plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/9A/87/039A87813E40F05E9BC5F896C6FB4828.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

394 lines
48 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="98A7268283D89F04023434E25ECA713A" ID-DOI="10.5281/zenodo.3710074" ID-GBIF-Dataset="f1bbeff3-8e8b-408c-a46c-11595dcf1560" ID-ISSN="1942-1354" ID-Zenodo-Dep="3710074" ID-ZooBank="3EB9D858-BE71-4F1F-BEBD-136898474B51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584124742059" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2018" docId="039A87813E40F05E9BC5F896C6FB4828" docLanguage="en" docName="InsectaMundi.0666.1-10.pdf.imf" docOrigin="Insecta Mundi 666" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Echinoaesalus Zelenka 1993" docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="FFA3FFF93E43F05B9B05F91BC4084D26" masterDocTitle="Generic changes in the stag beetle tribe Aesalini (Coleoptera: Lucanidae: Aesalinae) with the description of two new species" masterLastPageNumber="10" masterPageNumber="1" pageNumber="2" updateTime="1698728097052" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="8A169995ADAAF9F96A4CA9BB95E39F1B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7F16C863D1A0D1815ABE74F0584DBA40">
<mods:title id="ED7AC431202734D5ED87A60AA2A2D2A6">Generic changes in the stag beetle tribe Aesalini (Coleoptera: Lucanidae: Aesalinae) with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="0D5DE5390706400F808F4A510C4E4BD5" type="personal">
<mods:role id="EED8509B2A4FCB1FCF3A69ABABB76688">
<mods:roleTerm id="FE1A85274BF11452F05EFBD1D6523113">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15BFC1965CDD1EE22D2FF0A46D436C27">Paulsen, M. J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="92A5091697F35A3AECE93CA1205F51B7">text</mods:typeOfResource>
<mods:relatedItem id="578E8554943D4AE9E04338B4B2AA541F" type="host">
<mods:titleInfo id="5256075DB12E5A702EF68AEDD4B1DBD5">
<mods:title id="DF8D72F71CD8A756A8E03BDA9A50C239">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="5CB4F25CF95AAB4BE38F928BEAC8C856">
<mods:date id="547AEA74D400D12FBC37CF778B3390A8">2018</mods:date>
<mods:detail id="F3A4C9EEB5CEA145ED6FE4D48CC3B334" type="pubDate">
<mods:number id="E092F7E0926BFDF94724D918F94E5B8A">2018-10-31</mods:number>
</mods:detail>
<mods:detail id="228358C7BFAD07F09EA796A49216807D" type="issue">
<mods:number id="81041B186D40A451BBCC89E955D92DAC">666</mods:number>
</mods:detail>
<mods:extent id="A738BEB1C91B4C7A5F7295764C8D871C" unit="page">
<mods:start id="61DC94D89AC1C291FEEF91C1C8592792">1</mods:start>
<mods:end id="823948AEA48E82A9F73FC4A7E4AC3A5E">10</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4B665A29077C4C6AC9E97ADC8D8739C5">journal article</mods:classification>
<mods:identifier id="D287D8043B2DED656CF333E2C25A8B47" type="DOI">10.5281/zenodo.3710074</mods:identifier>
<mods:identifier id="65FC0F42E91B8897D66D9F3319064B63" type="GBIF-Dataset">f1bbeff3-8e8b-408c-a46c-11595dcf1560</mods:identifier>
<mods:identifier id="E29759241F48F3BFB356FFACBEE47AA0" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="D954AABDDBC3677FA8EF687B56EE65F2" type="Zenodo-Dep">3710074</mods:identifier>
<mods:identifier id="8B370B78E7E1AB7E96BDAD0F1D5694CF" type="ZooBank">3EB9D858-BE71-4F1F-BEBD-136898474B51</mods:identifier>
</mods:mods>
<treatment id="039A87813E40F05E9BC5F896C6FB4828" ID-DOI="http://doi.org/10.5281/zenodo.3716619" ID-GBIF-Taxon="162488155" ID-Zenodo-Dep="3716619" LSID="urn:lsid:plazi:treatment:039A87813E40F05E9BC5F896C6FB4828" httpUri="http://treatment.plazi.org/id/039A87813E40F05E9BC5F896C6FB4828" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
<subSubSection id="C329651C3E40F0589BC5F896C6404C83" box="[192,584,397,421]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8B8C36973E40F0589BC5F896C6404C83" blockId="3.[192,584,397,421]" box="[192,584,397,421]" pageId="3" pageNumber="2">
<heading id="D0C481FB3E40F0589BC5F896C6404C83" bold="true" box="[192,584,397,421]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<taxonomicName id="4C334D143E40F0589BC5F896C6404C83" ID-CoL="632YD" authority="Zelenka, 1993" authorityName="Zelenka" authorityYear="1993" box="[192,584,397,421]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589BC5F896C6404C83" bold="true" box="[192,584,397,421]" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F0589BC5F896C58F4C83" bold="true" box="[192,391,397,421]" italics="true" pageId="3" pageNumber="2">Echinoaesalus</emphasis>
<bibRefCitation id="EFA24B663E40F0589A8AF896C6404C83" author="Zelenka, W." box="[399,584,397,421]" pageId="3" pageNumber="2" pagination="235 - 237" refId="ref5647" refString="Zelenka, W. 1993. Echinoaesalus gen. n. eine neue lucaniden-Gattung aus Sudostasien (Coleoptera, Lucanidae). Koleopterologische Rundschau 63: 235 - 237." type="journal article" year="1993">Zelenka, 1993</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C329651C3E40F0589BC5F8D6C6944CC3" box="[192,668,461,485]" pageId="3" pageNumber="2" type="type_taxon">
<paragraph id="8B8C36973E40F0589BC5F8D6C6944CC3" blockId="3.[192,668,461,485]" box="[192,668,461,485]" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F0589BC5F8D6C57F4CC3" bold="true" box="[192,375,461,485]" pageId="3" pageNumber="2">
<typeStatus id="548888353E40F0589BC5F8D6C50C4CC3" box="[192,260,461,485]" pageId="3" pageNumber="2">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C334D143E40F0589A7BF8D6C6944CC3" authority="Zelenka." box="[382,668,461,485]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="B947EA853E40F0589A7BF8D6C6244CC3" box="[382,556,461,485]" italics="true" pageId="3" pageNumber="2">Aesalus jaechi</emphasis>
Zelenka.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C329651C3E40F05E9BEDF8E6C6FB4828" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="discussion">
<paragraph id="8B8C36973E40F0589BEDF8E6C67F4913" blockId="3.[192,1441,509,1270]" pageId="3" pageNumber="2">
The first aesaline from Southeast Asia was described by
<bibRefCitation id="EFA24B663E40F05898A4F8E6C0574F33" author="Krikken, J." box="[929,1119,509,533]" pageId="3" pageNumber="2" pagination="49 - 51" refId="ref5532" refString="Krikken, J. 1975. Aesalus timidus sp. nov., a remarkable addition to the Indonesian stag-beetle fauna. Entomologische Berichten 35: 49 - 51." type="journal article" year="1975">Krikken (1975)</bibRefCitation>
