278 lines
33 KiB
XML
278 lines
33 KiB
XML
<document id="2D9AE282BD11ECD7F04EA0D694CC9C78" ID-DOI="10.5194/fr-18-81-2015" ID-ISSN="2193-0074" ID-Zenodo-Dep="11585625" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1716844663622" checkinUser="felipe" docAuthor="Titus, A. L., Korn, D., Harrell, J. E. & Lambert, L. L." docDate="2015" docId="039287BEFC50FFA1FFA9FB2DFDAEFC11" docLanguage="en" docName="FossRec.18.2.81-104.pdf" docOrigin="Fossil Record 18 (2)" docSource="http://dx.doi.org/10.5194/fr-18-81-2015" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Delepinoceratidae Ruzhencev 1957" docType="treatment" docVersion="5" lastPageNumber="94" masterDocId="FFABFFC6FC5CFFACFFD5FFF5FFC9FF94" masterDocTitle="Late Viséan (late Mississippian) ammonoids from the Barnett Shale, Sierra Diablo Escarpment, Culberson County, Texas, USA" masterLastPageNumber="104" masterPageNumber="81" pageNumber="93" updateTime="1718146734426" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B017C1FD84C4321B8802AC7C869FEFDF">Late Viséan (late Mississippian) ammonoids from the Barnett Shale, Sierra Diablo Escarpment, Culberson County, Texas, USA</mods:title>
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<mods:name id="E6888DDA1E90F67F2A6FF13990A72723" type="personal">
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<mods:namePart id="D956AEED48075AB265CF84B96203FCC7">Titus, A. L.</mods:namePart>
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<mods:affiliation id="DC68B77F2C5EFE06ADFD5E5613B8D390">D. Korn & Grand Staircase-Escalante National Monument, 669 S. Highway 89 A, Kanab, UT 84741, USA</mods:affiliation>
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<mods:roleTerm id="931218167E572A98CF1D5A0FB682E26E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A816FA1ABEA133EEAE9C3C09EF1DB4A5">Korn, D.</mods:namePart>
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<mods:affiliation id="5E0C6F2DF5BCBBB883B2DC9B8FD33E09">J. E. Harrell & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43,</mods:affiliation>
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</mods:name>
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<mods:name id="128BC227CE6254B453037FD71BF8CF1C" type="personal">
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<mods:roleTerm id="CA555E7E569D287C604B27572F3F083E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4D1C108F273F2B15DBEF10AE5F5345FD">Harrell, J. E.</mods:namePart>
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<mods:affiliation id="D09704C5905039663B4CA9E7CE3950C1">L. L. Lambert & Noble Energy Inc., Houston, TX 77070, USA</mods:affiliation>
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<mods:roleTerm id="E86EB8264E9251D5A5EBFE203A039141">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D45ABB8C42527D4207F4181AEFC4EF39">Lambert, L. L.</mods:namePart>
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<mods:affiliation id="D1174248CD832C46ADCF2E9972D71EC7">Department of Geological Sciences, The University of Texas at San Antonio, One UTSA Circle, San Antonio, TX 78249, USA</mods:affiliation>
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<mods:typeOfResource id="828A8DC13BB352B7F8D1B36AD1E03435">text</mods:typeOfResource>
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<mods:title id="DB3EBADCE38387194B78A5C218EC70F2">Fossil Record</mods:title>
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<mods:part id="852E091A8A5881398534B9A88C3B272A">
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<mods:date id="B6F30B331A19D14E9A16224CF28002DC">2015</mods:date>
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<mods:number id="2116E186E7D6A5CC5AB105D9F40B6915">2015-06-04</mods:number>
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<mods:number id="D97F69B6BAAEC0255CEFEA7CD113EE08">18</mods:number>
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<mods:detail id="7FD4B0009A3474FD45927D72B3255583" type="issue">
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<mods:number id="1A34A2D9FB9DAA7A7523C98A79CA4C5E">2</mods:number>
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<mods:extent id="2D0BC99820FB2E6EC05A59BA39DD6B15" unit="page">
