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<mods:title id="D9EBC02DAC076806CCFE8A59C1FD5FE4">Geographic distribution of the hard ticks (Acari: Ixodida: Ixodidae) of the world by countries and territories</mods:title>
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<mods:namePart id="D5A214B014E9D68F5BA30A05E56804A4">Nava, Santiago</mods:namePart>
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<treatment id="03966A560F46C746BABF8FA9B03BF91D" ID-DOI="http://doi.org/10.5281/zenodo.7717670" ID-GBIF-Taxon="209449642" ID-Zenodo-Dep="7717670" LSID="urn:lsid:plazi:treatment:03966A560F46C746BABF8FA9B03BF91D" httpUri="http://treatment.plazi.org/id/03966A560F46C746BABF8FA9B03BF91D" lastPageNumber="46" pageId="45" pageNumber="46">
<subSubSection id="C32588CB0F46C746BABF8FA9B7D0FB79" box="[151,702,1178,1205]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8B80DB400F46C746BABF8FA9B7D0FB79" blockId="45.[151,1437,1178,1745]" box="[151,702,1178,1205]" pageId="45" pageNumber="46">
<emphasis id="B94B07520F46C746BABF8FA9B5D4FB79" bold="true" box="[151,186,1179,1205]" italics="true" pageId="45" pageNumber="46">41.</emphasis>
<taxonomicName id="4C3FA0C30F46C746BAE88FA8B7D4FB79" ID-CoL="CMDF" ID-ENA="659692" authority="(Lucas, 1846)" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[192,698,1178,1205]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">
<emphasis id="B94B07520F46C746BAE88FA8B77BFB79" bold="true" box="[192,533,1178,1205]" italics="true" pageId="45" pageNumber="46">Amblyomma flavomaculatum</emphasis>
(
<bibRefCitation id="EFAEA6B10F46C746B80B8FA9B7DFFB79" author="Lucas, H." box="[547,689,1179,1205]" pageId="45" pageNumber="46" pagination="53 - 64" refId="ref226806" refString="Lucas, H. (1846) Sur quelques especes nouvelles d' Ixodes qui vivent parasites sur les serpens (Boa constrictor et Python sebae) et sur l'ornythorhynque (Ornithorhynchus paradoxus, Blum). Annales de la Societe Entomologique de France, 4, 53 - 64." type="journal article" year="1846">Lucas, 1846</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32588CB0F46C746B8EB8FA9B672FAA5" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8B80DB400F46C746B8EB8FA9B672FAA5" blockId="45.[151,1437,1178,1745]" pageId="45" pageNumber="46">
Afrotropical: 1)
<collectingCountry id="F3289BD00F46C746B9528FA9B6AEFB79" box="[890,960,1179,1205]" name="Benin" pageId="45" pageNumber="46">Benin</collectingCountry>
, 2)
<collectingCountry id="F3289BD00F46C746B9C08FA9B1EFFB79" box="[1000,1153,1179,1205]" name="Burkina Faso" pageId="45" pageNumber="46">Burkina Faso</collectingCountry>
, 3)
<collectingCountry id="F3289BD00F46C746BE828FA9B04CFB79" box="[1194,1314,1179,1205]" name="Cameroon" pageId="45" pageNumber="46">Cameroon</collectingCountry>
, 4)
<collectingCountry id="F3289BD00F46C746BF628FA9B430FB15" name="Central African Republic" pageId="45" pageNumber="46">Central African Republic</collectingCountry>
, 5)
<collectingCountry id="F3289BD00F46C746BBA38F8DB4A8FB15" box="[395,454,1215,1241]" name="Chad" pageId="45" pageNumber="46">Chad</collectingCountry>
(south), 6)
<collectingCountry id="F3289BD00F46C746B8668F8DB68EFB15" box="[590,992,1215,1241]" name="Democratic Republic of the Congo" pageId="45" pageNumber="46">Democratic Republic of the Congo</collectingCountry>
, 7)
<collectingCountry id="F3289BD00F46C746BE268F8DB11EFB15" box="[1038,1136,1215,1241]" name="Ethiopia" pageId="45" pageNumber="46">Ethiopia</collectingCountry>
, 8)
<collectingCountry id="F3289BD00F46C746BEB58F8DB187FB15" box="[1181,1257,1215,1241]" name="Ghana" pageId="45" pageNumber="46">Ghana</collectingCountry>
