440 lines
56 KiB
XML
440 lines
56 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6485252" ID-GBIF-Dataset="3dfc64e7-8c23-456d-9678-1bb0858ac470" ID-GBIF-Taxon="150645358" ID-Zenodo-Dep="6485252" checkinTime="1545431109525" checkinUser="PlaziZenodoSync" docAuthor="Jan van Tol & André Günther" docDate="2018" docId="0395A91CAC78FF8895EEFC53FB08FDD3" docLanguage="en" docName="10.5281zenodo.1481114_vanTol_Guenther_2018.pdf" docOrigin="Odonatologica 47 (3)" docStyle="DocumentStyle{}" docTitle="Rhinocypha Rambur 1842" docType="treatment" docVersion="4" lastPageId="14" lastPageNumber="313" masterDocId="FFACD164AC71FF869443FFAFFF05FFBC" masterDocTitle="The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)" masterLastPageNumber="386" masterPageNumber="299" originalUpdateDomain="Boston" originalUpdateTime="1544520335069" originalUpdateUser="PlaziZenodoSync" pageId="9" pageNumber="308" updateTime="1650928538772" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jan van Tol</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>André Günther</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>47</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>299</mods:start>
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<mods:end>386</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.1481114</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3dfc64e7-8c23-456d-9678-1bb0858ac470</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1481114</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6485252" ID-GBIF-Taxon="150645358" ID-Zenodo-Dep="6485252" LSID="urn:lsid:plazi:treatment:0395A91CAC78FF8895EEFC53FB08FDD3" httpUri="http://treatment.plazi.org/id/0395A91CAC78FF8895EEFC53FB08FDD3" lastPageId="14" lastPageNumber="313" pageId="9" pageNumber="308">
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<subSubSection box="[429,766,1020,1048]" pageId="9" pageNumber="308" type="nomenclature">
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<paragraph blockId="9.[159,1037,983,1552]" box="[429,766,1020,1048]" pageId="9" pageNumber="308">
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<heading bold="true" box="[429,766,1020,1048]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="308" reason="4">
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<emphasis bold="true" box="[429,766,1020,1048]" pageId="9" pageNumber="308">
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Genus:
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<taxonomicName authority="Rambur" authorityName="Rambur" authorityYear="1842" box="[520,766,1020,1048]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="308" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[520,662,1020,1048]" italics="true" pageId="9" pageNumber="308">Rhinocypha</emphasis>
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Rambur
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="309" pageId="9" pageNumber="308" type="discussion">
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<paragraph blockId="9.[159,1037,983,1552]" box="[159,390,1066,1092]" pageId="9" pageNumber="308">
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<heading bold="true" box="[159,390,1066,1092]" fontSize="11" level="3" pageId="9" pageNumber="308" reason="6">
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<emphasis bold="true" box="[159,390,1066,1092]" pageId="9" pageNumber="308">Selected references</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="9.[159,1037,983,1552]" lastBlockId="10.[181,1037,253,369]" lastPageId="10" lastPageNumber="309" pageId="9" pageNumber="308">
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[159,283,1099,1123]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="308" phylum="Arthropoda" rank="genus">
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<emphasis box="[159,283,1099,1123]" italics="true" pageId="9" pageNumber="308">Rhinocypha</emphasis>
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</taxonomicName>
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. — RAMBUR (1842): 232–233 (original diagnosis; type species:
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<emphasis italics="true" pageId="9" pageNumber="308">
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<taxonomicName authorityName="Rambur" authorityYear="1842" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="308" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
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tincta
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</emphasis>
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Rambur [designated by KIRBY 1890: 114]) — SELYS (1853): sep. 59–64 (revision, downgraded to subgenus, new species described); SELYS (1854): 198–225 (monograph); SELYS (1869): sep. 18–21 (additions to revision); SELYS (1873a): 487–493, 507–519 (sep. 23–29, 43–55) (additions to revision, table with classification); SELYS (1873b): 614–615 (sep. 61–62) (descriptions of
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<emphasis italics="true" pageId="9" pageNumber="308">
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[911,1035,1252,1276]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="308" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
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monochroa
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</emphasis>
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and
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<emphasis box="[354,493,1283,1307]" italics="true" pageId="9" pageNumber="308">R. albistigma</emphasis>
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); SELYS (1879): sep. 39–51 (additions to revision); SELYS (1882): 21–22 (Philippines); KIRBY (1890): 112–114 (upgraded to genus; species included, selection of type species); SELYS (1891): 213–216 (Philippines, discussion
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<taxonomicName box="[299,436,1375,1398]" pageId="9" pageNumber="308">
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<emphasis box="[299,436,1375,1398]" italics="true" pageId="9" pageNumber="308">R. semitincta</emphasis>
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</taxonomicName>
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); RIS (1915): 87–89 (New Guinea); RIS (1916): 310–313 (revision species Celebes, key); FRASER (1926): 484 (
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<emphasis box="[753,844,1406,1429]" italics="true" pageId="9" pageNumber="308">R. tincta</emphasis>
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type locality Java [in error]); LIEFTINCK (1935): 175–177 (
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[622,746,1436,1460]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="308" phylum="Arthropoda" rank="genus">
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<emphasis box="[622,746,1436,1460]" italics="true" pageId="9" pageNumber="308">Rhinocypha</emphasis>
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</taxonomicName>
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species of Celebes); COW- LEY (1937): 1–18 (penis of
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<taxonomicName authorityName="" authorityYear="1937" box="[466,641,1467,1491]" class="Insecta" family="Chlorocyphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="308" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
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, group character); LIEFTINCK (1938): 49–70 (species of New Guinea described and illustrated); FRASER (1949): 10–16, figs. 1(4) (
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<taxonomicName authorityName="" authorityYear="1937" box="[285,461,1528,1552]" class="Insecta" family="Chlorocyphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="308" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
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revised, diagnoses of genera, key to genera); LAIdLAW (1950): 257–280 (classification of
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<taxonomicName authorityName="" authorityYear="1937" box="[547,722,253,277]" class="Insecta" family="Chlorocyphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
