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<document id="CE9A090096CA193B11E5FBF5FE462A95" ID-DOI="10.5281/zenodo.5321314" ID-ISSN="0374-1036" ID-Zenodo-Dep="5321314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630137080257" checkinUser="felipe" docAuthor="Gentili, Elio &amp; Fikaček, Martin" docDate="2009" docId="039387B6FFA915647FD4FD51FD42FBC7" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.607-623.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Laccobius (Glyptolaccobius) guttalis Gentili &amp; Fikaček 2009, sp. nov." docType="treatment" docVersion="4" lastPageNumber="612" masterDocId="FFAAFFCEFFAD15617E5CFFE8FFA7FFD4" masterDocTitle="Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae)" masterLastPageNumber="623" masterPageNumber="607" pageNumber="611" updateTime="1699163682439" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5C2F293A8CD287EEB3EDD5F14F4C8A74">Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae)</mods:title>
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<mods:name id="F1C3D728A892B6CDE22B48196AFD1445" type="personal">
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<mods:namePart id="21DAB4D7F9DF65F04ACBF0D2B83D312D">Gentili, Elio</mods:namePart>
<mods:affiliation id="8A8BE0A413647766E67A353B672C5DCE">Via San Gottardo 37, I- 21030 Varese-Rasa, Italy; e-mail: elio. gentili. 32 @ alice. it</mods:affiliation>
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<mods:namePart id="0085F7286E404C5E45920DE1287864AC">Fikaček, Martin</mods:namePart>
<mods:affiliation id="728DCB4FD3B37D91129D4806F174A06A">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ seznam. cz &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:title id="0C08201A6C4BC5D4DCB4831C4D282E4F">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="7EE001DDEF0779C1C5DD311D0875905A">
<mods:date id="F75C408603570AF0A6CB070D2D4106E1">2009</mods:date>
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<mods:number id="F6F098CD4C5768EFDDA88F0F58B5EF0F">2009-12-15</mods:number>
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<mods:number id="5721CCC1A99250E943A1928BB600CC5A">49</mods:number>
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<mods:identifier id="49D30F198019CBBFBBBD19C6C883C098" type="DOI">10.5281/zenodo.5321314</mods:identifier>
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<treatment id="039387B6FFA915647FD4FD51FD42FBC7" ID-DOI="http://doi.org/10.5281/zenodo.5342794" ID-Zenodo-Dep="5342794" LSID="urn:lsid:plazi:treatment:039387B6FFA915647FD4FD51FD42FBC7" httpUri="http://treatment.plazi.org/id/039387B6FFA915647FD4FD51FD42FBC7" lastPageId="5" lastPageNumber="612" pageId="4" pageNumber="611">
<subSubSection id="C320652BFFA915657FD4FD51FCD1FD06" box="[392,886,697,722]" pageId="4" pageNumber="611" type="nomenclature">
<paragraph id="8B8536A0FFA915657FD4FD51FCD1FD06" blockId="4.[392,886,697,722]" box="[392,886,697,722]" pageId="4" pageNumber="611">
<heading id="D0CD81CCFFA915657FD4FD51FCD1FD06" bold="true" box="[392,886,697,722]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="611" reason="3">
<emphasis id="B94EEAB2FFA915657FD4FD51FCD1FD06" bold="true" box="[392,886,697,722]" pageId="4" pageNumber="611">
<taxonomicName id="4C3A4D23FFA915657FD4FD51FCBDFD06" ID-CoL="3RLKW" authority="Gentili &amp; Fikaček, 2009" authorityName="Gentili &amp; Fikaček" authorityYear="2009" box="[392,794,697,722]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="611" phylum="Arthropoda" rank="species" species="guttalis" status="sp. nov." subGenus="Glyptolaccobius">
<emphasis id="B94EEAB2FFA915657FD4FD51FE5EFD06" bold="true" box="[392,505,697,722]" italics="true" pageId="4" pageNumber="611">Laccobius</emphasis>
(