, 9)
<collectingCountry id="F3289BD00F46C746BF3E8F8DB004FB15" box="[1302,1386,1215,1241]" name="Guinea-Bissau" pageId="45" pageNumber="46">Guinea</collectingCountry>
, 10)
<collectingCountry id="F3289BD00F46C746BABF8FD1B451FB31" box="[151,319,1251,1277]" name="Guinea-Bissau" pageId="45" pageNumber="46">Guinea-Bissau</collectingCountry>
, 11)
<collectingCountry id="F3289BD00F46C746BB5E8FD1B493FB31" box="[374,509,1251,1277]" name="Ivory Coast" pageId="45" pageNumber="46">Ivory Coast</collectingCountry>
, 12)
<collectingCountry id="F3289BD00F46C746B81C8FD1B7EEFB31" box="[564,640,1251,1277]" name="Kenya" pageId="45" pageNumber="46">Kenya</collectingCountry>
, 13)
<collectingCountry id="F3289BD00F46C746B8908FD1B783FB31" box="[696,749,1251,1277]" name="Serbia" pageId="45" pageNumber="46">Mali</collectingCountry>
(south), 14)
<collectingCountry id="F3289BD00F46C746B9578FD1B694FB31" box="[895,1018,1251,1277]" name="Mauritania" pageId="45" pageNumber="46">Mauritania</collectingCountry>
(south), 15)
<collectingCountry id="F3289BD00F46C746BEA38FD1B1A5FB31" box="[1163,1227,1251,1277]" name="Niger" pageId="45" pageNumber="46">Niger</collectingCountry>
(south), 16)
<collectingCountry id="F3289BD00F46C746BF748FD1B5D9FAED" name="Nigeria" pageId="45" pageNumber="46">Nigeria</collectingCountry>
, 17)
<collectingCountry id="F3289BD00F46C746BADA8E35B423FAED" box="[242,333,1287,1313]" name="Senegal" pageId="45" pageNumber="46">Senegal</collectingCountry>
, 18)
<collectingCountry id="F3289BD00F46C746BBAF8E35B775FAED" box="[391,539,1287,1313]" name="Sierra Leone" pageId="45" pageNumber="46">Sierra Leone</collectingCountry>
, 19)
<collectingCountry id="F3289BD00F46C746B87D8E35B786FAED" box="[597,744,1287,1313]" name="South Sudan" pageId="45" pageNumber="46">South Sudan</collectingCountry>
, 20)
<collectingCountry id="F3289BD00F46C746B90A8E35B605FAED" box="[802,875,1287,1313]" name="Sudan" pageId="45" pageNumber="46">Sudan</collectingCountry>
, 21)
<collectingCountry id="F3289BD00F46C746B98C8E35B162FAED" box="[932,1036,1287,1313]" name="Tanzania" pageId="45" pageNumber="46">Tanzania</collectingCountry>
, 22)
<collectingCountry id="F3289BD00F46C746BE6D8E35B1EFFAED" box="[1093,1153,1287,1313]" name="Togo" pageId="45" pageNumber="46">Togo</collectingCountry>
, 23)
<collectingCountry id="F3289BD00F46C746BE938E35B07BFAED" box="[1211,1301,1287,1313]" name="Uganda" pageId="45" pageNumber="46">Uganda</collectingCountry>
, 24)
<collectingCountry id="F3289BD00F46C746BF668E35B0F2FAED" box="[1358,1436,1287,1313]" name="Yemen" pageId="45" pageNumber="46">Yemen</collectingCountry>
(
<bibRefCitation id="EFAEA6B10F46C746BAB68E19B4DCFA89" author="Yeoman, G. H. &amp; Walker, J. B." box="[158,434,1323,1349]" pageId="45" pageNumber="46" refId="ref266297" refString="Yeoman, G. H. &amp; Walker, J. B. (1967) The ixodid ticks of Tanzania. A study of the zoogeography of the Ixodidae of an East African country. Commonwealth Institute of Entomology, London, 215 pp." type="book" year="1967">Yeoman &amp; Walker 1967</bibRefCitation>
,
<bibRefCitation id="EFAEA6B10F46C746BB948E19B71DFA89" author="Saratsiotis, A." box="[444,627,1323,1349]" pageId="45" pageNumber="46" pagination="476 - 495" refId="ref251938" refString="Saratsiotis, A. (1972) Contribution a l'etude morphologique et biologique du genre Aponomma Neumann, 1899 (Acariens, Ixodidea) [sic]. I. Ap. flavomaculatum (Lucas, 1846). II. Ap. latum (Koch, 1844). Acarologia, 13, 476 - 495." type="journal article" year="1972">Saratsiotis 1972</bibRefCitation>
,
