pedicel subglobular; antennomere 3 thinner and subequal in length to pedicel; antennomere 4 subequal in width and slightly longer than antennomere 3; antennomere 5 subequal in width and slightly shorter than antennomere 4; antennomere 6 slightly wider than antennomere 5; antennomere 7 slightly wid-er than 6, inner apex slightly produced; antennomeres 8–10 subsequently expanding in width, subequal in length, forming a distinctive club; antennomere 11 subequal in width and 3× length of antennomere 10.
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFC3FFFC6FD54E8973CCF925" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BB23682FFC3FFFC6FD54E897209F991" blockId="29.[151,1437,151,1797]" pageId="29" pageNumber="30">
<emphasis id="B979EA90FFC3FFFC6FD54E897014F9B5" bold="true" box="[199,315,1519,1545]" pageId="29" pageNumber="30">Remarks.</emphasis>
This is Broun species 2063, which was based on a single specimen from Paparoa, near Howick (
<bibRefCitation id="EF9C4B73FFC3FFFC6F8D4D757005F991" author="Broun, T." box="[159,298,1555,1581]" pageId="29" pageNumber="30" pagination="975 - 1504" refId="ref39241" refString="Broun, T. (1893) Manual of the New Zealand Coleoptera. Parts V-VII. Government Printer, Wellington, pp. XVII + 975 - 1504." type="book chapter" year="1893">Broun 1893</bibRefCitation>
). The 
<typeStatus id="54B68820FFC3FFFC6E7D4D7570FEF991" box="[367,465,1555,1581]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
was confirmed in the BMNH.
</paragraph>
<paragraph id="8BB23682FFC3FFFC6FD54D5173CCF925" blockId="29.[151,1437,151,1797]" pageId="29" pageNumber="30">
<emphasis id="B979EA90FFC3FFFC6FD54D5170ABF9EC" bold="true" box="[199,388,1590,1617]" pageId="29" pageNumber="30">Natural history.</emphasis>
This is a relatively rare and localized North Island species known from seven specimens. This species has been collected on foliage and using Malaise traps. It has also been collected by beating 
<taxonomicName id="4C0D4D01FFC3FFFC6BF84D3D71F7F925" class="Magnoliopsida" family="Violaceae" genus="Melicytus" kingdom="Plantae" order="Malpighiales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="ramiflorus">
<emphasis id="B979EA90FFC3FFFC6BF84D3D71F7F925" italics="true" pageId="29" pageNumber="30">Melicytus ramiflorus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0D4D01FFC3FFFC6E1D4D197306F925" box="[271,553,1663,1689]" class="Magnoliopsida" family="Corynocarpaceae" genus="Corynocarpus" kingdom="Plantae" order="Cucurbitales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="laevigatus">
<emphasis id="B979EA90FFC3FFFC6E1D4D197306F925" box="[271,553,1663,1689]" italics="true" pageId="29" pageNumber="30">Corynocarpus laevigatus</emphasis>
</taxonomicName>
J.R. &amp; G. Forst.
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFC3FFFC6FD54DC472B3F901" box="[199,924,1698,1725]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BB23682FFC3FFFC6FD54DC472B3F901" blockId="29.[151,1437,151,1797]" box="[199,924,1698,1725]" pageId="29" pageNumber="30">
<emphasis id="B979EA90FFC3FFFC6FD54DC4704EF900" bold="true" box="[199,353,1698,1724]" pageId="29" pageNumber="30">Distribution.</emphasis>
<materialsCitation id="3B653CDFFFC3FFFC6E754DC572B3F901" ID-GBIF-Occurrence="2989339304" box="[359,924,1699,1725]" country="New Zealand" location="North Island" municipality="Coromandel" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Auckland">
<emphasis id="B979EA90FFC3FFFC6E754DC57325F901" bold="true" box="[359,522,1699,1725]" pageId="29" pageNumber="30">
<location id="8ED26059FFC3FFFC6E754DC5732AF901" LSID="urn:lsid:plazi:treatment:03A48794FFC2FFFC6F854DF97467F8B9:8ED26059FFC3FFFC6E754DC5732AF901" box="[359,517,1699,1725]" municipality="Coromandel" name="North Island" pageId="29" pageNumber="30" stateProvince="Auckland">North Island</location>
:
</emphasis>
<collectingRegion id="49C9F860FFC3FFFC6D1D4DC57353F901" box="[527,636,1699,1725]" country="New Zealand" name="Auckland" pageId="29" pageNumber="30">Auckland</collectingRegion>
(AK), 
<collectingMunicipality id="6BD6ACF8FFC3FFFC6DDE4DC57279F901" box="[716,854,1699,1725]" pageId="29" pageNumber="30">Coromandel</collectingMunicipality>
(CL).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3176509FFC3FFFC6FD54DA17467F8B9" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BB23682FFC3FFFC6FD54DA17467F8B9" blockId="29.[151,1437,151,1797]" pageId="29" pageNumber="30">
<emphasis id="B979EA90FFC3FFFC6FD54DA170DDF95C" bold="true" box="[199,498,1734,1761]" pageId="29" pageNumber="30">Type material examined.</emphasis>
<materialsCitation id="3B653CDFFFC3FFFC6EE84DA17467F8B9" ID-GBIF-Occurrence="2988225324" collectionCode="BMNH" country="New Zealand" determinerName="J. C. Watt" location="New Zealand" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B68820FFC3FFFC6EE84DA1734AF95D" box="[506,613,1735,1761]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
, 
<specimenCount id="9D0BFD0BFFC3FFFC6D634DA17392F95D" box="[625,701,1735,1761]" pageId="29" pageNumber="30" type="female">female</specimenCount>
(
<collectionCode id="ED1CAE47FFC3FFFC6DDC4DA17206F95D" box="[718,809,1735,1761]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="29" pageNumber="30">BMNH</collectionCode>
): “2063 // Paparoa // 
<collectingCountry id="F31A7612FFC3FFFC6B0E4DA1759CF95D" box="[1052,1203,1735,1761]" name="New Zealand" pageId="29" pageNumber="30">New Zealand</collectingCountry>
/ Broun Coll. / Brit. Mus. / 1922-482 // HOLOTYPE [male symbol] / “ 
<taxonomicName id="4C0D4D01FFC3FFFC6DD94D8D7262F8B9" box="[715,845,1771,1797]" class="Insecta" family="Aderidae" genus="Xylophilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Xylophilus</taxonomicName>
” / antennalis / Broun / det. 
<determinerName id="7E6EDA46FFC3FFFC6B924D8D75C9F8B9" box="[1152,1254,1771,1797]" pageId="29" pageNumber="30">J.C. Watt</determinerName>
/ 1985”.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>