. The years
<date id="FF8D10573E40F0589FE9F8E5C1A84F30" box="[1260,1440,509,534]" pageId="3" pageNumber="2" value="1993" valueMax="1994">1993 and 1994</date>
were extremely active for species for the region, as Araya (1993),
<bibRefCitation id="EFA24B663E40F05898E0FB05C0C54F10" author="Araya, K. &amp; M. Kon &amp; Y. Johki" box="[997,1229,541,566]" pageId="3" pageNumber="2" pagination="93 - 107" refId="ref5223" refString="Araya, K., M. Kon, and Y. Johki. 1993. A new species of the genus Aesalus (Coleoptera, Lucanidae) from the Malay Peninsula. Elytra 21: 93 - 107." type="journal article" year="1993">Araya et al. (1993)</bibRefCitation>
, and
<bibRefCitation id="EFA24B663E40F0589E0AFB05C5044F70" author="Araya, K. &amp; M. Matsui &amp; J. Nabhitabhata &amp; S. Panha" pageId="3" pageNumber="2" pagination="73 - 77" refId="ref5263" refString="Araya, K., M. Matsui, J. Nabhitabhata, and S. Panha. 1994. A new bristly Aesalus from peninsular Thailand (Coleoptera, Lucanidae). Giornale Italiano di Entomologia 7 (36): 73 - 77." type="journal article" year="1994">Araya et al. (1994)</bibRefCitation>
named new species. At the same time,
<bibRefCitation id="EFA24B663E40F05899F8FB26C7BC4F70" author="Zelenka, W." box="[765,948,573,598]" pageId="3" pageNumber="2" pagination="235 - 237" refId="ref5647" refString="Zelenka, W. 1993. Echinoaesalus gen. n. eine neue lucaniden-Gattung aus Sudostasien (Coleoptera, Lucanidae). Koleopterologische Rundschau 63: 235 - 237." type="journal article" year="1993">Zelenka (1993)</bibRefCitation>
created the subgenus
<taxonomicName id="4C334D143E40F0589FCAFB26C1774F73" box="[1231,1407,573,597]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589FCAFB26C1774F73" box="[1231,1407,573,597]" italics="true" pageId="3" pageNumber="2">Echinoaesalus</emphasis>
</taxonomicName>
to accommodate three species of
<taxonomicName id="4C334D143E40F0589937FB46C6984F53" authorityName="Fabricius. Advances" authorityYear="1801" box="[562,656,605,629]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589937FB46C6984F53" box="[562,656,605,629]" italics="true" pageId="3" pageNumber="2">Aesalus</emphasis>
</taxonomicName>
from Southeast Asia, including two new species and
<taxonomicName id="4C334D143E40F0589E1AFB46C56C4FB3" authority="Krikken. The" authorityName="Krikken. The" authorityYear="2018" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="timidus" status="gen. nov.">
<emphasis id="B947EA853E40F0589E1AFB46C1A84F53" box="[1311,1440,605,629]" italics="true" pageId="3" pageNumber="2">A. timidus</emphasis>
Krikken. The
</taxonomicName>
<taxonomicNameLabel id="A27457FE3E40F0589A69FB65C5E74FB0" box="[364,495,638,662]" pageId="3" pageNumber="2" rank="genus">new genus</taxonomicNameLabel>
was differentiated from
<taxonomicName id="4C334D143E40F0589825FB66C7764FB3" authorityName="Fabricius. Advances" authorityYear="1801" box="[800,894,637,661]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589825FB66C7764FB3" box="[800,894,637,661]" italics="true" pageId="3" pageNumber="2">Aesalus</emphasis>
</taxonomicName>
based on the species’ convex form, protibiae with three teeth and lacking other minute serrations, middle and hind legs lacking strong external teeth, and lack of sexual dimorphism in the mandibles.
<bibRefCitation id="EFA24B663E40F0589877FBA6C0204FF0" author="Zelenka, W." box="[882,1064,701,726]" pageId="3" pageNumber="2" pagination="235 - 237" refId="ref5647" refString="Zelenka, W. 1993. Echinoaesalus gen. n. eine neue lucaniden-Gattung aus Sudostasien (Coleoptera, Lucanidae). Koleopterologische Rundschau 63: 235 - 237." type="journal article" year="1993">Zelenka (1993)</bibRefCitation>
also discussed the presence of weak depressions or complete sulci on the abdomen to receive the hind legs. However, two of the three species he included in
<taxonomicName id="4C334D143E40F0589ADEFBE6C6834E33" box="[475,651,765,789]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589ADEFBE6C6834E33" box="[475,651,765,789]" italics="true" pageId="3" pageNumber="2">Echinoaesalus</emphasis>
</taxonomicName>
, including the
<typeStatus id="548888353E40F058984FFBE5C7774E30" box="[842,895,766,790]" pageId="3" pageNumber="2">type</typeStatus>
species
<taxonomicName id="4C334D143E40F05898ECFBE6C0C84E33" authority="Zelenka" authorityName="Zelenka" box="[1001,1216,765,789]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="B947EA853E40F05898ECFBE6C05D4E33" box="[1001,1109,765,789]" italics="true" pageId="3" pageNumber="2">A. jaechi</emphasis>
Zelenka
</taxonomicName>
(
<figureCitation id="13082A123E40F0589FD7FBE6C1114E30" box="[1234,1305,765,790]" captionStart="Figures 1–2" captionStartId="4.[192,287,1819,1841]" captionTargetBox="[192,1407,276,1739]" captionTargetId="figure@4.[192,1440,261,1767]" captionTargetPageId="4" captionText="Figures 1–2. Zelenka holotypes. 1) Holotype of Echinoaesalus jaechi Zelenka. a) Dorsal habitus. b) Abdomen, ventral view. c) Labels. 2) Holotype female of Echinoaesalus sabahensis Zelenka. a) Dorsal habitus. b) Abdomen, ventral view, showing complete leg-shaped sulci. c) Labels. Photos courtesy H. Schillhammer (Naturhistorisches Museum, Vienna)." figureDoi="http://doi.org/10.5281/zenodo.3710076" httpUri="https://zenodo.org/record/3710076/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
), have the venter more or less unmodified and do not possess complete sulci (
<figureCitation id="13082A123E40F05898D4FA06C01E4E10" box="[977,1046,797,822]" captionStart="Figures 1–2" captionStartId="4.[192,287,1819,1841]" captionTargetBox="[192,1407,276,1739]" captionTargetId="figure@4.[192,1440,261,1767]" captionTargetPageId="4" captionText="Figures 1–2. Zelenka holotypes. 1) Holotype of Echinoaesalus jaechi Zelenka. a) Dorsal habitus. b) Abdomen, ventral view. c) Labels. 2) Holotype female of Echinoaesalus sabahensis Zelenka. a) Dorsal habitus. b) Abdomen, ventral view, showing complete leg-shaped sulci. c) Labels. Photos courtesy H. Schillhammer (Naturhistorisches Museum, Vienna)." figureDoi="http://doi.org/10.5281/zenodo.3710076" httpUri="https://zenodo.org/record/3710076/files/figure.png" pageId="3" pageNumber="2">Fig. 2</figureCitation>
).