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<mods:start id="5AA3A95E801AC590ACC2FD0B92E89464">81</mods:start>
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<mods:end id="95426293FF1AD06EAB7E248B3FA237E0">104</mods:end>
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<mods:url id="CC45B329A08A20156D17AFD4077B6FC9">http://dx.doi.org/10.5194/fr-18-81-2015</mods:url>
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<mods:classification id="D1C8CFBE05137839F11674031ACF8155">journal article</mods:classification>
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<mods:identifier id="BFB8FD747123D68CC9F8CB22821A8001" type="DOI">10.5194/fr-18-81-2015</mods:identifier>
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<mods:identifier id="51470A0E38954A3EFBB8ECBD4C2ADBE9" type="ISSN">2193-0074</mods:identifier>
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</mods:mods>
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<treatment id="039287BEFC50FFA1FFA9FB2DFDAEFC11" LSID="urn:lsid:plazi:treatment:039287BEFC50FFA1FFA9FB2DFDAEFC11" httpUri="http://treatment.plazi.org/id/039287BEFC50FFA1FFA9FB2DFDAEFC11" lastPageId="13" lastPageNumber="94" pageId="12" pageNumber="93">
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<subSubSection id="C3216523FC50FFA0FFA9FB2DFD82FB7B" box="[124,587,1240,1263]" pageId="12" pageNumber="93" type="nomenclature">
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<paragraph id="8B8436A8FC50FFA0FFA9FB2DFD82FB7B" blockId="12.[124,587,1240,1263]" box="[124,587,1240,1263]" pageId="12" pageNumber="93">
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<heading id="D0CC81C4FC50FFA0FFA9FB2DFD82FB7B" box="[124,587,1240,1263]" fontSize="10" level="2" pageId="12" pageNumber="93" reason="2">
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Family
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<taxonomicName id="4C3B4D2BFC50FFA0FF1BFB2DFD82FB7B" ID-CoL="92T" authority="Ruzhencev, 1957" authorityName="Ruzhencev" authorityYear="1957" box="[206,587,1240,1263]" class="Cephalopoda" family="Delepinoceratidae" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="family">
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Delepinoceratidae
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<bibRefCitation id="EFAA4B59FC50FFA0FE40FB2DFD82FB7B" author="Ruzhencev, V. E." box="[405,587,1240,1263]" pageId="12" pageNumber="93" pagination="49 - 64" refId="ref18794" refString="Ruzhencev, V. E.: Filogeneticheskaya sistema paleozoiskikh ammonoidei., Byulleten' Moskovskogo obshchestva ispytatelei prirody, otdelenie geologii, 32, 49 - 64, 1957." type="journal article" year="1957">Ruzhencev, 1957</bibRefCitation>
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</taxonomicName>
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</heading>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C3216523FC50FFA0FFA9FAEDFE23FA3B" pageId="12" pageNumber="93" type="diagnosis">
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<paragraph id="8B8436A8FC50FFA0FFA9FAEDFE23FA3B" blockId="12.[124,767,1304,1901]" pageId="12" pageNumber="93">
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Family diagnosis: Representatives of the superfamily
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<taxonomicName id="4C3B4D2BFC50FFA0FD14FAEDFF10FADB" authorityName="de Haan" authorityYear="1825" pageId="12" pageNumber="93" rank="superFamily" superFamily="Goniatitaceae">Goniatitaceae</taxonomicName>
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with advanced suture lines possessing a Y-shaped external lobe and a tectiform ventrolateral saddle. The adventive lobe and also the prongs of the external lobe show the tendency to become tridentate.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3216523FC50FFA1FFA9FA4DFDAEFC11" lastPageId="13" lastPageNumber="94" pageId="12" pageNumber="93" type="discussion">
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<paragraph id="8B8436A8FC50FFA0FFA9FA4DFCA9FADB" blockId="12.[124,767,1304,1901]" lastBlockId="12.[819,1462,1240,1901]" pageId="12" pageNumber="93">
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Discussion: There exist three diverging concepts about the classification of the advanced goniatitid species and genera.