<bibRefCitation id="EFAEA6B10F46C746B8568E19B64BFA89" author="Keirans, J. E." box="[638,805,1322,1349]" pageId="45" pageNumber="46" pagination="1 - 1785" refId="ref214582" refString="Keirans, J. E. (1985 b) George Henry Falkiner Nuttall and the Nuttall tick catalogue. United States Department of Agriculture, Agricultural Research Service, Miscellaneous Publication, (1438), 1 - 1785. https: // doi. org / 10.5962 / bhl. title. 65326" type="journal article" year="1985">
Keirans 1985
<bibRefCitation id="EFAEA6B10F46C746B93E8E18B64BFA89" author="Walker, J. B. &amp; Olwage, A." box="[790,805,1322,1349]" pageId="45" pageNumber="46" pagination="353 - 379" refId="ref263132" refString="Walker, J. B. &amp; Olwage, A. (1987) The tick vectors of Cowdria ruminantium (Ixodoidea, Ixodidae, genus Amblyomma) and their distribution. Onderstepoort Journal of Veterinary Research, 54, 353 - 379." type="journal article" year="1987">b</bibRefCitation>
</bibRefCitation>
, Jongejan
<emphasis id="B94B07520F46C746B9B28E19B6BCFA89" box="[922,978,1323,1349]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
1987,
<bibRefCitation id="EFAEA6B10F46C746BE368E19B19DFA89" author="Santos Dias, J. A. T." box="[1054,1267,1323,1349]" pageId="45" pageNumber="46" pagination="1 - 204" refId="ref251458" refString="Santos Dias, J. A. T. (1993 b) Contribuic &quot; o para o estudo da sistematica e taxonomia das especies do genero Aponomma Neumann, 1899 (Acarina-Ixodoidea). Estudos, Ensaios e Documentos, (157), 1 - 204." type="journal article" year="1993">
Santos Dias 1993
<bibRefCitation id="EFAEA6B10F46C746BECC8E19B19DFA89" author="Walker, J. B. &amp; Keirans, J. E. &amp; Horak, I. G." box="[1252,1267,1323,1349]" pageId="45" pageNumber="46" refId="ref263510" refString="Walker, J. B., Keirans, J. E. &amp; Horak, I. G. (2000) The genus Rhipicephalus (Acari: Ixodidae): a guide to the brown ticks of the world. Cambridge University Press, Cambridge, 643 pp. https: // doi. org / 10.1017 / CBO 9780511661754" type="book" year="2000">b</bibRefCitation>
</bibRefCitation>
, Terenius
<emphasis id="B94B07520F46C746BF4C8E19B0F3FA89" box="[1380,1437,1323,1349]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
2000,
<bibRefCitation id="EFAEA6B10F46C746BAF58E7DB40AFAA5" author="Morel, P. C." box="[221,356,1359,1385]" pageId="45" pageNumber="46" refId="ref232231" refString="Morel, P. C. (2003) Les tiques d'Afrique et du Bassin mediterraneen (1965 - 1995). CIRAD-EMVT, 1342 pp." type="book" year="2003">Morel 2003</bibRefCitation>
,
<bibRefCitation id="EFAEA6B10F46C746BB478E7DB77DFAA5" author="Burridge, M. J." box="[367,531,1359,1385]" pageId="45" pageNumber="46" refId="ref178986" refString="Burridge, M. J. (2011) Non-Native and Invasive Ticks. Threats to Human and Animal Health in the United States. University of Florida Press, Gainesville, 448 pp." type="book" year="2011">Burridge 2011</bibRefCitation>
,
<bibRefCitation id="EFAEA6B10F46C746B8368E7DB67FFAA5" author="Uilenberg, G. &amp; Estrada-Pena, A. &amp; Thal, J." box="[542,785,1359,1385]" pageId="45" pageNumber="46" pagination="1 - 40" refId="ref260920" refString="Uilenberg, G., Estrada-Pena, A. &amp; Thal, J. (2013) Ticks of the Central African Republic. Experimental and Applied Acarology, 60, 1 - 40. https: // doi. org / 10.1007 / s 10493 - 012 - 9605 - 2" type="journal article" year="2013">
Uilenberg
<emphasis id="B94B07520F46C746B8BD8E7DB7A3FAA5" box="[661,717,1359,1385]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32588CB0F46C746BAEF8E41B03BF91D" pageId="45" pageNumber="46" type="discussion">
<paragraph id="8B80DB400F46C746BAEF8E41B5A6FA7D" blockId="45.[151,1437,1178,1745]" pageId="45" pageNumber="46">
