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<document id="A02BEB90C6FBA96A064B2BB3439A6EBC" ID-CLB-Dataset="26396" ID-DOI="10.11646/zootaxa.4621.1.1" ID-GBIF-Dataset="a381190d-7f50-4a66-bc33-3a0417c9f89f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3257228" ID-ZooBank="3D77F329-9BB8-4231-90E4-6A9E0250FCDF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1561539959806" checkinUser="plazi" docAuthor="Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo &amp; Vieira, Juliana S." docDate="2019" docId="03978788B9A8FF18C1E36957AAD79BA3" docLanguage="en" docName="zootaxa.4621.1.1.pdf" docOrigin="Zootaxa 4621 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Blediini Adam 2001" docType="treatment" docVersion="13" lastPageNumber="222" masterDocId="FFAEFFF0B974FFC5C1746833AE7B997A" masterDocTitle="Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution" masterLastPageNumber="406" masterPageNumber="1" pageNumber="221" updateTime="1698737271145" updateUser="ExternalLinkService">
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<mods:title id="09459436AA78A61BAE6FEAE5480976E3">Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution</mods:title>
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Tribe
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Blediini
<bibRefCitation id="EFAF4B6FB9A8FF19C03F6957AF9A9804" author="Adam, L." box="[331,481,356,382]" pageId="220" pageNumber="221" refId="ref165998" refString="Adam, L. (2001) [systematic content] In: A satoraljaujhelyi Kazinczy Ferenc Muzeum fuzetei V, Satoraljaujhely. ANAdatok a Zempleni-hegyseg, a Hernad-volgy, a Bodrogkoz, a Retkoz es a Taktakoz holyvafaunajahoz (Coleoptera). In: Adam, L. &amp; Hegyessy, G. (Eds.). A satoraljaujhelyi Kazinczy Ferenc Muzeum fuzetei V, Satoraljaujhely, 249 pp." type="book" year="2001">Ádám, 2001</bibRefCitation>
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<emphasis id="B94AEA8CB9A8FF19C1E36987AEAD98B7" bold="true" box="[151,214,436,461]" pageId="220" pageNumber="221">0762.</emphasis>
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<emphasis id="B94AEA8CB9A8FF19C1A96987AF4F98B7" bold="true" box="[221,308,436,461]" italics="true" pageId="220" pageNumber="221">Bledius</emphasis>
Leach
<emphasis id="B94AEA8CB9A8FF19C0F26986AFE798B6" box="[390,412,437,460]" italics="true" pageId="220" pageNumber="221">in</emphasis>
<bibRefCitation id="EFAF4B6FB9A8FF19C0D76987ACE398B7" author="Samouelle, G." box="[419,664,436,461]" pageId="220" pageNumber="221" refId="ref232013" refString="Samouelle, G. (1819) The Entomologist's useful Compendium, or an Introduction to the Knowledge of British Insects: comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linne, and the modern Thomas Boys, London. 496 pp., 12 pls." type="book" year="1819">Samouelle, 1819: 174</bibRefCitation>
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(
<figureCitation id="13052A1BB9A8FF19C3B86987AD1598B7" box="[716,878,436,461]" captionStart="FIGURES 112–123" captionStartId="330.[151,264,1860,1883]" captionTargetBox="[155,1435,198,1832]" captionTargetId="figure@330.[150,1436,193,1840]" captionTargetPageId="330" captionText="FIGURES 112–123. Habitus; Osoriinae (112–118); Oxytelinae (119–123); 112, after Sharp 1887a; 113, after Bruch 1942; 114, after Sharp 1887a; 115, after Irmler 2003c; 116, after Sharp 1887a; 117, after Irmler 2003a; 118, after Sharp 1887a; 119, after Sharp 1887a; 120, after Caron &amp; Ribeiro-Costa 2007; 121, after Caron &amp; Ribeiro-Costa 2007; 122, after Sharp 1887a; 123, after Makranczy 2014; slightly modified. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3257252/files/figure.png" pageId="220" pageNumber="221">Figs. 120–121</figureCitation>
)
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<typeStatus id="5485883CB9A8FF19C1E369EFAEAA988E" box="[151,209,476,500]" pageId="220" pageNumber="221">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A8FF19C04369E8ACCE988E" authority="Panzer, 1799" authorityName="Panzer" authorityYear="1799" box="[311,693,475,500]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B94AEA8CB9A8FF19C04369E8AC669889" box="[311,541,475,500]" italics="true" pageId="220" pageNumber="221">Staphylinus armatus</emphasis>