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<bibRefCitation id="EFAA4B59FC50FFA0FFA9FA02FDD1F99A" author="Ruzhencev, V. E. & Bogoslovskaya, M. F." box="[124,536,1527,1550]" pageId="12" pageNumber="93" pagination="1 - 382" refId="ref18853" refString="Ruzhencev, V. E. and Bogoslovskaya, M. F.: Namyurskiy etap v evolyutsii ammonodey. Rannenamyurskiye ammonoidei., Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 133, 1 - 382, 1971." type="journal article" year="1971">Ruzhencev and Bogoslovskaya (1971</bibRefCitation>
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,
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<bibRefCitation id="EFAA4B59FC50FFA0FDF2FA02FD94F99A" author="Ruzhencev, V. E. & Bogoslovskaya, M. F." box="[551,605,1527,1550]" pageId="12" pageNumber="93" pagination="1 - 336" refId="ref18892" refString="Ruzhencev, V. E. and Bogoslovskaya, M. F.: Namyurskiy etap v evolyutsii ammonodey. Pozdnenamyurskiye ammonoidei, Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 167, 1 - 336, 1978." type="journal article" year="1978">1978</bibRefCitation>
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) separated the families
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<taxonomicName id="4C3B4D2BFC50FFA0FF0AF9E2FE69F9BA" authorityName="Ruzhencev" authorityYear="1957" box="[223,416,1559,1582]" class="Cephalopoda" family="Delepinoceratidae" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="family">Delepinoceratidae</taxonomicName>
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(including
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<taxonomicName id="4C3B4D2BFC50FFA0FDFCF9E2FD03F9BB" authorityName="Ruzhentzev" authorityYear="1956" box="[553,714,1559,1583]" class="Cephalopoda" family="Delepinoceratidae" genus="Platygoniatites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FDFCF9E2FD03F9BB" box="[553,714,1559,1583]" italics="true" pageId="12" pageNumber="93">Platygoniatites</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3B4D2BFC50FFA0FFA9F9C2FEDEF9DB" authorityName="Miller & Furnish" authorityYear="1954" box="[124,279,1591,1615]" class="Cephalopoda" family="Delepinoceratidae" genus="Delepinoceras" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FFA9F9C2FEDEF9DB" box="[124,279,1591,1615]" italics="true" pageId="12" pageNumber="93">Delepinoceras</emphasis>
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</taxonomicName>
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) and
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<taxonomicName id="4C3B4D2BFC50FFA0FE80F9C2FE34F9DA" box="[341,509,1591,1614]" class="Cephalopoda" family="Agathiceratidae" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="family">Agathiceratidae</taxonomicName>
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(including
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<taxonomicName id="4C3B4D2BFC50FFA0FDAFF9C2FD3EF9DB" authorityName="Librovitch" authorityYear="1957" box="[634,759,1591,1615]" class="Cephalopoda" family="Delepinoceratidae" genus="Dombarites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FDAFF9C2FD3EF9DB" box="[634,759,1591,1615]" italics="true" pageId="12" pageNumber="93">Dombarites</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B4D2BFC50FFA0FFA9F9A3FF22F9FA" baseAuthorityName="Renz" baseAuthorityYear="1910" box="[124,235,1622,1646]" class="Cephalopoda" family="Agathiceratidae" genus="Pericleites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FFA9F9A3FF22F9FA" box="[124,235,1622,1646]" italics="true" pageId="12" pageNumber="93">Pericleites</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B4D2BFC50FFA0FF28F9A3FE6CF9FA" authorityName="Rauser-Tschernoussowa" authorityYear="1928" box="[253,421,1622,1646]" class="Cephalopoda" family="Delepinoceratidae" genus="Proshumardites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FF28F9A3FE6CF9FA" box="[253,421,1622,1646]" italics="true" pageId="12" pageNumber="93">Proshumardites</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C3B4D2BFC50FFA0FE33F9A3FDAFF9FA" authorityName="Gemmellaro" authorityYear="1888" box="[486,614,1622,1646]" class="Cephalopoda" family="Agathiceratidae" genus="Agathiceras" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FE33F9A3FDAFF9FA" box="[486,614,1622,1646]" italics="true" pageId="12" pageNumber="93">Agathiceras</emphasis>
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</taxonomicName>
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) on the basis of the subdivision mode of the external lobe, thus delineating two independent evolutionary lineages.