Most records of
<taxonomicName id="4C3FA0C30F46C746BBAD8E41B7A3FA41" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[389,717,1395,1421]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">
<emphasis id="B94B07520F46C746BBAD8E41B7A3FA41" box="[389,717,1395,1421]" italics="true" pageId="45" pageNumber="46">Amblyomma flavomaculatum</emphasis>
</taxonomicName>
have been published under the name
<taxonomicName id="4C3FA0C30F46C746BEA88E41B5AFFA7C" class="Arachnida" family="Ixodidae" genus="Aponomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">
<emphasis id="B94B07520F46C746BEA88E41B5AFFA7C" italics="true" pageId="45" pageNumber="46">Aponomma flavomaculatum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B80DB400F46C746BAEF8E89B153F945" blockId="45.[151,1437,1178,1745]" pageId="45" pageNumber="46">
The geographic distribution of
<taxonomicName id="4C3FA0C30F46C746B8088E89B608FA19" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[544,870,1467,1493]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">
<emphasis id="B94B07520F46C746B8088E89B608FA19" box="[544,870,1467,1493]" italics="true" pageId="45" pageNumber="46">Amblyomma flavomaculatum</emphasis>
</taxonomicName>
is mostly based on
<bibRefCitation id="EFAEA6B10F46C746BE688E89B04DFA19" author="Santos Dias, J. A. T." box="[1088,1315,1467,1493]" pageId="45" pageNumber="46" pagination="1 - 204" refId="ref251458" refString="Santos Dias, J. A. T. (1993 b) Contribuic &quot; o para o estudo da sistematica e taxonomia das especies do genero Aponomma Neumann, 1899 (Acarina-Ixodoidea). Estudos, Ensaios e Documentos, (157), 1 - 204." type="journal article" year="1993">Santos Dias (1993b)</bibRefCitation>
and
<bibRefCitation id="EFAEA6B10F46C746BF708E89B58AFA35" author="Morel, P. C." pageId="45" pageNumber="46" refId="ref232231" refString="Morel, P. C. (2003) Les tiques d'Afrique et du Bassin mediterraneen (1965 - 1995). CIRAD-EMVT, 1342 pp." type="book" year="2003">Morel (2003)</bibRefCitation>
, who regarded
<taxonomicName id="4C3FA0C30F46C746BBBA8EEDB7F3FA34" authorityName="Koch" authorityYear="1844" box="[402,669,1503,1529]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="exornatum">
<emphasis id="B94B07520F46C746BBBA8EEDB7F3FA34" box="[402,669,1503,1529]" italics="true" pageId="45" pageNumber="46">Amblyomma exornatum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3FA0C30F46C746B8848EEDB6CCFA34" authorityName="Karsch" authorityYear="1879" box="[684,930,1503,1529]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="arcanum">
<emphasis id="B94B07520F46C746B8848EEDB6CCFA34" box="[684,930,1503,1529]" italics="true" pageId="45" pageNumber="46">Amblyomma arcanum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3FA0C30F46C746B9F28EEDB04FFA35" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[986,1313,1503,1529]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">
<emphasis id="B94B07520F46C746B9F28EEDB04FFA35" box="[986,1313,1503,1529]" italics="true" pageId="45" pageNumber="46">Amblyomma flavomaculatum</emphasis>
</taxonomicName>
(under the genus
<taxonomicName id="4C3FA0C30F46C746BAF78D31B431F9D1" box="[223,351,1539,1565]" class="Arachnida" family="Ixodidae" genus="Aponomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B94B07520F46C746BAF78D31B431F9D1" box="[223,351,1539,1565]" italics="true" pageId="45" pageNumber="46">Aponomma</emphasis>
</taxonomicName>
) as different species that are difficult to separate morphologically (see also