Panzer, 1799
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<taxonomicName id="4C3E4D1DB9A8FF19C1BB6A31AC2B9B61" ID-CoL="8XMS8" authority="Herman, 1972 a: 118" authorityName="Herman" authorityPageNumber="118" authorityYear="1972" box="[207,592,514,539]" class="Insecta" family="Staphylinidae" genus="Microbledius" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A8FF19C1BB6A31AF1F9B61" box="[207,356,514,539]" italics="true" pageId="220" pageNumber="221">Microbledius</emphasis>
<treatmentCitation id="0A9F108FB9A8FF19C01F6A30AC2B9B61" author="Herman, L. H." box="[363,592,515,539]" page="118" pageId="220" pageNumber="221" year="1972">
<bibRefCitation id="EFAF4B6FB9A8FF19C01F6A30AC2B9B61" author="Herman, L. H." box="[363,592,515,539]" pageId="220" pageNumber="221" pagination="111 - 254" refId="ref199394" refString="Herman, L. H. (1972 a) Revision of Bledius and related genera. Part I. The aequatorialis, mandibularis, and semiferrugineus groups and two new genera (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the American Museum of Natural History, 149 (2), 111 - 254." type="journal article" year="1972">Herman, 1972a: 118</bibRefCitation>
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[synonymized by
<treatmentCitation id="0A9F108FB9A8FF19C2546A30AD8E9B61" author="Herman, L. H." box="[800,1013,514,539]" page="295" pageId="220" pageNumber="221" year="1986">
<bibRefCitation id="EFAF4B6FB9A8FF19C2546A30AD8E9B61" author="Herman, L. H." box="[800,1013,514,539]" pageId="220" pageNumber="221" pagination="1 - 368" refId="ref199711" refString="Herman, L. H. (1986) Revision of Bledius. Part IV. Classification of species groups, phylogeny, natural history, and catalogue (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the American Museum of Natural History, 184 (1), 1 - 368." type="journal article" year="1986">Herman 1986: 295</bibRefCitation>
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]
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<typeStatus id="5485883CB9A8FF19C0736A19AF3A9B38" box="[263,321,554,578]" pageId="220" pageNumber="221">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A8FF19C0D36A1AAD2B9B38" authority="Herman, 1972" authorityName="Herman" authorityYear="1972" box="[423,848,553,578]" class="Insecta" family="Staphylinidae" genus="Microbledius" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="playanus">
<emphasis id="B94AEA8CB9A8FF19C0D36A1AACD39B38" box="[423,680,553,578]" italics="true" pageId="220" pageNumber="221">Microbledius playanus</emphasis>
Herman, 1972
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<treatmentCitationGroup id="AB2E11B0B9A8FF19C1E36A4BAFA79BEA" box="[151,476,631,656]" pageId="220" pageNumber="221">
<taxonomicName id="4C3E4D1DB9A8FF19C1E36A4BAFA79BEA" authority="Thomson, 1859: 43" authorityName="Thomson" authorityPageNumber="43" authorityYear="1859" box="[151,476,631,656]" class="Insecta" family="Staphylinidae" genus="Astycops" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A8FF19C1E36A4BAE809BF5" box="[151,251,632,655]" italics="true" pageId="220" pageNumber="221">Astycops</emphasis>
<treatmentCitation id="0A9F108FB9A8FF19C0756A44AFA79BEA" author="Thomson, C. G." box="[257,476,631,656]" page="43" pageId="220" pageNumber="221" year="1859">
<bibRefCitation id="EFAF4B6FB9A8FF19C0756A44AFA79BEA" author="Thomson, C. G." box="[257,476,631,656]" pageId="220" pageNumber="221" refId="ref241704" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. I. Berlingska Boktryckeriet, Lund, 290 pp." type="book" year="1859">Thomson, 1859: 43</bibRefCitation>
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<typeStatus id="5485883CB9A8FF19C0736AACAF3A9BCD" box="[263,321,671,695]" pageId="220" pageNumber="221">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A8FF19C0D36AADAD7E9BCD" authority="Gyllenhal, 1810" authorityName="Gyllenhal" authorityYear="1810" box="[423,773,670,695]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="talpa">
