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<document id="15C404207060E1A4746991410F6F15A9" ID-DOI="10.11646/zootaxa.4000.1.1" ID-GBIF-Dataset="507a3784-35f5-46f5-9492-e0884a502f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="289219" ID-ZooBank="00769C76-A2D4-4792-A5DD-5F2B7EA352CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461184674989" checkinUser="plazi" docAuthor="Allegro, Gianni &amp; Giachino, Pier Mauro" docDate="2015" docId="039087C7FFA1FF88E28C4BE2344F6DD8" docLanguage="en" docName="zt04000p048.pdf" docOrigin="Zootaxa 4000 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blennidus (Agraphoderus) pinguis Allegro &amp; Giachino 2011" docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FFA9FFBFFFAFFF86E21B4B7535296F7A" masterDocTitle="Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="15" updateTime="1698599231025" updateUser="plazi">
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<mods:title id="F8690BACBE05242557F1164004AF9159">Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)</mods:title>
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<mods:namePart id="A18359C293A534AF47E2610F270D2456">Allegro, Gianni</mods:namePart>
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<mods:namePart id="E88B26125D760EF7084FCB4FEBE05931">Giachino, Pier Mauro</mods:namePart>
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<mods:title id="1D66AF1C373D897D2C85BFDB215D099A">Zootaxa</mods:title>
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<mods:date id="5EB8657B033AD9AAF1B12C9A138E7867">2015</mods:date>
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<mods:number id="957D10393ED02A716307358A5530C0DA">4000</mods:number>
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<treatment id="039087C7FFA1FF88E28C4BE2344F6DD8" ID-DOI="http://doi.org/10.5281/zenodo.6096065" ID-GBIF-Taxon="127692768" ID-Zenodo-Dep="6096065" LSID="urn:lsid:plazi:treatment:039087C7FFA1FF88E28C4BE2344F6DD8" httpUri="http://treatment.plazi.org/id/039087C7FFA1FF88E28C4BE2344F6DD8" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="C323655AFFA1FF88E28C4BE2344C6FA9" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8B8636D1FFA1FF88E28C4BE236566FC8" blockId="14.[151,895,151,211]" box="[151,895,151,178]" pageId="14" pageNumber="15">
<heading id="D0CE81BDFFA1FF88E28C4BE236566FC8" bold="true" box="[151,895,151,178]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="4C394D52FFA1FF88E28C4BE236566FC8" ID-CoL="5WLZZ" authority="Allegro &amp; Giachino, 2011" authorityName="Allegro &amp; Giachino" authorityYear="2011" box="[151,895,151,178]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pinguis" subGenus="Agraphoderus">
<emphasis id="B94DEAC3FFA1FF88E28C4BE236566FC8" bold="true" box="[151,895,151,178]" pageId="14" pageNumber="15">
<emphasis id="B94DEAC3FFA1FF88E28C4BE2343A6FCB" bold="true" box="[151,275,151,177]" italics="true" pageId="14" pageNumber="15">Blennidus</emphasis>
(
<emphasis id="B94DEAC3FFA1FF88E33F4BE234F96FCB" bold="true" box="[292,464,151,177]" italics="true" pageId="14" pageNumber="15">Agraphoderus</emphasis>
)
<emphasis id="B94DEAC3FFA1FF88E3FA4BE237126FCB" bold="true" box="[481,571,151,177]" italics="true" pageId="14" pageNumber="15">pinguis</emphasis>
Allegro &amp; Giachino, 2011
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8636D1FFA1FF88E28C4BCF344C6FA9" blockId="14.[151,895,151,211]" box="[151,357,186,211]" pageId="14" pageNumber="15">
(
<figureCitation id="13022A54FFA1FF88E2844BCF35D56FA9" box="[159,252,186,211]" captionStart="FIGURES 36 39" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[248,1327,212,1904]" captionTargetId="figure@33.[231,1356,193,1932]" captionTargetPageId="33" captionText="FIGURES 36 39. Habitus of Blennidus: pascoensis male HT (36); phaenogonus male LT (37); pinguis male HT (38); procerus male HT (39)." httpUri="https://zenodo.org/record/289229/files/figure.png" pageId="14" pageNumber="15">Figs. 38</figureCitation>
,
<figureCitation id="13022A54FFA1FF88E3134BCF340F6FA9" box="[264,294,186,211]" captionStart="FIGURES 74 81" captionStartId="40.[151,264,1921,1944]" captionTargetBox="[164,1417,208,1893]" captionTargetId="figure@40.[151,1436,193,1900]" captionTargetPageId="40" captionText="FIGURES 74 81. Pronotum of Blennidus: pascoensis male HT (74); phaenogonus male LT (75); pinguis male HT (76); procerus male HT (77); pseudolaevis male HT (78); rectangulus male HT (79); refleximargo male HT (80); straneoi male HT (81)." httpUri="https://zenodo.org/record/289235/files/figure.png" pageId="14" pageNumber="15">76</figureCitation>
,
<figureCitation id="13022A54FFA1FF88E3294BCE34756FA9" box="[306,348,187,211]" captionStart="FIGURES 112 117" captionStartId="46.[151,262,1953,1976]" captionTargetBox="[230,1345,214,1916]" captionTargetId="figure@46.[213,1373,193,1932]" captionTargetPageId="46" captionText="FIGURES 112 117. Aedeagus (left lateral and dorsal aspects) of Blennidus: pinguis HT (112); procerus HT (113); pseudolaevis HT (114); rectangulus HT (115); refleximargo HT (116); straneoi HT (117)." httpUri="https://zenodo.org/record/289241/files/figure.png" pageId="14" pageNumber="15">112</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA1FF88E28C4A7737B56E62" box="[151,668,258,280]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8B8636D1FFA1FF88E28C4A7737B56E62" blockId="14.[151,668,258,280]" box="[151,668,258,280]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB2911FFFFA1FF88E28C4A7737B56E62" box="[151,668,258,280]" pageId="14" pageNumber="15">
<taxonomicName id="4C394D52FFA1FF88E28C4A7737B56E62" ID-CoL="5WLZZ" authority="Allegro &amp; Giachino, 2011b: 170" authorityName="Allegro &amp; Giachino" authorityPageNumber="170" authorityYear="2011" box="[151,668,258,280]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="B94DEAC3FFA1FF88E28C4A7734646E62" box="[151,333,258,280]" italics="true" pageId="14" pageNumber="15">Blennidus pinguis</emphasis>
<treatmentCitation id="0A9810C0FFA1FF88E3484A7737B56E62" author="Allegro" box="[339,668,258,280]" page="170" pageId="14" pageNumber="15" year="2011">
<bibRefCitation id="EFA84B20FFA1FF88E3484A7737456E62" author="Allegro" box="[339,620,258,280]" pageId="14" pageNumber="15" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino, 2011b</bibRefCitation>
: 170
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</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA1FF88E28C4A3034F46D68" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8B8636D1FFA1FF88E28C4A3037D56E24" blockId="14.[151,1436,325,674]" box="[151,764,325,350]" pageId="14" pageNumber="15">
<emphasis id="B94DEAC3FFA1FF88E28C4A30341C6E24" bold="true" box="[151,309,325,350]" pageId="14" pageNumber="15">
<typeStatus id="54828873FFA1FF88E28C4A3035FB6E24" box="[151,210,325,350]" pageId="14" pageNumber="15">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32E7641FFA1FF88E3274A33345B6E27" box="[316,370,326,349]" name="Peru" pageId="14" pageNumber="15">Peru</collectingCountry>
, Cajamarca, Michiquillay, m 4050.
</paragraph>
<paragraph id="8B8636D1FFA1FF88E2DC4A1C37036EDC" blockId="14.[151,1436,325,674]" pageId="14" pageNumber="15">
<emphasis id="B94DEAC3FFA1FF88E2DC4A1C349F6EF8" bold="true" box="[199,438,361,386]" pageId="14" pageNumber="15">Examined material.</emphasis>
HT ♂ (CGi); 10 PTT ♂ and 8 PTT ♀ (CGi, CAl, CCa, CMo, CMa,
<collectionCode id="ED28AE14FFA1FF88E6C24A1C30186EFB" box="[1241,1329,361,385]" pageId="14" pageNumber="15">MSNM</collectionCode>
,
<collectionCode id="ED28AE14FFA1FF88E7244A1C30BC6EFB" box="[1343,1429,361,385]" pageId="14" pageNumber="15">BMNH</collectionCode>
) (complete TS);
<specimenCount id="9D3FFD58FFA1FF88E35D4AFB34586EDF" box="[326,369,398,422]" pageId="14" pageNumber="15" type="generic">4 ♂</specimenCount>
and 3 ♀ (CMa).
</paragraph>
<paragraph id="8B8636D1FFA1FF88E2DC4AC434F46D68" blockId="14.[151,1436,325,674]" pageId="14" pageNumber="15">
<emphasis id="B94DEAC3FFA1FF88E2DC4AC4342F6EB0" bold="true" box="[199,262,433,458]" pageId="14" pageNumber="15">Note.</emphasis>
Species belonging to the
<taxonomicName id="4C394D52FFA1FF88E0524AC737F16EB3" box="[585,728,433,457]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA1FF88E0524AC737F16EB3" box="[585,728,433,457]" italics="true" pageId="14" pageNumber="15">B. orbicollis</emphasis>
</taxonomicName>
species group (along with
<taxonomicName id="4C394D52FFA1FF88E6324AC731896EB3" box="[1065,1184,434,457]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B94DEAC3FFA1FF88E6324AC731896EB3" box="[1065,1184,434,457]" italics="true" pageId="14" pageNumber="15">B. crassus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C394D52FFA1FF88E6AF4AC7306A6EB3" box="[1204,1347,433,457]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA1FF88E6AF4AC7306A6EB3" box="[1204,1347,433,457]" italics="true" pageId="14" pageNumber="15">B. orbicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C394D52FFA1FF88E79F4AC734016E97" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudolaevis">
<emphasis id="B94DEAC3FFA1FF88E79F4AC734016E97" italics="true" pageId="14" pageNumber="15">B. pseudolaevis</emphasis>
</taxonomicName>
) distinguished by the habitus as well as the distinctive and very similar morphology of aedeagus (
<bibRefCitation id="EFA84B20FFA1FF88E2854A8C34F86D68" author="Allegro" box="[158,465,505,530]" pageId="14" pageNumber="15" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino 2011b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA1FF88E2DC4968344F6DD8" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8B8636D1FFA1FF88E2DC496837B96D20" blockId="14.[151,1436,325,674]" pageId="14" pageNumber="15">
<emphasis id="B94DEAC3FFA1FF88E2DC496834496D4C" bold="true" box="[199,352,541,566]" pageId="14" pageNumber="15">Distribution.</emphasis>
All the recorded specimens of
<taxonomicName id="4C394D52FFA1FF88E0F8496B36726D4F" box="[739,859,542,565]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="B94DEAC3FFA1FF88E0F8496B36726D4F" box="[739,859,542,565]" italics="true" pageId="14" pageNumber="15">B. pinguis</emphasis>
</taxonomicName>
(M. Etonti legit) are from the Dept. Cajamarca (northern
<collectingCountry id="F32E7641FFA1FF88E31D493434126D23" box="[262,315,577,601]" name="Peru" pageId="14" pageNumber="15">Peru</collectingCountry>
) (
<bibRefCitation id="EFA84B20FFA1FF88E34A493437AD6D20" author="Allegro" box="[337,644,577,602]" pageId="14" pageNumber="15" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino 2011b</bibRefCitation>
).
</paragraph>
<paragraph id="8B8636D1FFA1FF88E2DC4910344F6DD8" blockId="14.[151,1436,325,674]" pageId="14" pageNumber="15">
<emphasis id="B94DEAC3FFA1FF88E2DC491034036D04" bold="true" box="[199,298,613,638]" pageId="14" pageNumber="15">Habitat.</emphasis>
The altitude data reported on the labels (
<quantity id="4CC19B34FFA1FF88E11F4910364D6D07" box="[772,868,613,638]" metricMagnitude="3" metricUnit="m" metricValue="4.05" pageId="14" pageNumber="15" unit="m" value="4050.0">4,050 m</quantity>
a.s.l.) suggest a preference of this species for the Andean grassland.
</paragraph>
</subSubSection>
</treatment>
</document>