<emphasis id="B94EEAB2FFA915657C54FD51FD1FFD06" bold="true" box="[520,696,697,722]" italics="true" pageId="4" pageNumber="611">Glyptolaccobius</emphasis>
)
<emphasis id="B94EEAB2FFA915657C94FD51FCBDFD06" bold="true" box="[712,794,697,722]" italics="true" pageId="4" pageNumber="611">guttalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57C9FFA915657D7CFD52FCD1FD06" box="[800,886,698,722]" pageId="4" pageNumber="611" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320652BFFA915657C0CFD30FD09FD3F" box="[592,686,728,747]" pageId="4" pageNumber="611" type="description">
<paragraph id="8B8536A0FFA915657C0CFD30FD09FD3F" blockId="4.[592,686,728,747]" box="[592,686,728,747]" pageId="4" pageNumber="611">
(
<figureCitation id="13012A25FFA915657C0AFD30FD35FD3F" box="[598,658,728,747]" captionStart="Figs" captionStartId="6.[151,190,1506,1525]" captionTargetBox="[161,1105,221,1387]" captionTargetId="figure-10@6.[151,1131,206,1387]" captionTargetPageId="6" captionText="Figs. 13. Aedeagus (a ventral view, b dorsal view, c lateral view). 1 L. guttalis sp. nov.; 2 L. hanka sp. nov.; 3 L. josefi sp. nov. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5321316" httpUri="https://zenodo.org/record/5321316/files/figure.png" pageId="4" pageNumber="611">Figs. 1</figureCitation>
,
<figureCitation id="13012A25FFA915657CC0FD30FD00FD3F" box="[668,679,728,747]" captionStart="Figs" captionStartId="12.[151,190,1531,1550]" captionTargetBox="[154,1111,222,1449]" captionTargetId="figure-10@12.[151,1117,219,1450]" captionTargetPageId="12" captionText="Figs.710. Habitus of Glyptolaccobius species.7 L. guttalis sp. nov., male, holotype; 8 L. moriyai Kamite, Ogata &amp; Hikida, 2007, female; 9 L. josefi sp. nov., male, paratype; 10 L. sipeki sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5321320" httpUri="https://zenodo.org/record/5321320/files/figure.png" pageId="4" pageNumber="611">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320652BFFA915657ECBFCEAFEEEFC9E" pageId="4" pageNumber="611" type="materials_examined">
<paragraph id="8B8536A0FFA915657ECBFCEAFCE9FCC1" blockId="4.[151,1126,769,842]" box="[151,846,769,789]" pageId="4" pageNumber="611">
<emphasis id="B94EEAB2FFA915657ECBFCEAFEB7FCC1" bold="true" box="[151,272,769,789]" pageId="4" pageNumber="611">
<typeStatus id="54818802FFA915657ECBFCEAFF63FCC1" box="[151,196,770,789]" pageId="4" pageNumber="611">Type</typeStatus>
locality.
</emphasis>
Central
<collectingCountry id="F32D7630FFA915657F05FCEAFE29FCC1" box="[345,398,770,789]" name="Nepal" pageId="4" pageNumber="611">Nepal</collectingCountry>
,
<collectingRegion id="49FEF842FFA915657FCBFCEAFE46FCC1" box="[407,481,770,789]" country="Nepal" name="Bagmati" pageId="4" pageNumber="611">Bagmati</collectingRegion>
, Sindhupalchok, Sarmatang,
<quantity id="4CC29B45FFA915657C86FCEAFCBCFCC1" box="[730,795,770,789]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="611" unit="m" value="2500.0">2500 m</quantity>
a.s.l..
</paragraph>
<paragraph id="8B8536A0FFA915657ECBFCF5FEEEFC9E" blockId="4.[151,1126,769,842]" pageId="4" pageNumber="611">
<emphasis id="B94EEAB2FFA915657ECBFCF5FEBDFCE4" bold="true" box="[151,282,796,816]" pageId="4" pageNumber="611">Type material.</emphasis>
<materialsCitation id="3B523CFDFFA915657F42FCF5FEEEFC9E" collectingDate="1989-06-03" collectingDateMax="1989-06-08" collectingDateMin="1989-06-03" collectionCode="NKME" collectorName="C. Holzschuh'" country="Nepal" elevation="2500" location="Sindhupalchok" pageId="4" pageNumber="611" specimenCount="1" stateProvince="Bagmati" typeStatus="holotype">
<typeStatus id="54818802FFA915657F42FCF5FED6FCFA" box="[286,369,797,816]" pageId="4" pageNumber="611" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED2BAE65FFA915657FC9FCF5FE7EFCE4" box="[405,473,797,816]" pageId="4" pageNumber="611">NKME</collectionCode>
): C.