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, series
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[801,925,253,277]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[801,925,253,277]" italics="true" pageId="10" pageNumber="309">Rhinocypha</emphasis>
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</taxonomicName>
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subdivid- ed, new genera erected); FRASER (1957): 70–74 (classification of
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<taxonomicName authorityName="" authorityYear="1937" box="[858,1033,284,308]" class="Insecta" family="Chlorocyphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
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, with key to genera); VAN TOL & GASSMANN (2007): 66–67, fig. 10 (
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<emphasis italics="true" pageId="10" pageNumber="309">
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[911,1035,314,338]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
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tincta
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</emphasis>
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group discussed); MICHALSKI (2012): 73–79 (summary taxa New Guinea).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="309" type="diagnosis">
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<paragraph blockId="10.[159,1037,405,1551]" box="[159,281,405,431]" pageId="10" pageNumber="309">
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<heading bold="true" box="[159,281,405,431]" fontSize="11" level="3" pageId="10" pageNumber="309" reason="6">
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<emphasis bold="true" box="[159,281,405,431]" pageId="10" pageNumber="309">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="10.[159,1037,405,1551]" box="[159,920,442,469]" pageId="10" pageNumber="309">Here we repeat the diagnosis as provided by FRASER (1949: 10 f.):</paragraph>
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<paragraph blockId="10.[159,1037,405,1551]" pageId="10" pageNumber="309">»Species varying from moderately large to rather small and slim stature. Wings of males, with few exceptions, coloured and marked with brilliant iridescent, vitreous and opaque fasciae or spot, the opaque areas often with hyaline vitreous spots or striae. Wings of females colourless or rarely with opaque tips, or, in old adults, often assuming an uniform brownish or yel- lowish tint. Shape of wings of males very variable, very narrow and elongate or broader, the hind wings often very broad and imparting an anisopterous character to the insects. Mesothoracic triangle variable, small and incon- spicuous or extending the whole length of dorsum and markedly dilated below. Body markings usually restricted and not ordinarily conspicuous, but occasionally blue, greenish yellow or even brick red on the male abdomen. MA entirely straight throughout its course and rather closely parallel with CuP to as far as wing margin: at least 2 intercalated veins between R2 and R3, the longer of which extends inwards to proximal of the pterostigma. Anal vein and CuP extending well beyond level of nodus; antenodals very numerous and always one or more secondaries between the two primaries.«</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="310" pageId="10" pageNumber="309" type="discussion">
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<paragraph blockId="10.[159,1037,405,1551]" pageId="10" pageNumber="309">
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LAIdLAW (1950: 270 ff.) split the genus as defined by Fraser, and limited the genus
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[281,418,1114,1140]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[281,418,1114,1140]" italics="true" pageId="10" pageNumber="309">Rhinocypha</emphasis>
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</taxonomicName>
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to those species with a short mesothoracic triangle, with brownish black areas in the wings of the males, and deeply tinged wings with milky-white patch at the apex of the hind wings in the female. Species with an enlarged mesothoracic triangle were assigned to
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<taxonomicName authority="Fraser" authorityName="Fraser" authorityYear="1928" box="[829,1033,1226,1252]" class="Insecta" family="Chlorocyphidae" genus="Calocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[829,951,1226,1252]" italics="true" pageId="10" pageNumber="309">Calocypha</emphasis>
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Fraser
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</taxonomicName>
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,
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<taxonomicName authority="Laidlaw" authorityName="Laidlaw" authorityYear="1950" box="[159,404,1264,1290]" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[159,300,1264,1290]" italics="true" pageId="10" pageNumber="309">Sundacypha</emphasis>
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Laidlaw
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</taxonomicName>
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,
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<taxonomicName authority="Fraser" authorityName="Fraser" box="[417,631,1264,1290]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[417,546,1264,1290]" italics="true" pageId="10" pageNumber="309">Heliocypha</emphasis>
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Fraser
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</taxonomicName>
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,
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<taxonomicName authority="Laidlaw" authorityName="Laidlaw" authorityYear="1950" box="[644,891,1264,1290]" class="Insecta" family="Chlorocyphidae" genus="Heterocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[644,789,1264,1290]" italics="true" pageId="10" pageNumber="309">Heterocypha</emphasis>
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Laidlaw
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</taxonomicName>
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and
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<taxonomicName authority="Laidlaw. At" authorityName="Laidlaw. At" authorityYear="1950" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="309">Aristocypha</emphasis>
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Laidlaw. At
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</taxonomicName>
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present there is some debate concerning the classification of the
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<taxonomicName authorityName="" authorityYear="1937" box="[236,429,1338,1364]" class="Insecta" family="Chlorocyphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
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, with, for example,
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<taxonomicName box="[662,791,1338,1364]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[662,791,1338,1364]" italics="true" pageId="10" pageNumber="309">Heliocypha</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Laidlaw. At" authorityYear="1950" box="[851,989,1338,1364]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[851,989,1338,1364]" italics="true" pageId="10" pageNumber="309">Aristocypha</emphasis>
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</taxonomicName>
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be- ing included as subgenera of
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[501,638,1376,1402]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis box="[501,638,1376,1402]" italics="true" pageId="10" pageNumber="309">Rhinocypha</emphasis>
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</taxonomicName>
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by some authorities (DIJKSTRA et al. 2014).
