treatments-xml/data/03/90/75/0390751B3D67FFC5FF61FABEFD7CBD21.xml

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<mods:title id="7FD8E95CB0877BC5B6D496588C60FFE6">Taxonomic revision of the Brazilian Atlantic Forest Atractus (Reptilia: Serpentes: Dipsadidae) 2364</mods:title>
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<treatment id="0390751B3D67FFC5FF61FABEFD7CBD21" ID-DOI="http://doi.org/10.5281/zenodo.5323920" ID-Zenodo-Dep="5323920" LSID="urn:lsid:plazi:treatment:0390751B3D67FFC5FF61FABEFD7CBD21" httpUri="http://treatment.plazi.org/id/0390751B3D67FFC5FF61FABEFD7CBD21" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C32397863D67FFC0FF61FABEFE21BF79" box="[151,449,1299,1325]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B86C40D3D67FFC0FF61FABEFE21BF79" blockId="11.[151,449,1299,1360]" box="[151,449,1299,1325]" pageId="11" pageNumber="12">
<heading id="D0CE73613D67FFC0FF61FABEFE21BF79" bold="true" box="[151,449,1299,1325]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B94D181F3D67FFC0FF61FABEFE21BF79" bold="true" box="[151,449,1299,1325]" pageId="11" pageNumber="12">
<taxonomicName id="4C39BF8E3D67FFC0FF61FABEFEBCBF79" authority="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite, 2010" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[151,348,1299,1325]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="francoi" status="sp. nov.">
<emphasis id="B94D181F3D67FFC0FF61FABEFEBCBF79" bold="true" box="[151,348,1299,1325]" italics="true" pageId="11" pageNumber="12">Atractus francoi</emphasis>
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<taxonomicNameLabel id="A27EA5643D67FFC0FE92FABEFE21BF79" box="[356,449,1299,1325]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C32397863D67FFC0FF61FA9BFD07BFE9" pageId="11" pageNumber="12" type="description">
<paragraph id="8B86C40D3D67FFC0FF61FA9BFE8BBF04" blockId="11.[151,449,1299,1360]" box="[151,363,1334,1360]" pageId="11" pageNumber="12">
<figureCitation id="1302D8883D67FFC0FF61FA9BFEFFBF04" box="[151,287,1334,1360]" captionStart="FIGURE 2" captionStartId="9.[151,255,1856,1880]" captionTargetBox="[155,1428,180,1813]" captionTargetId="figure-50@9.[151,1436,166,1832]" captionTargetPageId="9" captionText="FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. francoi sp. nov. (MNRJ 17537), (D) A. guentheri (BMNH 1946.1.1.76), and (E) A. maculatus (BMNH 1946.1.6.46)." figureDoi="http://doi.org/10.5281/zenodo.5315512" httpUri="https://zenodo.org/record/5315512/files/figure.png" pageId="11" pageNumber="12">Figs. 2B–C</figureCitation>
, 
<figureCitation id="1302D8883D67FFC0FEDDFA9BFEAFBF04" box="[299,335,1334,1360]" captionStart="FIGURE 3" captionStartId="10.[151,255,1944,1968]" captionTargetBox="[160,1430,181,1936]" captionTargetId="figure-82@10.[151,1436,146,1952]" captionTargetPageId="10" captionText="FIGURE 3. General view in life of (A, B) Atractus caete sp. nov. from Chã Preta, state of Alagoas, photo by G. Skuk, (C) A. francoi sp. nov. from Santa Virgínia, state of São Paulo, photo by P. Hartmann, (D) A. guentheri from Ilhéus, state of Bahia, photo by A.J.S. Argôlo, and (E) A. pantostictus from Palmas, state of Tocantis, photo by O.A.V. Marques." figureDoi="http://doi.org/10.5281/zenodo.5315516" httpUri="https://zenodo.org/record/5315516/files/figure.png" pageId="11" pageNumber="12">3C</figureCitation>
, 
<figureCitation id="1302D8883D67FFC0FEAAFA9BFE8BBF04" box="[348,363,1334,1360]" captionStart="FIGURE 5" captionStartId="13.[151,255,1619,1643]" captionTargetBox="[403,1186,196,1592]" captionTargetId="figure-158@13.[386,1201,163,1609]" captionTargetPageId="13" captionText="FIGURE 5. Dorsal (A) and lateral (B) views of head, and lateral view of body (C) of the holotype of Atractus francoi sp. nov. (MNRJ 17535). Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315522" httpUri="https://zenodo.org/record/5315522/files/figure.png" pageId="11" pageNumber="12">5</figureCitation>
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<taxonomicName id="4C39BF8E3D67FFC0FF61FA2BFEF7BFC9" box="[151,279,1414,1437]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B94D181F3D67FFC0FF61FA2BFF11BFC9" box="[151,241,1414,1437]" italics="true" pageId="11" pageNumber="12">Atractus</emphasis>
sp.
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– 
<bibRefCitation id="EFA8B9FC3D67FFC0FEC4FA2BFD4FBFC9" author="Marques, O. A. V. &amp; Eterovic, A. &amp; Sazima, I." box="[306,687,1414,1437]" pageId="11" pageNumber="12" refId="ref39360" refString="Marques, O. A. V., Eterovic, A. &amp; Sazima, I. (2004) Snakes of the Brazilian Rainforest: An Illustrate Field Guide for the Serra do Mar. Holos, Ribeirao Preto, Brazil, 205 pp." type="book" year="2004">
<collectingRegion id="49FD0AEF3D67FFC0FEC4FA2BFE72BFC9" box="[306,402,1414,1437]" country="Italy" name="Marche" pageId="11" pageNumber="12">Marques</collectingRegion>
, Eterovic &amp; Sazima, 2004
</bibRefCitation>
; Snakes of the Brazilian Atlantic Forest: An Illustrate Field Guide for the Serra do Mar. Holos, Ribeirão Preto, 1–205:69.
</paragraph>
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<materialsCitation id="3B51CE503D67FFC0FF61FA40FE7EBC0C" collectingDate="2006-07-17" collectionCode="MNRJ, DZUFRJ" collectorName="Adult &amp; M. C. Carlo &amp; Fazenda Recanto" country="Brazil" elevation="600" latitude="-22.833334" location="Serra do Piloto" longLatPrecision="1258" longitude="-44.05" municipality="Mangaratiba" pageId="11" pageNumber="12" specimenCode="MNRJ 17537, DZUFRJ 1742" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
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<typeStatus id="54827AAF3D67FFC0FF61FA40FEEDBC53" box="[151,269,1517,1543]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
:
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<collectorName id="26CCA1DB3D67FFC0FEECFA43FEBCBC5C" box="[282,348,1518,1544]" pageId="11" pageNumber="12">Adult</collectorName>
male, 
<specimenCode id="DB9F6C763D67FFC0FE5DFA43FDA9BC5C" box="[427,585,1518,1544]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 17537</specimenCode>
(formerly 
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), collected 
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<collectingDate id="EFC31B253D67FFC0FC08FA43FB74BC5C" box="[1022,1172,1518,1544]" pageId="11" pageNumber="12" value="2006-07-17">17 July 2006</collectingDate>
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by 
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, locality 
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, 
<location id="8EE692D63D67FFC0FE6DF9BBFDAFBC64" LSID="urn:lsid:plazi:treatment:0390751B3D67FFC5FF61FABEFD7CBD21:8EE692D63D67FFC0FE6DF9BBFDAFBC64" box="[411,591,1558,1584]" country="Brazil" latitude="-22.833334" longLatPrecision="1258" longitude="-44.05" municipality="Mangaratiba" name="Serra do Piloto" pageId="11" pageNumber="12" stateProvince="Rio de Janeiro">Serra do Piloto</location>
(
<geoCoordinate id="EE0DA2CA3D67FFC0FD94F9BBFD25BC64" box="[610,709,1558,1584]" degrees="22" direction="south" minutes="50" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="-22.833334">22°50’S</geoCoordinate>
, 
<geoCoordinate id="EE0DA2CA3D67FFC0FD22F9BBFCDFBC64" box="[724,831,1558,1584]" degrees="44" direction="west" minutes="03" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="-44.05">44°03’W</geoCoordinate>
, ca. 
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<elevation id="0014233E3D67FFC0FC8EF9BBFC28BC64" box="[888,968,1558,1584]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="600.0">600 m</elevation>
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), municipality of 
<collectingMunicipality id="6BE25E773D67FFC0FB69F9BBFAD6BC64" box="[1183,1334,1558,1584]" pageId="11" pageNumber="12">Mangaratiba</collectingMunicipality>
, state of 
<collectingRegion id="49FD0AEF3D67FFC0FF61F993FEA2BC0C" box="[151,322,1598,1624]" country="Brazil" name="Rio de Janeiro" pageId="11" pageNumber="12">Rio de Janeiro</collectingRegion>
, 
<collectingCountry id="F32E849D3D67FFC0FEB9F993FE7ABC0C" box="[335,410,1598,1624]" name="Brazil" pageId="11" pageNumber="12">Brazil</collectingCountry>
.
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</paragraph>
<paragraph id="8B86C40D3D67FFC0FF30F9C8FCFBBD1C" blockId="11.[151,1437,1517,1864]" pageId="11" pageNumber="12">
<materialsCitation id="3B51CE503D67FFC0FF30F9C8FD61BCFC" collectionCode="IBSP" collectorName="Six" elevation="600" latitude="-22.633333" location="Sao Jose do Barreiro" longLatPrecision="1259" longitude="-44.583332" pageId="11" pageNumber="12" specimenCode="IBSP 72654" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<emphasis id="B94D181F3D67FFC0FF30F9C8FE94BC2B" bold="true" box="[198,372,1637,1663]" pageId="11" pageNumber="12">
<typeStatus id="54827AAF3D67FFC0FF30F9C8FEA7BC2B" box="[198,327,1637,1663]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
: 
<collectorName id="26CCA1DB3D67FFC0FEBAF9C8FE94BC2B" box="[332,372,1637,1663]" pageId="11" pageNumber="12">Six</collectorName>
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n specimens: adult male (
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), municipality of 
<location id="8EE692D63D67FFC0FBD4F9CBFAC1BCD4" LSID="urn:lsid:plazi:treatment:0390751B3D67FFC5FF61FABEFD7CBD21:8EE692D63D67FFC0FBD4F9CBFAC1BCD4" box="[1058,1313,1638,1664]" latitude="-22.633333" longLatPrecision="1259" longitude="-44.583332" name="Sao Jose do Barreiro" pageId="11" pageNumber="12" stateProvince="Sao Paulo">São José do Barreiro</location>
(
<geoCoordinate id="EE0DA2CA3D67FFC0FAC0F9CBFA77BCD4" box="[1334,1431,1638,1664]" degrees="22" direction="south" minutes="38" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="-22.633333">22º38’S</geoCoordinate>
, 
<geoCoordinate id="EE0DA2CA3D67FFC0FF61F923FF1FBCFC" box="[151,255,1678,1704]" degrees="44" direction="west" minutes="35" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="-44.583332">44º35’W</geoCoordinate>
, ca. 
<quantity id="4CC169E83D67FFC0FECCF923FE6BBCFC" box="[314,395,1678,1704]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="600.0">
<elevation id="0014233E3D67FFC0FECCF923FE6BBCFC" box="[314,395,1678,1704]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="600.0">600 m</elevation>
</quantity>
), state of 
<collectingRegion id="49FD0AEF3D67FFC0FDF3F923FD61BCFC" box="[517,641,1678,1704]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="12">São Paulo</collectingRegion>
</materialsCitation>
; 
<materialsCitation id="3B51CE503D67FFC0FD67F923FA78BCAC" collectionCode="IBSP" collectorName="P. A. Hartmann &amp; Nucleo SantaVirginia" elevation="770" latitude="-23.233334" location="Sao Luis de Paraitinga" longLatPrecision="1256" longitude="-45.333332" municipality="Parque Estadual da Serra do Mar" pageId="11" pageNumber="12" specimenCode="IBSP 74724, IBSP 53924, 74723, IBSP 74648" specimenCount="1" specimenCount-juvenile="1" stateProvince="Sao Paulo" typeStatus="paratype">
an adult male (
<specimenCode id="DB9F6C763D67FFC0FCB1F923FC3FBCFC" box="[839,991,1678,1704]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="11" pageNumber="12">IBSP 74724</specimenCode>
), 
<specimenCount id="9D3F0F843D67FFC0FC03F923FB6ABCFC" box="[1013,1162,1678,1704]" pageId="11" pageNumber="12" type="juvenile">two juvenile</specimenCount>
females (
<specimenCode id="DB9F6C763D67FFC0FAF7F923FA77BCFC" box="[1281,1431,1678,1704]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="11" pageNumber="12">IBSP 53924</specimenCode>
, 
<specimenCode id="DB9F6C763D67FFC0FF61F91BFF04BC84" box="[151,228,1718,1744]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="11" pageNumber="12">74723</specimenCode>
), and adult female (
<specimenCode id="DB9F6C763D67FFC0FE3BF91BFD81BC84" box="[461,609,1718,1744]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="11" pageNumber="12">IBSP 74648</specimenCode>
), collected by 
<collectorName id="26CCA1DB3D67FFC0FCFBF91BFC25BC84" box="[781,965,1718,1744]" pageId="11" pageNumber="12">P. A. Hartmann</collectorName>
, 
<collectorName id="26CCA1DB3D67FFC0FC24F91BFB32BC84" box="[978,1234,1718,1744]" pageId="11" pageNumber="12">Núcleo SantaVirgínia</collectorName>
, 
<collectingMunicipality id="6BE25E773D67FFC0FB28F91BFEBBBCAC" pageId="11" pageNumber="12">Parque Estadual da Serra do Mar</collectingMunicipality>
, municipality of 
<location id="8EE692D63D67FFC0FDD1F973FCD4BCAC" LSID="urn:lsid:plazi:treatment:0390751B3D67FFC5FF61FABEFD7CBD21:8EE692D63D67FFC0FDD1F973FCD4BCAC" box="[551,820,1758,1784]" latitude="-23.233334" longLatPrecision="1256" longitude="-45.333332" municipality="Parque Estadual da Serra do Mar" name="Sao Luis de Paraitinga" pageId="11" pageNumber="12" stateProvince="Sao Paulo">São Luís de Paraitinga</location>
(
<geoCoordinate id="EE0DA2CA3D67FFC0FCB1F973FC47BCAC" box="[839,935,1758,1784]" degrees="23" direction="south" minutes="14" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="-23.233334">23º14’S</geoCoordinate>
, 
<geoCoordinate id="EE0DA2CA3D67FFC0FC43F973FBFEBCAC" box="[949,1054,1758,1784]" degrees="45" direction="west" minutes="20" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="-45.333332">45º20’W</geoCoordinate>
, ca. 
<quantity id="4CC169E83D67FFC0FBA0F973FB45BCAC" box="[1110,1189,1758,1784]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="11" pageNumber="12" unit="m" value="770.0">
<elevation id="0014233E3D67FFC0FBA0F973FB45BCAC" box="[1110,1189,1758,1784]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="11" pageNumber="12" unit="m" value="770.0">770 m</elevation>
</quantity>
), state of 
<collectingRegion id="49FD0AEF3D67FFC0FAEAF973FA78BCAC" box="[1308,1432,1758,1784]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="12">São Paulo</collectingRegion>
</materialsCitation>
; 
<materialsCitation id="3B51CE503D67FFC0FF61F8ABFCF7BD1C" collectingDate="1989-09" collectionCode="R" collectorName="Schasse &amp; Nova Friburgo" elevation="915" latitude="-22.266666" location="Rio de Janeiro" longLatPrecision="1260" longitude="-42.533333" pageId="11" pageNumber="12" specimenCode="MNRJ 17536" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
adult female (
<specimenCode id="DB9F6C763D67FFC0FEC9F8ABFE07BD74" box="[319,487,1798,1824]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 17536</specimenCode>
), collected on 
<date id="FF87E2CD3D67FFC0FD6BF8ABFCBFBD74" box="[669,863,1798,1824]" pageId="11" pageNumber="12" value="1989-09">
<collectingDate id="EFC31B253D67FFC0FD6BF8ABFCBFBD74" box="[669,863,1798,1824]" pageId="11" pageNumber="12" value="1989-09">September 1989</collectingDate>
</date>
by 
<collectionCode id="ED285CC83D67FFC0FC65F8ABFC4ABD74" box="[915,938,1798,1824]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="12">R</collectionCode>
. 
<collectorName id="26CCA1DB3D67FFC0FC4FF8ABFBF9BD74" box="[953,1049,1798,1824]" pageId="11" pageNumber="12">Schasse</collectorName>
, municipality of 
<collectorName id="26CCA1DB3D67FFC0FB1DF8ABFA7CBD74" box="[1259,1436,1798,1824]" pageId="11" pageNumber="12">Nova Friburgo</collectorName>
(
<geoCoordinate id="EE0DA2CA3D67FFC0FF56F883FEE0BD1C" box="[160,256,1838,1864]" degrees="22" direction="south" minutes="16" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="-22.266666">22º16’S</geoCoordinate>
, 
<geoCoordinate id="EE0DA2CA3D67FFC0FEFBF883FE96BD1C" box="[269,374,1838,1864]" degrees="42" direction="west" minutes="32" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="-42.533333">42º32’W</geoCoordinate>
, ca. 
<quantity id="4CC169E83D67FFC0FE5CF883FE18BD1C" box="[426,504,1838,1864]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="11" pageNumber="12" unit="m" value="915.0">
<elevation id="0014233E3D67FFC0FE5CF883FE18BD1C" box="[426,504,1838,1864]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="11" pageNumber="12" unit="m" value="915.0">915 m</elevation>
</quantity>
), state of 
<collectingRegion id="49FD0AEF3D67FFC0FD9CF883FCF7BD1C" box="[618,791,1838,1864]" country="Brazil" name="Rio de Janeiro" pageId="11" pageNumber="12">Rio de Janeiro</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32397863D67FFC7FF30F8D0FB72BE20" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8B86C40D3D67FFC7FF30F8D0FE84BBF0" blockId="11.[151,1436,1917,2024]" lastBlockId="12.[151,1436,152,2024]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B94D181F3D67FFC0FF30F8D0FEA8BDC3" bold="true" box="[198,328,1917,1943]" pageId="11" pageNumber="12">Diagnosis:</emphasis>
<taxonomicName id="4C39BF8E3D67FFC0FEA7F8D3FDF2BDC3" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[337,530,1918,1943]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="francoi">
<emphasis id="B94D181F3D67FFC0FEA7F8D3FDF2BDC3" box="[337,530,1918,1943]" italics="true" pageId="11" pageNumber="12">Atractus francoi</emphasis>
</taxonomicName>
is distinguished from all known congeners by the following combination of characters: (1) 17/17/17 smooth dorsal scale rows; (2) two postoculars; (3) moderate loreal; (4) temporals 1+2; (5) usually seven supralabials (six in one side), third and fourth contacting orbit; (6) seven infralabials, first four contacting chinshields; (7) eight or nine maxillary teeth; (8) two to four gular scale rows; (9) three or four preventrals; (10) 156–164 ventrals in females and 
<quantity id="4CC169E83D60FFC7FCDFFF6DFC4FBA8E" box="[809,943,192,218]" metricMagnitude="0" metricUnit="m" metricValue="4.1402" metricValueMax="4.1655999999999995" metricValueMin="4.1148" pageId="12" pageNumber="13" unit="in" value="163.0" valueMax="164.0" valueMin="162.0">162–164 in</quantity>
males; (11) 14–19 subcaudals in females and 
<quantity id="4CC169E83D60FFC7FF3EFF45FED0BB56" box="[200,304,232,258]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="7.62" metricValueMin="6.35" pageId="12" pageNumber="13" unit="in" value="27.5" valueMax="30.0" valueMin="25.0">25–30 in</quantity>
males; (12) dorsum uniformly black in adults and brown with scattered small black dots in juveniles; (13) venter beige anteriorly, becoming gradually black after 15 
<superScript id="7C4C69453D60FFC7FC21FEBDFC03BB4A" attach="left" box="[983,995,272,286]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
ventral in juveniles and adults; (14) moderate body size, females reaching 
<quantity id="4CC169E83D60FFC7FDADFE97FD5CBB00" box="[603,700,314,340]" metricMagnitude="-1" metricUnit="m" metricValue="4.7299999999999995" pageId="12" pageNumber="13" unit="mm" value="473.0">473 mm</quantity>
SVL and males 
<quantity id="4CC169E83D60FFC7FC75FE97FC04BB00" box="[899,996,314,340]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="13" unit="mm" value="420.0">420 mm</quantity>
SVL; (15) tail short in females (6.3– 8.6% SVL) and moderate (12.6–13.6% SVL) in males; (16) moderately bilobed, semicapitate, and semicalyculate hemipenis.
</paragraph>
<paragraph id="8B86C40D3D60FFC7FF30FE1FFCC1B9A8" blockId="12.[151,1436,152,2024]" pageId="12" pageNumber="13">
<emphasis id="B94D181F3D60FFC7FF30FE1FFE96BB98" bold="true" box="[198,374,434,460]" pageId="12" pageNumber="13">Comparisons:</emphasis>
Among congeners, 
<taxonomicName id="4C39BF8E3D60FFC7FD94FE1EFD38BB98" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[610,728,435,460]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="francoi">
<emphasis id="B94D181F3D60FFC7FD94FE1EFD38BB98" box="[610,728,435,460]" italics="true" pageId="12" pageNumber="13">A. francoi</emphasis>
</taxonomicName>
shares 17 dorsal scale rows, dorsal colour pattern uniformly black in adults, venter dark with posterior margin of ventral scales with pale (beige) pigmentation, seven to ten maxillary teeth, and semicapitate and semicalyculate hemipenis [unknown in 
<taxonomicName id="4C39BF8E3D60FFC7FBA6FDAEFB4EB848" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[1104,1198,515,540]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="caete">
<emphasis id="B94D181F3D60FFC7FBA6FDAEFB4EB848" box="[1104,1198,515,540]" italics="true" pageId="12" pageNumber="13">A. caete</emphasis>
</taxonomicName>
] only with 
<emphasis id="B94D181F3D60FFC7FAC0FDAEFEC6B810" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C39BF8E3D60FFC7FAC0FDAEFA77B848" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[1334,1431,515,540]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="caete">A. caete</taxonomicName>
, 
<taxonomicName id="4C39BF8E3D60FFC7FF61FD86FEC2B810" authorityName="Amaral" authorityYear="1930" box="[151,290,555,580]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="serranus">A. serranus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C39BF8E3D60FFC7FE92FD86FDEBB810" authorityName="Amaral" authorityYear="1926" box="[356,523,555,580]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="trihedrurus">
<emphasis id="B94D181F3D60FFC7FE92FD86FDEBB810" box="[356,523,555,580]" italics="true" pageId="12" pageNumber="13">A. trihedrurus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C39BF8E3D60FFC7FDEAFD86FD3CB810" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[540,732,555,580]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="francoi">
<emphasis id="B94D181F3D60FFC7FDEAFD86FD3CB810" box="[540,732,555,580]" italics="true" pageId="12" pageNumber="13">Atractus francoi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C39BF8E3D60FFC7FC8BFD86FC3EB810" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[893,990,555,580]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="caete">
<emphasis id="B94D181F3D60FFC7FC8BFD86FC3EB810" box="[893,990,555,580]" italics="true" pageId="12" pageNumber="13">A. caete</emphasis>
</taxonomicName>
by usually having seven supralabials (six in one side of 
<specimenCount id="9D3F0F843D60FFC7FE8DFDFFFDC2B838" box="[379,546,594,620]" pageId="12" pageNumber="13" type="generic">one specimen</specimenCount>
), eight or nine maxillary teeth, and juveniles dark brown with small black dots dorsally and having venter mostly brown (vs. usually six supralabials, seven maxillary teeth, and juvenile with dorsum uniformly black and venter mostly cream); from 
<taxonomicName id="4C39BF8E3D60FFC7FC6AFD0EFBC8B8E8" authorityName="Amaral" authorityYear="1930" box="[924,1064,675,700]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="serranus">
<emphasis id="B94D181F3D60FFC7FC6AFD0EFBC8B8E8" box="[924,1064,675,700]" italics="true" pageId="12" pageNumber="13">A. serranus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C39BF8E3D60FFC7FB9DFD0EFAF6B8E8" authorityName="Amaral" authorityYear="1926" box="[1131,1302,675,700]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="trihedrurus">
<emphasis id="B94D181F3D60FFC7FB9DFD0EFAF6B8E8" box="[1131,1302,675,700]" italics="true" pageId="12" pageNumber="13">A. trihedrurus</emphasis>
</taxonomicName>
by having females attain maturity below 
<quantity id="4CC169E83D60FFC7FE09FD67FD80B8B0" box="[511,608,714,740]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="500.0">500 mm</quantity>
SVL, 152–154 ventrals in males and 
<quantity id="4CC169E83D60FFC7FBD7FD67FB46B8B0" box="[1057,1190,714,740]" metricMagnitude="0" metricUnit="m" metricValue="4.1402" metricValueMax="4.1655999999999995" metricValueMin="4.1148" pageId="12" pageNumber="13" unit="in" value="163.0" valueMax="164.0" valueMin="162.0">162–164 in</quantity>
females, hemipenial lobes smaller than remaining capitulum, asulcate capitulum lacking medial and lobular crests, sulcus spermaticus narrow, basal region of hemipenis lacking moderate hooked spines, and barely distinct ontogenetic change of dorsal colouration (vs. mature females always more than 
<quantity id="4CC169E83D60FFC7FC40FCEFFBF7B908" box="[950,1047,834,860]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="500.0">500 mm</quantity>
SVL, 141–147 ventrals in males and 
<quantity id="4CC169E83D60FFC7FF3AFCC7FEB5B9D0" box="[204,341,874,900]" metricMagnitude="0" metricUnit="m" metricValue="3.9751" metricValueMax="4.1402" metricValueMin="3.81" pageId="12" pageNumber="13" unit="in" value="156.5" valueMax="163.0" valueMin="150.0">150–163 in</quantity>
females of 
<taxonomicName id="4C39BF8E3D60FFC7FE1CFCC6FD94B9D0" authorityName="Amaral" authorityYear="1930" box="[490,628,875,900]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="serranus">
<emphasis id="B94D181F3D60FFC7FE1CFCC6FD94B9D0" box="[490,628,875,900]" italics="true" pageId="12" pageNumber="13">A. serranus</emphasis>
</taxonomicName>
and 
<quantity id="4CC169E83D60FFC7FD43FCC7FCDDB9D0" box="[693,829,874,900]" metricMagnitude="0" metricUnit="m" metricValue="3.6322" metricValueMax="3.81" metricValueMin="3.4543999999999997" pageId="12" pageNumber="13" unit="in" value="143.0" valueMax="150.0" valueMin="136.0">136–150 in</quantity>
males and 
<quantity id="4CC169E83D60FFC7FC3BFCC7FBB5B9D0" box="[973,1109,874,900]" metricMagnitude="0" metricUnit="m" metricValue="3.8735" metricValueMax="4.0386" metricValueMin="3.7084" pageId="12" pageNumber="13" unit="in" value="152.5" valueMax="159.0" valueMin="146.0">146–159 in</quantity>
females of 
<taxonomicName id="4C39BF8E3D60FFC7FB1DFCC6FA75B9D0" authorityName="Amaral" authorityYear="1926" box="[1259,1429,875,900]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="trihedrurus">
<emphasis id="B94D181F3D60FFC7FB1DFCC6FA75B9D0" box="[1259,1429,875,900]" italics="true" pageId="12" pageNumber="13">A. trihedrurus</emphasis>
</taxonomicName>
, hemipenial lobes longer than remaining capitulum, asulcate capitulum with conspicuous medial and lobular crests, sulcus spermaticus expanded laterally, basal region of hemipenial with moderate hooked spines, and juveniles of 
<taxonomicName id="4C39BF8E3D60FFC7FEDEFC4EFE2EB9A8" authorityName="Amaral" authorityYear="1926" box="[296,462,995,1020]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="trihedrurus">
<emphasis id="B94D181F3D60FFC7FEDEFC4EFE2EB9A8" box="[296,462,995,1020]" italics="true" pageId="12" pageNumber="13">A. trihedrurus</emphasis>
</taxonomicName>
with banded colour pattern).
</paragraph>
<paragraph id="8B86C40D3D60FFC7FF30FBA6FB72BE20" blockId="12.[151,1436,152,2024]" pageId="12" pageNumber="13">
<taxonomicName id="4C39BF8E3D60FFC7FF30FBA6FE64BE70" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[198,388,1035,1060]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="francoi">
<emphasis id="B94D181F3D60FFC7FF30FBA6FE64BE70" box="[198,388,1035,1060]" italics="true" pageId="12" pageNumber="13">Atractus francoi</emphasis>
</taxonomicName>
occurrs sympatrically with 
<taxonomicName id="4C39BF8E3D60FFC7FD3CFBA6FCADBE70" baseAuthorityName="Jan" baseAuthorityYear="1862" box="[714,845,1035,1060]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="zebrinus">
<emphasis id="B94D181F3D60FFC7FD3CFBA6FCADBE70" box="[714,845,1035,1060]" italics="true" pageId="12" pageNumber="13">A. zebrinus</emphasis>
</taxonomicName>
along the coastal mountains of the states of 
<collectingRegion id="49FD0AEF3D60FFC7FAB9FBA7FF0CBE18" country="Brazil" name="Rio de Janeiro" pageId="12" pageNumber="13">Rio de Janeiro</collectingRegion>
and 
<collectingRegion id="49FD0AEF3D60FFC7FED3FB9FFE7EBE18" box="[293,414,1074,1100]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="13">São Paulo</collectingRegion>
, but differs from it by having the dorsum and venter mostly black (vs. dorsum beige to creamish red with wide transversal black blotches, venter immaculate creamish white).
</paragraph>
</subSubSection>
<subSubSection id="C32397863D60FFC5FF30FB2FFD09BFB1" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="8B86C40D3D60FFC7FF30FB2FFC54BDBC" blockId="12.[151,1436,152,2024]" pageId="12" pageNumber="13">
<emphasis id="B94D181F3D60FFC7FF30FB2FFDFFBEC8" bold="true" box="[198,543,1154,1180]" pageId="12" pageNumber="13">
Description of the 
<typeStatus id="54827AAF3D60FFC7FE5DFB2FFDFBBEC8" box="[427,539,1154,1180]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
:
</emphasis>
Adult male, SVL 
<quantity id="4CC169E83D60FFC7FD03FB2FFCB8BEC8" box="[757,856,1154,1180]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="13" unit="mm" value="420.0">420 mm</quantity>
, CL 
<quantity id="4CC169E83D60FFC7FC66FB2FFC01BEC8" box="[912,993,1154,1180]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="12" pageNumber="13" unit="mm" value="53.0">53 mm</quantity>
(12.6% SVL); body diameter 9.0 mm (2% SVL); head length 18.2 mm (4.3% SVL); head width 10.8 mm (59.3% head length); interorbital distance 6.9 mm; rostro-orbital distance 4.9 mm (70% interorbital distance); naso-orbital distance 3.7 mm; cervical constriction barely distinct; head slightly arched in lateral view, round in dorsal view; snout truncate in lateral view, round in dorsal view; rostral sub-triangular in frontal view, 3.0 mm wide, 1.0 mm high, little visible in dorsal view; internasal 1.3 mm long, as long as wide; internasal suture sinistral with respect to prefrontal suture; prefrontal 2.2 mm long, 3.5 mm wide; supraocular sub-rectangular, 1.9 mm long, 1.3 mm wide; frontal sub-pentagonal, 4.2 mm long, 4.4 mm wide; parietal 7.1 mm long, 4.0 mm wide; nasal divided; nostril located between prenasal and postnasal; prenasal 1.2 mm high, 0.6 mm long; postnasal 1.5 mm high, 0.9 mm long; loreal 2.5 mm long, 1.0 mm high, contacting second and third supralabials; eye diameter 2.0 mm; pupil subelliptical; two postoculars; upper postocular 1.1 mm high, 0.9 mm long, similar in height and slightly longer than lower postocular; temporals 1+2; anterior temporal 3.5 mm long, 1.3 mm high; upper posterior temporal fragmented into small occipital-like scales and elongate (on the left side), 5.1 mm long, 1.9 mm wide; six (left) and seven (rigth) supralabials, third and fourth contacting orbit; second supralabial higher than first and lower than third; sixth supralabial higher and seventh longer than remaining supralabials; symphisial triangular, 2.6 mm wide, 1.2 mm long; seven infralabials, first four contacting chinshields; first pair of infralabials in contact behind symphisial, preventing symphisial/chinshield contact; chinshields 5.1 mm long, 2.1 mm wide; two (left) and three (right) gular scale rows; two preventrals; 152 ventrals; 29 (left) and 30 (right) subcaudals; 17/17/17 smooth dorsal scale rows; dorsals lacking apical pits, supra-anal tubercles, and keels; eight dorsal scale rows at the level of second subcaudal; anal gland three subcaudals long; caudal spine short, robust, and rhomboid; retracted hemipenis extends to the level of 11 
<superScript id="7C4C69453D60FFC7FCCAF80EFCA8BDE5" attach="left" box="[828,840,1955,1969]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
and bifurcates at 10 
<superScript id="7C4C69453D60FFC7FBC8F80EFBAABDE5" attach="left" box="[1086,1098,1955,1969]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
subcaudal; six prediastemal and two postediastemal teeth; maxillary diastema moderate (
<figureCitation id="1302D8883D60FFC7FCA8F863FC45BDBC" box="[862,933,1998,2024]" captionStart="FIGURE 5" captionStartId="13.[151,255,1619,1643]" captionTargetBox="[403,1186,196,1592]" captionTargetId="figure-158@13.[386,1201,163,1609]" captionTargetPageId="13" captionText="FIGURE 5. Dorsal (A) and lateral (B) views of head, and lateral view of body (C) of the holotype of Atractus francoi sp. nov. (MNRJ 17535). Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315522" httpUri="https://zenodo.org/record/5315522/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF4694853D61FFC6FF61F9FEFDF5BCDE" ID-DOI="http://doi.org/10.5281/zenodo.5315522" ID-Zenodo-Dep="5315522" httpUri="https://zenodo.org/record/5315522/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,255,1619,1643]" targetBox="[403,1186,196,1592]" targetPageId="13">
<paragraph id="8B86C40D3D61FFC6FF61F9FEFDF5BCDE" blockId="13.[151,1430,1619,1674]" pageId="13" pageNumber="14">
<emphasis id="B94D181F3D61FFC6FF61F9FEFEF8BC3F" bold="true" box="[151,280,1619,1643]" pageId="13" pageNumber="14">FIGURE 5.</emphasis>
Dorsal (A) and lateral (B) views of head, and lateral view of body (C) of the holotype of 
<taxonomicName id="4C39BF8E3D61FFC6FB32F9FEFA92BC3E" authority="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite, 2010" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[1220,1394,1619,1642]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="francoi" status="sp. nov.">
<emphasis id="B94D181F3D61FFC6FB32F9FEFA92BC3E" box="[1220,1394,1619,1642]" italics="true" pageId="13" pageNumber="14">Atractus francoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EA5643D61FFC6FA8EF9FEFF25BCDE" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
(MNRJ 17535). Scale = 5 mm.