<emphasis id="B94AEA8CB9A8FF19C0D36AADAC319BCD" box="[423,586,670,695]" italics="true" pageId="220" pageNumber="221">Oxytelus talpa</emphasis>
Gyllenhal, 1810
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<treatmentCitationGroup id="AB2E11B0B9A8FF19C1E36AF5AC739BA4" box="[151,520,709,734]" pageId="220" pageNumber="221">
<taxonomicName id="4C3E4D1DB9A8FF19C1E36AF5AFA99BA4" ID-CoL="9L9J2" authority="Schiodte, 1866: 145" authorityName="Schiodte" authorityPageNumber="145" authorityYear="1866" box="[151,466,709,734]" class="Insecta" family="Staphylinidae" genus="Bargus" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A8FF19C1E36AF5AE939BA7" box="[151,232,710,733]" italics="true" pageId="220" pageNumber="221">Bargus</emphasis>
<treatmentCitation id="0A9F108FB9A8FF19C19B6AF6AFA99BA4" author="Schiodte, J. M. C." box="[239,466,709,734]" page="145" pageId="220" pageNumber="221" year="1866">
<bibRefCitation id="EFAF4B6FB9A8FF19C19B6AF6AFA99BA4" author="Schiodte, J. M. C." box="[239,466,709,734]" pageId="220" pageNumber="221" pagination="141 - 167" refId="ref236754" refString="Schiodte, J. M. C. (1866) De tunnelgravende Biller Bledius, Heterocerus, Dyschirius og deres danske Arter. Naturhistorisk Tidsskrift, 3 (4), 141 - 167." type="journal article" year="1866">Schiødte, 1866: 145</bibRefCitation>
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, 148
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<typeStatus id="5485883CB9A8FF19C0736ADEAF3A9A7F" box="[263,321,749,773]" pageId="220" pageNumber="221">Type</typeStatus>
species:
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<emphasis id="B94AEA8CB9A8FF19C1E36B20AE909A56" box="[151,235,787,812]" italics="true" pageId="220" pageNumber="221">Belidus</emphasis>
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species:
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Mulsant &amp; Rey, 1861
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<emphasis id="B94AEA8CB9A8FF19C1E36B52AE909A00" box="[151,235,865,890]" italics="true" pageId="220" pageNumber="221">Bledius</emphasis>
Leach
<emphasis id="B94AEA8CB9A8FF19C0496B51AF289A03" box="[317,339,866,889]" italics="true" pageId="220" pageNumber="221">in</emphasis>
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36BBBACF69ADA" box="[423,653,904,929]" italics="true" pageId="220" pageNumber="221">Staphylinus armatus</emphasis>
Panzer, 1799
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<emphasis id="B94AEA8CB9A8FF19C1E36B83AF779ABD" box="[151,268,944,967]" italics="true" pageId="220" pageNumber="221">Dicarenus</emphasis>
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<paragraph id="8B81369EB9A8FF19C0736BE4AD3B9A95" blockId="220.[151,1437,436,2021]" box="[263,832,982,1007]" pageId="220" pageNumber="221">
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36BE5ACE59A94" box="[423,670,982,1007]" italics="true" pageId="220" pageNumber="221">Staphylinus arenarius</emphasis>
Paykull, 1800
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<paragraph id="8B81369EB9A8FF19C1BB6BCDAD829D6C" blockId="220.[151,1437,436,2021]" box="[207,1017,1021,1046]" pageId="220" pageNumber="221">
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<emphasis id="B94AEA8CB9A8FF19C1BB6BCDAF4D9D6F" box="[207,310,1022,1045]" italics="true" pageId="220" pageNumber="221">Cotysops</emphasis>
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[synonymized by
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36C17ACE59D46" box="[423,670,1060,1085]" italics="true" pageId="220" pageNumber="221">Staphylinus arenarius</emphasis>
Paykull, 1800
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<emphasis id="B94AEA8CB9A8FF19C1E36C78AE969D1E" box="[151,237,1099,1124]" italics="true" pageId="220" pageNumber="221">Elbidus</emphasis>
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36C41AC179DF1" box="[423,620,1138,1163]" italics="true" pageId="220" pageNumber="221">Oxytelus bicornis</emphasis>
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<emphasis id="B94AEA8CB9A8FF19C1BB6CAAAF369DC8" box="[207,333,1177,1202]" italics="true" pageId="220" pageNumber="221">Neobledius</emphasis>
M.