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</paragraph>
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<paragraph blockId="10.[159,1037,405,1551]" pageId="10" pageNumber="309">
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The generic attribution of the species on Sulawesi is undisputed. All spe- cies are closely allied to the type species of the genus
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<emphasis box="[833,976,1488,1514]" italics="true" pageId="10" pageNumber="309">
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[833,973,1488,1514]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
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,
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</emphasis>
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viz.,
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<emphasis box="[159,368,1525,1551]" italics="true" pageId="10" pageNumber="309">
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[159,296,1525,1551]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="309" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
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tincta
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</emphasis>
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Rambur.
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</paragraph>
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<paragraph blockId="11.[159,1037,254,466]" box="[159,267,254,280]" pageId="11" pageNumber="310">
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<heading bold="true" box="[159,267,254,280]" fontSize="11" level="3" pageId="11" pageNumber="310" reason="6">
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<emphasis bold="true" box="[159,267,254,280]" pageId="11" pageNumber="310">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="11.[159,1037,254,466]" pageId="11" pageNumber="310">
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No descriptions of secondary genitalia or anal appendages are included, since these structures exhibit little variation in
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<taxonomicName authorityName="" authorityYear="1937" box="[726,915,328,354]" class="Insecta" family="Chlorocyphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="310" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
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(COWLEY 1937; LIEFTINCK 1938; LAIdLAW 1950), while all species of the
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<emphasis italics="true" pageId="11" pageNumber="310">
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[899,1036,365,391]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="310" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
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tincta
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</emphasis>
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group share a similar penis structure (‘type II’ of Cowley). Anal appendages are simple and lack diagnostic characters.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="313" pageId="11" pageNumber="310" type="key">
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<paragraph blockId="11.[159,1037,525,1555]" box="[214,981,525,554]" pageId="11" pageNumber="310">
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<emphasis bold="true" box="[214,981,525,554]" pageId="11" pageNumber="310">
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Key to the
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[344,486,526,554]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="310" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[344,486,526,554]" italics="true" pageId="11" pageNumber="310">Rhinocypha</emphasis>
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</taxonomicName>
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species of Sulawesi and adjacent islands
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</emphasis>
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</paragraph>
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<paragraph blockId="11.[159,1037,525,1555]" box="[159,1036,572,598]" pageId="11" pageNumber="310">1a – Males ........................................................................................................ 2</paragraph>
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<paragraph blockId="11.[159,1037,525,1555]" box="[159,1035,617,643]" pageId="11" pageNumber="310">1b – Females .................................................................................................... 10</paragraph>
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<paragraph blockId="11.[159,1037,525,1555]" box="[159,288,669,696]" pageId="11" pageNumber="310">2 – Males</paragraph>
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<paragraph blockId="11.[159,1037,525,1555]" pageId="11" pageNumber="310">
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2a – Head with paired blue or bluish white spot on frons (
|
||
<figureCitation box="[827,895,714,740]" captionStart="Figure 3" captionStartId="14.[159,227,1021,1046]" captionTargetBox="[234,961,653,982]" captionTargetId="figure@14.[234,961,653,982]" captionTargetPageId="14" captionText="Figure 3. Rhinocypha frontalis, male. Stream Sg. Tumpa kecil, North Sulawesi, Bogani-Nani-Wartabone NP. Photo: AG (10-viii-2011)" httpUri="https://zenodo.org/record/2200381/files/figure.png" pageId="11" pageNumber="310">Fig. 3</figureCitation>
|
||
); size of spots with substantial variation between populations (
|
||
<figureCitation captionStart="Figures 30–33" captionStartId="30.[159,238,1495,1520]" captionTargetBox="[177,1031,866,1468]" captionTargetPageId="30" captionText="Figures 30–33. Rhinocypha f. frontalis. Male head, dorsal view. 30 – Dumoga-Bone National Park; 31 – Sabang (Dampelas); 32 – Luwuk; 33 – Mokowu. Artwork by JvT" httpUri="https://zenodo.org/record/2200461/files/figure.png" pageId="11" pageNumber="310">Figs 30– 33</figureCitation>
|
||
,
|
||
<figureCitation box="[259,290,788,815]" captionStart="Figure 35" captionStartId="37.[651,719,1433,1458]" captionTargetBox="[169,619,1216,1538]" captionTargetId="figure@37.[156,627,1200,1576]" captionTargetPageId="37" captionText="Figure 35. Rhinocypha frontalis sulselensis ssp. nov. Male, head, dorsal view. Locality as in Fig. 34. Artwork by JvT" httpUri="https://zenodo.org/record/2200465/files/figure.png" pageId="11" pageNumber="310">35</figureCitation>
|
||
); median lobe of pronotum black; blue stripe on synthorax (
|
||
