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<document id="9DA409666179E5AA770903E27B57D769" ID-DOI="10.5281/zenodo.5329053" ID-ISSN="0374-1036" ID-Zenodo-Dep="5329053" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630160711340" checkinUser="felipe" docAuthor="Jákl, Stanislav" docDate="2011" docId="0393964FC137FFCECB071BB3FDF495C3" docLanguage="en" docName="ActaEntMusNatPra.2011.51.2.535-550.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 51 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thaumastopeus pugnator subsp. insulanus Jákl 2011, subsp. nov." docType="treatment" docVersion="4" lastPageNumber="549" masterDocId="FFAAEE37C13AFFC0CA571909FFD19650" masterDocTitle="Description of seven new taxa of Cetoniinae from Indonesia (Coleoptera: Scarabaeidae)" masterLastPageNumber="550" masterPageNumber="535" pageNumber="548" updateTime="1699169434667" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="57F7CFA4C30C6012535866E30679813F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FBF6441D2F53862FAAD226D3EA7AC5D9">
<mods:title id="DEAE43E1A725D7D2C8E6CE7D7D4F5A5C">Description of seven new taxa of Cetoniinae from Indonesia (Coleoptera: Scarabaeidae)</mods:title>
</mods:titleInfo>
<mods:name id="9F59E5D4F7FEF18945A0876A97B5D801" type="personal">
<mods:role id="A5BDFBB59B4177220BE3994AE3FB70D6">
<mods:roleTerm id="C64BC7A1BE1E1F3B7B5CF514C8D73FC1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="423D18B0F3E59EDFC60B37CC84D8A0E4">Jákl, Stanislav</mods:namePart>
</mods:name>
<mods:typeOfResource id="2732E5BDC3D352770A7878ED7205C1DB">text</mods:typeOfResource>
<mods:relatedItem id="46F0A96E9AA2278BF4F1E8D0F49A6352" type="host">
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<mods:title id="FFEE7B81D281466EA0BA1DD6510A26D9">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="5EB3A36A4CB36B1BCDCAF6BB490A198D">
<mods:date id="B56568D22B9E43E1F72E0F43EE3DE8D8">2011</mods:date>
<mods:detail id="06AFE2A328377C0928B1C9D26020ACA4" type="pubDate">
<mods:number id="62730989DAFC93B88817F952F739AE6E">2011-12-09</mods:number>
</mods:detail>
<mods:detail id="044C4B8AC0E90EAB22648D81B44EBF90" type="volume">
<mods:number id="0D0EF37DEB8C20D59FFDBACAC711BF01">51</mods:number>
</mods:detail>
<mods:detail id="3748DDC97DB01C9F0534FC3D9C5BCC7C" type="issue">
<mods:number id="2BFEF028FD69DCD6FE2BCE7AFF76C11D">2</mods:number>
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<mods:start id="C7F7D2168840E4F36313D876A1B71AC1">535</mods:start>
<mods:end id="9527204329D3218EC1DD07CAC1459AF4">550</mods:end>
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<mods:classification id="44B6D2E6D5F9CCFBB3FE401BAF3B360B">journal article</mods:classification>
<mods:identifier id="996011F5081FA86696D65F62F4170ABA" type="DOI">10.5281/zenodo.5329053</mods:identifier>
<mods:identifier id="FF40A2C02A8AE16BB1138FD5166B8B46" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="5D5A6924654E12C8C677956341C7AD58" type="Zenodo-Dep">5329053</mods:identifier>
</mods:mods>