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<bibRefCitation id="EFAA4B59FC50FFA0FD80F962FD33F93A" author="Leonova, T. B." box="[597,762,1687,1710]" pageId="12" pageNumber="93" pagination="1 - 114" refId="ref17986" refString="Leonova, T. B.: Permian Ammonoids: Classification and Phylogeny, Paleontol. J, Supplement, 36, 1 - 114, 2002." type="journal article" year="2002">Leonova (2002</bibRefCitation>
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,
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<bibRefCitation id="EFAA4B59FC50FFA0FFA9F942FF7BF95A" author="Leonova, T. B." box="[124,178,1719,1742]" pageId="12" pageNumber="93" pagination="1206 - 1312" refId="ref18014" refString="Leonova, T. B.: Permian ammonoids: Biostratigraphic, biogeographical, and ecological analysis, Paleontol. J., 45, 1206 - 1312, 2011." type="journal article" year="2011">2011</bibRefCitation>
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) doubted the assignment of
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<taxonomicName id="4C3B4D2BFC50FFA0FDD7F943FD4BF95A" authorityName="Gemmellaro" authorityYear="1888" box="[514,642,1718,1742]" class="Cephalopoda" family="Agathiceratidae" genus="Agathiceras" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FDD7F943FD4BF95A" box="[514,642,1718,1742]" italics="true" pageId="12" pageNumber="93">Agathiceras</emphasis>
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</taxonomicName>
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in the order
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<taxonomicName id="4C3B4D2BFC50FFA0FF7CF923FEE8F979" authorityName="Hyatt" authorityYear="1884" box="[169,289,1750,1773]" class="Cephalopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="order">Goniatitida</taxonomicName>
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and transferred it into the order
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<taxonomicName id="4C3B4D2BFC50FFA0FD42F923FF79F899" order="Tornoceratida" pageId="12" pageNumber="93" rank="order">Tornoceratida</taxonomicName>
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(which has usually only been accepted as a suborder within the
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<taxonomicName id="4C3B4D2BFC50FFA0FF24F8E3FEA7F8B9" authorityName="Hyatt" authorityYear="1884" box="[241,366,1814,1837]" class="Cephalopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="order">Goniatitida</taxonomicName>
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). Finally,
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<bibRefCitation id="EFAA4B59FC50FFA0FE0CF8E3FD06F8B9" author="Kullmann, J. & Wagner, R. H. & Winkler Prins, C." box="[473,719,1814,1837]" pageId="12" pageNumber="93" pagination="127 - 145" refId="ref17867" refString="Kullmann, J., Wagner, R. H., and Winkler Prins, C.: Significance for international correlation of the Perapertu Formation in northern Palencia, Cantabrian Mountains. Tectonic / stratigraphic context and description of Mississippian and Upper Bashkirian goniatites, Revista Espanola de Paleontologia, 22, 127 - 145, 2007." type="journal article" year="2007">Kullmann et al. (2007)</bibRefCitation>
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and
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<bibRefCitation id="EFAA4B59FC50FFA0FFA9F8C3FEF2F8D9" author="Kullmann, J." box="[124,315,1846,1869]" pageId="12" pageNumber="93" pagination="52 - 60" refId="ref17702" refString="Kullmann, J.: Goniatitoidea, in: Treatise on Invertebrate Paleontology. Part L. Mollusca 4 Revised. Volume 2: Carboniferous and Permian Ammonoidea (Goniatitida and Prolecanitida), edited by: Seldon, P. A., The University of Kansas Paleontological Institute, Lawrence, Kansas, 52 - 60, 2009 a." type="book chapter" year="2009">Kullmann (2009a</bibRefCitation>
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, b) separated
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<taxonomicName id="4C3B4D2BFC50FFA0FE00F8C3FD9CF8DA" authorityName="Gemmellaro" authorityYear="1888" box="[469,597,1846,1870]" class="Cephalopoda" family="Agathiceratidae" genus="Agathiceras" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FE00F8C3FD9CF8DA" box="[469,597,1846,1870]" italics="true" pageId="12" pageNumber="93">Agathiceras</emphasis>
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</taxonomicName>
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and closely related forms in the independent superfamily