<figureCitation box="[226,310,826,852]" captionStart="Figures 25–29" captionStartId="29.[159,238,1377,1402]" captionTargetBox="[159,1026,584,1344]" captionTargetId="figure@29.[212,988,914,1013]" captionTargetPageId="29" captionText="Figures 25–29. Rhinocypha f. frontalis. 25 – Male, head, frontal view. Sulawesi Utara, DumogaBone National Park, 22iv1985 [JvT 05948]; 26 – Male, pronotum, lef lateral view. Same locality [JvT 05949]; 27 – Male, synthorax, lef lateral view. Same locality [JvT 05948]; 28 – Male, abdomen, lef lateral view. Same locality, 03vi1985 [JvT 05970]; 29 – Female, head, dorsal view. Same locality, 20v1985 [JvT 05965]. Artwork by JvT" httpUri="https://zenodo.org/record/2200447/files/figure.png" pageId="11" pageNumber="310">Fig. 27</figureCitation>
|
||
) with distinct posterior projection over metepisternum, reaching posterior border of synthorax; ante-humeral stripe distinct in anterior part of mesepisternum (
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[723,960,901,927]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="310" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis box="[723,960,901,927]" italics="true" pageId="11" pageNumber="310">Rhinocypha frontalis</emphasis>
|
||
</taxonomicName>
|
||
complex) ................................................................................................... 3
|
||
</paragraph>
|
||
<paragraph blockId="11.[159,1037,525,1555]" pageId="11" pageNumber="310">
|
||
2b – Head with frons black, without pale spot; median lobe of prono- tum black, or with blue marking, usually extending on lateral cornerofanteriorlobe(
|
||
<emphasis box="[486,524,1058,1083]" italics="true" pageId="11" pageNumber="310">e.g.</emphasis>
|
||
,
|
||
<figureCitation box="[534,613,1058,1084]" captionStart="Figures 19–23" captionStartId="24.[159,238,1379,1404]" captionTargetBox="[688,1014,725,1343]" captionTargetPageId="24" captionText="Figures 19–23. Rhinocypha flavipoda sp. nov. Male, head in frontal view. Sulawesi Selatan, E. of Malili, 21x1993 [JvT 01458]; 20 – Male, pronotum, lef lateral view. Same locality [JvT 01458]; 21 – Male, synthorax,lef lateral view.Same locality [JvT 01458]; 22 – Male, abdomen, lef lateral view. Sulawesi Selatan, NNE of Malili, 02 v1991 [JvT 05919]; 23 – Female, head, dorsal view. Sulawesi Selatan, 45 km ENE of Malili, 19-x-1993 [JvT 01403]. Artwork by JvT" httpUri="https://zenodo.org/record/2200441/files/figure.png" pageId="11" pageNumber="310">Fig.20</figureCitation>
|
||
);bluestripeonsynthorax (
|
||
<emphasis box="[916,947,1058,1083]" italics="true" pageId="11" pageNumber="310">e.g</emphasis>
|
||
.,
|
||
<figureCitation box="[219,304,1095,1121]" captionStart="Figures 19–23" captionStartId="24.[159,238,1379,1404]" captionTargetBox="[688,1014,725,1343]" captionTargetPageId="24" captionText="Figures 19–23. Rhinocypha flavipoda sp. nov. Male, head in frontal view. Sulawesi Selatan, E. of Malili, 21x1993 [JvT 01458]; 20 – Male, pronotum, lef lateral view. Same locality [JvT 01458]; 21 – Male, synthorax,lef lateral view.Same locality [JvT 01458]; 22 – Male, abdomen, lef lateral view. Sulawesi Selatan, NNE of Malili, 02 v1991 [JvT 05919]; 23 – Female, head, dorsal view. Sulawesi Selatan, 45 km ENE of Malili, 19-x-1993 [JvT 01403]. Artwork by JvT" httpUri="https://zenodo.org/record/2200441/files/figure.png" pageId="11" pageNumber="310">Figs 21</figureCitation>
|
||
,
|
||
<figureCitation box="[315,345,1095,1122]" captionStart="Figures 61–65" captionStartId="58.[159,238,1379,1404]" captionTargetBox="[165,1027,647,1347]" captionTargetId="figure@58.[213,995,936,1049]" captionTargetPageId="58" captionText="Figures 61–65. Rhinocypha phantasma. All from Sulawesi Tengah. 61 – Male, head, frontal view. W of Luwuk, 14x1989 [JvT 05662]; 62 – Male, pronotum, lef lateral view. 28 km NE of Luwuk, 07x1989 [JvT 05442]; 63 – Male, synthorax, lef lateral view. Same locality [JvT 05440]; 64 – Male, abdomen, lef lateral view. Lo- cality as in Fig. 61 [JvT 05667]; 65 – Female, head, dorsal view. Locality as in Fig. 62 [JvT 05450]. Artwork by JvT" httpUri="https://zenodo.org/record/2200519/files/figure.png" pageId="11" pageNumber="310">63</figureCitation>
|
||
) dorsally straight, or with short projection over met- episternum, but with variation between populations; ante- humeral stripe absent .............................................................................. 5
|
||
</paragraph>
|
||
<paragraph blockId="11.[159,1037,525,1555]" pageId="11" pageNumber="310">
|
||
3a – Opaque dark patch in wing medially usually from level well basal to nodus (population of SW Sulawesi even from Ax5–6); blue markings on median lobe of pronotum relatively small; blue markings on frons variable, but usually much larger than in next species; hind wing length 23–30 mm; Sulawesi (
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[778,905,1371,1397]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="310" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis box="[778,905,1371,1397]" italics="true" pageId="11" pageNumber="310">R. frontalis</emphasis>
|
||
</taxonomicName>
|
||
) .............. 4
|
||
</paragraph>
|
||
<paragraph blockId="11.[159,1037,525,1555]" pageId="11" pageNumber="310">
|
||
3b – Opaque dark patch of wing only distal to nodus; blue markings on median lobe of pronotum extensive; blue markings on frons small; hind wing length 19–20 mm; Sangihe Is ...................... ... ...............................................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[781,942,1528,1554]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="310" phylum="Arthropoda" rank="species" species="sangihensis" status="sp. nov.">
|
||
<emphasis box="[781,942,1528,1554]" italics="true" pageId="11" pageNumber="310">R. sangihensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,1528,1554]" pageId="11" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
4a – Opaque dark patch distal to nodus in costal space, medially starting from level 5Ax anterior to nodus; wing length 24–25 mm ...................................................................
|
||
<emphasis box="[808,1036,328,354]" italics="true" pageId="12" pageNumber="311">
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[808,935,328,354]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="311" phylum="Arthropoda" rank="species" species="frontalis">R. frontalis</taxonomicName>
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[941,1036,328,354]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="311" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
4b – Opaque dark patch distal to Ax5 in costal space, medially basis just distal to fork of Rs, but leaving quadrangle free (transparent); wing length 27–28 mm ...........