<bibRefCitation id="EFA24B663E40F0589F2FFA06C0D44E10" author="Zelenka, W." box="[1066,1244,797,822]" pageId="3" pageNumber="2" pagination="56 - 61" refId="ref5677" refString="Zelenka, W. 1994. Zwei neue Echinoaesalus - Arten aus Sudostasien (Coleoptera, Lucanidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 46: 56 - 61." type="journal article" year="1994">Zelenka (1994)</bibRefCitation>
added two more new species and transferred
<taxonomicName id="4C334D143E40F058991FFA26C7234E73" authority="Araya et al." authorityName="Araya" box="[538,811,829,854]" class="Insecta" family="Lucanidae" genus="Strabaesalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="hidakai">
<emphasis id="B947EA853E40F058991FFA26C6914E73" box="[538,665,829,853]" italics="true" pageId="3" pageNumber="2">A. hidakai</emphasis>
Araya et al.
</taxonomicName>
to the genus while removing the presence of ventral depressions or sulci on the abdomen from the generic description (indicating that they could be absent or distinct). Later,
<bibRefCitation id="EFA24B663E40F0589AA3FA66C6574EB0" author="Krikken, J." box="[422,607,893,918]" pageId="3" pageNumber="2" pagination="275 - 284" refId="ref5562" refString="Krikken, J. 2008. Zelenkaesalus subgen. n. for three Southeast Asian species of Echinoaesalus Zelenka, including E. javanus sp. n. (Coleoptera: Lucanidae: Aesalinae). Koleopterologische Rundschau 78: 275 - 284." type="journal article" year="2008">Krikken (2008)</bibRefCitation>
created the subgenus
<taxonomicName id="4C334D143E40F0589874FA66C0154EB3" box="[881,1053,893,917]" pageId="3" pageNumber="2" rank="subGenus" subGenus="Zelenkaesalus">
<emphasis id="B947EA853E40F0589874FA66C0154EB3" box="[881,1053,893,917]" italics="true" pageId="3" pageNumber="2">Zelenkaesalus</emphasis>
</taxonomicName>
for
<taxonomicName id="4C334D143E40F0589F49FA66C0C44EB3" authorityName="Krikken. The" authorityYear="2018" box="[1100,1228,893,917]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="B947EA853E40F0589F49FA66C0C44EB3" box="[1100,1228,893,917]" italics="true" pageId="3" pageNumber="2">A. timidus</emphasis>
</taxonomicName>
(
<typeStatus id="548888353E40F0589FDEFA65C1194EB0" box="[1243,1297,894,918]" pageId="3" pageNumber="2">type</typeStatus>
species),
<taxonomicName id="4C334D143E40F0589E80FA65C5B84E93" authority="Zelenka" authorityName="Zelenka" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="B947EA853E40F0589E80FA65C54D4E93" italics="true" pageId="3" pageNumber="2">E. sabahensis</emphasis>
Zelenka
</taxonomicName>
(
<figureCitation id="13082A123E40F0589AC4FA86C60F4E90" box="[449,519,925,950]" captionStart="Figures 3–5" captionStartId="5.[192,287,605,627]" captionTargetBox="[192,1439,213,570]" captionTargetId="figure@5.[192,1440,213,571]" captionTargetPageId="5" captionText="Figures 3–5. Generic differences of ocular canthus/eye and clypeal apex. 3) Echinoaesalus spp. 4) Strabaesalus spp. 5) Zelenkaesalus spp." figureDoi="http://doi.org/10.5281/zenodo.3710078" httpUri="https://zenodo.org/record/3710078/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
), and one new species, uniting them for their distinctive, complete ventral sulci (
<figureCitation id="13082A123E40F0589A0EFAA6C55C4EF0" box="[267,340,957,982]" captionStart="Figures 3–5" captionStartId="5.[192,287,605,627]" captionTargetBox="[192,1439,213,570]" captionTargetId="figure@5.[192,1440,213,571]" captionTargetPageId="5" captionText="Figures 3–5. Generic differences of ocular canthus/eye and clypeal apex. 3) Echinoaesalus spp. 4) Strabaesalus spp. 5) Zelenkaesalus spp." figureDoi="http://doi.org/10.5281/zenodo.3710078" httpUri="https://zenodo.org/record/3710078/files/figure.png" pageId="3" pageNumber="2">Fig. 4</figureCitation>
).
<bibRefCitation id="EFA24B663E40F0589A6EFAA5C6534EF0" author="Huang, H. &amp; Y. Imura &amp; L. Wu" box="[363,603,957,982]" pageId="3" pageNumber="2" pagination="97 - 107" refId="ref5491" refString="Huang, H., Y. Imura, and L. Wu. 2011. Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane 12: 97 - 107." type="journal article" year="2011">Huang et al. (2011)</bibRefCitation>
described three new species from Borneo and Java, and included a key to the twelve species of
<taxonomicName id="4C334D143E40F058990FFAC6C6B24ED3" box="[522,698,989,1013]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F058990FFAC6C6B24ED3" box="[522,698,989,1013]" italics="true" pageId="3" pageNumber="2">Echinoaesalus</emphasis>
</taxonomicName>
then known.