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<taxonomicName id="4C3B4D2BFC50FFA0FD9CF8A3FD37F8F9" box="[585,766,1878,1901]" class="Cephalopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="superFamily" superFamily="Agathiceratoidea">Agathiceratoidea</taxonomicName>
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within the suborder
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<taxonomicName id="4C3B4D2BFC50FFA0FBD8FB2DFB41FB7B" authorityName="Hyatt" authorityYear="1884" box="[1037,1160,1240,1263]" class="Cephalopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="subOrder" subOrder="Goniatitina">Goniatitina</taxonomicName>
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. In Kullmann’s concept, the genera
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<taxonomicName id="4C3B4D2BFC50FFA0FC51FB0DFBC8FA84" authorityName="Librovitch" authorityYear="1957" box="[900,1025,1272,1296]" class="Cephalopoda" family="Delepinoceratidae" genus="Dombarites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
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<emphasis id="B94FEABAFC50FFA0FC51FB0DFBC8FA84" box="[900,1025,1272,1296]" italics="true" pageId="12" pageNumber="93">Dombarites</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3B4D2BFC50FFA0FBEEFB0DFB2AFA84" authorityName="Rauser-Tschernoussowa" authorityYear="1928" box="[1083,1251,1272,1296]" class="Cephalopoda" family="Delepinoceratidae" genus="Proshumardites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC50FFA0FBEEFB0DFB2AFA84" box="[1083,1251,1272,1296]" italics="true" pageId="12" pageNumber="93">Proshumardites</emphasis>
|
||
</taxonomicName>
|
||
are grouped in the subfamily
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FC7CFAEDFBF2FABB" baseAuthorityName="Kullmann" baseAuthorityYear="2009" box="[937,1083,1304,1327]" class="Cephalopoda" family="Delepinoceratidae" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="subFamily" subFamily="Dombaritinae">Dombaritinae</taxonomicName>
|
||
within the family
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FAC4FAEDFC92FADB" authorityName="Ruzhencev" authorityYear="1957" class="Cephalopoda" family="Delepinoceratidae" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="family">Delepinoceratidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8436A8FC50FFA0FC9BFAADFBA9F91A" blockId="12.[819,1462,1240,1901]" pageId="12" pageNumber="93">
|
||
If
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FCB3FAA2FC2FFAFB" ID-CoL="QLT" authorityName="Gemmellaro" authorityYear="1888" box="[870,998,1367,1391]" class="Cephalopoda" family="Agathiceratidae" genus="Agathiceras" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC50FFA0FCB3FAA2FC2FFAFB" box="[870,998,1367,1391]" italics="true" pageId="12" pageNumber="93">Agathiceras</emphasis>
|
||
</taxonomicName>
|
||
is excluded from the discussion of the phylogeny of the family
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FBF4FA8DFB6DFA1B" ID-CoL="623MK" authorityName="de Haan" authorityYear="1825" box="[1057,1188,1400,1423]" class="Cephalopoda" family="Goniatitidae" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="family">Goniatitidae</taxonomicName>
|
||
and its descendants (for instance, if the central or sub-central position of the siphuncle in the juvenile stage is been taken as a reason for separation on higher taxonomic levels), then the phylogenetic analysis (see below) speaks for the extension of the family
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FCE6F9E2FC3DF9BA" ID-CoL="92T" authorityName="Ruzhencev" authorityYear="1957" box="[819,1012,1559,1582]" class="Cephalopoda" family="Delepinoceratidae" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="family">Delepinoceratidae</taxonomicName>
|
||
to also include the subfamily
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FA95F9E2FBFCF9DA" authority="(Kullmann, 2009 a)" baseAuthorityName="Kullmann" baseAuthorityYear="2009" class="Cephalopoda" family="Delepinoceratidae" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="subFamily" subFamily="Dombaritinae">
|
||
Dombaritinae (
|
||