<collectingCountry id="F32D7630FFA915657C58FCF5FDEEFCE4" box="[516,585,797,816]" name="Nepal" pageId="4" pageNumber="611">NEPAL</collectingCountry>
,
<collectingRegion id="49FEF842FFA915657C0CFCF4FD3FFCFB" box="[592,664,796,815]" country="Nepal" name="Bagmati" pageId="4" pageNumber="611">Bagmati</collectingRegion>
/
<location id="8EE5607BFFA915657CFAFCF4FC81FCFB" LSID="urn:lsid:plazi:treatment:039387B6FFA915647FD4FD51FD42FBC7:8EE5607BFFA915657CFAFCF4FC81FCFB" box="[678,806,796,815]" country="Nepal" name="Sindhupalchok" pageId="4" pageNumber="611" stateProvince="Bagmati">Sindhupalchok</location>
/
<quantity id="4CC29B45FFA915657D68FCF4FCD1FCE4" box="[820,886,796,816]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="611" unit="m" value="2500.0">
<elevation id="0017D193FFA915657D68FCF4FCD1FCE4" box="[820,886,796,816]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="611" unit="m" value="2500.0">2500 m</elevation>
</quantity>
, Sarmatang /
<date id="FF841060FFA915657DB9FCF5FBC1FCE4" box="[997,1126,797,816]" pageId="4" pageNumber="611" value="1989-06-03" valueMax="1989-06-08" valueMin="1989-06-03">
<collectingDate id="EFC0E988FFA915657DB9FCF5FBC1FCE4" box="[997,1126,797,816]" pageId="4" pageNumber="611" value="1989-06-03" valueMax="1989-06-08" valueMin="1989-06-03">0308.VI.1989</collectingDate>
</date>
/ leg.
<collectorName id="26CF5376FFA915657E9BFCDFFEE2FC9E" box="[199,325,823,842]" pageId="4" pageNumber="611">C. Holzschuh</collectorName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C320652BFFA915647ECBFC8AFCB1FD07" lastPageId="5" lastPageNumber="612" pageId="4" pageNumber="611" type="description">
<paragraph id="8B8536A0FFA915657ECBFC8AFC19FC4F" blockId="4.[151,1127,866,1627]" pageId="4" pageNumber="611">
<emphasis id="B94EEAB2FFA915657ECBFC8AFE85FCAE" bold="true" box="[151,290,866,890]" pageId="4" pageNumber="611">Description.</emphasis>
Total length
<quantity id="4CC29B45FFA915657FF5FC8AFDAEFCAE" box="[425,521,866,890]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="4" pageNumber="611" unit="mm" value="2.15">2.15 mm</quantity>
, total width
<quantity id="4CC29B45FFA915657CD4FC8AFD4EFCAE" box="[648,745,866,890]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="611" unit="mm" value="1.25">1.25 mm</quantity>
. Maximum length / maximum width ratio 1.7. Body wide oval, convex, maximum width in anterior elytral third.
</paragraph>
<paragraph id="8B8536A0FFA915657EE9FC4BFCFAFAAE" blockId="4.[151,1127,866,1627]" pageId="4" pageNumber="611">
Head. Coloration entirely black, with rare and inconspicuous punctures; shining, smooth without microsculpture; systematic punctures indistinct; anterolateral branches of frontoclypeal suture conspicuous and punctate at 100×. Labrum black, trapezoid, without specula; anterior margin nearly straight, slightly emarginated medially; lateral margins oblique, posterior margin crescent-shaped. Eyes oblong, oblique, closest to each other posteriorly, not protruding laterad, interocular distance equal to 2.5× of width of one eye; postocular portion of tempora short.
<collectingRegion id="49FEF842FFA915657FEDFB8BFDACFBAE" box="[433,523,1123,1146]" country="Myanmar" name="Chin" pageId="4" pageNumber="611">Mentum</collectingRegion>
flat, smooth without punctures, emarginated anteriorly. Submentum and gula microgranulate. Maxillary palpi yellow-brown; palpomere 1 thin and short; palpomeres 2 and 3 nearly equal in length; palpomere 3 dilated at apex; palpomere 4 elongate, nearly twice as long as palpomere 3, asymmetrical, inner margin straight and outer margin convex, apex truncate. Eight antennomeres; scape longer than antennomeres 24 combined; pedicel cone-shaped; intermediate antennomeres (antennomeres 34) very short; cupule asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, bearing densely arranged setae, antennomeres 6 and 8 nearly equal in length, antennomere 6 wider, antennomere 7 shorter and narrower.