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36CF3ACC19DA3" box="[423,698,1216,1241]" italics="true" pageId="220" pageNumber="221">Neobledius karachiensis</emphasis>
<bibRefCitation id="EFAF4B6FB9A8FF19C3B46CF3ADA99DA3" author="Abdullah, M. &amp; Abdullah, A." box="[704,978,1216,1241]" pageId="220" pageNumber="221" pagination="1 - 3" refId="ref165554" refString="Abdullah, M. &amp; Abdullah, A. (1968) The discovery and probable phylogenetic significance of Lathrobium (Palaeolobrathium) whitei, a new subgenus and species of the Paederinae (Col., Staphylinidae) from Baltic amber. The Entomologist's Monthly Magazine, 104, 1 - 3, pl. 1." type="journal article" year="1968">Abdullah &amp; Qadri, 1968</bibRefCitation>
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<emphasis id="B94AEA8CB9A8FF19C1E36CD4AF349C7A" box="[151,335,1255,1280]" italics="true" pageId="220" pageNumber="221">Euceratobledius</emphasis>
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Olivier, 1811
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<emphasis id="B94AEA8CB9A8FF19C1E36D06AF419C34" box="[151,314,1333,1358]" italics="true" pageId="220" pageNumber="221">Hesperophilus</emphasis>
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36D6FACCF9C0F" box="[423,692,1372,1397]" italics="true" pageId="220" pageNumber="221">Staphylinus fracticornis</emphasis>
Paykull, 1790
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<emphasis id="B94AEA8CB9A8FF19C1BB6DB0AF459CE6" box="[207,318,1411,1436]" italics="true" pageId="220" pageNumber="221">Blediodes</emphasis>
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<treatmentCitation id="0A9F108FB9A8FF19C2386DB7AA569CE6" author="Herman, L. H." box="[844,1069,1411,1436]" page="375" pageId="220" pageNumber="221" year="1970">
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36D99ACCF9CB9" box="[423,692,1450,1475]" italics="true" pageId="220" pageNumber="221">Staphylinus fracticornis</emphasis>
Paykull, 1790
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<emphasis id="B94AEA8CB9A8FF19C1BB6DE2AF4B9C90" box="[207,304,1489,1514]" italics="true" pageId="220" pageNumber="221">Tadunus</emphasis>
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, 147 [synonymized by
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36DCBACCF9F6A" box="[423,692,1528,1553]" italics="true" pageId="220" pageNumber="221">Staphylinus fracticornis</emphasis>
Paykull, 1790
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<emphasis id="B94AEA8CB9A8FF19C1E36E2CAF239F42" box="[151,344,1567,1592]" italics="true" pageId="220" pageNumber="221">Psamathobledius</emphasis>
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36E75ACD39F24" box="[423,680,1606,1631]" italics="true" pageId="220" pageNumber="221">Bledius punctatissimus</emphasis>
LeConte, 1877
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<emphasis id="B94AEA8CB9A8FF19C1E36E5DAE889FFF" box="[151,243,1646,1669]" italics="true" pageId="220" pageNumber="221">Pucerus</emphasis>
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species:
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<emphasis id="B94AEA8CB9A8FF19C0D36EA7AC339FD6" box="[423,584,1684,1709]" italics="true" pageId="220" pageNumber="221">Bledius verres</emphasis>
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<emphasis id="B94AEA8CB9A8FF19C1E36E88AF489FAE" box="[151,307,1723,1748]" italics="true" pageId="220" pageNumber="221">Teratobledius</emphasis>
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<typeStatus id="5485883CB9A8FF19C0736ED0AF3A9F81" box="[263,321,1763,1787]" pageId="220" pageNumber="221">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A8FF19C0D36ED1ACA89F81" authority="Heer, 1839" authorityName="Heer" authorityYear="1839" box="[423,723,1762,1787]" class="Insecta" family="Staphylinidae" genus="Bledius" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B94AEA8CB9A8FF19C0D36ED1AC2B9F81" box="[423,592,1762,1787]" italics="true" pageId="220" pageNumber="221">Bledius tibialis</emphasis>
<bibRefCitation id="EFAF4B6FB9A8FF19C3236ED0ACA89F81" author="Heer, O." box="[599,723,1763,1787]" pageId="220" pageNumber="221" pagination="145 - 360" refId="ref198997" refString="Heer, O. (1839) Fauna coleopterorum Helvetica. Pars I, fasc. 2. Orelii, Fuesslini et Sociorum, Turici [Zurich], pp. 145 - 360." type="book chapter" year="1839">Heer, 1839</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9A8FF19C1E36F3AAF8E9E0A" blockId="220.[151,1437,436,2021]" pageId="220" pageNumber="221">71 species are included in this genus from Latin America. It is one of the super-large genera of the subfamily, its taxonomy is resolved only to species groups, keys to species level mostly missing [György Makranczy, personal communication].</paragraph>