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[683,931,446,472]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="311" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="ssp. nov." subSpecies="sulselensis">
|
||
<emphasis box="[683,931,446,472]" italics="true" pageId="12" pageNumber="311">R. frontalis sulselensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[938,1037,446,472]" pageId="12" pageNumber="311" rank="subSpecies">
|
||
<taxonomicName box="[938,1033,446,472]" pageId="12" pageNumber="311" rank="subSpecies" subSpecies="nov">ssp. nov</taxonomicName>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
5a – Mandibles with distinct blue spot, covering at least 50% of man- dible; blue stripe over synthorax with a ventral projection along the metapleural suture (
|
||
<figureCitation box="[496,627,570,596]" captionStart="Figures 39–44" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 39–44. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 – Male, head, frontal view. Bantimurung, 19iv1991 [JvT 05698]; 40 – Male, pronotum, lef lateral view. Bantimurung, 19–23iv1991 [JvT 05708]; 41 – Male, synthorax, lef lateral view. Bantimurung, 19iv1991 [JvT 05698]; 42 – Male, syn- thorax, lef lateral view. Madjene, 8–10xi1939 [JvT 05343]; 43 – Male, abdomen, lef lateral view. Bantimurung, 19iv1991 [JvT 06630]; 44 – Female, head, dorsal view. Bantimurung, 19iv1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="12" pageNumber="311">Figs 41–42</figureCitation>
|
||
) .................................
|
||
<taxonomicName box="[875,1036,570,596]" pageId="12" pageNumber="311">
|
||
<emphasis box="[875,1036,570,596]" italics="true" pageId="12" pageNumber="311">R. monochroa</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" box="[159,1035,613,640]" pageId="12" pageNumber="311">5b – Mandibles black, or with small pale (blue, or pale yellow) spot only ... 6</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" box="[159,1036,661,688]" pageId="12" pageNumber="311">
|
||
6a – Dorsum of abdomen pruinose (
|
||
<figureCitation box="[588,656,662,688]" captionStart="Figure 8" captionStartId="15.[828,896,1283,1308]" captionTargetBox="[159,791,247,1541]" captionTargetId="figure@15.[159,791,890,1216]" captionTargetPageId="15" captionText="Figure 8. Rhinocypha phantasma, female. Stream Sg. Wera above Saluopa waterfall, Tonusu, north of Lake Poso, Central Sulawesi. Photo: AG (21-viii-1999)" httpUri="https://zenodo.org/record/2200385/files/figure.png" pageId="12" pageNumber="311">Fig. 8</figureCitation>
|
||
) .............................
|
||
<taxonomicName box="[877,1036,662,688]" pageId="12" pageNumber="311">
|
||
<emphasis box="[877,1036,662,688]" italics="true" pageId="12" pageNumber="311">R. phantasma</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">6b – Dorsum of abdomen black, but may have well-defined blue annulae (rings) anteriorly on S4–S9 ....................................................... 7</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
7a – Distinct annulae anteriorly on S4–S9 of abdomen (
|
||
<figureCitation box="[804,892,791,817]" captionStart="Figure 10" captionStartId="16.[159,227,925,950]" captionText="Figure 10. Rhinocypha virgulata sp. nov., male. Stream Sg. Wera below Saluopa waterfall, Tonusu, north of Lake Poso, Central Sulawesi. Photo: AG (20-viii-1999)" pageId="12" pageNumber="311">Figs 10</figureCitation>
|
||
,
|
||
<figureCitation box="[902,931,791,817]" captionStart="Figures 78–82" captionStartId="77.[159,238,1407,1432]" captionTargetBox="[168,1023,585,1382]" captionTargetId="figure@77.[218,974,933,1038]" captionTargetPageId="77" captionText="Figures 78–82. Rhinocypha virgulata sp. nov. All from Sulawesi Selatan, Sg. Patea, river from Danau Matana to Danau Mahalona, 15-x-1993. 78 – Male, head, frontal view, [JvT 01370]; 79 – Male, pronotum, lef lateral view, [JvT 01371]; 80 – Male, synthorax, lef lateral view, [JvT 01370]; 81 – Male, abdomen, lef lateral view, [JvT 01373]; 82 – Female, head, dorsal view. [JvT 01377]. Artwork by JvT" httpUri="https://zenodo.org/record/2200549/files/figure.png" pageId="12" pageNumber="311">81</figureCitation>
|
||
) .. .....................................................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[807,942,828,854]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="311" phylum="Arthropoda" rank="species" species="virgulata" status="sp. nov.">
|
||
<emphasis box="[807,942,828,854]" italics="true" pageId="12" pageNumber="311">R. virgulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,828,854]" pageId="12" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">7b – Dorsum of abdomen black in all segments, no anterior annulae on S4–S9 ..................................................................................................... 8</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
8a – Pronotum black, without larger or smaller blue or creamish spots (
|
||
<figureCitation box="[226,311,995,1021]" captionStart="Figures 72–76" captionStartId="70.[159,238,1373,1398]" captionTargetBox="[167,1029,638,1339]" captionTargetId="figure@70.[234,948,945,1021]" captionTargetPageId="70" captionText="Figures 72–76. Rhinocypha togeanensis sp. nov. All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT" httpUri="https://zenodo.org/record/2200535/files/figure.png" pageId="12" pageNumber="311">Fig. 73</figureCitation>
|
||
); mesopreepisternum black ...................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[780,943,996,1021]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="311" phylum="Arthropoda" rank="species" species="togeanensis" status="sp. nov.">
|
||
<emphasis box="[780,943,996,1021]" italics="true" pageId="12" pageNumber="311">R. togeanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,995,1021]" pageId="12" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">8b – Pronotum with blue or creamish spots at least in latero-anterior corner of median lobe, and on lateral lobe; mesopreepisternum with blue marking ..................................................................................... 9</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
9a – Opaque parts of wings fully distal to nodus (
|
||
<figureCitation box="[716,799,1162,1189]" captionStart="Figure 54" captionStartId="51.[159,227,1489,1514]" captionTargetBox="[159,1036,868,1449]" captionTargetId="figure@51.[159,1036,868,1449]" captionTargetPageId="51" captionText="Figure 54. Rhinocypha pelengensis sp. nov. Male, holotype. Sulawesi Tengah, Banggai Arch., Peleng, 31-vii–05-viii-1941 [JvT 22905]. Photo: JvT" httpUri="https://zenodo.org/record/2200499/files/figure.png" pageId="12" pageNumber="311">Fig.54</figureCitation>