<bibRefCitation id="EFA24B663E40F0589858FAC5C0754ED3" author="Huang, H. &amp; C. - C. Chen" box="[861,1149,989,1014]" pageId="3" pageNumber="2" pagination="163 - 170" refId="ref5364" refString="Huang, H., and C. - C. Chen. 2015. Discovery of a second species of Aesalini from Taiwan, with description of the new species of the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae). Zootaxa 3920 (1): 163 - 170." type="journal article" year="2015">Huang and Chen (2015)</bibRefCitation>
described a new species from
<collectingCountry id="F32476073E40F0589A06FAE6C56B4933" box="[259,355,1021,1045]" name="Taiwan" pageId="3" pageNumber="2">Taiwan</collectingCountry>
. Huang et al. (2015) described one new Malaysian species and elevated
<taxonomicName id="4C334D143E40F0589FEBFAE5C5954913" authority="Huang and Imura" authorityName="Huang and Imura" class="Insecta" family="Lucanidae" genus="Strabaesalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="B947EA853E40F0589FEBFAE5C1A84933" box="[1262,1440,1021,1045]" italics="true" pageId="3" pageNumber="2">E. borneoensis</emphasis>
Huang and Imura
</taxonomicName>
to specific status.
</paragraph>
<paragraph id="8B8C36973E40F0589BEDFD25C1A849D0" blockId="3.[192,1441,509,1270]" pageId="3" pageNumber="2">
<bibRefCitation id="EFA24B663E40F0589BEDFD25C6054970" author="Huang, H. &amp; C. - C. Chen" box="[232,525,1085,1110]" pageId="3" pageNumber="2" pagination="472 - 503" refId="ref5418" refString="Huang, H., and C. - C. Chen. 2017. Chapter 11 - A review of the tribe Aesalini Macleay, 1819 from China. p. 472 - 503, In: H. Huang and C. - H. Chen (eds.). Stag beetles of China, Vol. 3. Formosa Ecological Company; Taipei, Taiwan. 524 p." type="book chapter" year="2017">Huang and Chen (2017)</bibRefCitation>
noted that
<taxonomicName id="4C334D143E40F058999FFD26C7424973" box="[666,842,1085,1109]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F058999FFD26C7424973" box="[666,842,1085,1109]" italics="true" pageId="3" pageNumber="2">Echinoaesalus</emphasis>
</taxonomicName>
was a composite of three disparate groups based on characters of the vestiture and female genitalia, but they refrained from formalizing these groups due to their inability to study the
<typeStatus id="548888353E40F0589920FD65C65249B0" box="[549,602,1150,1174]" pageId="3" pageNumber="2">type</typeStatus>
species,
<taxonomicName id="4C334D143E40F05899C0FD65C72649B3" box="[709,814,1149,1173]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="B947EA853E40F05899C0FD65C72649B3" box="[709,814,1149,1173]" italics="true" pageId="3" pageNumber="2">E. jaechi</emphasis>
</taxonomicName>
. Instead they characterized the three groups as the matsuii group, hidakai group, and subgenus
<emphasis id="B947EA853E40F05899DEFD86C7874993" box="[731,911,1181,1205]" italics="true" pageId="3" pageNumber="2">
<taxonomicName id="4C334D143E40F05899DEFD86C7824993" box="[731,906,1181,1205]" pageId="3" pageNumber="2" rank="subGenus" subGenus="Zelenkaesalus">Zelenkaesalus</taxonomicName>
.
</emphasis>
The characters separating these groups are robust and typically used to separate genera within
<taxonomicName id="4C334D143E40F058982CFDA6C78549F3" authorityName="MacLeay" authorityYear="1819" box="[809,909,1213,1237]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Aesalini">Aesalini</taxonomicName>
(
<tableCitation id="C6B1032C3E40F058989CFDA6C7F949F0" box="[921,1009,1213,1238]" captionStart="Table 1" captionStartId="3.[192,261,1366,1387]" captionTargetId="graphics@3.[192,1440,1412,1805]" captionTargetPageId="3" captionText="Table 1. Generic characters of groups previously comprising Echinoaesalus Zelenka." httpUri="http://table.plazi.org/id/DF4C661F3E40F0589BC5FC4DC06F484A" pageId="3" pageNumber="2" tableUuid="DF4C661F3E40F0589BC5FC4DC06F484A">Table 1</tableCitation>
). For this reason, after examination of detailed images of the Zelenka
<typeStatus id="548888353E40F0589955FDC5C69949D0" box="[592,657,1246,1270]" pageId="3" pageNumber="2">types</typeStatus>
in
<collectingRegion id="49F7F8753E40F05899B0FDC6C70549D3" box="[693,781,1245,1269]" country="Austria" name="Wien" pageId="3" pageNumber="2">Vienna</collectingRegion>
the three groups can be formally recognized as genera.