<paragraph id="8B81369EB9A8FF18C1E36F4DAFDC99A4" blockId="220.[151,1437,436,2021]" lastBlockId="221.[151,1437,158,729]" lastPageId="221" lastPageNumber="222" pageId="220" pageNumber="221">
Characters/Description:
<bibRefCitation id="EFAF4B6FB9A8FF19C0DC6F4DAC8C9EED" author="Lynch-Arribalzaga, F." box="[424,759,1918,1943]" pageId="220" pageNumber="221" pagination="5 - 392" refId="ref214921" refString="Lynch-Arribalzaga, F. (1884) Estafilinos de Buenos Aires. Boletin de la Academia Nacional de Ciencias en Cordoba, 7 (1 - 3), 5 - 392." type="journal article" year="1884">Lynch-Arribálzaga 1884: 351</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB9A8FF19C2E66F4DAAF19EED" author="Blackwelder, R. E." box="[914,1162,1918,1943]" pageId="220" pageNumber="221" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 57</bibRefCitation>
(characters in key), 112 (characters);
<bibRefCitation id="EFAF4B6FB9A8FF19C0176F96ACF19EC4" author="Coiffait, H. &amp; Saiz, F." box="[355,650,1957,1982]" pageId="220" pageNumber="221" pagination="339 - 468" refId="ref187580" refString="Coiffait, H. &amp; Saiz, F. (1968) Les Staphylinidae (sensu lato) du Chili. In: Deboutteville, C. &amp; Rapoport, E. (Eds.), Biologie de L'Amerique Australe. Volume IV. Etudes sur la faune du sol. Documents Biogeographiques et Ecologiques. Centre National de la Recherche Scientifique, Paris, pp. 339 - 468." type="book chapter" year="1968">Coiffait &amp; Sáiz 1968: 415</bibRefCitation>
(characters in key), 423 (characters);
<bibRefCitation id="EFAF4B6FB9A8FF19C54D6F95AB629EC4" author="Herman, L. H." box="[1081,1305,1957,1982]" pageId="220" pageNumber="221" pagination="343 - 454" refId="ref199309" refString="Herman, L. H. (1970 a) Phylogeny and reclassification of the genera of the rove-beetle subfamily Oxytelinae of the World (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History, 142 (5), 343 - 454." type="journal article" year="1970">Herman 1970a: 361</bibRefCitation>
(characters in key), 375 (characters);
<bibRefCitation id="EFAF4B6FB9A8FF19C09A6FFEACB79E9F" author="Herman, L. H." box="[494,716,1996,2021]" pageId="220" pageNumber="221" pagination="111 - 254" refId="ref199394" refString="Herman, L. H. (1972 a) Revision of Bledius and related genera. Part I. The aequatorialis, mandibularis, and semiferrugineus groups and two new genera (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the American Museum of Natural History, 149 (2), 111 - 254." type="journal article" year="1972">Herman 1972a: 153</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB9A8FF19C2116FFFAA639E9F" author="Saiz, F." box="[869,1048,1996,2021]" pageId="220" pageNumber="221" pagination="225 - 236" refId="ref231825" refString="Saiz, F. (1973 c) Bledius y Thinobius de Chile, con observaciones generales sobre el edeago (II. Contribucion). Anales del Museo de Historia Natural de Valparaiso, 6, 225 - 236." type="journal article" year="1973">Sáiz 1973c: 229</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB9A8FF19C5C56FFFABE79E9E" author="Navarrete-Heredia, J. L. &amp; Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[1201,1436,1996,2021]" pageId="220" pageNumber="221" refId="ref220178" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Universidad de Guadalajara y CONABIO, Mexico. xii + 401 pp." type="book" year="2002">
Navarrete-Heredia
<emphasis id="B94AEA8CB9A8FF19C4FC6FFEABE79E9E" box="[1416,1436,1997,2020]" italics="true" pageId="220" pageNumber="221">et</emphasis>
</bibRefCitation>
<emphasis id="B94AEA8CB9A9FF18C1BB68ADAE9799CD" box="[207,236,158,183]" italics="true" pageId="221" pageNumber="222">al.</emphasis>
2002: 230 (characters in key);
<bibRefCitation id="EFAF4B6FB9A9FF18C31268ADAD1D99CD" author="Makranczy, G." box="[614,870,158,183]" pageId="221" pageNumber="222" pagination="29 - 119" refId="ref215457" refString="Makranczy, G. (2006) Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici, 98, 29 - 119." type="journal article" year="2006">Makranczy 2006: 103</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB9A9FF18C52B68ACABE799CD" author="Newton, A. F. &amp; Peck, S. B." box="[1119,1436,158,183]" pageId="221" pageNumber="222" pagination="131 - 158" refId="ref220547" refString="Newton, A. F. &amp; Peck, S. B. (2006) Family Staphylinidae, the rove beetles. In: Peck, S. B. (Ed.), The beetles of the Galapagos Islands, Ecuador: evolution, ecology, and diversity (Insecta: Coleoptera). NRC Research Press, Ottawa, pp. 131 - 158." type="book chapter" year="2006">Newton &amp; Peck 2006: 145</bibRefCitation>
(characters in key).