</paragraph>
<paragraph id="8B8536A0FFA915647EE9FA6BFC30FEA7" blockId="4.[151,1127,866,1627]" lastBlockId="5.[113,1090,219,1044]" lastPageId="5" lastPageNumber="612" pageId="4" pageNumber="611">Thorax. Pronotum transverse, 0.35× as long as wide; black with yellowish lateral margins, yellow margin widened posteriorly and continuing as fine yellow strip along base; surface smooth, without microsculpture, with some sparse and faint punctures. Prosternum black, with longitudinal keel. Scutellar shield equilateral, black, bearing very small punctures. Mesoventrite with longitudinal keel simple, not tuberculate anteriorly. Elytra combined slightly elongate, ca 1.20× as long as wide; shining black with yellowish-brown lateral margins and apices, and pale yellow dots and stripes; 10 longitudinal rows of setiferous, more impressed punctures alternating with 10 rows (interstriae) of sparser and fainter punctures. Parasutural space covered by oblique lines reaching first punctural row in anterior fourth and then forming a flat and shining stripe separating suture from parasutural furrow, stripe and furrow disappearing just before elytral apex. Epipleura and pseudoepipleura oblique, nearly vertical. Metaventrite setiferous, with median longitudinal glabrous area posteriorly.</paragraph>
<paragraph id="8B8536A0FFA815647ECCFE94FF1DFD87" blockId="5.[113,1090,219,1044]" pageId="5" pageNumber="612">Legs. Fore coxae and trochanters granulate, pubescent; fore femora setiferous on basal third, smooth distally, with tibial groove; fore tibiae smooth, with stiff setae and two apical spurs. Middle femora with tibial grooves; ventral face of middle tibiae with three longitudinal rows of stiff setae and sulcus between two longitudinal ridges. Hind trochanters smooth and shining; hind femora smooth, with scarce punctures and tibial grooves; hind tibiae curved, ventral face with three longitudinal rows of stiff setae and longitudinal sulcus between two ridges.</paragraph>
<paragraph id="8B8536A0FFA815647ECCFDB3FC17FDA7" blockId="5.[113,1090,219,1044]" box="[144,944,603,628]" pageId="5" pageNumber="612">Abdomen. Ventrites 14 smooth, with lateral setae; ventrites 56 wrinkled.</paragraph>
<paragraph id="8B8536A0FFA815647ECCFD93FCB1FD07" blockId="5.[113,1090,219,1044]" pageId="5" pageNumber="612">
Aedeagus (
<figureCitation id="13012A25FFA815647F5FFD93FE99FD40" box="[259,318,635,660]" captionStart="Figs" captionStartId="6.[151,190,1506,1525]" captionTargetBox="[161,1105,221,1387]" captionTargetId="figure-10@6.[151,1131,206,1387]" captionTargetPageId="6" captionText="Figs. 13. Aedeagus (a ventral view, b dorsal view, c lateral view). 1 L. guttalis sp. nov.; 2 L. hanka sp. nov.; 3 L. josefi sp. nov. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5321316" httpUri="https://zenodo.org/record/5321316/files/figure.png" pageId="5" pageNumber="612">Fig. 1</figureCitation>
). Total length
<quantity id="4CC29B45FFA815647F8FFD93FD96FD47" box="[467,561,635,659]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="612" unit="mm" value="0.68">0.68 mm</quantity>
. Parameres slightly longer than phallobase, slightly overlapping apex of median lobe. Median lobe narrow, slightly widened in apical 0.4, with longitudinal excision ventrally; apical portion broadly rounded.
</paragraph>
</subSubSection>
<subSubSection id="C320652BFFA815647E2DFD33FC92FC40" pageId="5" pageNumber="612" type="diagnosis">
<paragraph id="8B8536A0FFA815647E2DFD33FC92FC40" blockId="5.[113,1090,219,1044]" pageId="5" pageNumber="612">
<emphasis id="B94EEAB2FFA815647E2DFD33FECDFD27" bold="true" box="[113,362,731,755]" pageId="5" pageNumber="612">Differential diagnosis.</emphasis>
<taxonomicName id="4C3A4D23FFA815647F33FD35FD95FD20" authority="Gentili &amp; Fikaček, 2009" authorityName="Gentili &amp; Fikaček" authorityYear="2009" box="[367,562,733,756]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="612" phylum="Arthropoda" rank="species" species="guttalis" status="sp. nov.">
<emphasis id="B94EEAB2FFA815647F33FD35FD95FD20" box="[367,562,733,756]" italics="true" pageId="5" pageNumber="612">Laccobius guttalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57C9FFA815647C64FD34FD2EFD27" box="[568,649,732,755]" pageId="5" pageNumber="612" rank="species">sp. nov.</taxonomicNameLabel>