</paragraph>
<caption id="DF4C661F3E40F0589BC5FC4DC06F484A" ID-Table-UUID="DF4C661F3E40F0589BC5FC4DC06F484A" box="[192,1127,1366,1388]" httpUri="http://table.plazi.org/id/DF4C661F3E40F0589BC5FC4DC06F484A" pageId="3" pageNumber="2" targetBox="[203,1419,1428,1802]" targetIsTable="true" targetPageId="3">
<paragraph id="8B8C36973E40F0589BC5FC4DC06F484A" blockId="3.[192,1127,1366,1388]" box="[192,1127,1366,1388]" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F0589BC5FC4DC528484D" bold="true" box="[192,288,1366,1387]" pageId="3" pageNumber="2">Table 1.</emphasis>
Generic characters of groups previously comprising
<taxonomicName id="4C334D143E40F0589867FC4DC06F484A" authority="Zelenka." authorityName="Zelenka." box="[866,1127,1366,1388]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589867FC4DC009484D" box="[866,1025,1366,1387]" italics="true" pageId="3" pageNumber="2">Echinoaesalus</emphasis>
Zelenka.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8B8C36973E40F0589BCEFCBFC0EE4A27" pageId="3" pageNumber="2">
<table id="F933C4373E400FA49BCEFC8FC1834A2C" box="[203,1419,1428,1802]" gridcols="5" gridrows="4" pageId="3" pageNumber="2">
<tr id="350334D53E400FA49BCEFC8FC18348EA" box="[203,1419,1428,1484]" gridrow="0" pageId="3" pageNumber="2">
<th id="76D25DA93E400FA49BCEFC8FC5D248EA" box="[203,474,1428,1484]" gridcol="0" gridrow="0" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F0589BCEFCBFC511489C" bold="true" box="[203,281,1444,1466]" pageId="3" pageNumber="2">Genus</emphasis>
</th>
<th id="76D25DA93E400FA49901FC8FC6BD48EA" box="[516,693,1428,1484]" gridcol="1" gridrow="0" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F0589901FCBFC67F489C" bold="true" box="[516,631,1444,1466]" pageId="3" pageNumber="2">Vestiture</emphasis>
</th>
<th id="76D25DA93E400FA499EBFC8FC7CA48EA" box="[750,962,1428,1484]" gridcol="2" gridrow="0" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F05899EBFCBFC7A4489C" bold="true" box="[750,940,1444,1466]" pageId="3" pageNumber="2">Ocular canthus</emphasis>
</th>
<th id="76D25DA93E400FA498D2FC8FC0A148EA" box="[983,1193,1428,1484]" gridcol="3" gridrow="0" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F05898D2FCBFC08F489C" bold="true" box="[983,1159,1444,1466]" pageId="3" pageNumber="2">Thoracic sulci</emphasis>
</th>
<th id="76D25DA93E400FA49FC4FC8FC18348EA" box="[1217,1419,1428,1484]" gridcol="4" gridrow="0" pageId="3" pageNumber="2">
<emphasis id="B947EA853E40F0589FC4FC8FC10A48EA" bold="true" box="[1217,1367,1428,1484]" pageId="3" pageNumber="2">Abdominal sulci</emphasis>
</th>
</tr>
<tr id="350334D53E400FA49BCEFCC7C1834B0A" box="[203,1419,1500,1580]" gridrow="1" pageId="3" pageNumber="2">
<th id="76D25DA93E400FA49BCEFCC7C5D24B0A" box="[203,474,1500,1580]" gridcol="0" gridrow="1" pageId="3" pageNumber="2">
<taxonomicName id="4C334D143E40F0589BCEFCC7C57648D4" box="[203,382,1500,1522]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589BCEFCC7C57648D4" bold="true" box="[203,382,1500,1522]" italics="true" pageId="3" pageNumber="2">Echinoaesalus</emphasis>
</taxonomicName>
(matsuii group of
<bibRefCitation id="EFA24B663E40F0589A8AFCE1C57A4B0A" author="Huang, H. &amp; C. - C. Chen" pageId="3" pageNumber="2" pagination="472 - 503" refId="ref5418" refString="Huang, H., and C. - C. Chen. 2017. Chapter 11 - A review of the tribe Aesalini Macleay, 1819 from China. p. 472 - 503, In: H. Huang and C. - H. Chen (eds.). Stag beetles of China, Vol. 3. Formosa Ecological Company; Taipei, Taiwan. 524 p." type="book chapter" year="2017">Huang and Chen 2017</bibRefCitation>
)
</th>
<td id="76D25DA93E400FA49901FCC7C6BD4B0A" box="[516,693,1500,1580]" gridcol="1" gridrow="1" pageId="3" pageNumber="2">All stick-like, scattered</td>
<td id="76D25DA93E400FA499EBFCC7C7CA4B0A" box="[750,962,1500,1580]" gridcol="2" gridrow="1" pageId="3" pageNumber="2">Strongly intruding; dorsal lobe of eye large (Fig. 3)</td>
<td id="76D25DA93E400FA498D2FCC7C0A14B0A" box="[983,1193,1500,1580]" gridcol="3" gridrow="1" pageId="3" pageNumber="2">Short, longitudinal</td>
<td id="76D25DA93E400FA49FC4FCC7C1834B0A" box="[1217,1419,1500,1580]" gridcol="4" gridrow="1" pageId="3" pageNumber="2">Absent (Fig. 1b)</td>
</tr>
<tr id="350334D53E400FA49BCEFF26C1834BAB" box="[203,1419,1597,1677]" gridrow="2" pageId="3" pageNumber="2">
<th id="76D25DA93E400FA49BCEFF26C5D24BAB" box="[203,474,1597,1677]" gridcol="0" gridrow="2" pageId="3" pageNumber="2">
<taxonomicName id="4C334D143E40F0589BCEFF26C5654B75" authorityName="Paulsen" authorityYear="2018" box="[203,365,1597,1619]" class="Insecta" family="Lucanidae" genus="Strabaesalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589BCEFF26C5654B75" bold="true" box="[203,365,1597,1619]" italics="true" pageId="3" pageNumber="2">Strabaesalus</emphasis>
</taxonomicName>
(hidakai group of
<bibRefCitation id="EFA24B663E40F0589A8AFF40C57A4BAB" author="Huang, H. &amp; C. - C. Chen" pageId="3" pageNumber="2" pagination="472 - 503" refId="ref5418" refString="Huang, H., and C. - C. Chen. 2017. Chapter 11 - A review of the tribe Aesalini Macleay, 1819 from China. p. 472 - 503, In: H. Huang and C. - H. Chen (eds.). Stag beetles of China, Vol. 3. Formosa Ecological Company; Taipei, Taiwan. 524 p." type="book chapter" year="2017">Huang and Chen 2017</bibRefCitation>
)
</th>
<td id="76D25DA93E400FA49901FF26C6BD4BAB" box="[516,693,1597,1677]" gridcol="1" gridrow="2" pageId="3" pageNumber="2">Some scale-like, clumped</td>
<td id="76D25DA93E400FA499EBFF26C7CA4BAB" box="[750,962,1597,1677]" gridcol="2" gridrow="2" pageId="3" pageNumber="2">Strongly intruding; dorsal lobe of eye narrow (Fig. 4)</td>