<treatment id="0393964FC137FFCECB071BB3FDF495C3" ID-DOI="http://doi.org/10.5281/zenodo.5397006" ID-Zenodo-Dep="5397006" LSID="urn:lsid:plazi:treatment:0393964FC137FFCECB071BB3FDF495C3" httpUri="http://treatment.plazi.org/id/0393964FC137FFCECB071BB3FDF495C3" lastPageId="14" lastPageNumber="549" pageId="13" pageNumber="548">
<subSubSection id="C32074D2C137FFCDCB071BB3FCB39484" box="[336,866,698,724]" pageId="13" pageNumber="548" type="nomenclature">
<paragraph id="8B852759C137FFCDCB071BB3FCB39484" blockId="13.[336,866,698,724]" box="[336,866,698,724]" pageId="13" pageNumber="548">
<heading id="D0CD9035C137FFCDCB071BB3FCB39484" bold="true" box="[336,866,698,724]" centered="true" fontSize="8" level="2" pageId="13" pageNumber="548" reason="3">
<emphasis id="B94EFB4BC137FFCDCB071BB3FCB39484" bold="true" box="[336,866,698,724]" pageId="13" pageNumber="548">
<taxonomicName id="4C3A5CDAC137FFCDCB071BB3FD0C9483" authority="Jákl, 2011" authorityName="Jákl" authorityYear="2011" box="[336,733,698,723]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="548" phylum="Arthropoda" rank="subSpecies" species="pugnator" status="subsp. nov." subSpecies="insulanus">
<emphasis id="B94EFB4BC137FFCDCB071BB3FD0C9483" bold="true" box="[336,733,698,723]" italics="true" pageId="13" pageNumber="548">Thaumastopeus pugnator insulanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D4630C137FFCDC8B31BB5FCB39484" box="[740,866,700,724]" pageId="13" pageNumber="548" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32074D2C137FFCDC8711BD3FD5D94BD" box="[550,652,730,749]" pageId="13" pageNumber="548" type="description">
<paragraph id="8B852759C137FFCDC8711BD3FD5D94BD" blockId="13.[550,652,730,749]" box="[550,652,730,749]" pageId="13" pageNumber="548">
(
<figureCitation id="13013BDCC137FFCDC87B1BD3FD5494BD" box="[556,645,730,749]" captionStart="Figs" captionStartId="11.[113,152,1590,1609]" captionTargetBox="[123,1059,221,1510]" captionTargetId="figure-11@11.[113,1070,210,1516]" captionTargetPageId="11" captionText="Figs. 6–7. 6 – Thaumastopeus striatus siberutanus subsp. nov.; 7 – T. pugnator insulanus subsp. nov. a – habitus dorsally; b – habitus laterally; c – habitus ventrally; d – aedeagus apically; e – aedeagus laterally. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.5329063" httpUri="https://zenodo.org/record/5329063/files/figure.png" pageId="13" pageNumber="548">Figs. 7a–c</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32074D2C137FFCDCA261A0DFE9D9526" pageId="13" pageNumber="548" type="materials_examined">
<paragraph id="8B852759C137FFCDCA261A0DFD5C9547" blockId="13.[113,1089,771,886]" box="[113,653,771,791]" pageId="13" pageNumber="548">
<emphasis id="B94EFB4BC137FFCDCA261A0DFF3B9547" bold="true" box="[113,234,771,791]" pageId="13" pageNumber="548">
<typeStatus id="548199FBC137FFCDCA261A0DFF4F9547" box="[113,158,772,791]" pageId="13" pageNumber="548">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32D67C9C137FFCDCAB81A0DFE959547" box="[239,324,772,791]" name="Indonesia" pageId="13" pageNumber="548">Indonesia</collectingCountry>
,
<collectingRegion id="49FEE9BBC137FFCDCB1A1A0DFDCC9547" box="[333,541,772,791]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="548">North Sumatra province</collectingRegion>
, Nias Island.
</paragraph>
<paragraph id="8B852759C137FFCDCA261A27FE9D9526" blockId="13.[113,1089,771,886]" pageId="13" pageNumber="548">
<emphasis id="B94EFB4BC137FFCDCA261A27FF259511" bold="true" box="[113,244,813,833]" pageId="13" pageNumber="548">Type material.</emphasis>
<materialsCitation id="3B522D04C137FFCDCAAF1A27FC0C9510" box="[248,989,808,833]" collectingDate="1993-10" collectionCode="NMPC" country="Indonesia" location="Nias island" pageId="13" pageNumber="548" specimenCount="1" typeStatus="holotype">
<typeStatus id="548199FBC137FFCDCAAF1A27FE9A956F" box="[248,331,814,833]" pageId="13" pageNumber="548" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED2BBF9CC137FFCDCB391A24FE619510" box="[366,432,813,832]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="548">NMPC</collectionCode>
):‘
<collectingCountry id="F32D67C9C137FFCDCBE81A24FDC69510" box="[447,535,813,832]" name="Indonesia" pageId="13" pageNumber="548">Indonesia</collectingCountry>
,
<location id="8EE57182C137FFCDC8481A24FDAE9510" LSID="urn:lsid:plazi:treatment:0393964FC137FFCECB071BB3FDF495C3:8EE57182C137FFCDC8481A24FDAE9510" box="[543,639,813,832]" country="Indonesia" name="Nias island" pageId="13" pageNumber="548">Nias island</location>
,
<date id="FF840199C137FFCDC8D11A27FD189511" box="[646,713,814,833]" pageId="13" pageNumber="548" value="1993-10">
<collectingDate id="EFC0F871C137FFCDC8D11A27FD189511" box="[646,713,814,833]" pageId="13" pageNumber="548" value="1993-10">X. 1993</collectingDate>
</date>
[handwritten], native collectors’
</materialsCitation>
.
<materialsCitation id="3B522D04C137FFCDC9B31A27FE28950B" collectingDate="1993-10" collectionCode="SJPC" country="Indonesia" location="Nias island" pageId="13" pageNumber="548" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="548199FBC137FFCDC9B31A27FBEA956F" box="[996,1083,814,833]" pageId="13" pageNumber="548" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D3CECD0C137FFCDCA261A4BFF50950B" box="[113,129,834,859]" pageId="13" pageNumber="548" type="female">♀</specimenCount>
(no. 1), same label data as holotype (
<collectionCode id="ED2BBF9CC137FFCDCB971A41FE22950B" box="[448,499,840,859]" country="0" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34278" lsid="urn:lsid:biocol.org:col:34278" name="Sergei J. Paramonov personal collection -- destroyed" pageId="13" pageNumber="548">SJPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B522D04C137FFCDC8561A4AFE999526" collectionCode="SJPC" country="Indonesia" location="Nias island" pageId="13" pageNumber="548" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
J (no. 2),
<specimenCount id="9D3CECD0C137FFCDC80D1A4BFDBB950B" box="[602,618,834,859]" pageId="13" pageNumber="548" type="female">♀</specimenCount>
(no. 3): ‘
<collectingCountry id="F32D67C9C137FFCDC8E11A41FCDE950B" box="[694,783,840,859]" name="Indonesia" pageId="13" pageNumber="548">Indonesia</collectingCountry>
,
<location id="8EE57182C137FFCDC94F1A41FCA8950B" LSID="urn:lsid:plazi:treatment:0393964FC137FFCECB071BB3FDF495C3:8EE57182C137FFCDC94F1A41FCA8950B" box="[792,889,840,859]" country="Indonesia" name="Nias island" pageId="13" pageNumber="548">Nias island</location>
, 1. 1995 [handwritten], native collectors’ (
<collectionCode id="ED2BBF9CC137FFCDCB581A6AFE939526" box="[271,322,867,886]" country="0" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34278" lsid="urn:lsid:biocol.org:col:34278" name="Sergei J. Paramonov personal collection -- destroyed" pageId="13" pageNumber="548">SJPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32074D2C137FFCECA261A87FDA09463" lastPageId="14" lastPageNumber="549" pageId="13" pageNumber="548" type="description">
<paragraph id="8B852759C137FFCDCA261A87FD679596" blockId="13.[113,1089,910,1638]" pageId="13" pageNumber="548">
<emphasis id="B94EFB4BC137FFCDCA261A87FF2D95F6" bold="true" box="[113,252,910,934]" pageId="13" pageNumber="548">Description.</emphasis>
<typeStatus id="548199FBC137FFCDCB541A87FEB795F6" box="[259,358,910,934]" pageId="13" pageNumber="548" type="holotype">Holotype</typeStatus>
length
<quantity id="4CC28ABCC137FFCDCBE01A87FDC895F6" box="[439,537,910,934]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="13" pageNumber="548" unit="mm" value="28.8">28.8 mm</quantity>
, maximum humeral width
<quantity id="4CC28ABCC137FFCDC96E1A87FC4A95F6" box="[825,923,910,934]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="13" pageNumber="548" unit="mm" value="11.4">11.4 mm</quantity>
. Narrow, rather parallel-sided; dark olive green with golden reflection.