|
||
); tibiae of mid and hind legs white ...............................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[784,942,1199,1225]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="311" phylum="Arthropoda" rank="species" species="pelengensis" status="sp. nov.">
|
||
<emphasis box="[784,942,1199,1225]" italics="true" pageId="12" pageNumber="311">R. pelengensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,1199,1225]" pageId="12" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
9b – Opaque parts of wings extending in middle of wing to distal side of quadrangle (
|
||
<figureCitation box="[445,529,1280,1306]" captionStart="Figure 18" captionStartId="19.[159,227,1520,1545]" captionTargetBox="[210,985,1045,1481]" captionTargetId="figure@19.[210,985,1045,1481]" captionTargetPageId="19" captionText="Figure 18. Rhinocypha flavipoda sp. nov. Male, holotype." httpUri="https://zenodo.org/record/2200439/files/figure.png" pageId="12" pageNumber="311">Fig. 18</figureCitation>
|
||
); tibiae of mid and hind legs typically pale yellow, but sometimes difficult to see in museum specimens ...................................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[803,942,1354,1380]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="311" phylum="Arthropoda" rank="species" species="flavipoda" status="sp. nov.">
|
||
<emphasis box="[803,942,1354,1380]" italics="true" pageId="12" pageNumber="311">R. flavipoda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,1355,1381]" pageId="12" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" box="[159,329,1403,1430]" pageId="12" pageNumber="311">10 – Females</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" pageId="12" pageNumber="311">
|
||
10a – Frons with paired pale spot (in some individual specimens rather small and indistinct) (
|
||
<figureCitation box="[568,656,1485,1511]" captionStart="Figures 25–29" captionStartId="29.[159,238,1377,1402]" captionTargetBox="[159,1026,584,1344]" captionTargetId="figure@29.[212,988,914,1013]" captionTargetPageId="29" captionText="Figures 25–29. Rhinocypha f. frontalis. 25 – Male, head, frontal view. Sulawesi Utara, DumogaBone National Park, 22iv1985 [JvT 05948]; 26 – Male, pronotum, lef lateral view. Same locality [JvT 05949]; 27 – Male, synthorax, lef lateral view. Same locality [JvT 05948]; 28 – Male, abdomen, lef lateral view. Same locality, 03vi1985 [JvT 05970]; 29 – Female, head, dorsal view. Same locality, 20v1985 [JvT 05965]. Artwork by JvT" httpUri="https://zenodo.org/record/2200447/files/figure.png" pageId="12" pageNumber="311">Figs 29</figureCitation>
|
||
,
|
||
<figureCitation box="[667,697,1485,1512]" captionStart="Figures 61–65" captionStartId="58.[159,238,1379,1404]" captionTargetBox="[165,1027,647,1347]" captionTargetId="figure@58.[213,995,936,1049]" captionTargetPageId="58" captionText="Figures 61–65. Rhinocypha phantasma. All from Sulawesi Tengah. 61 – Male, head, frontal view. W of Luwuk, 14x1989 [JvT 05662]; 62 – Male, pronotum, lef lateral view. 28 km NE of Luwuk, 07x1989 [JvT 05442]; 63 – Male, synthorax, lef lateral view. Same locality [JvT 05440]; 64 – Male, abdomen, lef lateral view. Lo- cality as in Fig. 61 [JvT 05667]; 65 – Female, head, dorsal view. Locality as in Fig. 62 [JvT 05450]. Artwork by JvT" httpUri="https://zenodo.org/record/2200519/files/figure.png" pageId="12" pageNumber="311">65</figureCitation>
|
||
) .......................................... 11
|
||
</paragraph>
|
||
<paragraph blockId="12.[159,1037,253,1556]" box="[159,1035,1529,1556]" pageId="12" pageNumber="311">10b – Frons black ............................................................................................ 15</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" box="[159,1035,254,280]" pageId="13" pageNumber="312">
|
||
11a – Anterior lobe of pronotum with paired median spot (
|
||
<emphasis box="[856,886,254,279]" italics="true" pageId="13" pageNumber="312">e.g</emphasis>
|
||
.,
|
||
<figureCitation box="[906,973,254,280]" captionStart="Figure 4" captionStartId="14.[159,227,1479,1504]" captionTargetBox="[234,961,1111,1440]" captionTargetId="figure@14.[234,961,1111,1440]" captionTargetPageId="14" captionText="Figure 4. Rhinocypha frontalis, female. Stream Sg. Tumpa, North Sulawesi, Bogani- Nani-Wartabone NP. Photo: AG (12-viii-2011)" httpUri="https://zenodo.org/record/2200383/files/figure.png" pageId="13" pageNumber="312">Fig. 4</figureCitation>
|
||
) .. 12
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">11b – Anterior lobe of pronotum black, or with small spot in lateral corners, which continues on median lobe, usually much larger than on anterior lobe ........................................................................... 14</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" box="[159,1035,428,454]" pageId="13" pageNumber="312">
|
||
12a – Mainland Sulawesi (
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[472,598,428,454]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="312" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis box="[472,598,428,454]" italics="true" pageId="13" pageNumber="312">R. frontalis</emphasis>
|
||
</taxonomicName>
|
||
) ........................................................ 13
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
12b – Sangihe islands (only one teneral and incomplete specimen available) ...............................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[781,942,511,537]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="312" phylum="Arthropoda" rank="species" species="sangihensis" status="sp. nov.">
|
||
<emphasis box="[781,942,511,537]" italics="true" pageId="13" pageNumber="312">R. sangihensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,512,538]" pageId="13" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
13a – Distinct transverse brown band on fore and hind wing, extreme distal part of wings somewhat opaque white; base to transverse band usually slightly enfumed (
|
||
<figureCitation box="[595,679,639,666]" captionStart="Figures 36–37" captionStartId="38.[159,238,1170,1195]" captionTargetBox="[469,1036,782,1539]" captionTargetPageId="38" captionText="Figures 36–37. Rhinocypha frontalis subspecies. Wings, female. 36 — Subspecies frontalis. Sulawesi Utara, surroundings Tondano, 09-vi-1940 [JvT 22866]; 37 — subspecies sulselensis. Sulawesi Selatan, Boeloedoea, vi-1935 [JvT 22891]. Both with same magnification. Photos: JvT" httpUri="https://zenodo.org/record/2200471/files/figure.png" pageId="13" pageNumber="312">Fig. 36</figureCitation>
|
||
); length of hind wing up to 26 mm ..............................................................................
|
||
<emphasis box="[889,1036,677,703]" italics="true" pageId="13" pageNumber="312">
|
||
R. f.
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[941,1036,677,703]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="312" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
13b – Transverse band on wings rather fuzzy, indistinct from approximately Px10 to just anterior to pterostigma (fore wing), or halfway pterostigma (hind wing); base enfumed dark brown (
|
||
<figureCitation box="[332,424,835,861]" captionStart="Figures 36–37" captionStartId="38.[159,238,1170,1195]" captionTargetBox="[469,1036,782,1539]" captionTargetPageId="38" captionText="Figures 36–37. Rhinocypha frontalis subspecies. Wings, female. 36 — Subspecies frontalis. Sulawesi Utara, surroundings Tondano, 09-vi-1940 [JvT 22866]; 37 — subspecies sulselensis. Sulawesi Selatan, Boeloedoea, vi-1935 [JvT 22891]. Both with same magnification. Photos: JvT" httpUri="https://zenodo.org/record/2200471/files/figure.png" pageId="13" pageNumber="312">Fig. 37</figureCitation>
|
||
); length of hind wing
|
||
<emphasis box="[721,746,836,861]" italics="true" pageId="13" pageNumber="312">ca</emphasis>
|
||
30 mm .............. ..................................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[693,942,872,898]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="312" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="sp. nov." subSpecies="sulselensis">
|
||
<emphasis box="[693,942,872,898]" italics="true" pageId="13" pageNumber="312">R. frontalis sulselensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,873,899]" pageId="13" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
14a – Lateral lobe of pronotum with pale, creamy white spot; scapus (base of antenna) pale, only interrupted in middle of this antennal segment (
|
||
<figureCitation box="[455,540,1000,1027]" captionStart="Figures 61–65" captionStartId="58.[159,238,1379,1404]" captionTargetBox="[165,1027,647,1347]" captionTargetId="figure@58.[213,995,936,1049]" captionTargetPageId="58" captionText="Figures 61–65. Rhinocypha phantasma. All from Sulawesi Tengah. 61 – Male, head, frontal view. W of Luwuk, 14x1989 [JvT 05662]; 62 – Male, pronotum, lef lateral view. 28 km NE of Luwuk, 07x1989 [JvT 05442]; 63 – Male, synthorax, lef lateral view. Same locality [JvT 05440]; 64 – Male, abdomen, lef lateral view. Lo- cality as in Fig. 61 [JvT 05667]; 65 – Female, head, dorsal view. Locality as in Fig. 62 [JvT 05450]. Artwork by JvT" httpUri="https://zenodo.org/record/2200519/files/figure.png" pageId="13" pageNumber="312">Fig. 65</figureCitation>
|
||
). Sulawesi, Buton .................
|
||
<taxonomicName box="[877,1036,1000,1026]" pageId="13" pageNumber="312">
|
||
<emphasis box="[877,1036,1000,1026]" italics="true" pageId="13" pageNumber="312">R. phantasma</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
14b – Pronotum black; scapus black (
|
||
<figureCitation box="[598,684,1046,1073]" captionStart="Figures 55–59" captionStartId="54.[159,238,1376,1401]" captionTargetBox="[171,1030,557,1334]" captionTargetId="figure@54.[172,1030,905,1005]" captionTargetPageId="54" captionText="Figures 55–59. Rhinocypha pelengensis sp. nov. All from Sulawesi Tengah. 55 – Male, head, frontal view. Pulau Peleng, 31-vii–05-viii-1941 [JvT 22905]; 56 – Male, pronotum, lef lateral view. Same locality [JvT 22904]; 57 – Male, synthorax, lef lateral view. Same locality [JvT 22906]; 58 – Male, abdomen, lef lateral view. Same locality [JvT 22898]; 59 – Female, head, dorsal view. Peleng Island, 25ii–03 -iii-1986 [JvT 33003]. Artwork by JvT" httpUri="https://zenodo.org/record/2200503/files/figure.png" pageId="13" pageNumber="312">Fig. 59</figureCitation>
|
||
). Banggai Islands only. ................................................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[784,942,1084,1110]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="312" phylum="Arthropoda" rank="species" species="pelengensis" status="sp. nov.">
|
||
<emphasis box="[784,942,1084,1110]" italics="true" pageId="13" pageNumber="312">R. pelengensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,1084,1110]" pageId="13" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
15a – Mandible largely bluish white, only black at base; gena with extensive yellow marking, which is connected with pale stripe along the eye, dorsally running to level of lateral ocelli (
|
||
<figureCitation box="[241,325,1249,1275]" captionStart="Figures 39–44" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 39–44. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 – Male, head, frontal view. Bantimurung, 19iv1991 [JvT 05698]; 40 – Male, pronotum, lef lateral view. Bantimurung, 19–23iv1991 [JvT 05708]; 41 – Male, synthorax, lef lateral view. Bantimurung, 19iv1991 [JvT 05698]; 42 – Male, syn- thorax, lef lateral view. Madjene, 8–10xi1939 [JvT 05343]; 43 – Male, abdomen, lef lateral view. Bantimurung, 19iv1991 [JvT 06630]; 44 – Female, head, dorsal view. Bantimurung, 19iv1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="13" pageNumber="312">Fig. 44</figureCitation>
|
||
) ..............................................................................
|
||
<taxonomicName box="[875,1036,1249,1275]" pageId="13" pageNumber="312">
|
||
<emphasis box="[875,1036,1249,1275]" italics="true" pageId="13" pageNumber="312">R. monochroa</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
15b – Mandible with pale spot covering 50% or less; yellow markings on gena and along eye small and narrow (
|
||
<emphasis box="[722,753,1334,1359]" italics="true" pageId="13" pageNumber="312">e.g</emphasis>
|
||
.,
|
||
<figureCitation box="[773,853,1333,1359]" captionStart="Figures 78–82" captionStartId="77.[159,238,1407,1432]" captionTargetBox="[168,1023,585,1382]" captionTargetId="figure@77.[218,974,933,1038]" captionTargetPageId="77" captionText="Figures 78–82. Rhinocypha virgulata sp. nov. All from Sulawesi Selatan, Sg. Patea, river from Danau Matana to Danau Mahalona, 15-x-1993. 78 – Male, head, frontal view, [JvT 01370]; 79 – Male, pronotum, lef lateral view, [JvT 01371]; 80 – Male, synthorax, lef lateral view, [JvT 01370]; 81 – Male, abdomen, lef lateral view, [JvT 01373]; 82 – Female, head, dorsal view. [JvT 01377]. Artwork by JvT" httpUri="https://zenodo.org/record/2200549/files/figure.png" pageId="13" pageNumber="312">Fig. 82</figureCitation>
|
||
) .................. 16
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
16a – Pronotum and mesopreepisternum black. Togian Islands ...
|
||
<emphasis box="[235,942,1424,1449]" italics="true" pageId="13" pageNumber="312">
|
||
..............................................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[780,942,1424,1449]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="312" phylum="Arthropoda" rank="species" species="togeanensis" status="sp. nov.">R. togeanensis</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[949,1037,1424,1450]" pageId="13" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="13.[159,1037,254,1534]" pageId="13" pageNumber="312">
|
||
16b – Pronotum with pale markings; mesopreepisternum creamy white (see
|
||
<figureCitation box="[359,438,1507,1533]" captionStart="Figure 11" captionStartId="16.[159,227,1254,1279]" captionTargetBox="[400,1038,734,1540]" captionTargetId="figure@16.[400,1035,734,1181]" captionTargetPageId="16" captionText="Figure 11. Rhinocypha virgulata sp. nov., female. Stream Sg. Wera below Saluopa waterfall, Tonusu, north of Lake Poso, Central Sulawesi. Photo: AG (20-viii- -1999)" httpUri="https://zenodo.org/record/2200401/files/figure.png" pageId="13" pageNumber="312">Fig. 11</figureCitation>
|
||
) ................................................................................ 17
|
||
</paragraph>
|
||
<paragraph blockId="14.[159,1037,254,624]" pageId="14" pageNumber="313">
|
||
17a – Median lobe of pronotum latero-anteriorly with large irregular spot, which continues in lateral corner of anterior lobe; lateral lobe of pronotum with large, irregular spot; sides of abdominal S2–S4 against dorsum with a longitudinal or L-shaped mark- ing (
|
||
<figureCitation box="[285,368,403,429]" captionStart="Figure 11" captionStartId="16.[159,227,1254,1279]" captionTargetBox="[400,1038,734,1540]" captionTargetId="figure@16.[400,1035,734,1181]" captionTargetPageId="16" captionText="Figure 11. Rhinocypha virgulata sp. nov., female. Stream Sg. Wera below Saluopa waterfall, Tonusu, north of Lake Poso, Central Sulawesi. Photo: AG (20-viii- -1999)" httpUri="https://zenodo.org/record/2200401/files/figure.png" pageId="14" pageNumber="313">Fig. 11</figureCitation>
|
||
) .............................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[807,942,403,429]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="313" phylum="Arthropoda" rank="species" species="virgulata" status="sp. nov.">
|
||
<emphasis box="[807,942,403,429]" italics="true" pageId="14" pageNumber="313">R. virgulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,403,429]" pageId="14" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="14.[159,1037,254,624]" pageId="14" pageNumber="313">
|
||
17b – Median lobe of pronotum latero-anteriorly with small pale spot; lateral lobe of pronotum with small spot; sides of abdom- inal segments against dorsum at most with a semicircular spot (there is a series of longitudinal markings close to ventral side of the tergites) .........................................................
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[803,942,597,623]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="313" phylum="Arthropoda" rank="species" species="flavipoda" status="sp. nov.">
|
||
<emphasis box="[803,942,597,623]" italics="true" pageId="14" pageNumber="313">R. flavipoda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[949,1037,597,623]" pageId="14" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |