treatments-xml/data/03/90/87/039087B9FF79FFF4FF4974EAFAC5FEB9.xml
2024-06-21 12:22:17 +02:00

375 lines
44 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="87000E89B9234A4797978FBC3496B49F" ID-CLB-Dataset="26556" ID-DOI="10.11646/zootaxa.4614.1.4" ID-GBIF-Dataset="2fdc41ca-97b1-4e90-bdd7-921797dff2c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3241154" ID-ZooBank="B0378DC5-5B3F-4E64-9ADA-B50408BCF374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559897220234" checkinUser="plazi" docAuthor="Zilberman, Bruno" docDate="2019" docId="039087B9FF79FFF4FF4974EAFAC5FEB9" docLanguage="en" docName="zootaxa.4614.1.4.pdf" docOrigin="Zootaxa 4614 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Termitozophilus belleae Zilberman 2019, sp. nov." docType="treatment" docUuid="ACC9637B-D917-4571-A3FA-FFD92A4F899B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="110" masterDocId="FFA9FFC1FF73FFFBFFDE7102FFCFFFF9" masterDocTitle="Synopsis of the genus Termitozophilus Silvestri (Coleoptera, Staphylinidae, Aleocharinae, Corotocini)" masterLastPageNumber="116" masterPageNumber="95" pageNumber="105" updateTime="1698733478518" updateUser="ExternalLinkService">
<mods:mods id="75F4B9EA134BB3E3A20CD458690BE8A6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="01FDC265F75C7376B527D2D17FFD71FA">
<mods:title id="B8100C7AC8A01A4D4FA48DBBA29A8011">Synopsis of the genus Termitozophilus Silvestri (Coleoptera, Staphylinidae, Aleocharinae, Corotocini)</mods:title>
</mods:titleInfo>
<mods:name id="687617777A3CD56239E12B2F2228DC90" type="personal">
<mods:role id="BD73EDBACBD32496799413463C665CE1">
<mods:roleTerm id="26199F8EC9756161B19FCCBF94ED6E4C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA6F26CEC0434F9C2C83EE37DDD0FB8E">Zilberman, Bruno</mods:namePart>
</mods:name>
<mods:typeOfResource id="234F5E4F11363D5298751F0FF69B4E15">text</mods:typeOfResource>
<mods:relatedItem id="B3AD3A9140C822D4937189A2D20AFD1F" type="host">
<mods:titleInfo id="2C30FB40FF806872E4C20E7664103563">
<mods:title id="153E9F81AC8C739DA8D3FE7CB306A402">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="36F5810E00F1E153C34E2094E82A17DF">
<mods:date id="79A79C9E27026015A2DD4CDBE4B07CAD">2019</mods:date>
<mods:detail id="64DAB8327F1D6FD1C109F68572152B42" type="pubDate">
<mods:number id="85DD4A1FB7E7EA735009934C9D83AB01">2019-06-07</mods:number>
</mods:detail>
<mods:detail id="4274BE06AD7F1C12378DEDA72F62E878" type="volume">
<mods:number id="FF357714B8615DE5D8CA15587971956F">4614</mods:number>
</mods:detail>
<mods:detail id="0C81B92EB1FC9B7F114ED010E6FEDF0D" type="issue">
<mods:number id="804F74E8FA28E5F09B772224D41DE6E1">1</mods:number>
</mods:detail>
<mods:extent id="FE22CC7135FE3CD03A7CC5A231F173D4" unit="page">
<mods:start id="71E709C855CB0158176C2C3A6768206C">95</mods:start>
<mods:end id="ED91A8F1AD1A1C09539CF6C8AEFBE3DC">116</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2CAA0AEF8D06B9851C57A378EB5633C3">journal article</mods:classification>
<mods:identifier id="BC4760C9DE57A7917AD691D28DFD24A3" type="CLB-Dataset">26556</mods:identifier>
<mods:identifier id="68F35FD4AE27C777B2C5621DB445DF68" type="DOI">10.11646/zootaxa.4614.1.4</mods:identifier>
<mods:identifier id="5F9A5B0D7CF9BF3740F223F6103F7A8C" type="GBIF-Dataset">2fdc41ca-97b1-4e90-bdd7-921797dff2c9</mods:identifier>
<mods:identifier id="61806F3F65920F3272D6BDFE8C75C5AC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="86C88A32DDF3F64E1F1276FADA80DEA1" type="Zenodo-Dep">3241154</mods:identifier>
<mods:identifier id="4E1E373BEA501AB83E2E8EDD45EB776D" type="ZooBank">B0378DC5-5B3F-4E64-9ADA-B50408BCF374</mods:identifier>
</mods:mods>
<treatment id="039087B9FF79FFF4FF4974EAFAC5FEB9" ID-GBIF-Taxon="157436063" LSID="urn:lsid:zoobank.org:act:ACC9637B-D917-4571-A3FA-FFD92A4F899B" httpUri="http://treatment.plazi.org/id/039087B9FF79FFF4FF4974EAFAC5FEB9" lastPageId="15" lastPageNumber="110" pageId="10" pageNumber="105">
<subSubSection id="C3236524FF79FFF1FF4974EAFDD6F9FB" box="[151,537,1512,1538]" pageId="10" pageNumber="105" type="nomenclature">
<paragraph id="8B8636AFFF79FFF1FF4974EAFDD6F9FB" blockId="10.[151,537,1512,1538]" box="[151,537,1512,1538]" pageId="10" pageNumber="105">
<heading id="D0CE81C3FF79FFF1FF4974EAFDD6F9FB" bold="true" box="[151,537,1512,1538]" fontSize="11" level="1" pageId="10" pageNumber="105" reason="1">
<emphasis id="B94DEABDFF79FFF1FF4974EAFDD6F9FB" bold="true" box="[151,537,1512,1538]" pageId="10" pageNumber="105">
<taxonomicName id="4C394D2CFF79FFF1FF4974EAFE7AF9FB" ID-CoL="8YBMG" authority="Zilberman, 2019" authorityName="Zilberman" authorityYear="2019" box="[151,437,1512,1538]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="105" phylum="Arthropoda" rank="species" species="belleae" status="sp. nov.">
<emphasis id="B94DEABDFF79FFF1FF4974EAFE7AF9FB" bold="true" box="[151,437,1512,1538]" italics="true" pageId="10" pageNumber="105">Termitozophilus belleae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E57C6FF79FFF1FE6274EAFDD6F9FB" box="[444,537,1512,1538]" pageId="10" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3236524FF79FFF0FF49772DFCEEFE54" lastPageId="11" lastPageNumber="106" pageId="10" pageNumber="105" type="materials_examined">
<paragraph id="8B8636AFFF79FFF1FF49772DFF21F94A" blockId="10.[151,1437,1583,2004]" pageId="10" pageNumber="105">
<materialsCitation id="3B513CF2FF79FFF1FF49772DFF22F969" ID-GBIF-Occurrence="2248478992" collectingDate="1976-02-15" collectionCode="MZSP" collectorName="Araujo" country="Brazil" location="Cuiaba" pageId="10" pageNumber="105" specimenCode="MZSP 21228, MZSP 6759" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<typeStatus id="5482880DFF79FFF1FF49772DFEE0F9BE" box="[151,303,1583,1607]" pageId="10" pageNumber="105" type="holotype">HOLOTYPE</typeStatus>
.
<collectingCountry id="F32E763FFF79FFF1FEE7772DFE4EF9B1" box="[313,385,1583,1608]" name="Brazil" pageId="10" pageNumber="105">Brazil</collectingCountry>
.
<collectingRegion id="49FDF84DFF79FFF1FE527732FDEFF9BE" box="[396,544,1584,1607]" country="Brazil" name="Mato Grosso" pageId="10" pageNumber="105">
<emphasis id="B94DEABDFF79FFF1FE527732FDEFF9BE" box="[396,544,1584,1607]" italics="true" pageId="10" pageNumber="105">Mato Grosso</emphasis>
</collectingRegion>
:
<location id="8EE66074FF79FFF1FDF1772DFDB1F9B1" LSID="urn:lsid:plazi:treatment:039087B9FF79FFF4FF4974EAFAC5FEB9:8EE66074FF79FFF1FDF1772DFDB1F9B1" box="[559,638,1583,1608]" country="Brazil" name="Cuiaba" pageId="10" pageNumber="105" stateProvince="Mato Grosso">Cuiabá</location>
(Estrada Cuiabá P. [Porto] Velho),
<collectingDate id="EFC3E987FF79FFF1FBFA772DFB52F9B1" box="[1060,1181,1583,1608]" pageId="10" pageNumber="105" value="1976-02-15">
<date id="FF87106FFF79FFF1FBFA772DFB52F9B1" box="[1060,1181,1583,1608]" pageId="10" pageNumber="105" value="1976-02-15">15/II/1976</date>
</collectingDate>
, R. [Renato] L. [Lion]
<collectorName id="26CC5379FF79FFF1FF497751FF2AF995" box="[151,229,1619,1644]" pageId="10" pageNumber="105">Araujo</collectorName>
col. (
<specimenCode id="DB9F9ED4FF79FFF1FEF37751FE06F992" box="[301,457,1619,1643]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="10" pageNumber="105" type="Museum">MZSP 21228</specimenCode>
/ [alternative label 21228-H]), host,
<taxonomicName id="4C394D2CFF79FFF1FCA97756FBA0F995" authorityName="Emerson" authorityYear="1949" box="[887,1135,1619,1644]" class="Insecta" family="Termitidae" genus="Cornitermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="105" phylum="Arthropoda" rank="species" species="silvestrii">
<emphasis id="B94DEABDFF79FFF1FCA97756FBA0F995" box="[887,1135,1619,1644]" italics="true" pageId="10" pageNumber="105">Cornitermes silvestrii</emphasis>
</taxonomicName>
(
<specimenCode id="DB9F9ED4FF79FFF1FB5D7751FAD9F995" box="[1155,1302,1619,1644]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="10" pageNumber="105" type="Museum">MZSP 6759</specimenCode>
), female in alcohol
</materialsCitation>
;
<materialsCitation id="3B513CF2FF79FFF1FF247775FF25F94A" ID-GBIF-Occurrence="2248478995" collectingDate="1976-02-15" collectionCode="MZSP" collectorName="Araujo" country="Brazil" location="Cuiaba" pageId="10" pageNumber="105" specimenCode="MZSP 21228" specimenCount="15" specimenCount-female="2" specimenCount-male="1" stateProvince="Mato Grosso" typeStatus="paratype">
<typeStatus id="5482880DFF79FFF1FF247775FE57F976" box="[250,408,1655,1679]" pageId="10" pageNumber="105" type="paratype">PARATYPES</typeStatus>
. same data of
<typeStatus id="5482880DFF79FFF1FD9F7775FD69F969" box="[577,678,1655,1680]" pageId="10" pageNumber="105" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D3FFD26FF79FFF1FD6D7775FC81F969" box="[691,846,1655,1680]" pageId="10" pageNumber="105" type="generic">12 specimens</specimenCount>
(
<specimenCount id="9D3FFD26FF79FFF1FC817775FC00F969" box="[863,975,1655,1680]" pageId="10" pageNumber="105" type="female">2 females</specimenCount>
and
<specimenCount id="9D3FFD26FF79FFF1FBD47775FB98F969" box="[1034,1111,1655,1680]" pageId="10" pageNumber="105" type="male">1 male</specimenCount>
dissected in slides,) (
<specimenCode id="DB9F9ED4FF79FFF1FA8F7775FF2DF94A" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="10" pageNumber="105" type="Museum">MZSP 21228</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8636AFFF79FFF1FF1977BDFEABF8BD" blockId="10.[151,1437,1583,2004]" pageId="10" pageNumber="105">Body moderately physogastric and strongly sclerotized; sclerotized areas mostly dark brown to black; sides of abdomen, tarsi and antennae brown; narrow posterior band of pronotum black. Sclerotized areas densely covered with long spine-like bristles (figs. 2936). Female - length ~2.7 mm (abdomen shrunk), male - length ~2.3 mm (abdomen shrunk)</paragraph>
<paragraph id="8B8636AFFF79FFF0FF19764DFCEEFE54" blockId="10.[151,1437,1583,2004]" lastBlockId="11.[151,1437,151,429]" lastPageId="11" lastPageNumber="106" pageId="10" pageNumber="105">
<emphasis id="B94DEABDFF79FFF1FF19764DFEC1F891" bold="true" box="[199,270,1871,1896]" pageId="10" pageNumber="105">Head.</emphasis>
Elongate, slightly longer than large, widest at ¾ anterior region of head and slightly narrowed on posterior 1/4; length of eyes occupying 1/3 of total head length; one setae behind each eye and a pair in medialposterior region of vertex (figs. 37, 38, 57).
<emphasis id="B94DEABDFF79FFF1FD687695FCFDF849" bold="true" box="[694,818,1943,1968]" pageId="10" pageNumber="105">Antennae:</emphasis>
scape slighter longer than any other antenommere; antennomeres IIX elongate, gradually decreasing in length; antennomere XI about twice longer than wide, slightly longer than IX (
<figureCitation id="13022A2AFF78FFF0FE6A7195FDC5FF49" box="[436,522,151,176]" captionStart="FIGURES 3745" captionStartId="14.[151,264,1455,1478]" captionTargetBox="[151,1433,193,1435]" captionTargetId="figure@14.[151,1436,193,1435]" captionTargetPageId="14" captionText="FIGURES 3745. Termitozophilus belleae sp. nov. 37, 38, head (ventral dorsal); 39, antenna; 40, pos-mentum; 41, prementum; 42, maxilla; 43, labrum; 44, 35, left and right mandibles (female, male). Scale bars: 0.05 mm, except figs. 37-38 = 0.1 mm, 39 = 0.5 mm." httpUri="https://zenodo.org/record/3241178/files/figure.png" pageId="11" pageNumber="106">Fig. 39</figureCitation>
,
<figureCitation id="13022A2AFF78FFF0FDC77195FDF6FF49" box="[537,569,151,176]" captionStart="FIGURES 5860" captionStartId="17.[151,264,1963,1986]" captionTargetId="figure@17.[175,1413,193,2008]" captionTargetPageId="17" captionText="FIGURES 5860. Antennae: 58, Termitozophilus laetus Silvestri, 1901; 59, Termitozophilus mirandus (Mann, 1923); 60, Termitozophilus belleae sp. nov. Scale bars = 0.5 mm, except fig. 60 = 0.1 mm." httpUri="https://zenodo.org/record/3404913/files/figure.png" pageId="11" pageNumber="106">60</figureCitation>
).
<emphasis id="B94DEABDFF78FFF0FD8E7195FC9AFF49" bold="true" box="[592,853,151,176]" pageId="11" pageNumber="106">Mouthparts. Labrum</emphasis>
more than twice larger than long; five main long bristles on each side, distributed in two diagonal rows, each with two bristles (D2 - D1/ M2 - M1) and one bristle (P1) alone placed at middle of labrum length; epipharynx covered with sensilla and numerous short setae (figs. 43, 62).
<emphasis id="B94DEABDFF78FFF0FF147006FE87FEE4" bold="true" box="[202,328,260,285]" pageId="11" pageNumber="106">Mandibles</emphasis>
with broad tooth in mesal median region; apical region broad in females and narrow in males (figs. 44, 45).
<emphasis id="B94DEABDFF78FFF0FF2D7025FEAAFEB9" bold="true" box="[243,357,295,320]" pageId="11" pageNumber="106">Maxillae:</emphasis>
galea elongate, slightly larger than lacinia, with small bristles at margins; lacinia narrow, as long as galea; palpomeres moderated sclerotized:
<collectionCode id="ED28AE6AFF78FFF0FD53704FFD6FFE9D" box="[653,672,333,356]" pageId="11" pageNumber="106">II</collectionCode>
and
<collectionCode id="ED28AE6AFF78FFF0FD08704FFD3BFE9D" box="[726,756,333,356]" pageId="11" pageNumber="106">IV</collectionCode>
elongate,
<collectionCode id="ED28AE6AFF78FFF0FCB9704FFC4CFE9D" box="[871,899,333,356]" pageId="11" pageNumber="106">III</collectionCode>
suboval (fig. 42).
<emphasis id="B94DEABDFF78FFF0FB8C704EFB7EFE9C" bold="true" box="[1106,1201,332,357]" pageId="11" pageNumber="106">Labium</emphasis>
with ligula widest at base with apex emarginated, resulting in two lobes (fig. 41); mentum and submentum fused, with two bristles on each side: pre-apical (
<emphasis id="B94DEABDFF78FFF0FE537097FE66FE55" box="[397,425,405,428]" italics="true" pageId="11" pageNumber="106">pa</emphasis>
) and proximal (
<emphasis id="B94DEABDFF78FFF0FD807097FDB7FE55" box="[606,632,405,428]" italics="true" pageId="11" pageNumber="106">px</emphasis>
) (figs. 40, 64).
</paragraph>
</subSubSection>
<caption id="DF466627FF78FFF0FF4976A5FE2CF824" httpUri="https://zenodo.org/record/3241172/files/figure.png" pageId="11" pageNumber="106" startId="11.[151,264,1959,1982]" targetBox="[350,1164,522,1915]" targetPageId="11">
<paragraph id="8B8636AFFF78FFF0FF4976A5FE2CF824" blockId="11.[151,1437,1959,2013]" pageId="11" pageNumber="106">
<emphasis id="B94DEABDFF78FFF0FF4976A5FE94F844" bold="true" box="[151,347,1959,1982]" pageId="11" pageNumber="106">FIGURES 2728.</emphasis>
<taxonomicName id="4C394D2CFF78FFF0FEB776AAFCD2F847" authority="(Mann, 1923)" authorityName="Mann" authorityYear="1923" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[361,797,1960,1982]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="mirandus">
<emphasis id="B94DEABDFF78FFF0FEB776AAFDB6F847" box="[361,633,1960,1982]" italics="true" pageId="11" pageNumber="106">Termitozophilus mirandus</emphasis>
(
<bibRefCitation id="EFA84B5EFF78FFF0FD5076AAFCDAF847" author="Mann, W. M." box="[654,789,1960,1982]" pageId="11" pageNumber="106" pagination="323 - 366" refId="ref6173" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from northern South America. Zoologica, 3 (17), 323 - 366." type="journal article" year="1923">Mann, 1923</bibRefCitation>
)
</taxonomicName>
, holotype.
<emphasis id="B94DEABDFF78FFF0FC4276AAFC74F844" bold="true" box="[924,955,1960,1981]" pageId="11" pageNumber="106">27,</emphasis>
head, dorsal view;
<emphasis id="B94DEABDFF78FFF0FB7E76AAFB70F844" bold="true" box="[1184,1215,1960,1981]" pageId="11" pageNumber="106">28,</emphasis>
head, ventral view. Photographs of Karolyn Darrow.
</paragraph>
</caption>
<caption id="DF466627FF7FFFF7FF49740EFF24FAB8" httpUri="https://zenodo.org/record/3241174/files/figure.png" pageId="12" pageNumber="107" startId="12.[151,264,1292,1315]" targetBox="[159,1424,193,1278]" targetPageId="12">
<paragraph id="8B8636AFFF7FFFF7FF49740EFF24FAB8" blockId="12.[151,1436,1292,1345]" pageId="12" pageNumber="107">
<emphasis id="B94DEABDFF7FFFF7FF49740EFE9BFADB" bold="true" box="[151,340,1292,1315]" pageId="12" pageNumber="107">FIGURES 2931.</emphasis>
<taxonomicName id="4C394D2CFF7FFFF7FE84740EFD84FADB" authority="Zilberman, 2019" authorityName="Zilberman" authorityYear="2019" box="[346,587,1292,1314]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="belleae" status="sp. nov.">
<emphasis id="B94DEABDFF7FFFF7FE84740EFD84FADB" box="[346,587,1292,1314]" italics="true" pageId="12" pageNumber="107">Termitozophilus belleae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E57C6FF7FFFF7FD8C740FFD6DFADB" box="[594,674,1293,1314]" pageId="12" pageNumber="107" rank="species">
<emphasis id="B94DEABDFF7FFFF7FD8C740FFD53FADB" bold="true" box="[594,668,1293,1314]" pageId="12" pageNumber="107">sp. nov</emphasis>
.
</taxonomicNameLabel>
, holotype, female.
<emphasis id="B94DEABDFF7FFFF7FCB8740EFC79FADB" box="[870,950,1292,1314]" italics="true" pageId="12" pageNumber="107">Habitus</emphasis>
: 30, lateral; 31, dorsal; 32, ventral. Scale bars = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3236524FF7FFFF7FF19746CFE9DF932" pageId="12" pageNumber="107" type="description">
<paragraph id="8B8636AFFF7FFFF7FF19746CFA57FA7F" blockId="12.[151,1437,1390,2027]" box="[199,1432,1390,1415]" pageId="12" pageNumber="107">
<emphasis id="B94DEABDFF7FFFF7FF19746CFEB6FA7E" bold="true" box="[199,377,1390,1415]" pageId="12" pageNumber="107">Thorax. Elytra</emphasis>
trapezoidal covered with long bristles (fig. 48).
<emphasis id="B94DEABDFF7FFFF7FC49746CFBD0FA7E" bold="true" box="[919,1055,1390,1415]" pageId="12" pageNumber="107">Hind wings</emphasis>
vestigial, with indistinct nervure.
</paragraph>
<paragraph id="8B8636AFFF7FFFF7FF197490FE9DF932" blockId="12.[151,1437,1390,2027]" pageId="12" pageNumber="107">
<emphasis id="B94DEABDFF7FFFF7FF197490FEF3FA52" bold="true" box="[199,316,1426,1451]" pageId="12" pageNumber="107">Abdomen</emphasis>
dorsal and ventral sclerites occupying almost whole abdominal segments, leaving only a few of membrane exposed; secondary sclerotization of tergites reaching inner paratergite margin; outer paratergites close to margin of sternites due to the latter secondary sclerotization; inner paratergites with two rows each with three to six bristles (figs. 2936); tergite I represented by a moderated sclerotized sclerite attached to metanotum, with median region slightly projected backwards (fig. 47); tergite VIII about twice as wide as long, with two rows, each with six main long bristles at median and apical regions; short bristles and pores cover whole surface (fig. 49); sternite IX in male in one piece of irregular shape, moderately sclerotized, with two bristles on apical region; in female, sternite IX is presented in a pair of hemiesternite laterally attached to tergite IX; tergite IX with six long bristles (fig. 51).
</paragraph>
</subSubSection>
<subSubSection id="C3236524FF7FFFF7FF1977D4FB7EF916" box="[199,1201,1750,1775]" pageId="12" pageNumber="107" type="etymology">
<paragraph id="8B8636AFFF7FFFF7FF1977D4FB7EF916" blockId="12.[151,1437,1390,2027]" box="[199,1201,1750,1775]" pageId="12" pageNumber="107">
<emphasis id="B94DEABDFF7FFFF7FF1977D4FE89F916" bold="true" box="[199,326,1750,1775]" pageId="12" pageNumber="107">Etymology</emphasis>
. The specific epithet is patronymic, in honor of my fiancée, Isabelle Amorim.
</paragraph>
</subSubSection>
<subSubSection id="C3236524FF7FFFF5FF1977F8FDFAF82D" lastPageId="14" lastPageNumber="109" pageId="12" pageNumber="107" type="discussion">
<paragraph id="8B8636AFFF7FFFF7FF1977F8FD2EF886" blockId="12.[151,1437,1390,2027]" pageId="12" pageNumber="107">
<emphasis id="B94DEABDFF7FFFF7FF1977F8FEF4F8EA" bold="true" box="[199,315,1786,1811]" pageId="12" pageNumber="107">Remarks.</emphasis>
<taxonomicName id="4C394D2CFF7FFFF7FE9977F8FD98F8EA" authorityName="Zilberman" authorityYear="2019" box="[327,599,1786,1811]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7FFFF7FE9977F8FD98F8EA" box="[327,599,1786,1811]" italics="true" pageId="12" pageNumber="107">Termitozophilus belleae</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C394D2CFF7FFFF7FD2077F9FCAFF8EA" authorityName="Silvestri" authorityYear="1901" box="[766,864,1786,1811]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7FFFF7FD2077F9FCAFF8EA" box="[766,864,1786,1811]" italics="true" pageId="12" pageNumber="107">T. laetus</emphasis>
</taxonomicName>
in the shape of antennomeres IVX, which are filiform; head, with one bristle behind each eye, as well a pair of bristles in posterior region of head; length of eyes is about 1/3 of total head length; and ligula is bifid.
<taxonomicName id="4C394D2CFF7FFFF7FD327640FC36F8A2" authorityName="Zilberman" authorityYear="2019" box="[748,1017,1858,1883]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7FFFF7FD327640FC36F8A2" box="[748,1017,1858,1883]" italics="true" pageId="12" pageNumber="107">Termitozophilus belleae</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C394D2CFF7FFFF7FB4C7641FAD6F8A2" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[1170,1305,1858,1883]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="mirandus">
<emphasis id="B94DEABDFF7FFFF7FB4C7641FAD6F8A2" box="[1170,1305,1858,1883]" italics="true" pageId="12" pageNumber="107">T. mirandus</emphasis>
</taxonomicName>
in the dark color of sclerites and the trapezoidal shape of elytra.
</paragraph>
<paragraph id="8B8636AFFF7FFFF7FF197688FBE5F812" blockId="12.[151,1437,1390,2027]" pageId="12" pageNumber="107">
The differences, between
<taxonomicName id="4C394D2CFF7FFFF7FE347688FD39F85A" authorityName="Zilberman" authorityYear="2019" box="[490,758,1930,1955]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7FFFF7FE347688FD39F85A" box="[490,758,1930,1955]" italics="true" pageId="12" pageNumber="107">Termitozophilus belleae</emphasis>
</taxonomicName>
as well as
<taxonomicName id="4C394D2CFF7FFFF7FCAA7689FC1EF85A" authorityName="Silvestri" authorityYear="1901" box="[884,977,1930,1955]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7FFFF7FCAA7689FC1EF85A" box="[884,977,1930,1955]" italics="true" pageId="12" pageNumber="107">T. laetus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C394D2CFF7FFFF7FBCC7689FB56F85A" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[1042,1177,1930,1955]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="mirandus">
T.
<emphasis id="B94DEABDFF7FFFF7FBF17688FB56F85A" box="[1071,1177,1930,1955]" italics="true" pageId="12" pageNumber="107">mirandus</emphasis>
</taxonomicName>
, are related to the host and locality.
<taxonomicName id="4C394D2CFF7FFFF7FEF476ACFDF8F83E" authorityName="Zilberman" authorityYear="2019" box="[298,567,1966,1991]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7FFFF7FEF476ACFDF8F83E" box="[298,567,1966,1991]" italics="true" pageId="12" pageNumber="107">Termitozophilus belleae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C394D2CFF7FFFF7FDAF76ADFD00F83E" authorityName="Silvestri" authorityYear="1901" box="[625,719,1966,1991]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7FFFF7FDAF76ADFD00F83E" box="[625,719,1966,1991]" italics="true" pageId="12" pageNumber="107">T. laetus</emphasis>
</taxonomicName>
are found in association with species of
<taxonomicName id="4C394D2CFF7FFFF7FB7976ADFAFCF83F" authorityName="Wasmann" authorityYear="1897" box="[1191,1331,1967,1990]" class="Insecta" family="Termitidae" genus="Cornitermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEABDFF7FFFF7FB7976ADFAFCF83F" box="[1191,1331,1967,1990]" italics="true" pageId="12" pageNumber="107">Cornitermes</emphasis>
</taxonomicName>
living in Cerrado, and
<taxonomicName id="4C394D2CFF7FFFF7FEF076D1FE7BF812" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[302,436,2002,2027]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="mirandus">
<emphasis id="B94DEABDFF7FFFF7FEF076D1FE7BF812" box="[302,436,2002,2027]" italics="true" pageId="12" pageNumber="107">T. mirandus</emphasis>
</taxonomicName>
is found with
<taxonomicName id="4C394D2CFF7FFFF7FD8B76D1FCF5F813" authorityName="Emerson" authorityYear="1925" box="[597,826,2003,2026]" class="Insecta" family="Termitidae" genus="Cornitermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="107" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B94DEABDFF7FFFF7FD8B76D1FCF5F813" box="[597,826,2003,2026]" italics="true" pageId="12" pageNumber="107">Cornitermes pugnax</emphasis>
</taxonomicName>
, occurring in forests.
</paragraph>
<caption id="DF466627FF7EFFF6FF4976B1FC2BF810" httpUri="https://zenodo.org/record/3241176/files/figure.png" pageId="13" pageNumber="108" startId="13.[151,264,1971,1994]" targetBox="[294,1290,193,1938]" targetPageId="13">
<paragraph id="8B8636AFFF7EFFF6FF4976B1FC2BF810" blockId="13.[151,1435,1971,2025]" pageId="13" pageNumber="108">
<emphasis id="B94DEABDFF7EFFF6FF4976B1FE9CF830" bold="true" box="[151,339,1971,1994]" pageId="13" pageNumber="108">FIGURES 3236.</emphasis>
<taxonomicName id="4C394D2CFF7EFFF6FE8776B6FD85F833" authority="Zilberman, 2019" authorityName="Zilberman" authorityYear="2019" box="[345,586,1972,1994]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="108" phylum="Arthropoda" rank="species" species="belleae" status="sp. nov.">
<emphasis id="B94DEABDFF7EFFF6FE8776B6FD85F833" box="[345,586,1972,1994]" italics="true" pageId="13" pageNumber="108">Termitozophilus belleae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E57C6FF7EFFF6FD8E76B6FD6EF833" box="[592,673,1972,1994]" pageId="13" pageNumber="108" rank="species">
<emphasis id="B94DEABDFF7EFFF6FD8E76B6FD54F830" bold="true" box="[592,667,1972,1993]" pageId="13" pageNumber="108">sp. nov</emphasis>
.
</taxonomicNameLabel>
, paratype, male. Habitus: 32, lateral; 33, ventral; 34, dorsal. Abdomen: 35, dorsal; 36, ventral. Scale bars: figs 32-33 = 0.2 mm, 34 = 0.1 mm, 35-36 = 0.2 mm.
</paragraph>
</caption>
<caption id="DF466627FF7DFFF5FF4974ADFE9BF9FA" httpUri="https://zenodo.org/record/3241178/files/figure.png" pageId="14" pageNumber="109" startId="14.[151,264,1455,1478]" targetBox="[151,1433,193,1435]" targetPageId="14">
<paragraph id="8B8636AFFF7DFFF5FF4974ADFE9BF9FA" blockId="14.[151,1436,1455,1539]" pageId="14" pageNumber="109">
<emphasis id="B94DEABDFF7DFFF5FF4974ADFE97FA3C" bold="true" box="[151,344,1455,1478]" pageId="14" pageNumber="109">FIGURES 3745.</emphasis>
<taxonomicName id="4C394D2CFF7DFFF5FEBC74B2FD97FA3F" authority="Zilberman, 2019" authorityName="Zilberman" authorityYear="2019" box="[354,600,1456,1478]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae" status="sp. nov.">
<emphasis id="B94DEABDFF7DFFF5FEBC74B2FD97FA3F" box="[354,600,1456,1478]" italics="true" pageId="14" pageNumber="109">Termitozophilus belleae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E57C6FF7DFFF5FDBD74B2FD77FA3C" box="[611,696,1456,1477]" pageId="14" pageNumber="109" rank="species">
<emphasis id="B94DEABDFF7DFFF5FDBD74B2FD77FA3C" bold="true" box="[611,696,1456,1477]" pageId="14" pageNumber="109">sp. nov.</emphasis>
</taxonomicNameLabel>
37, 38, head (ventral dorsal); 39, antenna; 40, pos-mentum; 41, prementum; 42, maxilla; 43, labrum; 44, 35, left and right mandibles (female, male). Scale bars: 0.05 mm, except figs. 37-38 = 0.1 mm, 39 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B8636AFFF7DFFF5FF197732FDF3F94C" blockId="14.[151,1437,1584,2005]" pageId="14" pageNumber="109">
<taxonomicName id="4C394D2CFF7DFFF5FF197732FE1DF9B0" authorityName="Zilberman" authorityYear="2019" box="[199,466,1584,1609]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7DFFF5FF197732FE1DF9B0" box="[199,466,1584,1609]" italics="true" pageId="14" pageNumber="109">Termitozophilus belleae</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="4C394D2CFF7DFFF5FCD37733FCA6F9B0" authorityName="Silvestri" authorityYear="1901" box="[781,873,1584,1609]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7DFFF5FCD37733FCA6F9B0" box="[781,873,1584,1609]" italics="true" pageId="14" pageNumber="109">T. laetus</emphasis>
</taxonomicName>
, by integument mostly dark brown, widest region of head in basal region, clos to foramen magnum, presence of strong secondary sclerotization of tergites, long spine-like bristles and trapezoidal shape of elytra. In addition,
<taxonomicName id="4C394D2CFF7DFFF5FC8F777BFC62F968" authorityName="Silvestri" authorityYear="1901" box="[849,941,1656,1681]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7DFFF5FC8F777BFC62F968" box="[849,941,1656,1681]" italics="true" pageId="14" pageNumber="109">T. laetus</emphasis>
</taxonomicName>
has a very elongate antennomere XI, which is about three times longer than wide.
</paragraph>
<paragraph id="8B8636AFFF7DFFF5FF1977C2FAACF8D8" blockId="14.[151,1437,1584,2005]" pageId="14" pageNumber="109">
The absence of suture separating mentum-submentum in
<taxonomicName id="4C394D2CFF7DFFF5FCBC77C2FBBEF920" authorityName="Zilberman" authorityYear="2019" box="[866,1137,1728,1753]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7DFFF5FCBC77C2FBBEF920" box="[866,1137,1728,1753]" italics="true" pageId="14" pageNumber="109">Termitozophilus belleae</emphasis>
</taxonomicName>
is unique in the genus, a possible convergent character shared with the other Corotocini groups. Since specimens of
<taxonomicName id="4C394D2CFF7DFFF5FB7777E7FAFFF904" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[1193,1328,1764,1789]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="mirandus">
<emphasis id="B94DEABDFF7DFFF5FB7777E7FAFFF904" box="[1193,1328,1764,1789]" italics="true" pageId="14" pageNumber="109">T. mirandus</emphasis>
</taxonomicName>
were not available for dissection, I propose to revise description of the genus provided in Seeverss (1957) monograph.
</paragraph>
<paragraph id="8B8636AFFF7DFFF5FF19762EFDFAF82D" blockId="14.[151,1437,1584,2005]" pageId="14" pageNumber="109">
Additional differences between
<taxonomicName id="4C394D2CFF7DFFF5FDF1762EFCF4F8BC" authorityName="Zilberman" authorityYear="2019" box="[559,827,1836,1861]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7DFFF5FDF1762EFCF4F8BC" box="[559,827,1836,1861]" italics="true" pageId="14" pageNumber="109">Termitozophilus belleae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C394D2CFF7DFFF5FCAD762FFC1FF8BC" authorityName="Silvestri" authorityYear="1901" box="[883,976,1836,1861]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7DFFF5FCAD762FFC1FF8BC" box="[883,976,1836,1861]" italics="true" pageId="14" pageNumber="109">T. laetus</emphasis>
</taxonomicName>
are the position of bristle P2 on labrum, which is placed before middle of labrum length in
<taxonomicName id="4C394D2CFF7DFFF5FD2A7653FCAAF890" authorityName="Zilberman" authorityYear="2019" box="[756,869,1872,1897]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7DFFF5FD2A7653FCAAF890" box="[756,869,1872,1897]" italics="true" pageId="14" pageNumber="109">T. belleae</emphasis>
</taxonomicName>
and after the middle in
<taxonomicName id="4C394D2CFF7DFFF5FB547653FB24F890" authorityName="Silvestri" authorityYear="1901" box="[1162,1259,1872,1897]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7DFFF5FB547653FB24F890" box="[1162,1259,1872,1897]" italics="true" pageId="14" pageNumber="109">T. laetus</emphasis>
</taxonomicName>
; chaetotaxy of mentum is composed of bristles
<emphasis id="B94DEABDFF7DFFF5FDD87677FDEDF875" box="[518,546,1909,1932]" italics="true" pageId="14" pageNumber="109">pa</emphasis>
and
<emphasis id="B94DEABDFF7DFFF5FD847677FDBBF875" box="[602,628,1909,1932]" italics="true" pageId="14" pageNumber="109">px</emphasis>
in
<taxonomicName id="4C394D2CFF7DFFF5FD447677FCC7F874" authorityName="Zilberman" authorityYear="2019" box="[666,776,1908,1933]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7DFFF5FD447677FCC7F874" box="[666,776,1908,1933]" italics="true" pageId="14" pageNumber="109">T. belleae</emphasis>
</taxonomicName>
and
<emphasis id="B94DEABDFF7DFFF5FC9E7677FC93F875" box="[832,860,1909,1932]" italics="true" pageId="14" pageNumber="109">ap</emphasis>
,
<emphasis id="B94DEABDFF7DFFF5FCB57677FC48F875" box="[875,903,1909,1932]" italics="true" pageId="14" pageNumber="109">pa</emphasis>
and
<emphasis id="B94DEABDFF7DFFF5FC617677FC15F875" box="[959,986,1909,1932]" italics="true" pageId="14" pageNumber="109">px</emphasis>
in
<taxonomicName id="4C394D2CFF7DFFF5FC217677FB93F874" authorityName="Silvestri" authorityYear="1901" box="[1023,1116,1908,1933]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7DFFF5FC217677FB93F874" box="[1023,1116,1908,1933]" italics="true" pageId="14" pageNumber="109">T. laetus</emphasis>
</taxonomicName>
; chaetotaxy of tergite IX, is composed of six bristles in
<taxonomicName id="4C394D2CFF7DFFF5FE0A769BFD8CF848" authorityName="Zilberman" authorityYear="2019" box="[468,579,1944,1969]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7DFFF5FE0A769BFD8CF848" box="[468,579,1944,1969]" italics="true" pageId="14" pageNumber="109">T. belleae</emphasis>
</taxonomicName>
. and five in
<taxonomicName id="4C394D2CFF7DFFF5FD09769BFCF9F848" authorityName="Silvestri" authorityYear="1901" box="[727,822,1944,1969]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7DFFF5FD09769BFCF9F848" box="[727,822,1944,1969]" italics="true" pageId="14" pageNumber="109">T. laetus</emphasis>
</taxonomicName>
; and tergite VIII is about twice wider than long in
<taxonomicName id="4C394D2CFF7DFFF5FA59769BFF27F82C" authorityName="Zilberman" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7DFFF5FA59769BFF27F82C" italics="true" pageId="14" pageNumber="109">T. belleae</emphasis>
</taxonomicName>
, and subquadrate in
<taxonomicName id="4C394D2CFF7DFFF5FE0E76BFFDE2F82C" authorityName="Silvestri" authorityYear="1901" box="[464,557,1980,2005]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="109" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94DEABDFF7DFFF5FE0E76BFFDE2F82C" box="[464,557,1980,2005]" italics="true" pageId="14" pageNumber="109">T. laetus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3236524FF7CFFF4FF19719AFAC5FEB9" pageId="15" pageNumber="110" type="reference_group">
<paragraph id="8B8636AFFF7CFFF4FF19719AFAC5FEB9" blockId="15.[151,1436,151,320]" pageId="15" pageNumber="110">
In
<taxonomicName id="4C394D2CFF7CFFF4FF3B7195FE3FFF49" ID-CoL="8YBMG" authorityName="Zilberman" authorityYear="2019" box="[229,496,151,176]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="110" phylum="Arthropoda" rank="species" species="belleae">
<emphasis id="B94DEABDFF7CFFF4FF3B7195FE3FFF49" box="[229,496,151,176]" italics="true" pageId="15" pageNumber="110">Termitozophilus belleae</emphasis>
</taxonomicName>
wings were absent. The presence or absence of wings in genera of Corotocini needs to be reexamine. In genus
<taxonomicName id="4C394D2CFF7CFFF4FE6271BFFCA3FF2C" ID-CoL="8XKKC" authority="Schiodte, 1853" authorityName="Schiodte" authorityYear="1853" box="[444,876,188,213]" class="Insecta" family="Staphylinidae" genus="Corotoca" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEABDFF7CFFF4FE6271BFFDEAFF2D" box="[444,549,189,212]" italics="true" pageId="15" pageNumber="110">Corotoca</emphasis>
<bibRefCitation id="EFA84B5EFF7CFFF4FDF271BEFD1CFF2C" author="Schiodte, J. M. C." box="[556,723,188,213]" pageId="15" pageNumber="110" pagination="101 - 103" refId="ref6283" refString="Schiodte, J. M. C. (1853) On some Staphylinidae, found in the nests of termites. Proceedings of the Zoological Society of London, 21, 101 - 103." type="journal article" year="1853">Schiødte, 1853</bibRefCitation>
(Corotocina)
</taxonomicName>
, there are some species with wings developed and other with wings reduced, as discussed in a previous paper (
<bibRefCitation id="EFA84B5EFF7CFFF4FC9371DDFBDEFF01" author="Zilberman, B." box="[845,1041,223,248]" pageId="15" pageNumber="110" pagination="547 - 560" refId="ref6516" refString="Zilberman, B. (2018) New species and synonymy in the genus Corotoca Schiodte, 1853 (Coleoptera, Aleocharinae, Corotocini). Zootaxa, 4434 (3), 547 - 560. https: // doi. org / 10.11646 / zootaxa. 4434.3.9" type="journal article" year="2018">Zilberman, 2018</bibRefCitation>
). In the genus
<taxonomicName id="4C394D2CFF7CFFF4FB6271DDFF1DFEE5" ID-CoL="8W476" authority="Mann, 1923" authorityName="Mann" authorityYear="1923" class="Insecta" family="Staphylinidae" genus="Thyreoxenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEABDFF7CFFF4FB6271DDFA85FF01" box="[1212,1354,223,248]" italics="true" pageId="15" pageNumber="110">Thyreoxenus</emphasis>
<bibRefCitation id="EFA84B5EFF7CFFF4FA8A71E2FF1DFEE5" author="Mann, W. M." pageId="15" pageNumber="110" pagination="323 - 366" refId="ref6173" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from northern South America. Zoologica, 3 (17), 323 - 366." type="journal article" year="1923">Mann, 1923</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFA84B5EFF7CFFF4FF037006FE18FEE4" author="Jacobson, H. R. &amp; Kistner, D. H. &amp; Pasteels, J. M." box="[221,471,260,285]" pageId="15" pageNumber="110" pagination="1 - 245" refId="ref6071" refString="Jacobson, H. R., Kistner, D. H. &amp; Pasteels, J. M. (1986) Generic revision, phylogenetic classification, and phylogeny of the termitophilous tribe Corotocini (Coleoptera: Staphylinidae). Sociobiology, 12 (1), 1 - 245." type="journal article" year="1986">
Jacobson
<emphasis id="B94DEABDFF7CFFF4FE927007FEB1FEE4" box="[332,382,260,285]" italics="true" pageId="15" pageNumber="110">et al</emphasis>
. (1986)
</bibRefCitation>
stated that the wings were present, but in the original description of the genus,
<bibRefCitation id="EFA84B5EFF7CFFF4FA857007FF2EFEB9" author="Mann, W. M." pageId="15" pageNumber="110" pagination="323 - 366" refId="ref6173" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from northern South America. Zoologica, 3 (17), 323 - 366." type="journal article" year="1923">Mann (1923)</bibRefCitation>
observed that in
<taxonomicName id="4C394D2CFF7CFFF4FE7C7025FCF3FEB9" ID-CoL="7BXHX" authority="Mann, 1923" authorityName="Mann" authorityYear="1923" box="[418,828,295,320]" class="Insecta" family="Staphylinidae" genus="Thyreoxenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="110" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B94DEABDFF7CFFF4FE7C7025FD65FEB9" box="[418,682,295,320]" italics="true" pageId="15" pageNumber="110">Thyreoxenus pulchellus</emphasis>
<bibRefCitation id="EFA84B5EFF7CFFF4FD6F702AFCF3FEB9" author="Mann, W. M." box="[689,828,296,320]" pageId="15" pageNumber="110" pagination="323 - 366" refId="ref6173" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from northern South America. Zoologica, 3 (17), 323 - 366." type="journal article" year="1923">Mann, 1923</bibRefCitation>
</taxonomicName>
, the wings were absent (confirmed here).
</paragraph>
</subSubSection>
</treatment>
</document>