<bibRefCitation id="EFAA4B59FC50FFA0FCBEF9C2FBE4F9DA" author="Kullmann, J." box="[875,1069,1591,1614]" pageId="12" pageNumber="93" pagination="52 - 60" refId="ref17702" refString="Kullmann, J.: Goniatitoidea, in: Treatise on Invertebrate Paleontology. Part L. Mollusca 4 Revised. Volume 2: Carboniferous and Permian Ammonoidea (Goniatitida and Prolecanitida), edited by: Seldon, P. A., The University of Kansas Paleontological Institute, Lawrence, Kansas, 52 - 60, 2009 a." type="book chapter" year="2009">Kullmann, 2009a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. This subfamily, however, also requires re-definition to include the genera
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FB38F9A3FAB8F9FA" authorityName="Wagner-Gentis" authorityYear="1980" box="[1261,1393,1622,1646]" class="Cephalopoda" family="Delepinoceratidae" genus="Revilloceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC50FFA0FB38F9A3FAB8F9FA" box="[1261,1393,1622,1646]" italics="true" pageId="12" pageNumber="93">Revilloceras</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FAAAF9A3FC52F91A" authorityName="Pereira de Sousa" authorityYear="1923" class="Cephalopoda" family="Goniatitidae" genus="Lusitanoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC50FFA0FAAAF9A3FC52F91A" italics="true" pageId="12" pageNumber="93">Lusitanoceras</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FC03F983FB90F91A" ID-CoL="62LH8" authorityName="Titus & Korn & Harrell & Lambert" authorityYear="2015" box="[982,1113,1654,1678]" class="Cephalopoda" family="Delepinoceratidae" genus="Choctawites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC50FFA0FC03F983FB90F91A" box="[982,1113,1654,1678]" italics="true" pageId="12" pageNumber="93">Choctawites</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8436A8FC50FFA1FC9BF962FD93FF73" blockId="12.[819,1462,1240,1901]" lastBlockId="13.[124,767,176,901]" lastPageId="13" lastPageNumber="94" pageId="12" pageNumber="93">
|
||
For resolving the phylogenetic relationships of the goniatitid ammonoid genera, we performed a cladistic analysis based on the character matrix of
|
||
<bibRefCitation id="EFAA4B59FC50FFA0FB75F923FAFCF979" author="Korn, D." box="[1184,1333,1750,1773]" pageId="12" pageNumber="93" pagination="177 - 199" refId="ref17094" refString="Korn, D.: Evolution of the Goniatitaceae and Visean-Namurian biogeography, Acta Palaeontol. Pol., 42, 177 - 199, 1997 a." type="journal article" year="1997">Korn (1997a)</bibRefCitation>
|
||
. In contrast to that analysis, we used
|
||
<emphasis id="B94FEABAFC50FFA0FB9BF903FAECF89A" box="[1102,1317,1782,1806]" italics="true" pageId="12" pageNumber="93">Progoniatites uncus</emphasis>
|
||
<bibRefCitation id="EFAA4B59FC50FFA0FAFAF903FB48F8B9" author="Korn, D. & Bockwinkel, J. & Ebbighausen, V." pageId="12" pageNumber="93" pagination="35 - 152" refId="ref17642" refString="Korn, D., Bockwinkel, J., and Ebbighausen, V.: The ammonoids from the Argiles de Teguentour of Oued Temertasset (early Late Tournaisian; Mouydir, Algeria), Foss. Rec., 13, 35 - 152, doi: 10.5194 / fr- 13 - 35 - 2010, 2010." type="journal article" year="2010">Korn, Bockwinkel and Ebbighausen, 2010</bibRefCitation>
|
||
as the outgroup species and included three more ammonoid species in the new analysis,
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FCE6F8A0FB82F8F9" ID-CoL="5XYLY" box="[819,1099,1877,1901]" class="Cephalopoda" family="Delepinoceratidae" genus="Choctawites" kingdom="Animalia" pageId="12" pageNumber="93" phylum="Mollusca" rank="species" species="choctawensis">
|
||
<emphasis id="B94FEABAFC50FFA0FCE6F8A0FB82F8F9" box="[819,1099,1877,1901]" italics="true" pageId="12" pageNumber="93">Choctawites choctawensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D2BFC50FFA0FB8FF8A0FA48F8F9" box="[1114,1409,1877,1901]" class="Cephalopoda" family="Delepinoceratidae" genus="Revilloceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="12" pageNumber="93" phylum="Mollusca" rank="species" species="granofalcatum">
|
||
<emphasis id="B94FEABAFC50FFA0FB8FF8A0FA48F8F9" box="[1114,1409,1877,1901]" italics="true" pageId="12" pageNumber="93">Revilloceras granofalcatum</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FFA9FF45FEB0FF5C" ID-CoL="378PV" authorityName="Ruzhentsev & Bogoslovskaya" authorityYear="1970" box="[124,377,176,200]" class="Cephalopoda" family="Delepinoceratidae" genus="Dombarites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="species" species="falcatoides">
|
||
<emphasis id="B94FEABAFC51FFA1FFA9FF45FEB0FF5C" box="[124,377,176,200]" italics="true" pageId="13" pageNumber="94">Dombarites falcatoides</emphasis>
|
||
</taxonomicName>
|
||
. We included a new character, the presence of shell constrictions, in the matrix.