</paragraph>
<paragraph id="8B852759C137FFCDCAC71AC7FF0D9216" blockId="13.[113,1089,910,1638]" pageId="13" pageNumber="548">Head. Dark green, shining, widest in apical third. Frons simply punctate. Clypeus with two densely wrinkled lateral furrows.Apex of clypeus deeply incised, its apical margin sharp but termination obtuse. Antenna moderately long, stalk black, club dark brown, setation brownish.</paragraph>
<paragraph id="8B852759C137FFCDCAC71D46FC959296" blockId="13.[113,1089,910,1638]" pageId="13" pageNumber="548">Pronotum. Olive to green with golden-metallic reflection. From base sharply narrowing to apex. Lateral margins with double emargination, first very mild in front of anterolateral margin, second deeper in front of posterolateral margin. Microreticulation densely developed throughout length. Apical half of lateral margins with fine wrinkles.</paragraph>
<paragraph id="8B852759C137FFCDCAC71DC7FE879356" blockId="13.[113,1089,910,1638]" pageId="13" pageNumber="548">Scutellum. Small, dark green, shining. Moderately narrowing to apex, here obtusely rounded. Impunctate.</paragraph>
<paragraph id="8B852759C137FFCDCAC71C07FC6393B6" blockId="13.[113,1089,910,1638]" pageId="13" pageNumber="548">Elytra. Olive to green with golden lustre. Basal half almost parallel-sided, apical half moderately narrowing toward apex. Apical two-thirds laterally wrinkled, wrinkles denser along apical third.Apex and apical calli also with moderately developed wrinkling. Each elytron with three to four indistinctly punctured lines. Fine microreticulation present throughout length. Basal half also with three to four irregularly shaped transverse impressions. Humeral calli small, apical calli not developed. Epipleura present only in basal half. Sutural ridge elevated in apical third, its termination rather obtuse, indistinctly protruding over apex.</paragraph>
<paragraph id="8B852759C137FFCDCAC71CE7FBCD9056" blockId="13.[113,1089,910,1638]" box="[144,1052,1518,1542]" pageId="13" pageNumber="548">Pygidium. Brownish with green-metallic lustre. Densely wrinkled throughout length.</paragraph>
<paragraph id="8B852759C137FFCECAC71F06FE569743" blockId="13.[113,1089,910,1638]" lastBlockId="14.[151,1127,219,915]" lastPageId="14" lastPageNumber="549" pageId="13" pageNumber="548">Venter. Abdomen brownish with green reflection. Each ventrite laterally wrinkled, middle part impunctate, but microreticulated. Metasternum green with mild lustre, laterally wrinkled, middle part only with microreticulation. Mesometasternal process long and robust, sharply pointed at apex, not reaching level of procoxae. Prosternum and mentum dark green, wrinkled, with blackish setation.</paragraph>
<paragraph id="8B852759C134FFCECAE21815FEDC9723" blockId="14.[151,1127,219,915]" pageId="14" pageNumber="549">Legs. Femora, tibiae and tarsi dark green, with mild lustre. Tibiae and tarsi elongated. Protibiae tridentate. Meso- and metatibiae with carina approximately between posterior third and fourth.</paragraph>
<paragraph id="8B852759C134FFCECAE21872FCDA97C3" blockId="14.[151,1127,219,915]" box="[181,779,379,403]" pageId="14" pageNumber="549">
Male genitalia. Similar to other subspecies (
<figureCitation id="13013BDCC134FFCEC8DC1872FD2E97C3" box="[651,767,379,403]" captionStart="Figs" captionStartId="11.[113,152,1590,1609]" captionTargetBox="[123,1059,221,1510]" captionTargetId="figure-11@11.[113,1070,210,1516]" captionTargetPageId="11" captionText="Figs. 6–7. 6 – Thaumastopeus striatus siberutanus subsp. nov.; 7 – T. pugnator insulanus subsp. nov. a – habitus dorsally; b – habitus laterally; c – habitus ventrally; d – aedeagus apically; e – aedeagus laterally. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.5329063" httpUri="https://zenodo.org/record/5329063/files/figure.png" pageId="14" pageNumber="549">Figs. 7d–e</figureCitation>
).