</paragraph>
<paragraph id="8B81369EB9A9FF18C1E368DEAA1198B2" blockId="221.[151,1437,158,729]" pageId="221" pageNumber="222">
Key:
<bibRefCitation id="EFAF4B6FB9A9FF18C1A068DFAC5A987F" author="Lynch-Arribalzaga, F." box="[212,545,236,261]" pageId="221" pageNumber="222" pagination="5 - 392" refId="ref214921" refString="Lynch-Arribalzaga, F. (1884) Estafilinos de Buenos Aires. Boletin de la Academia Nacional de Ciencias en Cordoba, 7 (1 - 3), 5 - 392." type="journal article" year="1884">Lynch-Arribálzaga 1884: 353</bibRefCitation>
(species of
<collectingRegion id="49FAF87CB9A9FF18C3DD68DEADD7987F" box="[681,940,236,261]" country="Argentina" name="Buenos Aires" pageId="221" pageNumber="222">Buenos Aires province</collectingRegion>
,
<collectingCountry id="F329760EB9A9FF18C2CC68DFAA56987F" box="[952,1069,236,261]" name="Argentina" pageId="221" pageNumber="222">Argentina</collectingCountry>
);
<bibRefCitation id="EFAF4B6FB9A9FF18C53468DFAB40987F" author="Bernhauer, M." box="[1088,1339,236,261]" pageId="221" pageNumber="222" pagination="229 - 264" refId="ref172508" refString="Bernhauer, M. (1927 b) Zur Staphylinidenfauna Sudamerikas, insbesondere Argentiniens (31. Beitrag). Archiv fur Naturgeschichte, 91 A (12), 229 - 264. [Note: nominal year of publication 1925]" type="journal article" year="1927">Bernhauer 1927b: 232</bibRefCitation>
(species of the
<collectingCountry id="F329760EB9A9FF18C06F6920AFEB9856" box="[283,400,275,300]" name="Argentina" pageId="221" pageNumber="222">Argentina</collectingCountry>
);
<bibRefCitation id="EFAF4B6FB9A9FF18C0D16920ACD49851" author="Blackwelder, R. E." box="[421,687,275,300]" pageId="221" pageNumber="222" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 113</bibRefCitation>
(species of the West Indies);
<bibRefCitation id="EFAF4B6FB9A9FF18C5736920AB499851" author="Coiffait, H. &amp; Saiz, F." box="[1031,1330,275,300]" pageId="221" pageNumber="222" pagination="339 - 468" refId="ref187580" refString="Coiffait, H. &amp; Saiz, F. (1968) Les Staphylinidae (sensu lato) du Chili. In: Deboutteville, C. &amp; Rapoport, E. (Eds.), Biologie de L'Amerique Australe. Volume IV. Etudes sur la faune du sol. Documents Biogeographiques et Ecologiques. Centre National de la Recherche Scientifique, Paris, pp. 339 - 468." type="book chapter" year="1968">Coiffait &amp; Sáiz 1968: 424</bibRefCitation>
(Chilean species);
<bibRefCitation id="EFAF4B6FB9A9FF18C04C6908AC6D9829" author="Herman, L. H." box="[312,534,314,339]" pageId="221" pageNumber="222" pagination="111 - 254" refId="ref199394" refString="Herman, L. H. (1972 a) Revision of Bledius and related genera. Part I. The aequatorialis, mandibularis, and semiferrugineus groups and two new genera (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the American Museum of Natural History, 149 (2), 111 - 254." type="journal article" year="1972">Herman 1972a: 154</bibRefCitation>
(species groups of the genus);
<bibRefCitation id="EFAF4B6FB9A9FF18C2056909AA5E9828" author="Saiz, F." box="[881,1061,314,339]" pageId="221" pageNumber="222" pagination="225 - 236" refId="ref231825" refString="Saiz, F. (1973 c) Bledius y Thinobius de Chile, con observaciones generales sobre el edeago (II. Contribucion). Anales del Museo de Historia Natural de Valparaiso, 6, 225 - 236." type="journal article" year="1973">Sáiz 1973c: 229</bibRefCitation>
(Chilean species);
<bibRefCitation id="EFAF4B6FB9A9FF18C58F6908AE909803" author="Herman, L. H." pageId="221" pageNumber="222" pagination="71 - 172" refId="ref199531" refString="Herman, L. H. (1976) Revision of Bledius and related genera part II, The armatus, basalis, and melanocephalus groups (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the American Museum of Natural History, 157 (2), 71 - 172." type="journal article" year="1976">Herman 1976: 77</bibRefCitation>
(species groups of the genus);
<bibRefCitation id="EFAF4B6FB9A9FF18C3396951AD7E9803" author="Herman, L. H." box="[589,773,353,377]" pageId="221" pageNumber="222" pagination="1 - 145" refId="ref199665" refString="Herman, L. H. (1983) Revision of Bledius. Part III. The annularis and e marginatus groups (Coleoptera, Oxytelinae). Bulletin of the American Museum of Natural History, 175 (1), 1 - 145." type="journal article" year="1983">Herman 1983: 8</bibRefCitation>
(species groups of the genus);
<bibRefCitation id="EFAF4B6FB9A9FF18C5126951AB419803" author="Herman, L. H." box="[1126,1338,353,378]" pageId="221" pageNumber="222" pagination="1 - 368" refId="ref199711" refString="Herman, L. H. (1986) Revision of Bledius. Part IV. Classification of species groups, phylogeny, natural history, and catalogue (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the American Museum of Natural History, 184 (1), 1 - 368." type="journal article" year="1986">Herman 1986: 107</bibRefCitation>
(species groups);
<bibRefCitation id="EFAF4B6FB9A9FF18C04E69BAAC0798DB" author="Newton, A. F. &amp; Peck, S. B." box="[314,636,392,417]" pageId="221" pageNumber="222" pagination="131 - 158" refId="ref220547" refString="Newton, A. F. &amp; Peck, S. B. (2006) Family Staphylinidae, the rove beetles. In: Peck, S. B. (Ed.), The beetles of the Galapagos Islands, Ecuador: evolution, ecology, and diversity (Insecta: Coleoptera). NRC Research Press, Ottawa, pp. 131 - 158." type="book chapter" year="2006">Newton &amp; Peck 2006: 145</bibRefCitation>
(species of the Galapagos Isl.);
<bibRefCitation id="EFAF4B6FB9A9FF18C57369BBABE798DB" author="Caron, E. &amp; Ribeiro-Costa, C. S." box="[1031,1436,392,417]" pageId="221" pageNumber="222" pagination="452 - 457" refId="ref181975" refString="Caron, E. &amp; Ribeiro-Costa, C. S. (2007) Bledius Leach from southern Brazil (Coleoptera, Staphylinidae, Oxytelinae). Revista Brasileira de Entomologia, 51 (4), 452 - 457. https: // doi. org / 10.1590 / s 0085 - 56262007000400008" type="journal article" year="2007">Caron &amp; Ribeiro-Costa 2007: 453</bibRefCitation>
(seashore of southern
<collectingCountry id="F329760EB9A9FF18C0B2699CAC7498B2" box="[454,527,431,456]" name="Brazil" pageId="221" pageNumber="222">Brazil</collectingCountry>
);
<bibRefCitation id="EFAF4B6FB9A9FF18C357699CAD5998B2" author="Castro, J. C. de &amp; Caron, E. &amp; Rosa, L. C. da" box="[547,802,431,456]" pageId="221" pageNumber="222" pagination="145 - 157" refId="ref183134" refString="Castro, J. C. de, Caron, E. &amp; Rosa, L. C. da. (2016) Update on the Brazilian coastal species of Bledius Leach (Coleoptera: Staphylinidae: Oxytelinae) with the description of two new species. Zootaxa, 4111 (2), 145 - 157. https: // doi. org / 10.11646 / zootaxa. 4111.2.3" type="journal article" year="2016">
Castro
<emphasis id="B94AEA8CB9A9FF18C3076983ACD098B2" box="[627,683,431,456]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
2016: 146
</bibRefCitation>
(Brazilian seashore species).
</paragraph>
<paragraph id="8B81369EB9A9FF18C1E369E5AF5E9894" blockId="221.[151,1437,158,729]" box="[151,293,470,495]" pageId="221" pageNumber="222">Distribution.</paragraph>
<paragraph id="8B81369EB9A9FF18C1DE69CDAAF29B47" blockId="221.[151,1437,158,729]" pageId="221" pageNumber="222">
West Indies:
<collectingCountry id="F329760EB9A9FF18C03169CEAFD49B6C" box="[325,431,509,534]" name="Bahamas" pageId="221" pageNumber="222">Bahamas</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C0CA69CEAF879B6C" box="[446,508,509,534]" name="Cuba" pageId="221" pageNumber="222">Cuba</collectingCountry>
, Cayman Is.,
<collectingCountry id="F329760EB9A9FF18C3D469CEAC869B6C" box="[672,765,509,534]" name="Jamaica" pageId="221" pageNumber="222">Jamaica</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C27869CEAD3C9B6C" box="[780,839,509,534]" name="Haiti" pageId="221" pageNumber="222">Haiti</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C22269CEAA3D9B6C" box="[854,1094,509,534]" name="Dominican Republic" pageId="221" pageNumber="222">Dominican Republic</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C52169CDAA9A9B6C" box="[1109,1249,509,534]" name="Puerto Rico" pageId="221" pageNumber="222">Puerto Rico</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C59B69CDAB689B6C" box="[1263,1299,510,534]" name="United States of America" pageId="221" pageNumber="222">US</collectingCountry>
Virgin Isl.,
<collectingCountry id="F329760EB9A9FF18C1BB6A17AFD49B47" box="[207,431,548,573]" name="Antigua and Barbuda" pageId="221" pageNumber="222">Antigua &amp; Barbuda</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C0CE6A16AC439B46" box="[442,568,549,572]" name="Montserrat" pageId="221" pageNumber="222">Montserrat</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C3376A17ACB69B47" box="[579,717,548,573]" name="Guadeloupe" pageId="221" pageNumber="222">Guadeloupe</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C3AC6A17AD329B47" box="[728,841,548,573]" name="Dominica" pageId="221" pageNumber="222">Dominica</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C2216A17ADCC9B47" box="[853,951,548,573]" name="Grenada" pageId="221" pageNumber="222">Grenada</collectingCountry>
, Trinidad-Tobago.
</paragraph>
<paragraph id="8B81369EB9A9FF18C1DE6A78AD3D9B19" blockId="221.[151,1437,158,729]" box="[170,838,587,612]" pageId="221" pageNumber="222">
Central America:
<collectingCountry id="F329760EB9A9FF18C0066A78AFB19B1E" box="[370,458,587,612]" name="Mexico" pageId="221" pageNumber="222">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C0A26A78AC299B1E" box="[470,594,587,612]" name="Guatemala" pageId="221" pageNumber="222">Guatemala</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C3296A78ACA19B1E" box="[605,730,587,612]" name="Costa Rica" pageId="221" pageNumber="222">Costa Rica</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C3916A7FAD3A9B19" box="[741,833,588,611]" name="Panama" pageId="221" pageNumber="222">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A9FF18C1DE6A41AFE99BC8" blockId="221.[151,1437,158,729]" pageId="221" pageNumber="222">
South America:
<collectingCountry id="F329760EB9A9FF18C0106A41AFAE9BF1" box="[356,469,626,651]" name="Colombia" pageId="221" pageNumber="222">Colombia</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C0966A41AC209BF1" box="[482,603,626,651]" name="Venezuela" pageId="221" pageNumber="222">Venezuela</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C3136A41AD6A9BF1" box="[615,785,626,651]" name="French Guiana" pageId="221" pageNumber="222">French Guiana</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C26A6A41AD059BF1" box="[798,894,626,651]" name="Ecuador" pageId="221" pageNumber="222">Ecuador</collectingCountry>
,
<collectingRegion id="49FAF87CB9A9FF18C2FE6A41AA7B9BF1" box="[906,1024,626,651]" country="Ecuador" name="Galapagos" pageId="221" pageNumber="222">Galapagos</collectingRegion>
Isl.,
<collectingCountry id="F329760EB9A9FF18C54F6A40AA0A9BF0" box="[1083,1137,627,650]" name="Peru" pageId="221" pageNumber="222">Peru</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C50A6A41AAA89BF1" box="[1150,1235,626,651]" name="Bolivia" pageId="221" pageNumber="222">Bolivia</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C5AB6A41AB5D9BF1" box="[1247,1318,626,651]" name="Brazil" pageId="221" pageNumber="222">Brazil</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C4476A40ABE39BF1" box="[1331,1432,627,651]" name="Uruguay" pageId="221" pageNumber="222">Uruguay</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C1BB6AAAAF749BC8" box="[207,271,665,690]" name="Chile" pageId="221" pageNumber="222">Chile</collectingCountry>
,
<collectingCountry id="F329760EB9A9FF18C06E6AAAAFF59BC8" box="[282,398,665,690]" name="Argentina" pageId="221" pageNumber="222">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A9FF18C1DE6AF3AAD79BA3" blockId="221.[151,1437,158,729]" box="[170,1196,704,729]" pageId="221" pageNumber="222">
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan,
<collectingRegion id="49FAF87CB9A9FF18C2286AF3ADC19BA3" box="[860,954,704,729]" country="Republic of the Congo" name="Orientale" pageId="221" pageNumber="222">Oriental</collectingRegion>
, Australian, Oceanic.
</paragraph>
</subSubSection>
</treatment>
</document>
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