|
||
</paragraph>
|
||
<paragraph id="8B8436A8FC51FFA1FF42FF05FEFBFD32" blockId="13.[124,767,176,901]" pageId="13" pageNumber="94">
|
||
The heuristic cladistics analysis performed with
|
||
<collectionCode id="ED2AAE6DFC51FFA1FD15FF05FD36FE93" box="[704,767,240,263]" pageId="13" pageNumber="94">PAST</collectionCode>
|
||
(
|
||
<bibRefCitation id="EFAA4B59FC51FFA1FF56FEE5FEA6FEB3" author="Hammer, O. & Harper, D. & Ryan, P." box="[131,367,272,295]" pageId="13" pageNumber="94" refId="ref16842" refString="Hammer, O., Harper, D., and Ryan, P.: PAST-PAlaeontological STatistics, ver. 1.89, University of Oslo, Oslo, 2009." type="book" year="2009">Hammer et al., 2009</bibRefCitation>
|
||
) led to 35 most parsimonious trees of 35 steps length, of which a majority consensus tree was formed (
|
||
<figureCitation id="13002A2DFC51FFA1FF02FEA5FEEEFEF3" box="[215,295,336,359]" captionStart="Figure 10" captionStartId="12.[124,191,1151,1173]" captionTargetBox="[265,1322,177,1120]" captionTargetId="graphics-34@12.[265,1322,177,1120]" captionTargetPageId="12" captionText="Figure 10. Cladogram of selected genera, represented by well-known species, of the superfamily Goniatitaceae." figureDoi="http://doi.org/10.5281/zenodo.11585645" httpUri="https://zenodo.org/record/11585645/files/figure.png" pageId="13" pageNumber="94">Fig. 10</figureCitation>
|
||
). This tree shows some unresolved branches but demonstrates that a crown group is well established with the genera
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FF20FE7AFEB1FE33" authorityName="Titus & Korn & Harrell & Lambert" authorityYear="2015" box="[245,376,399,423]" class="Cephalopoda" family="Delepinoceratidae" genus="Choctawites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FF20FE7AFEB1FE33" box="[245,376,399,423]" italics="true" pageId="13" pageNumber="94">Choctawites</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FE5DFE7AFDD7FE33" authorityName="Pereira de Sousa" authorityYear="1923" box="[392,542,399,423]" class="Cephalopoda" family="Goniatitidae" genus="Lusitanoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FE5DFE7AFDD7FE33" box="[392,542,399,423]" italics="true" pageId="13" pageNumber="94">Lusitanoceras</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FDFBFE7AFD7BFE33" authorityName="Wagner-Gentis" authorityYear="1980" box="[558,690,399,423]" class="Cephalopoda" family="Delepinoceratidae" genus="Revilloceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FDFBFE7AFD7BFE33" box="[558,690,399,423]" italics="true" pageId="13" pageNumber="94">Revilloceras</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FD17FE7AFF0FFE53" authorityName="Librovitch" authorityYear="1957" class="Cephalopoda" family="Delepinoceratidae" genus="Dombarites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FD17FE7AFF0FFE53" italics="true" pageId="13" pageNumber="94">Dombarites</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FF07FE5AFEB3FE53" authorityName="Rauser-Tschernoussowa" authorityYear="1928" box="[210,378,431,455]" class="Cephalopoda" family="Delepinoceratidae" genus="Proshumardites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FF07FE5AFEB3FE53" box="[210,378,431,455]" italics="true" pageId="13" pageNumber="94">Proshumardites</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FE50FE5AFDEFFE53" authorityName="Ruzhentzev" authorityYear="1956" box="[389,550,431,455]" class="Cephalopoda" family="Delepinoceratidae" genus="Platygoniatites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FE50FE5AFDEFFE53" box="[389,550,431,455]" italics="true" pageId="13" pageNumber="94">Platygoniatites</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FD88FE5AFD31FE53" authorityName="Miller & Furnish" authorityYear="1954" box="[605,760,431,455]" class="Cephalopoda" family="Delepinoceratidae" genus="Delepinoceras" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FD88FE5AFD31FE53" box="[605,760,431,455]" italics="true" pageId="13" pageNumber="94">Delepinoceras</emphasis>
|
||
</taxonomicName>
|
||
, which are all characterized by the apomorphies of a Y-shaped external lobe and a tendency towards a tectiform ventrolateral saddle. This clearly defined clade can best be described as the family
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FEB2FDDAFDE5FDD2" authorityName="Ruzhencev" authorityYear="1957" box="[359,556,559,582]" class="Cephalopoda" family="Delepinoceratidae" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="family">Delepinoceratidae</taxonomicName>
|
||
. One has, however, to take into account that a similar tendency of sutural development also occurs in the more conservative lineage leading to
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FF42FD7BFEE3FD32" authorityName="Ruzhentsev & Bogoslovskaya" authorityYear="1970" box="[151,298,654,678]" class="Cephalopoda" family="Goniatitidae" genus="Neogoniatites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FF42FD7BFEE3FD32" box="[151,298,654,678]" italics="true" pageId="13" pageNumber="94">Neogoniatites</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8436A8FC51FFA1FF42FD5BFDAEFC11" blockId="13.[124,767,176,901]" pageId="13" pageNumber="94">
|
||
The cladogram also shows that
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FE25FD5BFDBAFD52" authorityName="Titus & Korn & Harrell & Lambert" authorityYear="2015" box="[496,627,686,710]" class="Cephalopoda" family="Delepinoceratidae" genus="Choctawites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FE25FD5BFDBAFD52" box="[496,627,686,710]" italics="true" pageId="13" pageNumber="94">Choctawites</emphasis>
|
||
</taxonomicName>
|
||
is more plesiomorphic than the other of the genera, which are all defined by the presence of outer shell constrictions in the juvenile stage. The crown group is subdivided into two independent clades, one represented by the advanced
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FDF8FCD8FF5BFCF1" class="Cephalopoda" family="Delepinoceratidae" genus="Dombarites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="species" species="undefined-D">
|
||
<emphasis id="B94FEABAFC51FFA1FDF8FCD8FD63FCD1" box="[557,682,813,837]" italics="true" pageId="13" pageNumber="94">Dombarites</emphasis>
|
||
species
|
||
<emphasis id="B94FEABAFC51FFA1FFA9FCB8FF5BFCF1" box="[124,146,845,869]" italics="true" pageId="13" pageNumber="94">D</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94FEABAFC51FFA1FF47FCB8FF14FCF1" box="[146,221,845,869]" italics="true" pageId="13" pageNumber="94">. tectus</emphasis>
|
||
and
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FEC3FCB8FE77FCF1" authorityName="Rauser-Tschernoussowa" authorityYear="1928" box="[278,446,845,869]" class="Cephalopoda" family="Delepinoceratidae" genus="Proshumardites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FEC3FCB8FE77FCF1" box="[278,446,845,869]" italics="true" pageId="13" pageNumber="94">Proshumardites</emphasis>
|
||
</taxonomicName>
|
||
and the other represented by the genera
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FF24FC98FE5BFC11" authorityName="Ruzhentzev" authorityYear="1956" box="[241,402,877,901]" class="Cephalopoda" family="Delepinoceratidae" genus="Platygoniatites" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FF24FC98FE5BFC11" box="[241,402,877,901]" italics="true" pageId="13" pageNumber="94">Platygoniatites</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B4D2BFC51FFA1FE10FC98FDA9FC11" authorityName="Miller & Furnish" authorityYear="1954" box="[453,608,877,901]" class="Cephalopoda" family="Delepinoceratidae" genus="Delepinoceras" kingdom="Animalia" pageId="13" pageNumber="94" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B94FEABAFC51FFA1FE10FC98FDA9FC11" box="[453,608,877,901]" italics="true" pageId="13" pageNumber="94">Delepinoceras</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |