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<document id="F391FD84FBA7D7E7302232F78C214E3A" ID-CLB-Dataset="57608" ID-DOI="10.11646/zootaxa.5276.1.1" ID-GBIF-Dataset="590e0e3a-f3ea-4420-b976-b668987049e6" ID-ISSN="1175­5334" ID-Zenodo-Dep="7907294" ID-ZooBank="92300500-BB24-45B0-8ADD-977C3220A069" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1682929863379" checkinUser="plazi" docAuthor="Camargo, Alexssandro, Vieira, Rodrigo &amp; Rafael, José Albertino" docDate="2023" docId="03A4878EEE78380ABEE1FAFAF3CAFE27" docLanguage="en" docName="zootaxa.5276.1.1.pdf" docOrigin="Zootaxa 5276" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Catostola indecisa Camargo, Vieira &amp; Rafael, 2023, sp. nov." docType="treatment" docUuid="66EF012B-E1CB-4838-B318-AD8432744A8B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="66" masterDocId="FF9DFFF6EE423848BE76FFD5F253FFD2" masterDocTitle="Taxonomic review of Ctenodontina Enderlein, 1914 with the revalidation of Catostola Hull, 1958 (Diptera: Asilidae: Asilinae) and description of a new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="59" updateTime="1699515144633" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="085083BD323C012519080F0E4B79C3F0">Taxonomic review of Ctenodontina Enderlein, 1914 with the revalidation of Catostola Hull, 1958 (Diptera: Asilidae: Asilinae) and description of a new species</mods:title>
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<mods:namePart id="77741200FC641DBD7D89DB1B03D812DB">Camargo, Alexssandro</mods:namePart>
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<mods:namePart id="F74FCC4A4ECAEA4DDD57446A32BD1954">Vieira, Rodrigo</mods:namePart>
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<mods:namePart id="28C7F3D0440B65A9125B91428CE714AF">Rafael, José Albertino</mods:namePart>
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<mods:title id="CDD0878D905E78088006173E01512315">Zootaxa</mods:title>
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<treatment id="03A4878EEE78380ABEE1FAFAF3CAFE27" ID-DOI="http://doi.org/10.5281/zenodo.7907348" ID-GBIF-Taxon="209588236" ID-Zenodo-Dep="7907348" LSID="urn:lsid:zoobank.org:act:66EF012B-E1CB-4838-B318-AD8432744A8B" httpUri="http://treatment.plazi.org/id/03A4878EEE78380ABEE1FAFAF3CAFE27" lastPageId="66" lastPageNumber="66" pageId="58" pageNumber="59">
<subSubSection id="C3176513EE783872BEE1FAFAF38AFA98" box="[151,473,1327,1354]" pageId="58" pageNumber="59" type="nomenclature">
<paragraph id="8BB23698EE783872BEE1FAFAF38AFA98" blockId="58.[151,1086,1327,1426]" box="[151,473,1327,1354]" pageId="58" pageNumber="59">
<heading id="D0FA81F4EE783872BEE1FAFAF38AFA98" bold="true" box="[151,473,1327,1354]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
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<taxonomicName id="4C0D4D1BEE783872BEE1FAFAF327FA98" ID-CoL="BDNLW" authorityName="Carmago &amp; Vieira &amp; Rafael" authorityYear="2023" box="[151,372,1327,1354]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE783872BEE1FAFAF327FA98" bold="true" box="[151,372,1327,1354]" italics="true" pageId="58" pageNumber="59">Catostola indecisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A57F1EE783872BF0DFAE5F38AFA98" box="[379,473,1328,1354]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BB23698EE783872BEE1FA81F302FABC" blockId="58.[151,1086,1327,1426]" box="[151,337,1364,1390]" pageId="58" pageNumber="59">
(
<figureCitation id="13362A1DEE783872BEE9FA81F372FABC" box="[159,289,1364,1390]" captionStart-0="FIGURE 38" captionStart-1="FIGURE 39" captionStart-2="FIGURE 40" captionStart-3="FIGURE 41" captionStart-4="FIGURE 42" captionStartId-0="60.[151,250,1943,1967]" captionStartId-1="61.[151,250,1877,1901]" captionStartId-2="62.[151,250,1881,1905]" captionStartId-3="63.[151,250,1885,1909]" captionStartId-4="64.[151,250,1854,1878]" captionTargetBox-0="[203,1385,181,1919]" captionTargetBox-1="[225,1362,181,1853]" captionTargetBox-2="[219,1369,193,1846]" captionTargetBox-3="[216,1368,184,1859]" captionTargetBox-4="[233,1354,181,1830]" captionTargetId-0="figure-21@60.[203,1385,181,1919]" captionTargetId-1="figure-17@61.[225,1362,181,1853]" captionTargetId-2="figure-21@62.[219,1369,181,1857]" captionTargetId-3="figure-17@63.[216,1371,181,1861]" captionTargetId-4="figure-21@64.[233,1354,181,1830]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionTargetPageId-2="62" captionTargetPageId-3="63" captionTargetPageId-4="64" captionText-0="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." captionText-1="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." captionText-2="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." captionText-3="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." captionText-4="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7907408" figureDoi-1="http://doi.org/10.5281/zenodo.7907412" figureDoi-2="http://doi.org/10.5281/zenodo.7907414" figureDoi-3="http://doi.org/10.5281/zenodo.7907418" figureDoi-4="http://doi.org/10.5281/zenodo.7907420" httpUri-0="https://zenodo.org/record/7907408/files/figure.png" httpUri-1="https://zenodo.org/record/7907412/files/figure.png" httpUri-2="https://zenodo.org/record/7907414/files/figure.png" httpUri-3="https://zenodo.org/record/7907418/files/figure.png" httpUri-4="https://zenodo.org/record/7907420/files/figure.png" pageId="58" pageNumber="59">Figs 3842</figureCitation>
,
<figureCitation id="13362A1DEE783872BF5AFA81F31AFABC" box="[300,329,1364,1390]" captionStart="FIGURE 44" captionStartId="66.[151,250,1682,1706]" captionTargetBox="[151,1435,525,1658]" captionTargetId="figure-172@66.[151,1435,525,1658]" captionTargetPageId="66" captionText="FIGURE 44. Distribution map of Catostola species: Catostola baleta (Walker, 1849) comb. nov., Catostola complicata (James, 1953) comb. nov., Catostola indecisa sp. nov., and Catostola martini (Fisher, 1992) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.7907427" httpUri="https://zenodo.org/record/7907427/files/figure.png" pageId="58" pageNumber="59">44</figureCitation>
)
</paragraph>
<paragraph id="8BB23698EE783872BEE1FAADF66DFA40" blockId="58.[151,1086,1327,1426]" box="[151,1086,1400,1426]" pageId="58" pageNumber="59">
ZooBank:
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urn:lsid:zoobank.org:act:
<uuid id="FFAB0C4DEE783872BC56FAADF66DFA40" box="[544,1086,1400,1426]" pageId="58" pageNumber="59">66EF012B-E1CB-4838-B318-AD8432744A8B</uuid>
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</subSubSection>
<subSubSection id="C3176513EE783872BEE1FA15F0B8F960" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="8BB23698EE783872BEE1FA15F0B8F960" blockId="58.[151,1437,1472,2038]" pageId="58" pageNumber="59">
<emphasis id="B979EA8AEE783872BEE1FA15F341FA08" bold="true" box="[151,274,1472,1498]" pageId="58" pageNumber="59">Diagnosis.</emphasis>
Face yellowish brown pruinose; mystacal macrosetae yellowish white, except by a few dorsal black macrosetae (
<figureCitation id="13362A1DEE783872BF50FA31F3E2FA2C" box="[294,433,1508,1534]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="58" pageNumber="59">Fig. 38CD</figureCitation>
); postpedicel dark reddish; second article of stylus black, dark reddish at base, long and slender, slightly enlarged subapically (
<figureCitation id="13362A1DEE783872BC3CF9DDF086F9F0" box="[586,725,1544,1570]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="58" pageNumber="59">Fig. 39AB</figureCitation>
); fore femur yellow, mid femur yellow with black apical third, hind femur black (
<figureCitation id="13362A1DEE783872BF1DF9F9F385F994" box="[363,470,1580,1606]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="58" pageNumber="59">Fig. 38A</figureCitation>
); male terminalia shining black (
<figureCitation id="13362A1DEE783872BD38F9F9F18BF994" box="[846,984,1580,1606]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="58" pageNumber="59">Fig. 39EG</figureCitation>
); S8 with dark reddish long macrosetae and setae at posterior margin, lateral to mid-posterior projection and on apex of mid-posterior projection (
<figureCitation id="13362A1DEE783872BB42F985F2FCF95C" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="58" pageNumber="59">Fig. 39E, G</figureCitation>
); apical epandrial finger-like projection pointed distally; gonocoxite tapering distally and curving upwards with apical half very narrow and apex rounded (
<figureCitation id="13362A1DEE783872BC0FF94DF08CF960" box="[633,735,1688,1714]" captionStart="FIGURE 40" captionStartId="62.[151,250,1881,1905]" captionTargetBox="[219,1369,193,1846]" captionTargetId="figure-21@62.[219,1369,181,1857]" captionTargetPageId="62" captionText="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="58" pageNumber="59">Fig. 40E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE783809BEB1F969F052FA6D" lastPageId="65" lastPageNumber="66" pageId="58" pageNumber="59" type="description">
<paragraph id="8BB23698EE783873BEB1F969F760FF2B" blockId="58.[151,1437,1472,2038]" lastBlockId="59.[151,1437,150,1977]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<emphasis id="B979EA8AEE783872BEB1F969F0E6F907" bold="true" box="[199,693,1723,1750]" pageId="58" pageNumber="59">
Description.
<typeStatus id="54B6883AEE783872BF14F969F39CF904" box="[354,463,1724,1750]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="13362A1DEE783872BC54F96EF0FBF907" box="[546,680,1723,1749]" captionStart-0="FIGURE 38" captionStart-1="FIGURE 39" captionStart-2="FIGURE 40" captionStart-3="FIGURE 41" captionStartId-0="60.[151,250,1943,1967]" captionStartId-1="61.[151,250,1877,1901]" captionStartId-2="62.[151,250,1881,1905]" captionStartId-3="63.[151,250,1885,1909]" captionTargetBox-0="[203,1385,181,1919]" captionTargetBox-1="[225,1362,181,1853]" captionTargetBox-2="[219,1369,193,1846]" captionTargetBox-3="[216,1368,184,1859]" captionTargetId-0="figure-21@60.[203,1385,181,1919]" captionTargetId-1="figure-17@61.[225,1362,181,1853]" captionTargetId-2="figure-21@62.[219,1369,181,1857]" captionTargetId-3="figure-17@63.[216,1371,181,1861]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionTargetPageId-2="62" captionTargetPageId-3="63" captionText-0="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." captionText-1="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." captionText-2="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." captionText-3="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7907408" figureDoi-1="http://doi.org/10.5281/zenodo.7907412" figureDoi-2="http://doi.org/10.5281/zenodo.7907414" figureDoi-3="http://doi.org/10.5281/zenodo.7907418" httpUri-0="https://zenodo.org/record/7907408/files/figure.png" httpUri-1="https://zenodo.org/record/7907412/files/figure.png" httpUri-2="https://zenodo.org/record/7907414/files/figure.png" httpUri-3="https://zenodo.org/record/7907418/files/figure.png" pageId="58" pageNumber="59">Figs 3841</figureCitation>
).
</emphasis>
Head (
<figureCitation id="13362A1DEE783872BD7DF969F1CBF904" box="[779,920,1724,1750]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="58" pageNumber="59">Fig. 38CD</figureCitation>
): eyes black; face yellowish brown pruinose; mystacal macrosetae yellowish white, except by a few dorsal black macrosetae (
<figureCitation id="13362A1DEE783872BD8AF935F6D0F928" box="[1020,1155,1760,1786]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="58" pageNumber="59">Fig. 38CD</figureCitation>
); subvibrissal macrosetae and setae yellowish white; gena yellowish pruinose; palpus black with two white apical macrosetae and black setae, except at base ventrally, yellowish white setose; proboscis black, labial setae dark yellow, proboscial setae whitish; antennal socket reddish black, brownish pruinose; frons brownish pruinose; orbital setae black with 12 yellowish mixed; ocellar tubercle black, brown pruinose dorsally with four proclinate black setae; vertex brown pruinose; occiput yellowish brown pruinose dorsally becoming white laterally; 56 black postocular macrosetae (proximal ones dark yellow), other dorsal occipital setae whitish; lateral occipital and lower occipital setae white, slightly branched. Antenna: scape and pedicel black with dark reddish apex, black setose and sparsely brown pruinose; postpedicel dark reddish, sparsely brown pruinose with 45 short yellowish setae dorsally at base (left postpedicel with an anomalous black macrosetae dorsally); first article of stylus dark reddish, minute, second article black, dark reddish at base, long and slender, slightly enlarged subapically ending in a dark reddish tip (
<figureCitation id="13362A1DEE793873BAEAFF0AF775FF2B" box="[1180,1318,223,249]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Fig. 39AB</figureCitation>
).
</paragraph>
<paragraph id="8BB23698EE793873BEB1FED6F6CAFDEF" blockId="59.[151,1437,150,1977]" pageId="59" pageNumber="60">
Thorax (
<figureCitation id="13362A1DEE793873BF5EFED6F3E0FECF" box="[296,435,259,285]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="59" pageNumber="60">Fig. 38AB</figureCitation>
): yellowish brown; antepronotum and postpronotum grey pruinose laterally; scutum with a pair of dark brown pruinose paramedian stripes rounded anteriorly, laterally, and fading after postsutural spots; pre and postsutural spots and prescutellar spots dark brown pruinose; median stripe brown pruinose; triangular apruinose black spot at posterior scutal margin; postalar wall and scutellum brownish grey pruinose (
<figureCitation id="13362A1DEE793873BB52FEBAF7DCFE5B" box="[1316,1423,367,393]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Fig. 39D</figureCitation>
); pleura grey pruinose, except anterodorsal part of proepimeron and anepisternum brownish grey pruinose. Chaetotaxy: antepronotal macrosetae and setae yellowish; 2 notopleural; 2 supra alar; 1 postalar and 5 pairs of dorsocentral postsutural black macrosetae; scutum with short black acrostichal setae; scutellum with short sparse black discal and marginal scutellar setae (
<figureCitation id="13362A1DEE793873BFADFE2AF06FFDCB" box="[475,572,511,537]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Fig.39D</figureCitation>
); anepisternum with short, thin yellowish and black setae dorsally; katatergite with whitish macrosetae and setae; remainder of pleura with very sparse, thin whitish setae.
</paragraph>
<paragraph id="8BB23698EE793873BEB1FD92F01EFD1F" blockId="59.[151,1437,150,1977]" pageId="59" pageNumber="60">
Wings (
<figureCitation id="13362A1DEE793873BF52FD92F3C2FDB3" box="[292,401,583,609]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Fig. 39C</figureCitation>
): yellowish translucent; veins dark brown;
<emphasis id="B979EA8AEE793873BDE3FD92F1FDFDB5" box="[917,942,583,615]" italics="true" pageId="59" pageNumber="60">
R
<subScript id="178934DDEE793873BDD0FD8CF1FDFDB5" attach="left" box="[934,942,601,615]" fontSize="6" pageId="59" pageNumber="60">4</subScript>
</emphasis>
slightly rounded in the junction with
<emphasis id="B979EA8AEE793873BB0DFD92F7C7FDB5" box="[1403,1428,583,615]" italics="true" pageId="59" pageNumber="60">
R
<subScript id="178934DDEE793873BBFAFD8CF7C7FDB5" attach="left" box="[1420,1428,601,615]" fontSize="6" pageId="59" pageNumber="60">5</subScript>
</emphasis>
; bifurcation of
<emphasis id="B979EA8AEE793873BF40FDBEF31CFD59" box="[310,335,619,651]" italics="true" pageId="59" pageNumber="60">
R
<subScript id="178934DDEE793873BF31FDA8F31CFD59" attach="left" box="[327,335,637,651]" fontSize="6" pageId="59" pageNumber="60">4</subScript>
</emphasis>
and
<emphasis id="B979EA8AEE793873BFF3FDBEF3CDFD59" box="[389,414,619,651]" italics="true" pageId="59" pageNumber="60">
R
<subScript id="178934DDEE793873BFE0FDA8F3CDFD59" attach="left" box="[406,414,637,651]" fontSize="6" pageId="59" pageNumber="60">5</subScript>
</emphasis>
beyond discal cell at a distance equivalent to the length of two crossvein
<emphasis id="B979EA8AEE793873BAB8FDBEF6A5FD56" box="[1230,1270,619,644]" italics="true" pageId="59" pageNumber="60">r-m</emphasis>
; crossvein
<emphasis id="B979EA8AEE793873BB02FDBEF7CFFD56" box="[1396,1436,619,644]" italics="true" pageId="59" pageNumber="60">r-m</emphasis>
in the middle of discal cell; cells
<emphasis id="B979EA8AEE793873BC77FD5AF04EFD7D" box="[513,541,655,687]" italics="true" pageId="59" pageNumber="60">
m
<subScript id="178934DDEE793873BC63FD74F04EFD7D" attach="left" box="[533,541,673,687]" fontSize="6" pageId="59" pageNumber="60">3</subScript>
</emphasis>
and
<emphasis id="B979EA8AEE793873BC27FD5AF02AFD7A" box="[593,633,655,680]" italics="true" pageId="59" pageNumber="60">cua</emphasis>
closed before wing margin. Halteres: dark reddish yellow, apex of steam and base of knob dark brown ventrally.
</paragraph>
<paragraph id="8BB23698EE793873BEB1FD02F69AFB8B" blockId="59.[151,1437,150,1977]" pageId="59" pageNumber="60">
Legs (
<figureCitation id="13362A1DEE793873BF7CFD02F322FD23" box="[266,369,727,753]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="59" pageNumber="60">Fig. 38A</figureCitation>
): coxae and trochanters black brownish grey pruinose; fore femur yellow (slightly dark at extreme apex), mid femur yellow with black apical third; hind femur black, slightly dark reddish anteroventrally on basal third; fore tibia yellow, only slightly reddish apically; mid tibia yellow with an anterior dark yellow stripe and black apex; hind tibia black with a yellow spot posterodorsally on basal half; tarsi black, except first tarsomere of fore tarsi dark reddish yellow. Chaetotaxy: coxae with yellowish white macrosetae and setae; hind trochanter with 3 posterior yellow macrosetae; fore femur with 1112 ventral yellowish macrosetae; mid femur with 910 ventral (yellowish), 5 anteroventral, 3 anterior, 1 posterodorsal preapical and 3 posterior (all yellow) black macrosetae; hind femur with 2 anteroventral preapical, 2 anterior (proximal yellow), 1 anterodorsal preapical, 1 posterodorsal preapical black macrosetae; femora black setose anterodorsally and yellowish posteroventrally; tibiae with black macrosetae and setae, except by a few yellow setae ventrally on fore tibia; tarsi with black macrosetae and setae; brush setae golden reddish yellow; pulvilli dark yellow; empodium dark reddish; claws black, dark reddish basally.
</paragraph>
<paragraph id="8BB23698EE793873BEB1FBB6F2B0FADF" blockId="59.[151,1437,150,1977]" pageId="59" pageNumber="60">
Abdomen (
<figureCitation id="13362A1DEE793873BF31FBB6F3E0FBAF" box="[327,435,1123,1149]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="59" pageNumber="60">Fig. 38A</figureCitation>
): black, T17 dark brown pruinose dorsally and grey laterally and at posterior corners; T1 with 34 yellowish white marginal lateral macrosetae and a tuft of long yellowish white setae; T23 with 45 yellowish white lateral marginal macrosetae; T57 only with long white setae at lateral posterior margin; other short appressed dorsal setae black, becoming yellowish laterally; sternites greyish white pruinose; sternites sparsely white setose.
</paragraph>
<paragraph id="8BB23698EE793873BEB1FAC2F083F94B" blockId="59.[151,1437,150,1977]" pageId="59" pageNumber="60">
Terminalia (
<figureCitation id="13362A1DEE793873BF3BFAC2F386FAE3" box="[333,469,1303,1329]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Figs 39EG</figureCitation>
,
<figureCitation id="13362A1DEE793873BFA8FAC2F07AFAE3" box="[478,553,1303,1329]" captionStart-0="FIGURE 40" captionStart-1="FIGURE 41" captionStartId-0="62.[151,250,1881,1905]" captionStartId-1="63.[151,250,1885,1909]" captionTargetBox-0="[219,1369,193,1846]" captionTargetBox-1="[216,1368,184,1859]" captionTargetId-0="figure-21@62.[219,1369,181,1857]" captionTargetId-1="figure-17@63.[216,1371,181,1861]" captionTargetPageId-0="62" captionTargetPageId-1="63" captionText-0="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." captionText-1="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7907414" figureDoi-1="http://doi.org/10.5281/zenodo.7907418" httpUri-0="https://zenodo.org/record/7907414/files/figure.png" httpUri-1="https://zenodo.org/record/7907418/files/figure.png" pageId="59" pageNumber="60">4041</figureCitation>
): shining black, except apical half of gonocoxite and dorsal margin of gonostylus and anterior mid-ventral part of S8 dark reddish; phallus dark yellow; T8 black setose (
<figureCitation id="13362A1DEE793873BA1EFAEEF6BFFA87" box="[1128,1260,1339,1365]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Figs 39EF</figureCitation>
,
<figureCitation id="13362A1DEE793873BA81FAEEF774FA87" box="[1271,1319,1339,1365]" captionStart="FIGURE 40" captionStartId="62.[151,250,1881,1905]" captionTargetBox="[219,1369,193,1846]" captionTargetId="figure-21@62.[219,1369,181,1857]" captionTargetPageId="62" captionText="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="59" pageNumber="60">40F</figureCitation>
); S8 black setose, mid-posterior projection rectangular with a tuft of long dark reddish black macrosetae at the apex (
<figureCitation id="13362A1DEE793873BB1CFA8AF2B8FA4F" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Figs 39E, G</figureCitation>
,
<figureCitation id="13362A1DEE793873BE8EFA56F37EFA4F" box="[248,301,1411,1437]" captionStart="FIGURE 40" captionStartId="62.[151,250,1881,1905]" captionTargetBox="[219,1369,193,1846]" captionTargetId="figure-21@62.[219,1369,181,1857]" captionTargetPageId="62" captionText="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="59" pageNumber="60">40D</figureCitation>
); cercus yellowish setose; epandrium with black setae at anterior dorsal margin and 34 long black macrosetae at anterior ventral margin (
<figureCitation id="13362A1DEE793873BC3BFA72F085FA13" box="[589,726,1447,1473]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Fig. 39EG</figureCitation>
), epandrial apical finger-like projection with short dark yellow setae apically and pointed distally (
<figureCitation id="13362A1DEE793873BC47FA1EF0CEFA37" box="[561,669,1483,1509]" captionStart="FIGURE 39" captionStartId="61.[151,250,1877,1901]" captionTargetBox="[225,1362,181,1853]" captionTargetId="figure-17@61.[225,1362,181,1853]" captionTargetPageId="61" captionText="FIGURE 39. Catostola indecisa sp. nov., holotype male. A. Postpedicel and stylus, anterior; B. Same, dorsal; C. Wing, dorsal; D. Scutellum, dorsal; E. Terminalia, lateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="59" pageNumber="60">Figs 39E</figureCitation>
,
<figureCitation id="13362A1DEE793873BCDDFA1EF08EFA37" box="[683,733,1483,1509]" captionStart="FIGURE 40" captionStartId="62.[151,250,1881,1905]" captionTargetBox="[219,1369,193,1846]" captionTargetId="figure-21@62.[219,1369,181,1857]" captionTargetPageId="62" captionText="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="59" pageNumber="60">40E</figureCitation>
); hypandrium with a shallow indentation at posterior margin (
<figureCitation id="13362A1DEE793873BEE9FA3AF35AF9DB" box="[159,265,1519,1545]" captionStart="FIGURE 40" captionStartId="62.[151,250,1881,1905]" captionTargetBox="[219,1369,193,1846]" captionTargetId="figure-21@62.[219,1369,181,1857]" captionTargetPageId="62" captionText="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="59" pageNumber="60">Fig. 40G</figureCitation>
); gonocoxite tapering distally and curving upwards with apical half very narrow and apex rounded (
<figureCitation id="13362A1DEE793873BB19FA3AF35FF9FF" captionStart="FIGURE 41" captionStartId="63.[151,250,1885,1909]" captionTargetBox="[216,1368,184,1859]" captionTargetId="figure-17@63.[216,1371,181,1861]" captionTargetPageId="63" captionText="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="59" pageNumber="60">Fig. 41A, DE</figureCitation>
), apex curved outwards in dorsal and ventral views (
<figureCitation id="13362A1DEE793873BD25F9C6F189F9FF" box="[851,986,1555,1581]" captionStart="FIGURE 41" captionStartId="63.[151,250,1885,1909]" captionTargetBox="[216,1368,184,1859]" captionTargetId="figure-17@63.[216,1371,181,1861]" captionTargetPageId="63" captionText="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="59" pageNumber="60">Fig. 41BC</figureCitation>
); gonostylus wide with apex conical and blade-like in lateral view (
<figureCitation id="13362A1DEE793873BFCDF9E2F04DF983" box="[443,542,1591,1617]" captionStart="FIGURE 41" captionStartId="63.[151,250,1885,1909]" captionTargetBox="[216,1368,184,1859]" captionTargetId="figure-17@63.[216,1371,181,1861]" captionTargetPageId="63" captionText="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="59" pageNumber="60">Fig. 41F</figureCitation>
); ejaculatory apodeme fan-shaped, directed posteriorly; phallus with prongs thin and long (
<figureCitation id="13362A1DEE793873BF70F98EF3DCF9A7" box="[262,399,1627,1653]" captionStart="FIGURE 41" captionStartId="63.[151,250,1885,1909]" captionTargetBox="[216,1368,184,1859]" captionTargetId="figure-17@63.[216,1371,181,1861]" captionTargetPageId="63" captionText="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="59" pageNumber="60">Fig. 41A, G</figureCitation>
); cercus short with apex rounded; subepandrial sclerite short with apex rounded and ventrally with an inverted Y-shaped projection (
<figureCitation id="13362A1DEE793873BC35F9AAF097F94B" box="[579,708,1663,1689]" captionStart="FIGURE 41" captionStartId="63.[151,250,1885,1909]" captionTargetBox="[216,1368,184,1859]" captionTargetId="figure-17@63.[216,1371,181,1861]" captionTargetPageId="63" captionText="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="59" pageNumber="60">Fig. 41HI</figureCitation>
).
</paragraph>
<paragraph id="8BB23698EE793873BEB1F976F0C8F96F" blockId="59.[151,1437,150,1977]" box="[199,667,1698,1725]" pageId="59" pageNumber="60">
<emphasis id="B979EA8AEE793873BEB1F976F377F96F" bold="true" box="[199,292,1699,1725]" pageId="59" pageNumber="60">Length.</emphasis>
Body: 18.5 mm; wing: 12.3 mm.
</paragraph>
<paragraph id="8BB23698EE793873BEB1F912F1B3F86B" blockId="59.[151,1437,150,1977]" pageId="59" pageNumber="60">
<emphasis id="B979EA8AEE793873BEB1F912F3C4F932" bold="true" box="[199,407,1734,1761]" pageId="59" pageNumber="60">
Female (
<figureCitation id="13362A1DEE793873BF47F913F3D9F932" box="[305,394,1734,1760]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42</figureCitation>
).
</emphasis>
Similar to male except terminalia (
<figureCitation id="13362A1DEE793873BD4DF912F196F933" box="[827,965,1735,1761]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42AF</figureCitation>
); T7 shining black and black setose; T8 shining black and black setose; cercus with yellowish setae apically; S8 and hypoproct with dark reddish setae; keel of genital fork and hypoproct with dark reddish spines (
<figureCitation id="13362A1DEE793873BD65F8DAF1CCF8FB" box="[787,927,1807,1833]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42AC</figureCitation>
); spermathecae rod-like (
<figureCitation id="13362A1DEE793873BACBF8DAF775F8FB" box="[1213,1318,1807,1833]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42G</figureCitation>
), reaching abdominal segment 6; spermathecal pump about 3 times smaller than length of spermathecal duct, slightly turgid (
<figureCitation id="13362A1DEE793873BEE9F882F373F8A3" box="[159,288,1879,1905]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42GI</figureCitation>
); spermathecal ducts ending separately at genital fork (
<figureCitation id="13362A1DEE793873BDF3F882F656F8A3" box="[901,1029,1879,1905]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42GI</figureCitation>
); apodeme of genital fork connected laterally with lateral sclerites (
<figureCitation id="13362A1DEE793873BF9EF8AEF034F847" box="[488,615,1915,1941]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42GI</figureCitation>
), slightly enlarged at mid-length (
<figureCitation id="13362A1DEE793873BDA9F8AEF639F847" box="[991,1130,1915,1941]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42GH</figureCitation>
); lateral sclerites connected with T9+10; apodeme of genital fork without a mid-dorsal ridge (
<figureCitation id="13362A1DEE793873BD00F84AF187F86B" box="[886,980,1951,1977]" captionStart="FIGURE 42" captionStartId="64.[151,250,1854,1878]" captionTargetBox="[233,1354,181,1830]" captionTargetId="figure-21@64.[233,1354,181,1830]" captionTargetPageId="64" captionText="FIGURE 42. Catostola indecisa sp. nov., paratype female. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Dissected terminalia, lateral; E. Same, dorsal; F. Same, ventral; G. Spermathecae, dorsal; H. genital fork, detail, dorsal; I. Same, lateral. Abbreviations: acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="59" pageNumber="60">Fig. 42I</figureCitation>
).
</paragraph>
<caption id="DF726610EE7E3874BEE1F842F0D3F801" ID-DOI="http://doi.org/10.5281/zenodo.7907408" ID-Zenodo-Dep="7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="60" pageNumber="61" startId="60.[151,250,1943,1967]" targetBox="[203,1385,181,1919]" targetPageId="60" targetType="figure">
<paragraph id="8BB23698EE7E3874BEE1F842F0D3F801" blockId="60.[151,1436,1943,2004]" pageId="60" pageNumber="61">
<emphasis id="B979EA8AEE7E3874BEE1F842F372F87D" bold="true" box="[151,289,1943,1967]" pageId="60" pageNumber="61">FIGURE 38.</emphasis>
<taxonomicName id="4C0D4D1BEE7E3874BF5EF84DF3B5F862" box="[296,486,1944,1968]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE7E3874BF5EF84DF3B5F862" box="[296,486,1944,1968]" italics="true" pageId="60" pageNumber="61">Catostola indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE7E3874BF98F84DF06CF862" bold="true" box="[494,575,1944,1968]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="A24A57F1EE7E3874BF98F84DF06CF862" box="[494,575,1944,1968]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B979EA8AEE7E3874BC9AF842F156F87D" bold="true" box="[748,773,1943,1967]" pageId="60" pageNumber="61">A.</emphasis>
Habitus, lateral (insert: labels, not to scale);
<emphasis id="B979EA8AEE7E3874BAA5F84DF6B9F862" bold="true" box="[1235,1258,1944,1968]" pageId="60" pageNumber="61">B.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE7E3874BBF2F842F7CFF87D" bold="true" box="[1412,1436,1943,1967]" pageId="60" pageNumber="61">C.</emphasis>
Head, frontal;
<emphasis id="B979EA8AEE7E3874BF5CF869F310F806" bold="true" box="[298,323,1980,2004]" pageId="60" pageNumber="61">D.</emphasis>
Same, lateral. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF726610EE7F3875BEE1F880F1BFF80B" ID-DOI="http://doi.org/10.5281/zenodo.7907412" ID-Zenodo-Dep="7907412" httpUri="https://zenodo.org/record/7907412/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1877,1901]" targetBox="[225,1362,181,1853]" targetPageId="61" targetType="figure">
<paragraph id="8BB23698EE7F3875BEE1F880F1BFF80B" blockId="61.[151,1437,1877,2010]" pageId="61" pageNumber="62">
<emphasis id="B979EA8AEE7F3875BEE1F880F373F8BF" bold="true" box="[151,288,1877,1901]" pageId="61" pageNumber="62">FIGURE 39.</emphasis>
<taxonomicName id="4C0D4D1BEE7F3875BF53F880F3B1F8BF" box="[293,482,1877,1901]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE7F3875BF53F880F3B1F8BF" box="[293,482,1877,1901]" italics="true" pageId="61" pageNumber="62">Catostola indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE7F3875BF9EF883F06BF8BC" bold="true" box="[488,568,1878,1902]" pageId="61" pageNumber="62">
<taxonomicNameLabel id="A24A57F1EE7F3875BF9EF883F06BF8BC" box="[488,568,1878,1902]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B979EA8AEE7F3875BC97F880F0A9F8BF" bold="true" box="[737,762,1877,1901]" pageId="61" pageNumber="62">A.</emphasis>
Postpedicel and stylus, anterior;
<emphasis id="B979EA8AEE7F3875BA3FF883F633F8BC" bold="true" box="[1097,1120,1878,1902]" pageId="61" pageNumber="62">B.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE7F3875BA80F880F75DF8BF" bold="true" box="[1270,1294,1877,1901]" pageId="61" pageNumber="62">C.</emphasis>
Wing, dorsal;
<emphasis id="B979EA8AEE7F3875BEE1F8AFF2E3F840" bold="true" box="[151,176,1914,1938]" pageId="61" pageNumber="62">D.</emphasis>
Scutellum, dorsal;
<emphasis id="B979EA8AEE7F3875BF05F8AFF3D9F840" bold="true" box="[371,394,1914,1938]" pageId="61" pageNumber="62">E.</emphasis>
Terminalia, lateral;
<emphasis id="B979EA8AEE7F3875BC23F8AFF03BF840" bold="true" box="[597,616,1914,1938]" pageId="61" pageNumber="62">F.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE7F3875BC8BF8ACF14BF843" bold="true" box="[765,792,1913,1937]" pageId="61" pageNumber="62">G.</emphasis>
Same, ventral.
<emphasis id="B979EA8AEE7F3875BDC2F8ACF606F843" bold="true" box="[948,1109,1913,1937]" pageId="61" pageNumber="62">Abbreviations:</emphasis>
cerc: cercus; epand: epandrium; f a styl: first article of stylus; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; pped: postpedicel; s a styl: second article of stylus; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF726610EE7C3876BEE1F88CF36EF80F" ID-DOI="http://doi.org/10.5281/zenodo.7907414" ID-Zenodo-Dep="7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="62" pageNumber="63" startId="62.[151,250,1881,1905]" targetBox="[219,1369,193,1846]" targetPageId="62" targetType="figure">
<paragraph id="8BB23698EE7C3876BEE1F88CF36EF80F" blockId="62.[151,1437,1881,2013]" pageId="62" pageNumber="63">
<emphasis id="B979EA8AEE7C3876BEE1F88CF373F8A3" bold="true" box="[151,288,1881,1905]" pageId="62" pageNumber="63">FIGURE 40.</emphasis>
<taxonomicName id="4C0D4D1BEE7C3876BF50F88CF3B0F8A3" box="[294,483,1881,1905]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE7C3876BF50F88CF3B0F8A3" box="[294,483,1881,1905]" italics="true" pageId="62" pageNumber="63">Catostola indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE7C3876BF9CF88CF069F8A3" bold="true" box="[490,570,1881,1905]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A24A57F1EE7C3876BF9CF88CF069F8A3" box="[490,570,1881,1905]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B979EA8AEE7C3876BC92F88CF0AEF8A3" bold="true" box="[740,765,1881,1905]" pageId="62" pageNumber="63">A.</emphasis>
Terminalia, dorsal;
<emphasis id="B979EA8AEE7C3876BDBEF88CF18CF8A3" bold="true" box="[968,991,1881,1905]" pageId="62" pageNumber="63">B.</emphasis>
Same, ventral;
<emphasis id="B979EA8AEE7C3876BA08F88CF6C5F8A3" bold="true" box="[1150,1174,1881,1905]" pageId="62" pageNumber="63">C.</emphasis>
Same, lateral;
<emphasis id="B979EA8AEE7C3876BB58F88CF714F8A3" bold="true" box="[1326,1351,1881,1905]" pageId="62" pageNumber="63">D.</emphasis>
Sternite 8, ventral;
<emphasis id="B979EA8AEE7C3876BF72F8A8F348F847" bold="true" box="[260,283,1917,1941]" pageId="62" pageNumber="63">E.</emphasis>
Epandrium, lateral;
<emphasis id="B979EA8AEE7C3876BF9CF8A8F3AEF847" bold="true" box="[490,509,1917,1941]" pageId="62" pageNumber="63">F.</emphasis>
Tergite 8, dorsal;
<emphasis id="B979EA8AEE7C3876BCC2F8A8F09DF847" bold="true" box="[692,718,1917,1941]" pageId="62" pageNumber="63">G.</emphasis>
Hypandrium, ventral.
<emphasis id="B979EA8AEE7C3876BDC5F8A8F607F847" bold="true" box="[947,1108,1917,1941]" pageId="62" pageNumber="63">Abbreviations:</emphasis>
cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF726610EE7D3877BEE1F888F61CF833" ID-DOI="http://doi.org/10.5281/zenodo.7907418" ID-Zenodo-Dep="7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,250,1885,1909]" targetBox="[216,1368,184,1859]" targetPageId="63" targetType="figure">
<paragraph id="8BB23698EE7D3877BEE1F888F61CF833" blockId="63.[151,1437,1885,2018]" pageId="63" pageNumber="64">
<emphasis id="B979EA8AEE7D3877BEE1F888F372F8A7" bold="true" box="[151,289,1885,1909]" pageId="63" pageNumber="64">FIGURE 41.</emphasis>
<taxonomicName id="4C0D4D1BEE7D3877BF5EF88BF3B5F8A4" box="[296,486,1886,1910]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE7D3877BF5EF88BF3B5F8A4" box="[296,486,1886,1910]" italics="true" pageId="63" pageNumber="64">Catostola indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE7D3877BF98F88BF013F8A4" bold="true" box="[494,576,1886,1910]" pageId="63" pageNumber="64">
<taxonomicNameLabel id="A24A57F1EE7D3877BF98F88BF013F8A4" box="[494,576,1886,1910]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B979EA8AEE7D3877BC9BF888F155F8A7" bold="true" box="[749,774,1885,1909]" pageId="63" pageNumber="64">A.</emphasis>
Gonopods + phallus, lateral;
<emphasis id="B979EA8AEE7D3877BA41F88BF61CF8A4" bold="true" box="[1079,1103,1886,1910]" pageId="63" pageNumber="64">B.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE7D3877BA9FF888F752F8A7" bold="true" box="[1257,1281,1885,1909]" pageId="63" pageNumber="64">C.</emphasis>
Same, ventral;
<emphasis id="B979EA8AEE7D3877BEE1F857F2E3F848" bold="true" box="[151,176,1922,1946]" pageId="63" pageNumber="64">D.</emphasis>
Gonopods, lateral;
<emphasis id="B979EA8AEE7D3877BF0BF857F3C7F848" bold="true" box="[381,404,1922,1946]" pageId="63" pageNumber="64">E.</emphasis>
Gonocoxite, lateral;
<emphasis id="B979EA8AEE7D3877BC06F857F0D0F848" bold="true" box="[624,643,1922,1946]" pageId="63" pageNumber="64">F.</emphasis>
Gonostylus, lateral;
<emphasis id="B979EA8AEE7D3877BD2AF854F125F84B" bold="true" box="[860,886,1921,1945]" pageId="63" pageNumber="64">G.</emphasis>
Phallus, lateral;
<emphasis id="B979EA8AEE7D3877BA50F857F613F848" bold="true" box="[1062,1088,1922,1946]" pageId="63" pageNumber="64">H.</emphasis>
Cercus and subepandrial sclerite, lateral;
<emphasis id="B979EA8AEE7D3877BE95F873F2A0F86C" bold="true" box="[227,243,1958,1982]" pageId="63" pageNumber="64">I.</emphasis>
Same, ventral.
<emphasis id="B979EA8AEE7D3877BFF8F870F07CF86F" bold="true" box="[398,559,1957,1981]" pageId="63" pageNumber="64">Abbreviations:</emphasis>
cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF726610EE023808BEE1F8EBF3D9F834" ID-DOI="http://doi.org/10.5281/zenodo.7907420" ID-Zenodo-Dep="7907420" httpUri="https://zenodo.org/record/7907420/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,250,1854,1878]" targetBox="[233,1354,181,1830]" targetPageId="64" targetType="figure">
<paragraph id="8BB23698EE023808BEE1F8EBF3D9F834" blockId="64.[151,1437,1854,2022]" pageId="64" pageNumber="65">
<emphasis id="B979EA8AEE023808BEE1F8EBF377F884" bold="true" box="[151,292,1854,1878]" pageId="64" pageNumber="65">FIGURE 42.</emphasis>
<taxonomicName id="4C0D4D1BEE023808BF58F8EBF3BCF884" box="[302,495,1854,1878]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE023808BF58F8EBF3BCF884" box="[302,495,1854,1878]" italics="true" pageId="64" pageNumber="65">Catostola indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE023808BF8FF8EBF01DF884" bold="true" box="[505,590,1854,1878]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="A24A57F1EE023808BF8FF8EBF01DF884" box="[505,590,1854,1878]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female.
<emphasis id="B979EA8AEE023808BD60F8EBF17DF884" bold="true" box="[790,814,1854,1878]" pageId="64" pageNumber="65">A.</emphasis>
Terminalia, lateral;
<emphasis id="B979EA8AEE023808BA71F8EBF64DF884" bold="true" box="[1031,1054,1854,1878]" pageId="64" pageNumber="65">B.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE023808BAB6F8EBF68AF884" bold="true" box="[1216,1241,1854,1878]" pageId="64" pageNumber="65">C.</emphasis>
Same, ventral;
<emphasis id="B979EA8AEE023808BBF2F8EBF7CFF884" bold="true" box="[1412,1436,1854,1878]" pageId="64" pageNumber="65">D.</emphasis>
Dissected terminalia, lateral;
<emphasis id="B979EA8AEE023808BFBFF8B7F3B3F8A8" bold="true" box="[457,480,1890,1914]" pageId="64" pageNumber="65">E.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE023808BCF6F8B7F0C0F8A8" bold="true" box="[640,659,1890,1914]" pageId="64" pageNumber="65">F.</emphasis>
Same, ventral;
<emphasis id="B979EA8AEE023808BD4DF8B7F106F8A8" bold="true" box="[827,853,1890,1914]" pageId="64" pageNumber="65">G.</emphasis>
Spermathecae, dorsal;
<emphasis id="B979EA8AEE023808BA3CF8B7F636F8A8" bold="true" box="[1098,1125,1890,1914]" pageId="64" pageNumber="65">H.</emphasis>
genital fork, detail, dorsal;
<emphasis id="B979EA8AEE023808BBFAF8B7F7CFF8A8" bold="true" box="[1420,1436,1890,1914]" pageId="64" pageNumber="65">I.</emphasis>
Same, lateral.
<emphasis id="B979EA8AEE023808BF5AF853F39EF84C" bold="true" box="[300,461,1926,1950]" pageId="64" pageNumber="65">Abbreviations:</emphasis>
acc gl: accessory gland; apod: apodeme of genital fork; cerc: cercus; gen fk: genital fork; lat sc: lateral sclerite of genital fork; spe du: spermathecal duct; spe pu: spermathecal pump; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF726610EE033809BEE1FAEEF178FA81" ID-DOI="http://doi.org/10.5281/zenodo.7907423" ID-Zenodo-Dep="7907423" box="[151,811,1339,1363]" httpUri="https://zenodo.org/record/7907423/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1339,1363]" targetBox="[151,1435,181,1314]" targetPageId="65" targetType="figure">
<paragraph id="8BB23698EE033809BEE1FAEEF178FA81" blockId="65.[151,811,1339,1363]" box="[151,811,1339,1363]" pageId="65" pageNumber="66">
<emphasis id="B979EA8AEE033809BEE1FAEEF373FA81" bold="true" box="[151,288,1339,1363]" pageId="65" pageNumber="66">FIGURE 43.</emphasis>
Distribution map of
<taxonomicName id="4C0D4D1BEE033809BF82FAEEF099FA81" authority="Hull, 1958" authorityName="Hull" authorityYear="1958" box="[500,714,1339,1363]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus" status="stat. nov.">
<emphasis id="B979EA8AEE033809BF82FAEEF004FA81" box="[500,599,1339,1363]" italics="true" pageId="65" pageNumber="66">Catostola</emphasis>
<bibRefCitation id="EF9C4B69EE033809BC2BFAEEF099FA81" author="Hull, F. M." box="[605,714,1339,1363]" pageId="65" pageNumber="66" pagination="317 - 324" refId="ref35129" refString="Hull, F. M. (1958) New genera of robber flies (Diptera, Asilidae). Revista Brasileira de Biologia, 18 (3), 317 - 324." type="journal article" year="1958">Hull, 1958</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A24A57F1EE033809BCA6FAEEF178FA81" box="[720,811,1339,1363]" pageId="65" pageNumber="66">stat. rev.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB23698EE033809BEB1FA54F052FA6D" blockId="65.[151,1437,1409,2014]" pageId="65" pageNumber="66">
<emphasis id="B979EA8AEE033809BEB1FA54F36CFA49" bold="true" box="[199,319,1409,1435]" pageId="65" pageNumber="66">Variation.</emphasis>
Face pruinosity golden yellow; postpedicel black; apex of fore femur black; apex of fore tibia and first tarsomere of fore leg black.
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE033809BEB1FA1CF6D9F8F5" pageId="65" pageNumber="66" type="etymology">
<paragraph id="8BB23698EE033809BEB1FA1CF03FF9F9" blockId="65.[151,1437,1409,2014]" pageId="65" pageNumber="66">
<emphasis id="B979EA8AEE033809BEB1FA1CF31DFA31" bold="true" box="[199,334,1481,1507]" pageId="65" pageNumber="66">Etymology.</emphasis>
<taxonomicName id="4C0D4D1BEE033809BF21FA1CF3E7FA31" box="[343,436,1481,1507]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="indecisa">
<emphasis id="B979EA8AEE033809BF21FA1CF3E7FA31" box="[343,436,1481,1507]" italics="true" pageId="65" pageNumber="66">indecisa</emphasis>
</taxonomicName>
, from Latin:
<emphasis id="B979EA8AEE033809BC26FA1CF0EBFA31" box="[592,696,1481,1507]" italics="true" pageId="65" pageNumber="66">indecisus</emphasis>
: undecided, not settled or determined. The name makes allusion to the similarity of this species with
<taxonomicName id="4C0D4D1BEE033809BC43FA38F0CEF9D5" baseAuthorityName="Walker" baseAuthorityYear="1849" box="[565,669,1517,1543]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="baleta">
<emphasis id="B979EA8AEE033809BC43FA38F0CEF9D5" box="[565,669,1517,1543]" italics="true" pageId="65" pageNumber="66">C. baleta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D1BEE033809BCA3FA38F11AF9D5" baseAuthorityName="Fisher" baseAuthorityYear="1992" box="[725,841,1517,1543]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="B979EA8AEE033809BCA3FA38F11AF9D5" box="[725,841,1517,1543]" italics="true" pageId="65" pageNumber="66">C. martini</emphasis>
</taxonomicName>
, being
<taxonomicName id="4C0D4D1BEE033809BDE9FA38F64EF9D5" box="[927,1053,1517,1543]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE033809BDE9FA38F64EF9D5" box="[927,1053,1517,1543]" italics="true" pageId="65" pageNumber="66">C. indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE033809BA50FA38F6D3F9D5" bold="true" box="[1062,1152,1517,1543]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A24A57F1EE033809BA50FA38F6D3F9D5" box="[1062,1152,1517,1543]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, an intermediate, hard to identify among its closely related species.
</paragraph>
<paragraph id="8BB23698EE033809BEB1F9E0F6D9F8F5" blockId="65.[151,1437,1409,2014]" pageId="65" pageNumber="66">
<emphasis id="B979EA8AEE033809BEB1F9E0F383F99D" bold="true" box="[199,464,1589,1615]" pageId="65" pageNumber="66">Taxonomic discussion.</emphasis>
The most similar species are
<taxonomicName id="4C0D4D1BEE033809BD65F9E0F198F99D" baseAuthorityName="Walker" baseAuthorityYear="1849" box="[787,971,1589,1615]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="baleta" status="comb. nov.">
<emphasis id="B979EA8AEE033809BD65F9E0F198F99D" box="[787,971,1589,1615]" italics="true" pageId="65" pageNumber="66">Catostola baleta</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE033809BDA7F9E0F603F99D" bold="true" box="[977,1104,1589,1615]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A24A57F1EE033809BDA7F9E0F603F99D" box="[977,1104,1589,1615]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C0D4D1BEE033809BAFCF9E0F71DF99D" baseAuthorityName="Fisher" baseAuthorityYear="1992" box="[1162,1358,1589,1615]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="martini" status="comb. nov.">
<emphasis id="B979EA8AEE033809BAFCF9E0F71DF99D" box="[1162,1358,1589,1615]" italics="true" pageId="65" pageNumber="66">Catostola martini</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE033809BB22F9E0F29BF9A1" bold="true" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A24A57F1EE033809BB22F9E0F29BF9A1" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, It can be differentiated from the first by the entire black hind femur and by the shape of epandrial finger-like projection with the apex being constricted earlier (
<figureCitation id="13362A1DEE033809BCB4F9A8F177F945" box="[706,804,1661,1687]" captionStart="FIGURE 40" captionStartId="62.[151,250,1881,1905]" captionTargetBox="[219,1369,193,1846]" captionTargetId="figure-21@62.[219,1369,181,1857]" captionTargetPageId="62" captionText="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="65" pageNumber="66">Fig. 40E</figureCitation>
) and the shape of gonocoxite which has the posterior half narrower and the gonostylus slightly wider (
<figureCitation id="13362A1DEE033809BCF9F974F147F969" box="[655,788,1697,1723]" captionStart="FIGURE 41" captionStartId="63.[151,250,1885,1909]" captionTargetBox="[216,1368,184,1859]" captionTargetId="figure-17@63.[216,1371,181,1861]" captionTargetPageId="63" captionText="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="65" pageNumber="66">Fig. 41EF</figureCitation>
). From the second only characters of the male terminalia are useful to make the distinction. The epandrial finger-like projection is more indented ventrally sub-apically in
<taxonomicName id="4C0D4D1BEE033809BEE1F93CF359F8D1" baseAuthorityName="Fisher" baseAuthorityYear="1992" box="[151,266,1769,1795]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="martini" status="comb. nov.">
<emphasis id="B979EA8AEE033809BEE1F93CF359F8D1" box="[151,266,1769,1795]" italics="true" pageId="65" pageNumber="66">C. martini</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE033809BF67F93CF3C1F8D1" bold="true" box="[273,402,1769,1795]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A24A57F1EE033809BF67F93CF3C1F8D1" box="[273,402,1769,1795]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13362A1DEE033809BFD7F93CF057F8D1" box="[417,516,1769,1795]" captionStart="FIGURE 20" captionStartId="35.[151,250,1891,1915]" captionTargetBox="[241,1347,191,1863]" captionTargetId="figure-17@35.[238,1350,181,1867]" captionTargetPageId="35" captionText="FIGURE 20. Catostola martini (Fisher, 1992) comb. nov., paratype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907370" httpUri="https://zenodo.org/record/7907370/files/figure.png" pageId="65" pageNumber="66">Fig. 20E</figureCitation>
) than
<taxonomicName id="4C0D4D1BEE033809BC3CF93CF09BF8D1" box="[586,712,1769,1795]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE033809BC3CF93CF09BF8D1" box="[586,712,1769,1795]" italics="true" pageId="65" pageNumber="66">C. indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE033809BCB9F93CF17AF8D1" bold="true" box="[719,809,1769,1795]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A24A57F1EE033809BCB9F93CF17AF8D1" box="[719,809,1769,1795]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13362A1DEE033809BD4EF93CF1CEF8D1" box="[824,925,1769,1795]" captionStart="FIGURE 40" captionStartId="62.[151,250,1881,1905]" captionTargetBox="[219,1369,193,1846]" captionTargetId="figure-21@62.[219,1369,181,1857]" captionTargetPageId="62" captionText="FIGURE 40. Catostola indecisa sp. nov., holotype male. A. Terminalia, dorsal; B. Same, ventral; C. Same, lateral; D. Sternite 8, ventral; E. Epandrium, lateral; F. Tergite 8, dorsal; G. Hypandrium, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907414" httpUri="https://zenodo.org/record/7907414/files/figure.png" pageId="65" pageNumber="66">Fig. 40E</figureCitation>
), the gonocoxite is much narrower (
<figureCitation id="13362A1DEE033809BB47F93CF7C7F8D1" box="[1329,1428,1769,1795]" captionStart="FIGURE 41" captionStartId="63.[151,250,1885,1909]" captionTargetBox="[216,1368,184,1859]" captionTargetId="figure-17@63.[216,1371,181,1861]" captionTargetPageId="63" captionText="FIGURE 41. Catostola indecisa sp. nov., holotype male. A. Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907418" httpUri="https://zenodo.org/record/7907418/files/figure.png" pageId="65" pageNumber="66">Fig. 41E</figureCitation>
) while in
<taxonomicName id="4C0D4D1BEE033809BE8FF8D8F33FF8F5" baseAuthorityName="Fisher" baseAuthorityYear="1992" box="[249,364,1805,1831]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="martini" status="comb. nov.">
<emphasis id="B979EA8AEE033809BE8FF8D8F33FF8F5" box="[249,364,1805,1831]" italics="true" pageId="65" pageNumber="66">C. martini</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE033809BF05F8D8F3A7F8F5" bold="true" box="[371,500,1805,1831]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A24A57F1EE033809BF05F8D8F3A7F8F5" box="[371,500,1805,1831]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
it only becomes extremely narrow sub-apically (
<figureCitation id="13362A1DEE033809BA6EF8D8F62DF8F5" box="[1048,1150,1805,1831]" captionStart="FIGURE 21" captionStartId="36.[151,250,1892,1916]" captionTargetBox="[226,1364,181,1866]" captionTargetId="figure-21@36.[215,1372,181,1869]" captionTargetPageId="36" captionText="FIGURE 21. Catostola martini(Fisher, 1992) comb. nov., paratype male.Gonopods + phallus, lateral; B. Same, dorsal; C. Same, ventral; D. Gonopods, lateral; E. Gonocoxite, lateral; F. Gonostylus, lateral; G. Phallus, lateral; H. Cercus and subepandrial sclerite, lateral; I. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; ph: phallus; subepand scl: subepandrial sclerite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907372" httpUri="https://zenodo.org/record/7907372/files/figure.png" pageId="65" pageNumber="66">Fig. 21E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE033809BEB1F8E4F18BF899" box="[199,984,1841,1867]" pageId="65" pageNumber="66" type="distribution">
<paragraph id="8BB23698EE033809BEB1F8E4F18BF899" blockId="65.[151,1437,1409,2014]" box="[199,984,1841,1867]" pageId="65" pageNumber="66">
<emphasis id="B979EA8AEE033809BEB1F8E4F398F899" bold="true" box="[199,459,1841,1867]" pageId="65" pageNumber="66">
Distribution (
<figureCitation id="13362A1DEE033809BF1EF8E4F3EDF899" box="[360,446,1841,1867]" captionStart="FIGURE 44" captionStartId="66.[151,250,1682,1706]" captionTargetBox="[151,1435,525,1658]" captionTargetId="figure-172@66.[151,1435,525,1658]" captionTargetPageId="66" captionText="FIGURE 44. Distribution map of Catostola species: Catostola baleta (Walker, 1849) comb. nov., Catostola complicata (James, 1953) comb. nov., Catostola indecisa sp. nov., and Catostola martini (Fisher, 1992) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.7907427" httpUri="https://zenodo.org/record/7907427/files/figure.png" pageId="65" pageNumber="66">Fig. 44</figureCitation>
).
</emphasis>
<collectingCountry id="F31A7608EE033809BFA4F8E4F01FF899" box="[466,588,1841,1867]" name="Costa Rica" pageId="65" pageNumber="66">Costa Rica</collectingCountry>
(
<collectingRegion id="49C9F87AEE033809BC2DF8E4F089F899" box="[603,730,1841,1867]" country="Costa Rica" name="Puntarenas" pageId="65" pageNumber="66">Puntarenas</collectingRegion>
,
<collectingRegion id="49C9F87AEE033809BC93F8E4F163F899" box="[741,816,1841,1867]" country="Costa Rica" name="Limon" pageId="65" pageNumber="66">Limón</collectingRegion>
and
<collectingRegion id="49C9F87AEE033809BD10F8E4F19FF899" box="[870,972,1841,1867]" country="Costa Rica" name="San Jose" pageId="65" pageNumber="66">San Jose</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE03380ABEB1F880F3CAFE27" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="8BB23698EE03380ABEB1F880F3CAFE27" blockId="65.[151,1437,1409,2014]" lastBlockId="66.[151,1437,150,504]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<emphasis id="B979EA8AEE033809BEB1F880F3E1F8BD" bold="true" box="[199,434,1877,1903]" pageId="65" pageNumber="66">Material examined.</emphasis>
<materialsCitation id="3B653CC5EE033809BFCCF880F604F809" collectingDate="2003-06-21" collectingDateMax="2003-07-07" collectingDateMin="2003-06-21" collectionCode="MNCR, INBIO" collectorName="K. Caballero" country="Costa Rica" county="Corcovado" elevation="201" latitude="8.453333" location="Cerro Puma" longLatPrecision="21" longitude="-83.49305" municipality="Sector La Leona" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<emphasis id="B979EA8AEE033809BFCCF880F0B0F8BD" bold="true" box="[442,739,1877,1903]" pageId="65" pageNumber="66">
<typeStatus id="54B6883AEE033809BFCCF880F07AF8BD" box="[442,553,1877,1903]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F31A7608EE033809BC40F880F0B0F8BD" box="[566,739,1877,1903]" name="Costa Rica" pageId="65" pageNumber="66">COSTA RICA</collectingCountry>
</emphasis>
, Prov.[incia]
<emphasis id="B979EA8AEE033809BD08F880F655F8BD" bold="true" box="[894,1030,1877,1903]" pageId="65" pageNumber="66">
<collectingRegion id="49C9F87AEE033809BD08F880F655F8BD" box="[894,1030,1877,1903]" country="Costa Rica" name="Puntarenas" pageId="65" pageNumber="66">Puntarenas</collectingRegion>
</emphasis>
, P.N.[Parque Nacional]
<collectingCounty id="62D34E14EE033809BB6CF880F7CBF8BD" box="[1306,1432,1877,1903]" pageId="65" pageNumber="66">Corcovado</collectingCounty>
,
<collectingMunicipality id="6BD6ACE2EE033809BEE1F8ACF307F846" box="[151,340,1913,1940]" pageId="65" pageNumber="66">Sector La Leona</collectingMunicipality>
,
<location id="8ED26043EE033809BF29F8ACF3B5F846" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE033809BF29F8ACF3B5F846" box="[351,486,1913,1940]" country="Costa Rica" county="Corcovado" latitude="8.453333" longLatPrecision="21" longitude="-83.49305" municipality="Sector La Leona" name="Cerro Puma" pageId="65" pageNumber="66" stateProvince="Puntarenas">Cerro Puma</location>
<quantity id="4CF59B7DEE033809BF98F8ACF039F846" box="[494,618,1913,1940]" metricMagnitude="2" metricUnit="m" metricValue="2.01" metricValueMax="3.02" metricValueMin="1.0" pageId="65" pageNumber="66" unit="m" value="201.0" valueMax="302.0" valueMin="100.0">
<elevation id="0020D1ABEE033809BF98F8ACF039F846" box="[494,618,1913,1940]" metricMagnitude="2" metricUnit="m" metricValue="2.01" metricValueMax="3.02" metricValueMin="1.0" pageId="65" pageNumber="66" unit="m" value="201.0" valueMax="302.0" valueMin="100.0">100-302 m</elevation>
</quantity>
,
<date id="FFB31058EE033809BC00F8ACF131F841" box="[630,866,1913,1940]" pageId="65" pageNumber="66" value="2003-06-21" valueMax="2003-07-07" valueMin="2003-06-21">
<collectingDate id="EFF7E9B0EE033809BC00F8ACF131F841" box="[630,866,1913,1940]" pageId="65" pageNumber="66" value="2003-06-21" valueMax="2003-07-07" valueMin="2003-06-21">21 JUN7 JUL 2003</collectingDate>
</date>
,
<collectorName id="26F8534EEE033809BD18F8AFF653F841" box="[878,1024,1913,1940]" pageId="65" pageNumber="66">K. Caballero</collectorName>
,
<collectorName id="26F8534EEE033809BA7DF8ACF634F841" box="[1035,1127,1913,1939]" pageId="65" pageNumber="66">Libre, L</collectorName>
_S_[Lambert Sur] 267700_ 518900 [
<geoCoordinate id="EE39505FEE033809BE80F84BF323F86B" box="[246,368,1950,1977]" degrees="8" direction="north" minutes="27" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="12" value="8.453333">8°2712″N</geoCoordinate>
<geoCoordinate id="EE39505FEE033809BF02F84BF051F86B" box="[372,514,1950,1977]" degrees="83" direction="west" minutes="29" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="35" value="-83.49305">83°2935″W</geoCoordinate>
]#74483 / bar code INB0003734727 INBIOCRI COSTA RICA /
<typeStatus id="54B6883AEE033809BAB5F848F778F865" box="[1219,1323,1949,1975]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C0D4D1BEE033809BB59F848F2A7F809" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="indecisa" status="sp. nov.">
<emphasis id="B979EA8AEE033809BB59F848F2A7F809" italics="true" pageId="65" pageNumber="66">Catostola indecisa</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE033809BE89F814F311F809" bold="true" box="[255,322,1985,2011]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A24A57F1EE033809BE89F814F311F809" box="[255,322,1985,2011]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
<collectorName id="26F8534EEE033809BF3BF814F3E5F809" box="[333,438,1985,2011]" pageId="65" pageNumber="66">Camargo</collectorName>
,
<collectorName id="26F8534EEE033809BFB2F814F05BF809" box="[452,520,1985,2011]" pageId="65" pageNumber="66">Vieira</collectorName>
&amp;
<collectorName id="26F8534EEE033809BC42F814F02EF809" box="[564,637,1985,2011]" pageId="65" pageNumber="66">Rafael</collectorName>
(
<figureCitation id="13362A1DEE033809BCE6F814F0AAF809" box="[656,761,1985,2011]" captionStart="FIGURE 38" captionStartId="60.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-21@60.[203,1385,181,1919]" captionTargetPageId="60" captionText="FIGURE 38. Catostola indecisa sp. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907408" httpUri="https://zenodo.org/record/7907408/files/figure.png" pageId="65" pageNumber="66">Fig. 38A</figureCitation>
[insert])
<emphasis id="B979EA8AEE033809BD2BF817F604F809" bold="true" box="[861,1111,1985,2014]" pageId="65" pageNumber="66">
(♁
<collectionCode id="ED1CAE5DEE033809BDF0F814F18EF809" box="[902,989,1985,2011]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="65" pageNumber="66">MNCR</collectionCode>
(
<collectionCode id="ED1CAE5DEE033809BD99F814F617F809" box="[1007,1092,1985,2011]" pageId="65" pageNumber="66">INBIO</collectionCode>
))
</emphasis>
</materialsCitation>
.
<emphasis id="B979EA8AEE033809BA1EF814F6B4F809" bold="true" box="[1128,1255,1985,2011]" pageId="65" pageNumber="66">
<typeStatus id="54B6883AEE033809BA1EF814F6B1F809" box="[1128,1250,1985,2011]" pageId="65" pageNumber="66" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Amubri,
<quantity id="4CF59B7DEE033809BB2FF814F7C4F80E" box="[1369,1431,1985,2012]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="65" pageNumber="66" unit="m" value="70.0">70 m</quantity>
<materialsCitation id="3B653CC5EE03380ABBEEF817F330FF2A" collectingDate="1993-06-06" collectingDateMax="1993-06-11" collectingDateMin="1993-06-06" collectionCode="INBIO, MNCR" collectorName="Gallardo. &amp; Lambert Sur" country="Costa Rica" lastPageId="66" lastPageNumber="67" latitude="9.517777" location="Talamanca" longLatPrecision="21" longitude="-82.953606" municipality="Prov." pageId="65" pageNumber="66" specimenCode="CRI001" specimenCount="1" specimenCount-female="1" stateProvince="Limon">
<location id="8ED26043EE03380ABBEEF817F340FF63" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE03380ABBEEF817F340FF63" country="Costa Rica" lastPageId="66" lastPageNumber="67" latitude="9.517777" longLatPrecision="21" longitude="-82.953606" municipality="Prov." name="Talamanca" pageId="65" pageNumber="66" stateProvince="Limon">,Talamanca</location>
,
<collectingMunicipality id="6BD6ACE2EE00380ABF69FF42F30FFF63" box="[287,348,151,177]" pageId="66" pageNumber="67">Prov.</collectingMunicipality>
[incia]
<emphasis id="B979EA8AEE00380ABFDCFF43F3AAFF63" bold="true" box="[426,505,150,177]" pageId="66" pageNumber="67">
<collectingRegion id="49C9F87AEE00380ABFDCFF43F3AAFF63" box="[426,505,150,177]" country="Costa Rica" name="Limon" pageId="66" pageNumber="67">Limón</collectingRegion>
</emphasis>
,
<emphasis id="B979EA8AEE00380ABC7EFF43F0E7FF62" bold="true" box="[520,692,150,176]" pageId="66" pageNumber="67">
<collectingCountry id="F31A7608EE00380ABC7EFF43F0E7FF62" box="[520,692,150,176]" name="Costa Rica" pageId="66" pageNumber="67">COSTA RICA</collectingCountry>
</emphasis>
.
<date id="FFB31058EE00380ABCB5FF42F138FF63" box="[707,875,151,177]" pageId="66" pageNumber="67" value="1993-06-06" valueMax="1993-06-11" valueMin="1993-06-06">
<collectingDate id="EFF7E9B0EE00380ABCB5FF42F138FF63" box="[707,875,151,177]" pageId="66" pageNumber="67" value="1993-06-06" valueMax="1993-06-11" valueMin="1993-06-06">611 Jun 1993</collectingDate>
</date>
, G.
<collectorName id="26F8534EEE00380ABDECFF42F650FF63" box="[922,1027,151,177]" pageId="66" pageNumber="67">Gallardo.</collectorName>
L-S-[
<collectorName id="26F8534EEE00380ABA35FF42F684FF63" box="[1091,1239,151,177]" pageId="66" pageNumber="67">Lambert Sur</collectorName>
] 385500, 578050 [
<geoCoordinate id="EE39505FEE00380ABEE8FF69F375FF05" box="[158,294,188,215]" degrees="09" direction="north" minutes="31" orientation="latitude" pageId="66" pageNumber="67" precision="15" seconds="04" value="9.517777">09°3104″N</geoCoordinate>
<geoCoordinate id="EE39505FEE00380ABF59FF69F3EDFF05" box="[303,446,188,215]" degrees="82" direction="west" minutes="57" orientation="longitude" pageId="66" pageNumber="67" precision="15" seconds="13" value="-82.953606">82°5713″W</geoCoordinate>
] / bar code
<collectingCountry id="F31A7608EE00380ABC3DFF6EF0BDFF07" box="[587,750,187,213]" name="Costa Rica" pageId="66" pageNumber="67">COSTA RICA</collectingCountry>
<collectionCode id="ED1CAE5DEE00380ABC8EFF6EF115FF07" box="[760,838,187,213]" pageId="66" pageNumber="67">INBIO</collectionCode>
<specimenCode id="DBAB9EE3EE00380ABD26FF6EF1FAFF07" box="[848,937,187,213]" collectionCode="INBIO" pageId="66" pageNumber="67">CRI001</specimenCode>
174886 /
<taxonomicName id="4C0D4D1BEE00380ABA57FF6EF6EAFF07" authorityName=", Enderlein" authorityYear="1914" box="[1057,1209,187,213]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B979EA8AEE00380ABA57FF6EF6EAFF07" box="[1057,1209,187,213]" italics="true" pageId="66" pageNumber="67">Ctenodontina</emphasis>
</taxonomicName>
det. EMFisher
<emphasis id="B979EA8AEE00380ABB06FF69F330FF2A" bold="true" pageId="66" pageNumber="67">
(
<specimenCount id="9D0BFD11EE00380ABB0CFF69F7CEFF0A" box="[1402,1437,188,216]" pageId="66" pageNumber="67" type="female">1♀</specimenCount>
<collectionCode id="ED1CAE5DEE00380ABEE1FF0BF2BDFF2A" box="[151,238,222,248]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="66" pageNumber="67">MNCR</collectionCode>
(
<collectionCode id="ED1CAE5DEE00380ABE8AFF0BF302FF2A" box="[252,337,222,248]" pageId="66" pageNumber="67">INBIO</collectionCode>
))
</emphasis>
</materialsCitation>
;
<materialsCitation id="3B653CC5EE00380ABF07FF0AF3C5FE92" collectingDate="1993-12-27" collectionCode="INBIO, MNCR" collectorName="Zumbado" country="Costa Rica" county="Punta" elevation="30" latitude="8.618889" location="Rio Rincon" longLatPrecision="21" longitude="-83.48555" municipality="Prov." pageId="66" pageNumber="67" specimenCode="CRI001" specimenCount="1" specimenCount-female="1">
<location id="8ED26043EE00380ABF07FF0AF3A0FF2B" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE00380ABF07FF0AF3A0FF2B" box="[369,499,223,249]" country="Costa Rica" county="Punta" latitude="8.618889" longLatPrecision="21" longitude="-83.48555" municipality="Prov." name="Rio Rincon" pageId="66" pageNumber="67">Río Rincón</location>
,
<location id="8ED26043EE00380ABF8BFF0AF07CFF2B" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE00380ABF8BFF0AF07CFF2B" box="[509,559,223,249]" country="Costa Rica" county="Punta" latitude="8.618889" longLatPrecision="21" longitude="-83.48555" municipality="Prov." name="Pen." pageId="66" pageNumber="67">Pen.</location>
[ínsula]
<location id="8ED26043EE00380ABCFCFF0AF0EBFF2B" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE00380ABCFCFF0AF0EBFF2B" box="[650,696,223,249]" country="Costa Rica" county="Punta" latitude="8.618889" longLatPrecision="21" longitude="-83.48555" municipality="Prov." name="Osa" pageId="66" pageNumber="67">Osa</location>
,
<collectingMunicipality id="6BD6ACE2EE00380ABCB4FF0AF0ACFF2B" box="[706,767,223,249]" pageId="66" pageNumber="67">Prov.</collectingMunicipality>
[incia]
<emphasis id="B979EA8AEE00380ABD3AFF0AF1B4FF2B" bold="true" box="[844,999,223,249]" pageId="66" pageNumber="67">
<collectingCounty id="62D34E14EE00380ABD3AFF0AF1C4FF2B" box="[844,919,223,249]" pageId="66" pageNumber="67">Punta</collectingCounty>
[renas]
</emphasis>
,
<collectingCountry id="F31A7608EE00380ABD82FF0AF6CBFF2B" box="[1012,1176,223,249]" name="Costa Rica" pageId="66" pageNumber="67">COSTA RICA</collectingCountry>
,
<quantity id="4CF59B7DEE00380ABAD4FF0AF68FFF2B" box="[1186,1244,223,249]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="66" pageNumber="67" unit="m" value="30.0">
<elevation id="0020D1ABEE00380ABAD4FF0AF68FFF2B" box="[1186,1244,223,249]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="66" pageNumber="67" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<date id="FFB31058EE00380ABA91FF0AF721FF2B" box="[1255,1394,223,249]" pageId="66" pageNumber="67" value="1993-12-27">
<collectingDate id="EFF7E9B0EE00380ABA91FF0AF721FF2B" box="[1255,1394,223,249]" pageId="66" pageNumber="67" value="1993-12-27">27 Dic 1993</collectingDate>
</date>
, M.
<collectorName id="26F8534EEE00380ABEE1FED6F356FECF" box="[151,261,259,285]" pageId="66" pageNumber="67">Zumbado</collectorName>
, L S 519700_286000 [
<geoCoordinate id="EE39505FEE00380ABF89FED1F0D4FECD" box="[511,647,260,287]" degrees="08" direction="north" minutes="37" orientation="latitude" pageId="66" pageNumber="67" precision="15" seconds="08" value="8.618889">08°3708″N</geoCoordinate>
<geoCoordinate id="EE39505FEE00380ABCF8FED1F14FFECD" box="[654,796,260,287]" degrees="83" direction="west" minutes="29" orientation="longitude" pageId="66" pageNumber="67" precision="15" seconds="08" value="-83.48555">83°2908″W</geoCoordinate>
] # 2667 / bar code
<collectingCountry id="F31A7608EE00380ABD99FED6F6DDFECF" box="[1007,1166,259,285]" name="Costa Rica" pageId="66" pageNumber="67">COSTA RICA</collectingCountry>
<collectionCode id="ED1CAE5DEE00380ABAE3FED6F6B0FECF" box="[1173,1251,259,285]" pageId="66" pageNumber="67">INBIO</collectionCode>
<specimenCode id="DBAB9EE3EE00380ABA9FFED6F711FECF" box="[1257,1346,259,285]" collectionCode="INBIO" pageId="66" pageNumber="67">CRI001</specimenCode>
725936
<emphasis id="B979EA8AEE00380ABEE1FEFDF3C5FE92" bold="true" box="[151,406,294,324]" pageId="66" pageNumber="67">
(
<specimenCount id="9D0BFD11EE00380ABED7FEFDF297FE96" box="[161,196,296,324]" pageId="66" pageNumber="67" type="female">1♀</specimenCount>
<collectionCode id="ED1CAE5DEE00380ABEBCFEF3F372FE92" box="[202,289,294,320]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="66" pageNumber="67">MNCR</collectionCode>
(
<collectionCode id="ED1CAE5DEE00380ABF58FEF3F3D0FE92" box="[302,387,294,320]" pageId="66" pageNumber="67">INBIO</collectionCode>
))
</emphasis>
</materialsCitation>
;
<materialsCitation id="3B653CC5EE00380ABFD5FEF2F6D0FE5A" collectingDate="1990-10" collectionCode="INBIO, MNCR" collectorName="E. M. Fisher" country="Costa Rica" county="Punt." elevation="50" latitude="8.478611" location="Est." longLatPrecision="21" longitude="-83.58917" municipality="Prov." pageId="66" pageNumber="67" specimenCount="1">
<location id="8ED26043EE00380ABFD5FEF2F39DFE93" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE00380ABFD5FEF2F39DFE93" box="[419,462,295,321]" country="Costa Rica" county="Punt." latitude="8.478611" longLatPrecision="21" longitude="-83.58917" municipality="Prov." name="Est." pageId="66" pageNumber="67">Est.</location>
[ación]
<collectorName id="26F8534EEE00380ABC55FEF2F0F7FE93" box="[547,676,295,321]" pageId="66" pageNumber="67">Sirena, P.N.</collectorName>
[Parque Nacional]
<location id="8ED26043EE00380ABD0EFEF2F1A5FE93" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE00380ABD0EFEF2F1A5FE93" box="[888,1014,295,321]" country="Costa Rica" county="Punt." latitude="8.478611" longLatPrecision="21" longitude="-83.58917" municipality="Prov." name="Corcovado" pageId="66" pageNumber="67">Corcovado</location>
,
<quantity id="4CF59B7DEE00380ABA76FEF2F633FE93" box="[1024,1120,295,321]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="66" pageNumber="67" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
<elevation id="0020D1ABEE00380ABA76FEF2F633FE93" box="[1024,1120,295,321]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="66" pageNumber="67" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0-100 m</elevation>
</quantity>
.
<collectingMunicipality id="6BD6ACE2EE00380ABA1CFEF2F6F4FE93" box="[1130,1191,295,321]" pageId="66" pageNumber="67">Prov.</collectingMunicipality>
[incia]
<emphasis id="B979EA8AEE00380ABA85FEF2F7C6FE93" bold="true" box="[1267,1429,295,321]" pageId="66" pageNumber="67">
<collectingCounty id="62D34E14EE00380ABA85FEF2F766FE93" box="[1267,1333,295,321]" pageId="66" pageNumber="67">Punt.</collectingCounty>
[arenas]
</emphasis>
,
<collectingCountry id="F31A7608EE00380ABEE1FE9EF36CFEB7" box="[151,319,331,357]" name="Costa Rica" pageId="66" pageNumber="67">COSTA RICA</collectingCountry>
.
<collectorName id="26F8534EEE00380ABF3AFE9EF39AFEB7" box="[332,457,331,357]" pageId="66" pageNumber="67">C. Saborio</collectorName>
,
<date id="FFB31058EE00380ABFA0FE9EF010FEB7" box="[470,579,331,357]" pageId="66" pageNumber="67" value="1990-10">
<collectingDate id="EFF7E9B0EE00380ABFA0FE9EF010FEB7" box="[470,579,331,357]" pageId="66" pageNumber="67" value="1990-10">Oct 1990</collectingDate>
</date>
, L-S- 270500-508300 [
<geoCoordinate id="EE39505FEE00380ABD38FE99F185FEB5" box="[846,982,332,359]" degrees="08" direction="north" minutes="28" orientation="latitude" pageId="66" pageNumber="67" precision="15" seconds="43" value="8.478611">08°2843″N</geoCoordinate>
<geoCoordinate id="EE39505FEE00380ABDA9FE99F63DFEB5" box="[991,1134,332,359]" degrees="83" direction="west" minutes="35" orientation="longitude" pageId="66" pageNumber="67" precision="15" seconds="21" value="-83.58917">83°3521″W</geoCoordinate>
] / bar code
<collectingCountry id="F31A7608EE00380ABA8FFE9EF7CFFEB7" box="[1273,1436,331,357]" name="Costa Rica" pageId="66" pageNumber="67">COSTA RICA</collectingCountry>
<collectionCode id="ED1CAE5DEE00380ABEE1FEBAF2B6FE5B" box="[151,229,367,393]" pageId="66" pageNumber="67">INBIO</collectionCode>
CRI000 062067 /
<taxonomicName id="4C0D4D1BEE00380ABFCAFEBAF0CBFE5B" box="[444,664,367,393]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B979EA8AEE00380ABFCAFEBAF007FE5B" box="[444,596,367,393]" italics="true" pageId="66" pageNumber="67">Ctenodontina</emphasis>
sp. A
</taxonomicName>
det
<collectorName id="26F8534EEE00380ABCBDFEBAF101FE5B" box="[715,850,367,393]" pageId="66" pageNumber="67">E.M. Fisher</collectorName>
04
<emphasis id="B979EA8AEE00380ABD09FEA5F6D0FE5A" bold="true" box="[895,1155,366,396]" pageId="66" pageNumber="67">
(1♁
<collectionCode id="ED1CAE5DEE00380ABDC2FEBBF658FE5A" box="[948,1035,366,392]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="66" pageNumber="67">MNCR</collectionCode>
(
<collectionCode id="ED1CAE5DEE00380ABA6AFEBBF622FE5A" box="[1052,1137,366,392]" pageId="66" pageNumber="67">INBIO</collectionCode>
))
</emphasis>
</materialsCitation>
;
<materialsCitation id="3B653CC5EE00380ABAE5FEBAF116FE7E" collectingDate="1990-06" collectionCode="MNCR, INBIO" collectorName="N. Obando" country="Costa Rica" location="Same" pageId="66" pageNumber="67" specimenCount="1" specimenCount-female="1">
<location id="8ED26043EE00380ABAE5FEBAF682FE5B" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE00380ABAE5FEBAF682FE5B" box="[1171,1233,367,393]" country="Costa Rica" name="Same" pageId="66" pageNumber="67">Same</location>
data,
<collectorName id="26F8534EEE00380ABB6FFEBAF7CAFE5B" box="[1305,1433,367,393]" pageId="66" pageNumber="67">N. Obando</collectorName>
,
<date id="FFB31058EE00380ABEE1FE46F352FE7F" box="[151,257,403,429]" pageId="66" pageNumber="67" value="1990-06">
<collectingDate id="EFF7E9B0EE00380ABEE1FE46F352FE7F" box="[151,257,403,429]" pageId="66" pageNumber="67" value="1990-06">Jun 1990</collectingDate>
</date>
, / bar code CRI000 644140
<emphasis id="B979EA8AEE00380ABC35FE41F116FE7E" bold="true" box="[579,837,402,432]" pageId="66" pageNumber="67">
(
<specimenCount id="9D0BFD11EE00380ABC3AFE41F03CFE62" box="[588,623,404,432]" pageId="66" pageNumber="67" type="female">1♀</specimenCount>
<collectionCode id="ED1CAE5DEE00380ABC01FE47F09DFE7E" box="[631,718,402,428]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="66" pageNumber="67">MNCR</collectionCode>
(
<collectionCode id="ED1CAE5DEE00380ABCA8FE47F160FE7E" box="[734,819,402,428]" pageId="66" pageNumber="67">INBIO</collectionCode>
))
</emphasis>
</materialsCitation>
.
<materialsCitation id="3B653CC5EE00380ABD22FE47F3C1FE26" collectionCode="NHM" country="Costa Rica" latitude="9.789167" location="Jan." longLatPrecision="21" longitude="-84.126114" pageId="66" pageNumber="67" specimenCount="1" stateProvince="San Jose">
<emphasis id="B979EA8AEE00380ABD22FE47F614FE7E" bold="true" box="[852,1095,402,428]" pageId="66" pageNumber="67">Additional material.</emphasis>
<collectorName id="26F8534EEE00380ABA39FE46F755FE7F" box="[1103,1286,403,429]" pageId="66" pageNumber="67">Monte Redondo</collectorName>
[
<geoCoordinate id="EE39505FEE00380ABB62FE41F7CFFE7D" box="[1300,1436,404,431]" degrees="09" direction="north" minutes="47" orientation="latitude" pageId="66" pageNumber="67" precision="15" seconds="21" value="9.789167">09°4721″N</geoCoordinate>
<geoCoordinate id="EE39505FEE00380ABEE1FE6DF37BFE01" box="[151,296,440,467]" degrees="84" direction="west" minutes="07" orientation="longitude" pageId="66" pageNumber="67" precision="15" seconds="34" value="-84.126114">84°0734″W</geoCoordinate>
],
<location id="8ED26043EE00380ABF4FFE62F335FE03" LSID="urn:lsid:plazi:treatment:03A4878EEE78380ABEE1FAFAF3CAFE27:8ED26043EE00380ABF4FFE62F335FE03" box="[313,358,439,465]" country="Costa Rica" latitude="9.789167" longLatPrecision="21" longitude="-84.126114" name="Jan." pageId="66" pageNumber="67" stateProvince="San Jose">Jan.</location>
03 /
<emphasis id="B979EA8AEE00380ABFE0FE63F012FE02" bold="true" box="[406,577,438,464]" pageId="66" pageNumber="67">
<collectingCountry id="F31A7608EE00380ABFE0FE63F012FE02" box="[406,577,438,464]" name="Costa Rica" pageId="66" pageNumber="67">COSTA RICA</collectingCountry>
</emphasis>
[
<emphasis id="B979EA8AEE00380ABC26FE63F0E5FE03" bold="true" box="[592,694,438,465]" pageId="66" pageNumber="67">
<collectingRegion id="49C9F87AEE00380ABC26FE63F0E5FE03" box="[592,694,438,465]" country="Costa Rica" name="San Jose" pageId="66" pageNumber="67">San Jose</collectingRegion>
</emphasis>
],
<collectorName id="26F8534EEE00380ABCBBFE62F641FE03" box="[717,1042,439,465]" pageId="66" pageNumber="67">C.F. Underwood. Per Janson.</collectorName>
/
<collectorName id="26F8534EEE00380ABA50FE62F70EFE03" box="[1062,1373,439,465]" pageId="66" pageNumber="67">C.J. Wainwright Collection.</collectorName>
B.M. 1948-488.
<emphasis id="B979EA8AEE00380ABF67FE09F3C1FE26" bold="true" box="[273,402,474,504]" pageId="66" pageNumber="67">
(1♁
<collectionCode id="ED1CAE5DEE00380ABF32FE0FF3D9FE26" box="[324,394,474,500]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="66" pageNumber="67" type="Herbarium">NHM</collectionCode>
)
</emphasis>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>