</paragraph>
</caption>
<paragraph id="8B86C40D3D61FFC6FF30F916FD95BD21" blockId="13.[151,1436,1723,2029]" pageId="13" pageNumber="14">
<emphasis id="B94D181F3D61FFC6FF30F916FE6CBC81" bold="true" box="[198,396,1723,1749]" pageId="13" pageNumber="14">Maxillary arch:</emphasis>
Arched in dorsal view, with six prediastemal and two postdiastemal teeth; prediastemal teeth large, similar in size, curved posteriorly, angular in cross section, robust at base, narrower on the apices; first five teeth generally closely spaced, space between fifth and sixth tooth moderate; maxillary diastema moderate; postdiastemal teeth about half size of prediastemal teeth; lateral process of maxilla moderately developed, lacking posterior projections.
</paragraph>
<paragraph id="8B86C40D3D61FFC5FF30F82EFD95BB03" blockId="13.[151,1436,1723,2029]" lastBlockId="14.[151,1436,152,1909]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B94D181F3D61FFC6FF30F82EFE0EBDC9" bold="true" box="[198,494,1923,1949]" pageId="13" pageNumber="14">Colour in preservative:</emphasis>
Dorsum of head dark brown, except for pale brown spots on distal portion of prefrontals; head dark brown, except for irregular pale brown dots covering anterior portion of loreal and fourth to sixth supralabials; symphisial and first three infralabials spotted with brown, remaining infralabials, chinshields, and preventrals beige coloured; gulars with dark brown dots; venter uniform beige anteriorly, with brown blotches on lateral portions of ventrals from 15 
<superScript id="7C4C69453D62FFC5FCAEFF6DFC84BA9A" attach="left" box="[856,868,192,206]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
scale; lateral blotches increase progressively in size, collapsing at level of 20 
<superScript id="7C4C69453D62FFC5FE00FF46FDE2BAAD" attach="left" box="[502,514,235,249]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
ventral; posterior to this point venter becomes uniform brown, with only the posterior margin pale (creamish brown); tail brown with beige dots on posterior suture of subcaudals; dorsum of body uniformly dark brown (
<figureCitation id="1302D8883D62FFC5FDFDFE90FD87BB03" box="[523,615,317,343]" captionStart="FIGURE 2" captionStartId="9.[151,255,1856,1880]" captionTargetBox="[155,1428,180,1813]" captionTargetId="figure-50@9.[151,1436,166,1832]" captionTargetPageId="9" captionText="FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. francoi sp. nov. (MNRJ 17537), (D) A. guentheri (BMNH 1946.1.1.76), and (E) A. maculatus (BMNH 1946.1.6.46)." figureDoi="http://doi.org/10.5281/zenodo.5315512" httpUri="https://zenodo.org/record/5315512/files/figure.png" pageId="14" pageNumber="15">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8B86C40D3D62FFC5FF30FEC8FA62BB9B" blockId="14.[151,1436,152,1909]" pageId="14" pageNumber="15">
<emphasis id="B94D181F3D62FFC5FF30FEC8FD79BB2B" bold="true" box="[198,665,357,383]" pageId="14" pageNumber="15">Juvenile colouration in preservative:</emphasis>
Dorsum of body dark brown with small (one scale long) black dots concentrated on paravertebral region, occasionally in a fragmented barely distinct vertebral black line; first four scale rows lacking marks on both sides of body; venter similar to adults with slightly paler colour.
</paragraph>
<paragraph id="8B86C40D3D62FFC5FF30FE70FB74B813" blockId="14.[151,1436,152,1909]" pageId="14" pageNumber="15">
<emphasis id="B94D181F3D62FFC5FF30FE70FDFDBBA3" bold="true" box="[198,541,477,503]" pageId="14" pageNumber="15">Juvenile colouration in life:</emphasis>
Dorsum of head almost black, with reddish brown blotches concentrated on anterolateral portion of prefrontals and temporal region; lower margins of supralabials beige; dorsal ground colour of body and tail dark reddish brown with small fragmented black dots (
<figureCitation id="1302D8883D62FFC5FBDCFD80FB67B813" box="[1066,1159,557,583]" captionStart="FIGURE 3" captionStartId="10.[151,255,1944,1968]" captionTargetBox="[160,1430,181,1936]" captionTargetId="figure-82@10.[151,1436,146,1952]" captionTargetPageId="10" captionText="FIGURE 3. General view in life of (A, B) Atractus caete sp. nov. from Chã Preta, state of Alagoas, photo by G. Skuk, (C) A. francoi sp. nov. from Santa Virgínia, state of São Paulo, photo by P. Hartmann, (D) A. guentheri from Ilhéus, state of Bahia, photo by A.J.S. Argôlo, and (E) A. pantostictus from Palmas, state of Tocantis, photo by O.A.V. Marques." figureDoi="http://doi.org/10.5281/zenodo.5315516" httpUri="https://zenodo.org/record/5315516/files/figure.png" pageId="14" pageNumber="15">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8B86C40D3D62FFC5FF30FDF8FF2CBEF4" blockId="14.[151,1436,152,1909]" pageId="14" pageNumber="15">
<emphasis id="B94D181F3D62FFC5FF30FDF8FCD0B83B" bold="true" box="[198,816,597,623]" pageId="14" pageNumber="15">
Hemipenis morphology (everted organs 
<emphasis id="B94D181F3D62FFC5FD22FDF8FD04B83B" bold="true" box="[724,740,597,623]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2):
</emphasis>
Retracted organ bifurcates at level of eighth and extends to level of ninth subcaudal. Everted hemipenis moderately bilobed, semicapitate, semicalyculate; lobes distinct and restricted to distal portion of capitulum; lobes sub-cylindrical, of similar size, with round apices; lobes slightly centrifugally oriented, with length similar to remaining capitulum; lobes and capitulum covered with small papillate calyces; vertical walls of calyces well defined, not arranged in distinct calyculate flounces on both sides of organ; asulcate side of capitulum with irregular calyces lacking lobular and medial crests; capitular groove indistinct on the sulcate side and well defined on the asulcate side of hemipenis; capitulum located just above sulcus spermaticus bifurcation and slightly smaller than hemipenial body; sulcus spermaticus bifurcates at about middle of organ; sulcus spermaticus branches centrifugally oriented, running to tip of lobes; margins of sulcus spermaticus stout and narrow, bordered with papillae from the base to the apices of lobes; hemipenial body sub-cylindrical, covered with moderate hooked spines on laterodistal region of the sulcate side of hemipenis; basal portion of the hemipenial body with small spines only, lacking moderate hooked spines; basal naked pocket restricted to basal portion of hemipenial body; proximal region of hemipenis with an evident constriction with uniformly scattered longitudinal plicae and diffuse spinules (
<figureCitation id="1302D8883D62FFC5FA9BFBF3FF5FBEF4" captionStart="FIGURE 6" captionStartId="15.[151,255,1793,1817]" captionTargetBox="[151,1436,189,1767]" captionTargetId="figure-65@15.[151,1436,189,1767]" captionTargetPageId="15" captionText="FIGURE 6. Sulcate (left) and asulcate (right) sides of the hemipenis of (A) Atractus francoi (IBSP 72654), (B) A. guentheri (MNRJ 6710), (C) A. maculatus (MNRJ 14197), (D) A. potschi (MZUEFS 682), (E) A. pantostictus (MNRJ 13195), and (F) A. paraguaguayensis (MNRJ 10123). Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315524" httpUri="https://zenodo.org/record/5315524/files/figure.png" pageId="14" pageNumber="15">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8B86C40D3D62FFC5FF30FB00FD09BFB1" blockId="14.[151,1436,152,1909]" pageId="14" pageNumber="15">
<emphasis id="B94D181F3D62FFC5FF30FB00FEA5BE93" bold="true" box="[198,325,1197,1223]" pageId="14" pageNumber="15">Variation:</emphasis>
Largest male 
<quantity id="4CC169E83D62FFC5FE06FB03FDB3BE9C" box="[496,595,1198,1224]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="14" pageNumber="15" unit="mm" value="420.0">420 mm</quantity>
SVL, 
<quantity id="4CC169E83D62FFC5FD55FB03FD17BE9C" box="[675,759,1198,1224]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="14" pageNumber="15" unit="mm" value="53.0">53 mm</quantity>
CL; largest female 
<quantity id="4CC169E83D62FFC5FC13FB03FBA7BE9C" box="[997,1095,1198,1224]" metricMagnitude="-1" metricUnit="m" metricValue="4.75" pageId="14" pageNumber="15" unit="mm" value="475.0">475 mm</quantity>
SVL, 
<quantity id="4CC169E83D62FFC5FB6FFB03FB0CBE9C" box="[1177,1260,1198,1224]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="mm" value="40.0">40 mm</quantity>
CL; tail 12.6– 13.6% (
<emphasis id="B94D181F3D62FFC5FF02FB7BFEE3BEBB" box="[244,259,1238,1263]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2) SVL in males, 6.3–8.6% (
<emphasis id="B94D181F3D62FFC5FD99FB7BFD9CBEBB" box="[623,636,1238,1263]" italics="true" pageId="14" pageNumber="15">x¯</emphasis>
= 7.6; SD = 3.5; 
<emphasis id="B94D181F3D62FFC5FCBFFB7BFCB8BEBB" box="[841,856,1238,1263]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 4) SVL in females; 152–154 (
<emphasis id="B94D181F3D62FFC5FB22FB7BFB03BEBB" box="[1236,1251,1238,1263]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2) ventrals in males, 162–164 (
<emphasis id="B94D181F3D62FFC5FE97FB53FE8DBF43" box="[353,365,1278,1303]" italics="true" pageId="14" pageNumber="15">x¯</emphasis>
= 161; SD = 1; 
<emphasis id="B94D181F3D62FFC5FDC2FB53FDA3BF43" box="[564,579,1278,1303]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 3) in females; 25–30 (
<emphasis id="B94D181F3D62FFC5FC90FB53FC93BF43" box="[870,883,1278,1303]" italics="true" pageId="14" pageNumber="15">x¯</emphasis>
= 27.2; SD = 2.1; 
<emphasis id="B94D181F3D62FFC5FBA1FB53FB86BF43" box="[1111,1126,1278,1303]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 3) subcaudals in males, 14–19 (
<emphasis id="B94D181F3D62FFC5FF07FA8BFF1EBF6B" box="[241,254,1318,1343]" italics="true" pageId="14" pageNumber="15">x¯</emphasis>
= 16.9; SD = 2.1; 
<emphasis id="B94D181F3D62FFC5FE2CFA8BFE09BF6B" box="[474,489,1318,1343]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 4) in females; 6 (
<emphasis id="B94D181F3D62FFC5FD26FA8BFD3FBF6B" box="[720,735,1318,1343]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 1 side) or 7 (
<emphasis id="B94D181F3D62FFC5FC7DFA8BFC7ABF6B" box="[907,922,1318,1343]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 13 sides) supralabials; 2 (
<emphasis id="B94D181F3D62FFC5FB21FA8BFB06BF6B" box="[1239,1254,1318,1343]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2 sides), 3 (
<emphasis id="B94D181F3D62FFC5FA7BFA8BFA7CBF6B" box="[1421,1436,1318,1343]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 8 sides), or 4 (
<emphasis id="B94D181F3D62FFC5FEA9FAE3FE8EBF33" box="[351,366,1358,1383]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 3 sides) gular scale rows; 2 (
<emphasis id="B94D181F3D62FFC5FD1EFAE3FD17BF33" box="[744,759,1358,1383]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 1) or 3 (
<emphasis id="B94D181F3D62FFC5FC8BFAE3FC6CBF33" box="[893,908,1358,1383]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 6) preventrals; 8 (
<emphasis id="B94D181F3D62FFC5FB72FAE3FB73BF33" box="[1156,1171,1358,1383]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 6 sides) or 9 (
<emphasis id="B94D181F3D62FFC5FAAAFAE3FA8BBF33" box="[1372,1387,1358,1383]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2 sides) maxillary teeth; 8–10 (
<emphasis id="B94D181F3D62FFC5FE06FADBFE1CBFDB" box="[496,508,1398,1423]" italics="true" pageId="14" pageNumber="15">x¯</emphasis>
= 8.9; SD = 0.7; 
<emphasis id="B94D181F3D62FFC5FD3BFADBFD3CBFDB" box="[717,732,1398,1423]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 14 sides) dorsal scale rows at the level of second subcaudal; 7.2–9.2 mm (
<emphasis id="B94D181F3D62FFC5FE90FA0CFE93BFEE" box="[358,371,1441,1466]" italics="true" pageId="14" pageNumber="15">x¯</emphasis>
= 8.6; SD = 0.9; 
<emphasis id="B94D181F3D62FFC5FDB4FA0CFDB1BFEE" box="[578,593,1441,1466]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 4) body diameter; hemipenis bifurcates at level of seventh to 10 
<superScript id="7C4C69453D62FFC5FAABFA30FA89BFFF" attach="left" box="[1373,1385,1437,1451]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
and extends to level of eighth to 11 
<superScript id="7C4C69453D62FFC5FDF7FA65FDEDBF82" attach="left" box="[513,525,1480,1494]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
(
<emphasis id="B94D181F3D62FFC5FDE8FA61FDCDBFB1" box="[542,557,1484,1509]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2) subcaudal.
</paragraph>
</subSubSection>
<subSubSection id="C32397863D62FFC5FF30FA5EFD2BBC09" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8B86C40D3D62FFC5FF30FA5EFD2BBC09" blockId="14.[151,1436,152,1909]" pageId="14" pageNumber="15">
<emphasis id="B94D181F3D62FFC5FF30FA5EFEB7BC59" bold="true" box="[198,343,1523,1549]" pageId="14" pageNumber="15">Etymology:</emphasis>
This species is dedicated to our friend Dr. Francisco Luís Franco, Curator of Herpetology at Instituto Butantan in 
<collectingCountry id="F32E849D3D62FFC5FE60F9B6FE00BC61" box="[406,480,1563,1589]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
, for his contribution to the systematics of South American snakes, and also for calling our attention to this undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="C32397863D62FFC5FF30F9C6FB90BC81" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8B86C40D3D62FFC5FF30F9C6FB90BC81" blockId="14.[151,1436,152,1909]" pageId="14" pageNumber="15">
<emphasis id="B94D181F3D62FFC5FF30F9C6FE89BCD1" bold="true" box="[198,361,1643,1669]" pageId="14" pageNumber="15">Distribution:</emphasis>
Highlands of the Serra da 
<taxonomicName id="4C39BF8E3D62FFC5FD58F9C6FCEEBCD1" authorityName="Piza" authorityYear="1943" box="[686,782,1643,1669]" class="Arachnida" family="Gonyleptidae" genus="Bocaina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Bocaina</taxonomicName>
and Serra dos Órgãos, in Southeastern 
<collectingCountry id="F32E849D3D62FFC5FB1FF9C6FAD0BCD1" box="[1257,1328,1643,1669]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
between the states of 
<collectingRegion id="49FD0AEF3D62FFC5FED9F93EFE3ABCF9" box="[303,474,1683,1709]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="15">Rio de Janeiro</collectingRegion>
and 
<collectingRegion id="49FD0AEF3D62FFC5FDE1F93EFD71BCF9" box="[535,657,1683,1709]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="15">São Paulo</collectingRegion>
, ranging from Nova Friburgo to São Luís do Paraitinga. 
<taxonomicName id="4C39BF8E3D62FFC5FACFF939FF0CBC81" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="francoi">
<emphasis id="B94D181F3D62FFC5FACFF939FF0CBC81" italics="true" pageId="14" pageNumber="15">Atractus francoi</emphasis>
</taxonomicName>
inhabits Lower Montane Rainforest between 
<quantity id="4CC169E83D62FFC5FCF2F916FC77BC81" box="[772,919,1723,1749]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="14" pageNumber="15" unit="m" value="800.0" valueMax="1000.0" valueMin="600.0">600–1000 m</quantity>
elevation (
<figureCitation id="1302D8883D62FFC5FBECF916FB82BC81" box="[1050,1122,1723,1749]" captionStart="FIGURE 7" captionStartId="16.[151,255,1228,1252]" captionTargetBox="[160,1429,159,1181]" captionTargetId="figure-256@16.[155,1432,131,1204]" captionTargetPageId="16" captionText="FIGURE 7. Geographical distribution of Atractus francoi, A. serranus, and A. trihedrurus." figureDoi="http://doi.org/10.5281/zenodo.5315528" httpUri="https://zenodo.org/record/5315528/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32397863D62FFC5FF30F94EFD7CBD21" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B86C40D3D62FFC5FF30F94EFD7CBD21" blockId="14.[151,1436,152,1909]" pageId="14" pageNumber="15">
<emphasis id="B94D181F3D62FFC5FF30F94EFEA1BCA9" bold="true" box="[198,321,1763,1789]" pageId="14" pageNumber="15">Remarks:</emphasis>
<bibRefCitation id="EFA8B9FC3D62FFC5FEBAF94EFDB2BCA9" author="Marques, O. A. V. &amp; Eterovic, A. &amp; Sazima, I." box="[332,594,1763,1789]" pageId="14" pageNumber="15" refId="ref39360" refString="Marques, O. A. V., Eterovic, A. &amp; Sazima, I. (2004) Snakes of the Brazilian Rainforest: An Illustrate Field Guide for the Serra do Mar. Holos, Ribeirao Preto, Brazil, 205 pp." type="book" year="2004">
<collectingRegion id="49FD0AEF3D62FFC5FEBAF94EFE53BCA9" box="[332,435,1763,1789]" country="Italy" name="Marche" pageId="14" pageNumber="15">Marques</collectingRegion>
<emphasis id="B94D181F3D62FFC5FE4AF949FE13BCA9" box="[444,499,1764,1789]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2004)
</bibRefCitation>
identified and illustrate a juvenile female of 
<taxonomicName id="4C39BF8E3D62FFC5FB81F949FB10BCA9" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" box="[1143,1264,1764,1789]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="francoi">
<emphasis id="B94D181F3D62FFC5FB81F949FB10BCA9" box="[1143,1264,1764,1789]" italics="true" pageId="14" pageNumber="15">A. francoi</emphasis>
</taxonomicName>
(IBSP 74723, actually 
<typeStatus id="54827AAF3D62FFC5FF0CF8A6FE81BD71" box="[250,353,1803,1829]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
) as possibly an undescribed species (= 
<taxonomicName id="4C39BF8E3D62FFC5FCDDF8A1FC58BD71" box="[811,952,1803,1829]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B94D181F3D62FFC5FCDDF8A1FC6EBD71" box="[811,910,1804,1829]" italics="true" pageId="14" pageNumber="15">Atractus</emphasis>
sp.
</taxonomicName>
). A female (MNRJ 
<quantity id="4CC169E83D62FFC5FB69F8A6FAB6BD71" box="[1183,1366,1803,1829]" metricMagnitude="1" metricUnit="m" metricValue="1.7536473" pageId="14" pageNumber="15" unit="mm" value="17536.473">17536, 473 mm</quantity>
SVL, CL 
<quantity id="4CC169E83D62FFC5FF30F89EFEFDBD19" box="[198,285,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="30.0">30 mm</quantity>
) layed 
<specimenCount id="9D3F0F843D62FFC5FE8EF89EFE3BBD19" box="[376,475,1843,1869]" pageId="14" pageNumber="15" type="egg">six eggs</specimenCount>
in captivity (R. Fernandes pers. observ.), which corroborates that females of 
<taxonomicName id="4C39BF8E3D62FFC5FA75F899FF0CBD21" authorityName="Passos &amp; Fernandes &amp; Bérnils &amp; De Moura-Leite" authorityYear="2010" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="francoi">
<emphasis id="B94D181F3D62FFC5FA75F899FF0CBD21" italics="true" pageId="14" pageNumber="15">A. francoi</emphasis>
</taxonomicName>
attain maturity below 
<quantity id="4CC169E83D62FFC5FE02F8F6FDB6BD21" box="[500,598,1883,1909]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="mm" value="500.0">500 mm</quantity>
SVL.
</paragraph>
</subSubSection>
</treatment>
</document>