</paragraph>
<paragraph id="8B852759C134FFCECAC01892FDA09463" blockId="14.[151,1127,219,915]" pageId="14" pageNumber="549">
<emphasis id="B94EFB4BC134FFCECAC01892FEAF97E3" bold="true" box="[151,382,411,435]" pageId="14" pageNumber="549">Sexual dimorphism.</emphasis>
Both female
<typeStatus id="548199FBC134FFCEC8431895FDAB97E4" box="[532,634,412,436]" pageId="14" pageNumber="549" type="paratype">paratypes</typeStatus>
are bigger, 29.0 and
<quantity id="4CC28ABCC134FFCEC9321892FC1B97E3" box="[869,970,411,435]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="14" pageNumber="549" unit="mm" value="32.5">32.5 mm</quantity>
. Punctation of the pronotum is almost the same as in the
<typeStatus id="548199FBC134FFCEC83118B2FD129783" box="[614,707,443,467]" pageId="14" pageNumber="549" type="holotype">holotype</typeStatus>
male, but punctation of elytra is more developed. Both specimens have eight punctate lines on each elytron. The body is wider and much more robust. Also legs are more robust, especially the protibia. The abdomen is more arched, without a distinct abdominal furrow.
</paragraph>
</subSubSection>
<subSubSection id="C32074D2C134FFCECAC01B32FBC29544" pageId="14" pageNumber="549" type="diagnosis">
<paragraph id="8B852759C134FFCECAC01B32FBC29544" blockId="14.[151,1127,219,915]" pageId="14" pageNumber="549">
<emphasis id="B94EFB4BC134FFCECAC01B32FE469403" bold="true" box="[151,407,571,595]" pageId="14" pageNumber="549">Differential diagnosis.</emphasis>
<taxonomicName id="4C3A5CDAC134FFCECBF61B34FCF69404" authority="Jákl, 2011" authorityName="Jákl" authorityYear="2011" box="[417,807,573,596]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="subSpecies" species="pugnator" status="subsp. nov." subSpecies="insulanus">
<emphasis id="B94EFB4BC134FFCECBF61B34FCF69404" box="[417,807,573,596]" italics="true" pageId="14" pageNumber="549">Thaumastopeus pugnator insulanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D4630C134FFCEC9651B32FC7F9403" box="[818,942,571,595]" pageId="14" pageNumber="549" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
differs from the nominotypical subspecies by the much less developed punctation of pronotum and elytra, shining appearence, body slender and more parallel-sided, and also by smaller size. From
<taxonomicName id="4C3A5CDAC134FFCECAC01B94FD4F94E3" authority="Allard, 1995" authorityName="Allard" authorityYear="1995" box="[151,670,667,692]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="subSpecies" species="pugnator" subSpecies="arrowi">
<emphasis id="B94EFB4BC134FFCECAC01B94FDD494E4" box="[151,517,669,692]" italics="true" pageId="14" pageNumber="549">Thaumastopeus pugnator arrowi</emphasis>
<bibRefCitation id="EFAB5AA8C134FFCEC8591B92FD4F94E3" author="ALLARD V." box="[526,670,667,691]" pageId="14" pageNumber="549" refId="ref7487" refString="ALLARD V. 1995: Les Coleopteres du Monde. The Beetles of the World. 24: Schizorhinini 2. Sciences Nat, Venette, 136 pp + 15 pls." type="book" year="1995">Allard, 1995</bibRefCitation>
</taxonomicName>
, described from northern
<collectingCountry id="F32D67C9C134FFCEC9901B92FBFA94E3" box="[967,1067,667,691]" name="Thailand" pageId="14" pageNumber="549">Thailand</collectingCountry>
,
<taxonomicNameLabel id="A27D4630C134FFCECE6E1B95FED89483" pageId="14" pageNumber="549" rank="subSpecies">new subspecies</taxonomicNameLabel>
differs by the presence of elytral punctured lines (
<taxonomicName id="4C3A5CDAC134FFCEC97D1BB4FC299484" authorityName="Allard" authorityYear="1995" box="[810,1016,700,724]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="subSpecies" species="pugnator" subSpecies="arrowi">
<emphasis id="B94EFB4BC134FFCEC97D1BB4FCE89484" box="[810,825,701,724]" italics="true" pageId="14" pageNumber="549">T</emphasis>
.
<emphasis id="B94EFB4BC134FFCEC9101BB4FC299484" box="[839,1016,701,724]" italics="true" pageId="14" pageNumber="549">pugnator arrowi</emphasis>
</taxonomicName>
has elytra only indistinctly punctate) and by the punctation of the pronotum, which is finer than in the nominotypical subspecies, but still present (almost glabrous in
<taxonomicName id="4C3A5CDAC134FFCEC9601BF4FBD59544" authorityName="Allard" authorityYear="1995" box="[823,1028,764,788]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="subSpecies" species="pugnator" subSpecies="arrowi">
<emphasis id="B94EFB4BC134FFCEC9601BF4FC979544" box="[823,838,765,788]" italics="true" pageId="14" pageNumber="549">T</emphasis>
.
<emphasis id="B94EFB4BC134FFCEC9041BF4FBD59544" box="[851,1028,765,788]" italics="true" pageId="14" pageNumber="549">pugnator arrowi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C32074D2C134FFCECAC01A15FE979524" pageId="14" pageNumber="549" type="etymology">
<paragraph id="8B852759C134FFCECAC01A15FE979524" blockId="14.[151,1127,219,915]" pageId="14" pageNumber="549">
<emphasis id="B94EFB4BC134FFCECAC01A15FEC69564" bold="true" box="[151,279,796,820]" pageId="14" pageNumber="549">Etymology.</emphasis>
The name emphasizes that this is the first population of
<taxonomicName id="4C3A5CDAC134FFCEC90D1A14FBB79564" authorityName="Heller" authorityYear="1899" box="[858,1126,797,820]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="pugnator">
<emphasis id="B94EFB4BC134FFCEC90D1A14FBB79564" box="[858,1126,797,820]" italics="true" pageId="14" pageNumber="549">Thaumastopeus pugnator</emphasis>
</taxonomicName>
discovered on a small island off the continental mainland or large islands of the Great Sundas (
<collectingRegion id="49FEE9BBC134FFCECAC81A55FF2D9524" box="[159,252,860,884]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="549">Sumatra</collectingRegion>
,
<collectingRegion id="49FEE9BBC134FFCECB511A55FEEB9524" box="[262,314,860,884]" country="Indonesia" name="Jawa Timur" pageId="14" pageNumber="549">Java</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32074D2C134FFCECAC01A72FDF495C3" box="[151,549,891,915]" pageId="14" pageNumber="549" type="distribution">
<paragraph id="8B852759C134FFCECAC01A72FDF495C3" blockId="14.[151,1127,219,915]" box="[151,549,891,915]" pageId="14" pageNumber="549">
<emphasis id="B94EFB4BC134FFCECAC01A72FEFB95C3" bold="true" box="[151,298,891,915]" pageId="14" pageNumber="549">Distribution.</emphasis>
Nias Island,
<collectingCountry id="F32D67C9C134FFCECBE11A72FDF095C3" box="[438,545,891,915]" name="Indonesia" pageId="14" pageNumber="549">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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