<td id="76D25DA93E400FA498D2FF26C0A14BAB" box="[983,1193,1597,1677]" gridcol="3" gridrow="2" pageId="3" pageNumber="2">Short, transverse</td>
<td id="76D25DA93E400FA49FC4FF26C1834BAB" box="[1217,1419,1597,1677]" gridcol="4" gridrow="2" pageId="3" pageNumber="2">Absent</td>
</tr>
<tr id="350334D53E400FA49BCEFF85C1834A2C" box="[203,1419,1694,1802]" gridrow="3" pageId="3" pageNumber="2">
<th id="76D25DA93E400FA49BCEFF85C5D24A2C" box="[203,474,1694,1802]" gridcol="0" gridrow="3" pageId="3" pageNumber="2">
<taxonomicName id="4C334D143E40F0589BCEFFD2C5734BF9" authorityName="Krikken" authorityYear="2008" box="[203,379,1737,1759]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E40F0589BCEFFD2C5734BF9" bold="true" box="[203,379,1737,1759]" italics="true" pageId="3" pageNumber="2">Zelenkaesalus</emphasis>
</taxonomicName>
</th>
<td id="76D25DA93E400FA49901FF85C6BD4A2C" box="[516,693,1694,1802]" gridcol="1" gridrow="3" pageId="3" pageNumber="2">Some scale-like, clumped</td>
<td id="76D25DA93E400FA499EBFF85C7CA4A2C" box="[750,962,1694,1802]" gridcol="2" gridrow="3" pageId="3" pageNumber="2">Almost obsolete; eye without distinct dorsal lobe (Fig. 5)</td>
<td id="76D25DA93E400FA498D2FF85C0A14A2C" box="[983,1193,1694,1802]" gridcol="3" gridrow="3" pageId="3" pageNumber="2">Complete (leg-shaped)</td>
<td id="76D25DA93E400FA49FC4FF85C1834A2C" box="[1217,1419,1694,1802]" gridcol="4" gridrow="3" pageId="3" pageNumber="2">Complete (leg-shaped; Fig. 2b)</td>
</tr>
</table>
</paragraph>
<caption id="DF4C661F3E47F05F9BC5FE00C5844AAE" ID-DOI="http://doi.org/10.5281/zenodo.3710076" ID-Zenodo-Dep="3710076" httpUri="https://zenodo.org/record/3710076/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,287,1819,1841]" targetBox="[192,1407,276,1739]" targetPageId="4">
<paragraph id="8B8C36973E47F05F9BC5FE00C5844AAE" blockId="4.[192,1440,1819,1928]" pageId="4" pageNumber="3">
<emphasis id="B947EA853E47F05F9BC5FE00C55F4A16" bold="true" box="[192,343,1819,1841]" pageId="4" pageNumber="3">Figures 1–2.</emphasis>
Zelenka holotypes.
<emphasis id="B947EA853E47F05F993CFE00C6584A17" bold="true" box="[569,592,1819,1841]" pageId="4" pageNumber="3">1)</emphasis>
Holotype of
<taxonomicName id="4C334D143E47F05F99E7FE00C03A4A14" authority="Zelenka." authorityName="Zelenka." box="[738,1074,1819,1842]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="B947EA853E47F05F99E7FE00C7C24A16" box="[738,970,1819,1840]" italics="true" pageId="4" pageNumber="3">Echinoaesalus jaechi</emphasis>
Zelenka.
</taxonomicName>
<emphasis id="B947EA853E47F05F9F3FFE00C05B4A17" bold="true" box="[1082,1107,1819,1841]" pageId="4" pageNumber="3">a)</emphasis>
Dorsal habitus.
<emphasis id="B947EA853E47F05F9E15FE00C1204A17" bold="true" box="[1296,1320,1819,1841]" pageId="4" pageNumber="3">b)</emphasis>
Abdomen, ventral view.
<emphasis id="B947EA853E47F05F9A53FE23C5654A68" bold="true" box="[342,365,1848,1870]" pageId="4" pageNumber="3">c)</emphasis>
Labels.
<emphasis id="B947EA853E47F05F9ACFFE23C5E84A68" bold="true" box="[458,480,1848,1870]" pageId="4" pageNumber="3">2)</emphasis>
Holotype female of
<taxonomicName id="4C334D143E47F05F99BEFE23C0364A68" authority="Zelenka." authorityName="Zelenka." box="[699,1086,1848,1870]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="B947EA853E47F05F99BEFE23C7D04A6B" box="[699,984,1848,1869]" italics="true" pageId="4" pageNumber="3">Echinoaesalus sabahensis</emphasis>
Zelenka.
</taxonomicName>
<emphasis id="B947EA853E47F05F9F41FE23C0544A68" bold="true" box="[1092,1116,1848,1870]" pageId="4" pageNumber="3">a)</emphasis>
Dorsal habitus.
<emphasis id="B947EA853E47F05F9E17FE23C1234A68" bold="true" box="[1298,1323,1848,1870]" pageId="4" pageNumber="3">b)</emphasis>
Abdomen, ventral view, showing complete leg-shaped sulci.
<emphasis id="B947EA853E47F05F99E7FE4EC6F14A4D" bold="true" box="[738,761,1877,1899]" pageId="4" pageNumber="3">c)</emphasis>
Labels. Photos courtesy H. Schillhammer (Naturhistorisches Museum, Vienna).
</paragraph>
</caption>
<caption id="DF4C661F3E46F05E9BC5FB46C5E84FA9" ID-DOI="http://doi.org/10.5281/zenodo.3710078" ID-Zenodo-Dep="3710078" httpUri="https://zenodo.org/record/3710078/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,287,605,627]" targetBox="[192,1439,213,570]" targetPageId="5">
<paragraph id="8B8C36973E46F05E9BC5FB46C5E84FA9" blockId="5.[192,1440,605,655]" pageId="5" pageNumber="4">
<emphasis id="B947EA853E46F05E9BC5FB46C55F4F54" bold="true" box="[192,343,605,627]" pageId="5" pageNumber="4">Figures 3–5.</emphasis>
Generic differences of ocular canthus/eye and clypeal apex.
<emphasis id="B947EA853E46F05E98F2FB46C0064F55" bold="true" box="[1015,1038,605,627]" pageId="5" pageNumber="4">3)</emphasis>
<taxonomicName id="4C334D143E46F05E9F13FB46C0BD4F54" box="[1046,1205,605,626]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E46F05E9F13FB46C0BD4F54" box="[1046,1205,605,626]" italics="true" pageId="5" pageNumber="4">Echinoaesalus</emphasis>
</taxonomicName>
spp.
<emphasis id="B947EA853E46F05E9FF4FB46C1004F55" bold="true" box="[1265,1288,605,627]" pageId="5" pageNumber="4">4)</emphasis>
<taxonomicName id="4C334D143E46F05E9E15FB46C1A84F54" authorityName="Paulsen" authorityYear="2018" box="[1296,1440,605,626]" class="Insecta" family="Lucanidae" genus="Strabaesalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E46F05E9E15FB46C1A84F54" box="[1296,1440,605,626]" italics="true" pageId="5" pageNumber="4">Strabaesalus</emphasis>
</taxonomicName>
spp.
<emphasis id="B947EA853E46F05E9BF1FB62C5024FA9" bold="true" box="[244,266,633,655]" pageId="5" pageNumber="4">5)</emphasis>
<taxonomicName id="4C334D143E46F05E9A17FB61C5A54FA9" authorityName="Krikken" authorityYear="2008" box="[274,429,634,655]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E46F05E9A17FB61C5A54FA9" box="[274,429,634,655]" italics="true" pageId="5" pageNumber="4">Zelenkaesalus</emphasis>
</taxonomicName>
spp.
</paragraph>
</caption>
<paragraph id="8B8C36973E46F05E9BEDFBDDC5164E26" blockId="5.[192,1440,710,768]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BEDFBDDC6094FF9" authority="Zelenka" authorityName="Zelenka" box="[232,513,710,735]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E46F05E9BEDFBDDC5904FF8" box="[232,408,710,734]" italics="true" pageId="5" pageNumber="4">Echinoaesalus</emphasis>
Zelenka
</taxonomicName>
(matsuii group of
<bibRefCitation id="EFA24B663E46F05E99DAFBDCC7F94FF9" author="Huang, H. &amp; C. - C. Chen" box="[735,1009,711,735]" pageId="5" pageNumber="4" pagination="472 - 503" refId="ref5418" refString="Huang, H., and C. - C. Chen. 2017. Chapter 11 - A review of the tribe Aesalini Macleay, 1819 from China. p. 472 - 503, In: H. Huang and C. - H. Chen (eds.). Stag beetles of China, Vol. 3. Formosa Ecological Company; Taipei, Taiwan. 524 p." type="book chapter" year="2017">Huang and Chen 2017</bibRefCitation>
) is restricted to the following seven species:
</paragraph>
<paragraph id="8B8C36973E46F05E9BC5FA02C0864E17" blockId="5.[192,1166,793,817]" box="[192,1166,793,817]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BC5FA02C61E4E16" authority="Huang and Imura" authorityName="Huang and Imura" box="[192,534,793,817]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="arayai">
<emphasis id="B947EA853E46F05E9BC5FA02C5394E16" box="[192,305,793,816]" italics="true" pageId="5" pageNumber="4">E. arayai</emphasis>
Huang and Imura
</taxonomicName>
<collectingCountry id="F32476073E46F05E9835FA02C7CF4E17" box="[816,967,793,817]" name="Malaysia" pageId="5" pageNumber="4">MALAYSIA</collectingCountry>
:
<collectingRegion id="49F7F8753E46F05E98D4FA02C0164E17" box="[977,1054,793,817]" country="Malaysia" name="Sabah" pageId="5" pageNumber="4">Sabah</collectingRegion>
(Borneo)
</paragraph>
<paragraph id="8B8C36973E46F05E9BC5FA5BC0444E71" blockId="5.[192,1100,831,855]" box="[192,1100,831,855]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BC5FA5BC5A24E71" authority="Zelenka" authorityName="Zelenka" authorityYear="1993" box="[192,426,831,855]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="barriesi">
<emphasis id="B947EA853E46F05E9BC5FA5BC5494E71" box="[192,321,831,855]" italics="true" pageId="5" pageNumber="4">E. barriesi</emphasis>
Zelenka
</taxonomicName>
<collectingCountry id="F32476073E46F05E9835FA24C7DD4E71" box="[816,981,831,855]" name="Indonesia" pageId="5" pageNumber="4">INDONESIA</collectingCountry>
:
<collectingRegion id="49F7F8753E46F05E98E5FA24C0444E71" box="[992,1100,831,855]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="4">Sumatra</collectingRegion>
</paragraph>
<paragraph id="8B8C36973E46F05E9BC5FA7DC7AB4E58" blockId="5.[192,931,869,894]" box="[192,931,869,894]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BC5FA7DC6064E58" authority="Huang and Chen" authorityName="Huang and Chen" box="[192,526,869,894]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="chungi">
<emphasis id="B947EA853E46F05E9BC5FA7DC53D4E5B" box="[192,309,869,893]" italics="true" pageId="5" pageNumber="4">E. chungi</emphasis>
Huang and Chen
</taxonomicName>
<collectingCountry id="F32476073E46F05E9835FA7DC7AB4E58" box="[816,931,870,894]" name="Taiwan" pageId="5" pageNumber="4">TAIWAN</collectingCountry>
</paragraph>
<paragraph id="8B8C36973E46F05E9BC5FA97C7CD4E82" blockId="5.[192,965,908,932]" box="[192,965,908,932]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BC5FA97C5E54E82" authority="(Araya et al.)" baseAuthorityName="Araya" box="[192,493,908,932]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="dharma">
<emphasis id="B947EA853E46F05E9BC5FA97C54C4E82" box="[192,324,908,932]" italics="true" pageId="5" pageNumber="4">E. dharma</emphasis>
(Araya et al.)
</taxonomicName>
<collectingCountry id="F32476073E46F05E9835FA97C7CD4E82" box="[816,965,908,932]" name="Thailand" pageId="5" pageNumber="4">THAILAND</collectingCountry>
</paragraph>
<paragraph id="8B8C36973E46F05E9BC5FAA8C0474EEC" blockId="5.[192,1103,946,970]" box="[192,1103,946,970]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BC5FAA8C59B4EEC" authority="Zelenka" authorityName="Zelenka" box="[192,403,946,970]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="B947EA853E46F05E9BC5FAA8C5224EEC" box="[192,298,946,970]" italics="true" pageId="5" pageNumber="4">E. jaechi</emphasis>
Zelenka
</taxonomicName>
<collectingCountry id="F32476073E46F05E9835FAA9C7DD4EEC" box="[816,981,946,970]" name="Indonesia" pageId="5" pageNumber="4">INDONESIA</collectingCountry>
:
<collectingRegion id="49F7F8753E46F05E98E5FAA9C0474EEC" box="[992,1103,946,970]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="4">Sulawesi</collectingRegion>
</paragraph>
<paragraph id="8B8C36973E46F05E9BC5FAC2C0274ED7" blockId="5.[192,1071,985,1009]" box="[192,1071,985,1009]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BC5FAC2C5AB4ED7" authority="(Araya)" baseAuthorityName="Araya" baseAuthorityYear="1993" box="[192,419,985,1009]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="matsuii">
<emphasis id="B947EA853E46F05E9BC5FAC2C5484ED6" box="[192,320,985,1008]" italics="true" pageId="5" pageNumber="4">E. matsuii</emphasis>
(Araya)
</taxonomicName>
<collectingCountry id="F32476073E46F05E9835FAC2C7CF4ED7" box="[816,967,985,1009]" name="Malaysia" pageId="5" pageNumber="4">MALAYSIA</collectingCountry>
:
<collectingRegion id="49F7F8753E46F05E98D4FAC2C0274ED7" box="[977,1071,985,1009]" country="Malaysia" name="Pahang" pageId="5" pageNumber="4">Pahang</collectingRegion>
</paragraph>
<paragraph id="8B8C36973E46F05E9BC5FAE4C0364931" blockId="5.[192,1086,1023,1047]" box="[192,1086,1023,1047]" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9BC5FAE4C58F4931" authority="(Araya)" baseAuthorityName="Araya" baseAuthorityYear="1993" box="[192,391,1023,1047]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="yongi">
<emphasis id="B947EA853E46F05E9BC5FAE4C52C4930" box="[192,292,1023,1046]" italics="true" pageId="5" pageNumber="4">E. yongi</emphasis>
(Araya)
</taxonomicName>
<collectingCountry id="F32476073E46F05E9835FAE4C7CF4931" box="[816,967,1023,1047]" name="Malaysia" pageId="5" pageNumber="4">MALAYSIA</collectingCountry>
:
<collectingRegion id="49F7F8753E46F05E98D4FAE4C0364931" box="[977,1086,1023,1047]" country="Malaysia" name="Selangor" pageId="5" pageNumber="4">Selangor</collectingRegion>
</paragraph>
<paragraph id="8B8C36973E46F05E9BEDFD2BC6FB4828" blockId="5.[192,1441,1072,1294]" pageId="5" pageNumber="4">
Among Southeast Asian taxa the genus
<taxonomicName id="4C334D143E46F05E99CAFD2BC777496E" box="[719,895,1072,1096]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E46F05E99CAFD2BC777496E" box="[719,895,1072,1096]" italics="true" pageId="5" pageNumber="4">Echinoaesalus</emphasis>
</taxonomicName>
can be distinguished easily by the scattered, stick-like bristles on the elytra (
<figureCitation id="13082A123E46F05E9940FD4AC692494F" box="[581,666,1105,1129]" captionStart="Figures 1–2" captionStartId="4.[192,287,1819,1841]" captionTargetBox="[192,1407,276,1739]" captionTargetId="figure@4.[192,1440,261,1767]" captionTargetPageId="4" captionText="Figures 1–2. Zelenka holotypes. 1) Holotype of Echinoaesalus jaechi Zelenka. a) Dorsal habitus. b) Abdomen, ventral view. c) Labels. 2) Holotype female of Echinoaesalus sabahensis Zelenka. a) Dorsal habitus. b) Abdomen, ventral view, showing complete leg-shaped sulci. c) Labels. Photos courtesy H. Schillhammer (Naturhistorisches Museum, Vienna)." figureDoi="http://doi.org/10.5281/zenodo.3710076" httpUri="https://zenodo.org/record/3710076/files/figure.png" pageId="5" pageNumber="4">Fig. 1a</figureCitation>
; clumps of broad scales are absent). The ventral surface is relatively unmodified and lacks the complete sulci to receive the middle or hindlegs found in
<emphasis id="B947EA853E46F05E9FE8FD69C1A849AC" box="[1261,1440,1138,1162]" italics="true" pageId="5" pageNumber="4">
<taxonomicName id="4C334D143E46F05E9FE8FD69C19549AC" authorityName="Krikken" authorityYear="2008" box="[1261,1437,1138,1162]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">Zelenkaesalus</taxonomicName>
.
</emphasis>
Instead the metasternum has only a short, longitudinal pit laterally (that may receive the tarsi only), and the abdomen is unmodified (
<figureCitation id="13082A123E46F05E9959FDAFC6BB49EA" box="[604,691,1204,1228]" captionStart="Figures 1–2" captionStartId="4.[192,287,1819,1841]" captionTargetBox="[192,1407,276,1739]" captionTargetId="figure@4.[192,1440,261,1767]" captionTargetPageId="4" captionText="Figures 1–2. Zelenka holotypes. 1) Holotype of Echinoaesalus jaechi Zelenka. a) Dorsal habitus. b) Abdomen, ventral view. c) Labels. 2) Holotype female of Echinoaesalus sabahensis Zelenka. a) Dorsal habitus. b) Abdomen, ventral view, showing complete leg-shaped sulci. c) Labels. Photos courtesy H. Schillhammer (Naturhistorisches Museum, Vienna)." figureDoi="http://doi.org/10.5281/zenodo.3710076" httpUri="https://zenodo.org/record/3710076/files/figure.png" pageId="5" pageNumber="4">Fig. 1b</figureCitation>
). The ocular canthus is long and the eye has a large, ovoid, dorsal portion that cannot be obscured by the pronotum (
<figureCitation id="13082A123E46F05E9872FDCEC7B549CB" box="[887,957,1237,1261]" captionStart="Figures 3–5" captionStartId="5.[192,287,605,627]" captionTargetBox="[192,1439,213,570]" captionTargetId="figure@5.[192,1440,213,571]" captionTargetPageId="5" captionText="Figures 3–5. Generic differences of ocular canthus/eye and clypeal apex. 3) Echinoaesalus spp. 4) Strabaesalus spp. 5) Zelenkaesalus spp." figureDoi="http://doi.org/10.5281/zenodo.3710078" httpUri="https://zenodo.org/record/3710078/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
). The clypeus is more or less triangular but not distinctly acuminate medially (
<figureCitation id="13082A123E46F05E9998FDEDC6EC4828" box="[669,740,1270,1294]" captionStart="Figures 3–5" captionStartId="5.[192,287,605,627]" captionTargetBox="[192,1439,213,570]" captionTargetId="figure@5.[192,1440,213,571]" captionTargetPageId="5" captionText="Figures 3–5. Generic differences of ocular canthus/eye and clypeal apex. 3) Echinoaesalus spp. 4) Strabaesalus spp. 5) Zelenkaesalus spp." figureDoi="http://doi.org/10.5281/zenodo.3710078" httpUri="https://zenodo.org/record/3710078/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink