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<document id="7137872BF27E14C143722FAD9C66C85E" ID-CLB-Dataset="36817" ID-DOI="10.11646/zootaxa.3835.1.1" ID-GBIF-Dataset="7d11dc81-2b61-40fb-8eb1-fcdfb1547668" ID-ISSN="1175-5326" ID-Zenodo-Dep="286701" ID-ZooBank="5BBF2D80-F682-49E7-9B32-64E75ACAA6CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460455174602" checkinUser="plazi" docAuthor="Sartori, Michel" docDate="2014" docId="03A087A0432B4A05FF7AE8A290875C07" docLanguage="en" docName="zt03835p032.pdf" docOrigin="Zootaxa 3835 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Compsoneuriella langensis Braasch &amp; Boonsoong 2010, comb. nov." docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="FF99FFD843314A1FFFEDE915942F5F7D" masterDocTitle="The concept of Compsoneuria Eaton, 1881 revisited in light of historical and new material from the Sunda Islands (Ephemeroptera: Heptageniidae: Ecdyonurinae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="27" updateTime="1698338546610" updateUser="ExternalLinkService">
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<mods:title id="72509C765A0A38F5A931F71E343B8F35">The concept of Compsoneuria Eaton, 1881 revisited in light of historical and new material from the Sunda Islands (Ephemeroptera: Heptageniidae: Ecdyonurinae)</mods:title>
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<mods:namePart id="6F5CBDB02DD71F9CD93609F193CEF6EB">Sartori, Michel</mods:namePart>
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<mods:date id="9E3E760EAD313FAF3EDF66BB547935BC">2014</mods:date>
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<treatment id="03A087A0432B4A05FF7AE8A290875C07" ID-DOI="http://doi.org/10.5281/zenodo.6124753" ID-GBIF-Taxon="127668973" ID-Zenodo-Dep="6124753" LSID="urn:lsid:plazi:treatment:03A087A0432B4A05FF7AE8A290875C07" httpUri="http://treatment.plazi.org/id/03A087A0432B4A05FF7AE8A290875C07" lastPageNumber="27" pageId="26" pageNumber="27">
<subSubSection id="C313653D432B4A05FF7AE8A297C85EAF" box="[151,999,439,466]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BB636B6432B4A05FF7AE8A297C85EAF" blockId="26.[151,999,439,466]" box="[151,999,439,466]" pageId="26" pageNumber="27">
<heading id="D0FE81DA432B4A05FF7AE8A297C85EAF" bold="true" box="[151,999,439,466]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B97DEAA4432B4A05FF7AE8A297C85EAF" bold="true" box="[151,999,439,466]" pageId="26" pageNumber="27">
<taxonomicName id="4C094D35432B4A05FF7AE8A297765EAF" authority="Braasch &amp; Boonsoong, 2010" authorityName="Braasch &amp; Boonsoong" authorityYear="2010" box="[151,857,439,466]" class="Insecta" family="Heptageniidae" genus="Compsoneuriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="langensis" status="comb. nov.">
<emphasis id="B97DEAA4432B4A05FF7AE8A295F05EAC" bold="true" box="[151,479,439,465]" italics="true" pageId="26" pageNumber="27">Compsoneuriella langensis</emphasis>
(
<bibRefCitation id="EF984B47432B4A05FE03E8AD977E5EAF" author="Braasch, D. &amp; Boonsoong, B." box="[494,849,440,466]" firstAuthor="Braasch" journalOrPublisher="Zootaxa" pageId="26" pageNumber="27" pagination="1 - 26" part="2610" refId="ref15537" refString="Braasch, D. &amp; Boonsoong, B. (2010) A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia. Zootaxa, 2610, 1 - 26." title="A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia" type="journal article" year="2010">Braasch &amp; Boonsoong, 2010</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A24E57DF432B4A05FC8CE8AD97C85EAF" box="[865,999,440,466]" pageId="26" pageNumber="27">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C313653D432B4A05FF7AE8EA900B5D69" box="[151,1060,510,532]" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="8BB636B6432B4A05FF7AE8EA900B5D69" blockId="26.[151,1060,510,532]" box="[151,1060,510,532]" pageId="26" pageNumber="27">
<taxonomicName id="4C094D35432B4A05FF7AE8EA96905D69" ID-CoL="5ZS79" authority="Braasch &amp; Boonsoong, 2010" authorityName="Braasch &amp; Boonsoong" authorityYear="2010" box="[151,703,510,532]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="langensis">
<emphasis id="B97DEAA4432B4A05FF7AE8EA95BD5D69" box="[151,402,510,532]" italics="true" pageId="26" pageNumber="27">Compsoneuria langensis</emphasis>
<bibRefCitation id="EF984B47432B4A05FE75E8EB96905D69" author="Braasch, D. &amp; Boonsoong, B." box="[408,703,510,532]" firstAuthor="Braasch" journalOrPublisher="Zootaxa" pageId="26" pageNumber="27" pagination="1 - 26" part="2610" refId="ref15537" refString="Braasch, D. &amp; Boonsoong, B. (2010) A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia. Zootaxa, 2610, 1 - 26." title="A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia" type="journal article" year="2010">Braasch &amp; Boonsoong, 2010</bibRefCitation>
</taxonomicName>
, Zootaxa 2610, p. 7 (adult, nymph)
</paragraph>
</subSubSection>
<subSubSection id="C313653D432B4A05FF7AEB5497585D00" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BB636B6432B4A05FF7AEB5497585D00" blockId="26.[151,1436,577,890]" pageId="26" pageNumber="27">
<emphasis id="B97DEAA4432B4A05FF7AEB5495AA5D27" bold="true" box="[151,389,577,602]" pageId="26" pageNumber="27">Material examined.</emphasis>
<materialsCitation id="3B613CEB432B4A05FE7CEB5497585D00" ID-GBIF-Occurrence="4037950307" collectingDate="2003-04-03" collectingDateMax="2003-04-25" collectingDateMin="2003-04-03" collectionCode="MZL" collectorName="D. Braasch" country="Thailand" elevation="605" latitude="19.574116" location="Soppong" longLatPrecision="1" longitude="98.278786" municipality="Mae Hong Son Province" pageId="26" pageNumber="27" specimenCount="2">
<specimenCount id="9D0FFD3F432B4A05FE7CEB5496285D27" box="[401,519,577,602]" pageId="26" pageNumber="27">2 nymphs</specimenCount>
,
<collectingCountry id="F31E7626432B4A05FDFBEB5496535D27" box="[534,636,577,602]" name="Thailand" pageId="26" pageNumber="27">Thailand</collectingCountry>
,
<collectingMunicipality id="6BD2ACCC432B4A05FD61EB57979D5D27" box="[652,946,577,602]" pageId="26" pageNumber="27">Mae Hong Son Province</collectingMunicipality>
,
<location id="8ED6606D432B4A05FC2FEB5490075D24" LSID="urn:lsid:plazi:treatment:03A087A0432B4A05FF7AE8A290875C07:8ED6606D432B4A05FC2FEB5490075D24" box="[962,1064,577,601]" country="Thailand" latitude="19.574116" longLatPrecision="1" longitude="98.278786" municipality="Mae Hong Son Province" name="Soppong" pageId="26" pageNumber="27">Soppong</location>
,
<location id="8ED6606D432B4A05FBD5EB57912F5D27" LSID="urn:lsid:plazi:treatment:03A087A0432B4A05FF7AE8A290875C07:8ED6606D432B4A05FBD5EB57912F5D27" box="[1080,1280,577,602]" country="Thailand" latitude="19.574116" longLatPrecision="1" longitude="98.278786" municipality="Mae Hong Son Province" name="Nam Lang River" pageId="26" pageNumber="27">Nam Lang River</location>
,
<quantity id="4CF19B53432B4A05FAFDEB54914F5D24" box="[1296,1376,577,602]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="26" pageNumber="27" unit="m" value="605.0">
<elevation id="0024D185432B4A05FAFDEB54914F5D24" box="[1296,1376,577,602]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="26" pageNumber="27" unit="m" value="605.0">605 m</elevation>
</quantity>
.,
<geoCoordinate id="EE3D5071432B4A05FA98EB5495245D00" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="1" value="19.574116">19° 34.447 N</geoCoordinate>
,
<geoCoordinate id="EE3D5071432B4A05FEFBEB73959A5D00" box="[278,437,613,638]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="1" value="98.278786">98° 16.727 E</geoCoordinate>
,
<date id="FFB71076432B4A05FE2CEB7096455D03" box="[449,618,613,638]" pageId="26" pageNumber="27" value="2003-04-03" valueMax="2003-04-25" valueMin="2003-04-03">
<collectingDate id="EFF3E99E432B4A05FE2CEB7096455D03" box="[449,618,613,638]" pageId="26" pageNumber="27" value="2003-04-03" valueMax="2003-04-25" valueMin="2003-04-03">325. IV. 2003</collectingDate>
</date>
,
<collectorName id="26FC5360432B4A05FD98EB7396DE5D03" box="[629,753,613,638]" pageId="26" pageNumber="27">D. Braasch</collectorName>
leg. [
<collectionCode id="ED18AE73432B4A05FCC2EB73975F5D00" box="[815,880,614,637]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="26" pageNumber="27">MZL</collectionCode>
]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C313653D432B4A05FF2AEB9C90875C07" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BB636B6432B4A05FF2AEB9C90875C07" blockId="26.[151,1436,577,890]" pageId="26" pageNumber="27">
<emphasis id="B97DEAA4432B4A05FF2AEB9C95635DDF" bold="true" box="[199,332,649,674]" pageId="26" pageNumber="27">Discussion.</emphasis>
The species has been adequately described by
<bibRefCitation id="EF984B47432B4A05FCB3EB9C909E5DDF" author="Braasch, D. &amp; Boonsoong, B." box="[862,1201,649,674]" firstAuthor="Braasch" journalOrPublisher="Zootaxa" pageId="26" pageNumber="27" pagination="1 - 26" part="2610" refId="ref15537" refString="Braasch, D. &amp; Boonsoong, B. (2010) A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia. Zootaxa, 2610, 1 - 26." title="A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia" type="journal article" year="2010">Braasch &amp; Boonsoong (2010)</bibRefCitation>
, and differs from the two previous ones by its labrum less expended laterally (
<figureCitation id="13322A33432B4A05FCCCEBB897595DBB" box="[801,886,685,710]" captionStart="FIGURES 68 77" captionStartId="24.[151,264,1889,1912]" captionTargetBox="[192,1377,230,1849]" captionTargetId="figure@24.[167,1403,193,1865]" captionTargetPageId="24" captionText="FIGURES 68 77. Compsoneuriella spp, nymphal structures. Figs 68 - 70: Compsoneuriella sp. 1 from Thailand. Fig. 68: Outline of the labium (left side paraglossa only drawn); Fig. 69: Bristles on the upper face of hind femora; Fig. 70: Posterior margin of tergite IV. Figs 71 - 73: Compsoneuriella langensis. Fig. 71: Outline of the labrum; Fig. 72: Outline of the glossae; Fig. 73: Outline of gill VII. Figs 74 - 77: Compsoneuriella tagbanua. Fig. 74: Outline of the labrum; Fig. 75: Outline of the glossae; Fig. 76: Detail of the left mandible; Fig. 77: Scattered setae on the ventral side of the galea." httpUri="https://zenodo.org/record/286715/files/figure.png" pageId="26" pageNumber="27">Fig. 71</figureCitation>
), by the shape of the glossae, with inner margin regularly convex and outer margin slightly concave (
<figureCitation id="13322A33432B4A05FD18EBC497655D97" box="[757,842,721,746]" captionStart="FIGURES 68 77" captionStartId="24.[151,264,1889,1912]" captionTargetBox="[192,1377,230,1849]" captionTargetId="figure@24.[167,1403,193,1865]" captionTargetPageId="24" captionText="FIGURES 68 77. Compsoneuriella spp, nymphal structures. Figs 68 - 70: Compsoneuriella sp. 1 from Thailand. Fig. 68: Outline of the labium (left side paraglossa only drawn); Fig. 69: Bristles on the upper face of hind femora; Fig. 70: Posterior margin of tergite IV. Figs 71 - 73: Compsoneuriella langensis. Fig. 71: Outline of the labrum; Fig. 72: Outline of the glossae; Fig. 73: Outline of gill VII. Figs 74 - 77: Compsoneuriella tagbanua. Fig. 74: Outline of the labrum; Fig. 75: Outline of the glossae; Fig. 76: Detail of the left mandible; Fig. 77: Scattered setae on the ventral side of the galea." httpUri="https://zenodo.org/record/286715/files/figure.png" pageId="26" pageNumber="27">Fig. 72</figureCitation>
), the inner margin of the fore femora with a row of spines and fine and long setae, the presence of microdenticles on the posterior margin of the abdominal tergites, and by the presence of 2 subapical teeth on the tarsal claw. Contrary to what has been previously illustrated (
<bibRefCitation id="EF984B47432B4A05FF73EA28966D5C2B" author="Braasch, D. &amp; Boonsoong, B." box="[158,578,829,854]" journalOrPublisher="Zootaxa" pageId="26" pageNumber="27" pagination="1 - 26" part="2610" refId="ref15537" refString="Braasch, D. &amp; Boonsoong, B. (2010) A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia. Zootaxa, 2610, 1 - 26." title="A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia" type="journal article" year="2010">
Braasch &amp; Boonsoong 2010:
<figureCitation id="13322A33432B4A05FE02EA28966D5C2B" box="[495,578,829,854]" captionStart="FIGURES 43 47" captionStartId="20.[151,264,1857,1880]" captionTargetBox="[212,1359,203,1824]" captionTargetId="figure@20.[195,1371,193,1837]" captionTargetPageId="20" captionText="FIGURES 43 47. Nymphal maxillae details of Compsoneuria spectabilis. Fig. 43: SEM view of the dentisetae, proximal dentiseta in red, distal dentisetae in yellow, with first one fimbriate; Fig 44: Same view in optical microscope; Fig. 45: SEM view of fimbriate scattered setae on the ventral face of the galea; Fig. 46: Same view in optical microscope; Fig. 47: SEM view of the comb-shape setae on the crown of the galea; abbreviations: dp: proximal dentiseta, dd 1, first distal dentiseta, dd 2 second distal dentiseta." httpUri="https://zenodo.org/record/286711/files/figure.png" pageId="26" pageNumber="27">Fig. 45</figureCitation>
</bibRefCitation>
), gill VII is not obtusely rounded but is apically narrowly pointed (
<figureCitation id="13322A33432B4A05FAD6EA2891A05C2B" box="[1339,1423,829,854]" captionStart="FIGURES 68 77" captionStartId="24.[151,264,1889,1912]" captionTargetBox="[192,1377,230,1849]" captionTargetId="figure@24.[167,1403,193,1865]" captionTargetPageId="24" captionText="FIGURES 68 77. Compsoneuriella spp, nymphal structures. Figs 68 - 70: Compsoneuriella sp. 1 from Thailand. Fig. 68: Outline of the labium (left side paraglossa only drawn); Fig. 69: Bristles on the upper face of hind femora; Fig. 70: Posterior margin of tergite IV. Figs 71 - 73: Compsoneuriella langensis. Fig. 71: Outline of the labrum; Fig. 72: Outline of the glossae; Fig. 73: Outline of gill VII. Figs 74 - 77: Compsoneuriella tagbanua. Fig. 74: Outline of the labrum; Fig. 75: Outline of the glossae; Fig. 76: Detail of the left mandible; Fig. 77: Scattered setae on the ventral side of the galea." httpUri="https://zenodo.org/record/286715/files/figure.png" pageId="26" pageNumber="27">Fig. 73</figureCitation>
). Finally the overall body coloration and pattern is not as contrasted as in the previous species.
</paragraph>
</subSubSection>
</treatment>
</document>