belongs to a group of species with curved hind tibiae, longitudinal rows of elytral punctures and black elytral surface with yellowish dots and stripes (together with
<taxonomicName id="4C3A4D23FFA815647FEAFCF5FDAEFCE0" authority="Gentili &amp; Fikaček, 2009" authorityName="Gentili &amp; Fikaček" authorityYear="2009" box="[438,521,797,820]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="612" phylum="Arthropoda" rank="species" species="josefi" status="sp. nov.">
<emphasis id="B94EEAB2FFA815647FEAFCF5FDAEFCE0" box="[438,521,797,820]" italics="true" pageId="5" pageNumber="612">L. josefi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57C9FFA815647C53FCF4FDC4FCE7" box="[527,611,796,819]" pageId="5" pageNumber="612" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C3A4D23FFA815647C30FCF5FD6AFCE0" authorityName="Gentili" authorityYear="1995" box="[620,717,797,820]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="612" phylum="Arthropoda" rank="species" species="munus">
<emphasis id="B94EEAB2FFA815647C30FCF5FD6AFCE0" box="[620,717,797,820]" italics="true" pageId="5" pageNumber="612">L. munus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D23FFA815647CA2FCF5FCD7FCE0" authorityName="Gentili" authorityYear="1995" box="[766,880,797,820]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="612" phylum="Arthropoda" rank="species" species="sharmai">
<emphasis id="B94EEAB2FFA815647CA2FCF5FCD7FCE0" box="[766,880,797,820]" italics="true" pageId="5" pageNumber="612">L. sharmai</emphasis>
</taxonomicName>
). It differs from the other three species by the presence of transverse lines in the anterior part of the parasutural stripe and the shape of the aedeagus, with the apical portion of the median lobe swollen and the median lobe bearing a longitudinal ventral excavation (
<figureCitation id="13012A25FFA815647CBAFC93FC82FC40" box="[742,805,891,916]" captionStart="Figs" captionStartId="6.[151,190,1506,1525]" captionTargetBox="[161,1105,221,1387]" captionTargetId="figure-10@6.[151,1131,206,1387]" captionTargetPageId="6" captionText="Figs. 13. Aedeagus (a ventral view, b dorsal view, c lateral view). 1 L. guttalis sp. nov.; 2 L. hanka sp. nov.; 3 L. josefi sp. nov. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5321316" httpUri="https://zenodo.org/record/5321316/files/figure.png" pageId="5" pageNumber="612">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320652BFFA815647E2DFC74FEC1FC27" pageId="5" pageNumber="612" type="etymology">
<paragraph id="8B8536A0FFA815647E2DFC74FF54FC07" blockId="5.[113,1090,219,1044]" pageId="5" pageNumber="612">
<emphasis id="B94EEAB2FFA815647E2DFC74FF55FC60" bold="true" box="[113,242,924,948]" pageId="5" pageNumber="612">Etymology.</emphasis>
From a drop (Latin:
<emphasis id="B94EEAB2FFA815647F86FC75FDB6FC60" box="[474,529,925,948]" italics="true" pageId="5" pageNumber="612">gutta</emphasis>
), alluding to the preferred seepage habitat of
<taxonomicName id="4C3A4D23FFA815647A58FC75FF4BFC00" authorityName="Gentili" authorityYear="1989" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="623" phylum="Arthropoda" rank="subGenus" subGenus="Glyptolaccobius">
<emphasis id="B94EEAB2FFA815647A58FC75FF4BFC00" italics="true" pageId="5" pageNumber="612">Glyptolaccobius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8536A0FFA815647E2DFC33FEC1FC27" blockId="5.[113,1090,219,1044]" box="[113,358,987,1011]" pageId="5" pageNumber="612">
<emphasis id="B94EEAB2FFA815647E2DFC33FF48FC27" bold="true" box="[113,239,987,1011]" pageId="5" pageNumber="612">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C320652BFFA815647E2DFC13FD42FBC7" box="[113,741,1019,1044]" pageId="5" pageNumber="612" type="distribution">
<paragraph id="8B8536A0FFA815647E2DFC13FD42FBC7" blockId="5.[113,1090,219,1044]" box="[113,741,1019,1044]" pageId="5" pageNumber="612">
<emphasis id="B94EEAB2FFA815647E2DFC13FEA3FBC7" bold="true" box="[113,260,1019,1043]" pageId="5" pageNumber="612">Distribution.</emphasis>
Known only from the
<typeStatus id="54818802FFA815647FABFC14FD82FBC0" box="[503,549,1020,1044]" pageId="5" pageNumber="612">type</typeStatus>
locality in
<collectingCountry id="F32D7630FFA815647CC2FC13FD46FBC7" box="[670,737,1019,1043]" name="Nepal" pageId="5" pageNumber="612">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>