<taxonomicName id="4C3FA0C30F46C746BE8F8D31B0F3F9D0" authorityName="Karsch" authorityYear="1879" box="[1191,1437,1539,1565]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="arcanum">
<emphasis id="B94B07520F46C746BE8F8D31B0F3F9D0" box="[1191,1437,1539,1565]" italics="true" pageId="45" pageNumber="46">Amblyomma arcanum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3FA0C30F46C746BAE08D15B4BAF98C" authorityName="Koch" authorityYear="1844" box="[200,468,1575,1601]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="exornatum">
<emphasis id="B94B07520F46C746BAE08D15B4BAF98C" box="[200,468,1575,1601]" italics="true" pageId="45" pageNumber="46">Amblyomma exornatum</emphasis>
</taxonomicName>
) and treat records of
<taxonomicName id="4C3FA0C30F46C746B8E28D15B6B8F98C" authorityName="Koch" authorityYear="1844" box="[714,982,1575,1601]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="exornatum">
<emphasis id="B94B07520F46C746B8E28D15B6B8F98C" box="[714,982,1575,1601]" italics="true" pageId="45" pageNumber="46">Amblyomma exornatum</emphasis>
</taxonomicName>
from northern sub-Saharan countries as
<emphasis id="B94B07520F46C746BABF8D79B487F9A9" box="[151,489,1611,1637]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="4C3FA0C30F46C746BABF8D79B48BF9A9" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[151,485,1611,1637]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">Amblyomma flavomaculatum</taxonomicName>
.
</emphasis>
The presence of
<taxonomicName id="4C3FA0C30F46C746B89B8D79B692F9A9" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[691,1020,1611,1637]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">
<emphasis id="B94B07520F46C746B89B8D79B692F9A9" box="[691,1020,1611,1637]" italics="true" pageId="45" pageNumber="46">Amblyomma flavomaculatum</emphasis>
</taxonomicName>
in
<collectingCountry id="F3289BD00F46C746BE0D8D79B11DF9A9" box="[1061,1139,1611,1637]" name="Yemen" pageId="45" pageNumber="46">Yemen</collectingCountry>
is based on B ö hme
<emphasis id="B94B07520F46C746BF4A8D79B0F3F9A9" box="[1378,1437,1611,1637]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
(1989) and
<bibRefCitation id="EFAEA6B10F46C746BB308D5DB495F945" author="Wassef, H. I. &amp; Gallagher, M. D." box="[280,507,1647,1673]" pageId="45" pageNumber="46" pagination="63 - 88" refId="ref264487" refString="Wassef, H. I., B ¸ ttiker, W. &amp; Gallagher, M. D. (1997) Further records of ticks (Acari: Argasidae and Ixodidae) from the Arab Peninsula. Fauna of Saudi Arabia, 16, 63 - 88." type="journal article" year="1997">
Wassef
<emphasis id="B94B07520F46C746BB478D5DB4C9F945" box="[367,423,1647,1673]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
(1997)
</bibRefCitation>
, records that are treated as provisionally valid here.
</paragraph>
<paragraph id="8B80DB400F46C746BAEF8DA1B03BF91D" blockId="45.[151,1437,1178,1745]" pageId="45" pageNumber="46">
Several records of
<taxonomicName id="4C3FA0C30F46C746BBB58DA1B78BF961" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[413,741,1683,1709]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="flavomaculatum">
<emphasis id="B94B07520F46C746BBB58DA1B78BF961" box="[413,741,1683,1709]" italics="true" pageId="45" pageNumber="46">Amblyomma flavomaculatum</emphasis>
</taxonomicName>
collected outside the Afrotropical Zoogeographic
<collectingRegion id="49FB15A20F46C746BF098DA1B01CF961" box="[1313,1394,1683,1709]" country="Iraq" pageId="45" pageNumber="46">Region</collectingRegion>
are from imported hosts, but there is no indication that this species has established itself outside the Afrotropics.
</paragraph>
</subSubSection>
</treatment>
</document>