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<document id="D1F167888976E113718C7ACD77078306" ID="10.11646/zootaxa.4144.1.6" ID-DOI="10.11646/zootaxa.4144.1.6" ID-GBIF-Dataset="1f1ca849-1a3e-4a79-bc65-01478389b4ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="264333" ID-ZooBank="C81927A3-6CC0-43CC-B6DE-E8ACF5D93F96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469515026979" checkinUser="plazi" docAuthor="Bekchiev, Rostislav &amp; Hlaváč, Peter" docDate="2016" docId="038E87B70D4D6921FF52C897FAECFBBF" docLanguage="en" docName="zootaxa.4144.1.6.pdf" docOrigin="Zootaxa 4144 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bryaxis biokovensis Bekchiev &amp; Hlaváč, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="113" masterDocId="FFB7FFCF0D496925FFC5CA40FF8AFD04" masterDocTitle="New and interesting species of the genus Bryaxis Kugelann, 1794 (Coleoptera, Staphylinidae, Pselaphinae) from Croatia" masterLastPageNumber="116" masterPageNumber="109" pageNumber="113" updateTime="1698366187038" updateUser="plazi">
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<mods:title id="70C1ABE62E822E6A28829C25EEEBBB31">New and interesting species of the genus Bryaxis Kugelann, 1794 (Coleoptera, Staphylinidae, Pselaphinae) from Croatia</mods:title>
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<mods:name id="DA4DFD7FA40EB52A28BD72D69EDBDBC0" type="personal">
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<mods:namePart id="4F87036A2587BAC1041EDCA0598C692F">Bekchiev, Rostislav</mods:namePart>
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<mods:namePart id="B24F7C79B90165DF1B642DB08AC4662D">Hlaváč, Peter</mods:namePart>
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<mods:title id="7A1064A411963EA099A2C8EADEF761F7">Zootaxa</mods:title>
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<mods:date id="C3F48E93239B90C2BE8642D3331F3E93">2016</mods:date>
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<mods:number id="2C8D449B31C156C3D4047E42203BFF18">4144</mods:number>
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<treatment id="038E87B70D4D6921FF52C897FAECFBBF" ID-DOI="http://doi.org/10.5281/zenodo.6081989" ID-GBIF-Taxon="123207641" ID-Zenodo-Dep="6081989" LSID="urn:lsid:plazi:treatment:038E87B70D4D6921FF52C897FAECFBBF" httpUri="http://treatment.plazi.org/id/038E87B70D4D6921FF52C897FAECFBBF" lastPageNumber="113" pageId="4" pageNumber="113">
<subSubSection id="C33D652A0D4D6921FF52C897FF61FE17" pageId="4" pageNumber="113" type="nomenclature">
<paragraph id="8B9836A10D4D6921FF52C897FE66FFF6" blockId="4.[151,492,727,787]" box="[151,492,727,754]" pageId="4" pageNumber="113">
<heading id="D0D081CD0D4D6921FF52C897FE66FFF6" bold="true" box="[151,492,727,754]" fontSize="11" level="1" pageId="4" pageNumber="113" reason="1">
<emphasis id="B953EAB30D4D6921FF52C897FE66FFF6" bold="true" box="[151,492,727,754]" pageId="4" pageNumber="113">
<taxonomicName id="4C274D220D4D6921FF52C897FE0DFFF5" ID-CoL="68T42" box="[151,391,727,753]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="species" species="biokovensis" status="sp. nov.">
<emphasis id="B953EAB30D4D6921FF52C897FE0DFFF5" bold="true" box="[151,391,727,753]" italics="true" pageId="4" pageNumber="113">Bryaxis biokovensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057C80D4D6921FE4BC898FE66FFF6" box="[398,492,728,754]" pageId="4" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9836A10D4D6921FF52C8BAFF61FE17" blockId="4.[151,492,727,787]" box="[151,235,762,787]" pageId="4" pageNumber="113">
(
<figureCitation id="131C2A240D4D6921FF5AC8BAFF68FE17" box="[159,226,762,787]" captionStart="FIGURE 3" captionStartId="6.[151,250,1971,1993]" captionTargetBox="[240,1358,206,1929]" captionTargetId="figure@6.[222,1368,193,1940]" captionTargetPageId="6" captionText="FIGURE 3. Bryaxis biokovensis sp. nov. a. Habitus, male (scale bar— 1 mm). b. scape and pedicel, dorsal view (scale bar— 0.1 mm). c, d. Aedeagus, ventral (c) and lateral (d) view (scale bar— 0.2 mm)." httpUri="https://zenodo.org/record/264336/files/figure.png" pageId="4" pageNumber="113">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33D652A0D4D6921FF52C902FD40FE7B" pageId="4" pageNumber="113" type="materials_examined">
<paragraph id="8B9836A10D4D6921FF52C902FD40FE7B" blockId="4.[151,1437,834,1723]" pageId="4" pageNumber="113">
<materialsCitation id="3B4F3CFC0D4D6921FF52C902FD4DFE7B" ID-GBIF-Occurrence="1288049019" collectingDate="2014-11-10" collectionCode="PCPH" collectorName="P. Hlavac" country="Croatia" county="Ledenica" location="Material" municipality="Biokovo" pageId="4" pageNumber="113" specimenCount="1" typeStatus="holotype">
<emphasis id="B953EAB30D4D6921FF52C902FDF8FE5F" bold="true" box="[151,626,834,859]" pageId="4" pageNumber="113">
<location id="8EF8607A0D4D6921FF52C902FF75FE5F" LSID="urn:lsid:plazi:treatment:038E87B70D4D6921FF52C897FAECFBBF:8EF8607A0D4D6921FF52C902FF75FE5F" box="[151,255,834,859]" country="Croatia" county="Ledenica" municipality="Biokovo" name="Material" pageId="4" pageNumber="113">Material</location>
examined.
<typeStatus id="549C88030D4D6921FE48C902FE74FE5F" box="[397,510,834,859]" pageId="4" pageNumber="113" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F33076310D4D6921FDC8C902FDE6FE5F" box="[525,620,834,859]" name="Croatia" pageId="4" pageNumber="113">Croatia</collectingCountry>
:
</emphasis>
1 Ƌ:
<collectingCountry id="F33076310D4D6921FD7FC902FC85FE5F" box="[698,783,834,859]" name="Croatia" pageId="4" pageNumber="113">Croatia</collectingCountry>
,
<collectingMunicipality id="6BFCACDB0D4D6921FCD8C902FC0AFE5F" box="[797,896,834,859]" pageId="4" pageNumber="113">Biokovo</collectingMunicipality>
, near špilja
<collectingCounty id="62F94E2D0D4D6921FBD5C902FBF0FE5F" box="[1040,1146,834,859]" pageId="4" pageNumber="113">Ledenica</collectingCounty>
, sifting in forest close to cave entrance,
<collectingDate id="EFDDE9890D4D6921FEFBC927FE34FE7B" box="[318,446,871,895]" pageId="4" pageNumber="113" value="2014-11-10">
<date id="FF9910610D4D6921FEFBC927FE34FE7B" box="[318,446,871,895]" pageId="4" pageNumber="113" value="2014-11-10">11.10.2014</date>
</collectingDate>
, leg.
<collectorName id="26D253770D4D6921FE3CC927FDE9FE7B" box="[505,611,870,895]" pageId="4" pageNumber="113">P. Hlaváč</collectorName>
(
<collectionCode id="ED36AE640D4D6921FDB7C927FD35FE7B" box="[626,703,871,895]" pageId="4" pageNumber="113">PCPH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D652A0D4D6921FF02C9CAFD18FB2F" pageId="4" pageNumber="113" type="description">
<paragraph id="8B9836A10D4D6921FF02C9CAFD4EFB03" blockId="4.[151,1437,834,1723]" pageId="4" pageNumber="113">
<emphasis id="B953EAB30D4D6921FF02C9CAFED3FEA7" bold="true" box="[199,345,906,931]" pageId="4" pageNumber="113">Description.</emphasis>
Body (
<figureCitation id="131C2A240D4D6921FE77C9CAFE7DFEA7" box="[434,503,906,931]" captionStart="FIGURE 3" captionStartId="6.[151,250,1971,1993]" captionTargetBox="[240,1358,206,1929]" captionTargetId="figure@6.[222,1368,193,1940]" captionTargetPageId="6" captionText="FIGURE 3. Bryaxis biokovensis sp. nov. a. Habitus, male (scale bar— 1 mm). b. scape and pedicel, dorsal view (scale bar— 0.1 mm). c, d. Aedeagus, ventral (c) and lateral (d) view (scale bar— 0.2 mm)." httpUri="https://zenodo.org/record/264336/files/figure.png" pageId="4" pageNumber="113">Fig. 3</figureCitation>
a) pale reddish-brown, antennae, maxillary palpi and legs slightly lighter, length
<quantity id="4CDF9B440D4D6921FF52C9EEFF77FEC2" box="[151,253,942,967]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="4" pageNumber="113" unit="mm" value="1.56">1.56 mm</quantity>
, max width
<quantity id="4CDF9B440D4D6921FE4DC9EEFE65FEC2" box="[392,495,942,967]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="113" unit="mm" value="0.62">0.62 mm</quantity>
. Pubescence on head, pronotum and elytra gold, short, dense, and suberected. Head 0.8 times as long as wide. Frons with, shallow, triangular impression between antennal tubercles; rostrum oval anteriorly. Vertex flat, with thin median longitudinal carina, reaching frontal triangular impression; frontal fovea absent, vertexal foveae very small, their diameter smaller than the distance between them. Eyes with medium size, with about 15 facets each. Maxillary palpi normally developed, with small, sparse tubercles on external part of apex of palpomeres II and III, terminal palpomere
<quantity id="4CDF9B440D4D6921FD1CCE23FCB4F97E" box="[729,830,1123,1147]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="113" unit="mm" value="0.2">0.20 mm</quantity>
long, about 0.33 times as long as wide. Gular region flat, without modifications. Antennae about
<quantity id="4CDF9B440D4D6921FD51CEC6FD71F99A" box="[660,763,1158,1183]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="4" pageNumber="113" unit="mm" value="0.63">0.63 mm</quantity>
long, scape (
<figureCitation id="131C2A240D4D6921FC52CEC6FC51F99B" box="[919,987,1158,1183]" captionStart="FIGURE 3" captionStartId="6.[151,250,1971,1993]" captionTargetBox="[240,1358,206,1929]" captionTargetId="figure@6.[222,1368,193,1940]" captionTargetPageId="6" captionText="FIGURE 3. Bryaxis biokovensis sp. nov. a. Habitus, male (scale bar— 1 mm). b. scape and pedicel, dorsal view (scale bar— 0.1 mm). c, d. Aedeagus, ventral (c) and lateral (d) view (scale bar— 0.2 mm)." httpUri="https://zenodo.org/record/264336/files/figure.png" pageId="4" pageNumber="113">Fig. 3</figureCitation>
b) with very small, flat tubercle on the apex, 2.402.80 times as long as pedicel; pedicel simple (
<figureCitation id="131C2A240D4D6921FCDFCEEAFCD6F9C7" box="[794,860,1194,1219]" captionStart="FIGURE 3" captionStartId="6.[151,250,1971,1993]" captionTargetBox="[240,1358,206,1929]" captionTargetId="figure@6.[222,1368,193,1940]" captionTargetPageId="6" captionText="FIGURE 3. Bryaxis biokovensis sp. nov. a. Habitus, male (scale bar— 1 mm). b. scape and pedicel, dorsal view (scale bar— 0.1 mm). c, d. Aedeagus, ventral (c) and lateral (d) view (scale bar— 0.2 mm)." httpUri="https://zenodo.org/record/264336/files/figure.png" pageId="4" pageNumber="113">Fig. 3</figureCitation>
b), 1.5 times as long as wide; antennomeres IIIVII elongate, III 1.6 times as long as IV, IVV subequal in length, slightly longer than subequal antennomeres VI and VII, VIII smallest, as long as wide, IX quadrate, about 1.25 times as wide as VIII, X 0.7 times as long as wide and 1.25 times as long as IX, terminal antennomere largest, fusiform, about twice as long as wide. Pronotum 0.9 as long as wide and 1.35 times as long as head, widest in anterior third, with thin antebasal sulcus connecting large lateral foveae, median fovea absent. Elytra 1.06 times as wide as long, slightly convex, with two basal foveae and entire sutural striae, discal striae absent. Legs slender, tibia I with shalow preapical indentation on inner margin, II simple, III with small internal apical spur. Aedeagus (
<figureCitation id="131C2A240D4D6921FD77CFE6FD7CF8BB" box="[690,758,1446,1471]" captionStart="FIGURE 3" captionStartId="6.[151,250,1971,1993]" captionTargetBox="[240,1358,206,1929]" captionTargetId="figure@6.[222,1368,193,1940]" captionTargetPageId="6" captionText="FIGURE 3. Bryaxis biokovensis sp. nov. a. Habitus, male (scale bar— 1 mm). b. scape and pedicel, dorsal view (scale bar— 0.1 mm). c, d. Aedeagus, ventral (c) and lateral (d) view (scale bar— 0.2 mm)." httpUri="https://zenodo.org/record/264336/files/figure.png" pageId="4" pageNumber="113">Fig. 3</figureCitation>
cd) elongate,
<quantity id="4CDF9B440D4D6921FC60CFE7FB87F8BA" box="[933,1037,1447,1471]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="113" unit="mm" value="0.39">0.39 mm</quantity>
long, 2,80 times as long as wide, strongly narrowed to apex of apical lobe, with three short preapical setae, endophallus with two thin, separated, equal, apically sharply pointed sclerotized pieces.
</paragraph>
<paragraph id="8B9836A10D4D6921FF02CC52FD18FB2F" blockId="4.[151,1437,834,1723]" box="[199,658,1554,1579]" pageId="4" pageNumber="113">
<emphasis id="B953EAB30D4D6921FF02CC52FE3DFB2F" bold="true" box="[199,439,1554,1579]" pageId="4" pageNumber="113">Sexual dimorphism.</emphasis>
Females unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33D652A0D4D6921FF02CC76FB9EFB77" pageId="4" pageNumber="113" type="diagnosis">
<paragraph id="8B9836A10D4D6921FF02CC76FB9EFB77" blockId="4.[151,1437,834,1723]" pageId="4" pageNumber="113">
<emphasis id="B953EAB30D4D6921FF02CC76FE45FB4B" bold="true" box="[199,463,1590,1615]" pageId="4" pageNumber="113">Differential diagnosis.</emphasis>
<taxonomicName id="4C274D220D4D6921FE13CC77FD3DFB4B" box="[470,695,1590,1615]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis id="B953EAB30D4D6921FE13CC77FD3DFB4B" box="[470,695,1590,1615]" italics="true" pageId="4" pageNumber="113">Bryaxis biokovensis</emphasis>
</taxonomicName>
is very distinctive in the shape of the scape, gular region and the aedeagus, and at present cannot be clearly assigned to any other known species.
</paragraph>
</subSubSection>
<subSubSection id="C33D652A0D4D6921FF02CC3EFB3EFB93" box="[199,1204,1662,1687]" pageId="4" pageNumber="113" type="biology_ecology">
<paragraph id="8B9836A10D4D6921FF02CC3EFB3EFB93" blockId="4.[151,1437,834,1723]" box="[199,1204,1662,1687]" pageId="4" pageNumber="113">
<emphasis id="B953EAB30D4D6921FF02CC3EFEADFB93" bold="true" box="[199,295,1662,1687]" pageId="4" pageNumber="113">Biology.</emphasis>
Found from leaf litter samples in forest litter near the entrance of cave Ledenica.
</paragraph>
</subSubSection>
<subSubSection id="C33D652A0D4D6921FF02CCE2FAECFBBF" box="[199,1382,1698,1723]" pageId="4" pageNumber="113" type="etymology">
<paragraph id="8B9836A10D4D6921FF02CCE2FAECFBBF" blockId="4.[151,1437,834,1723]" box="[199,1382,1698,1723]" pageId="4" pageNumber="113">
<emphasis id="B953EAB30D4D6921FF02CCE2FEC4FBBF" bold="true" box="[199,334,1698,1723]" pageId="4" pageNumber="113">Etymology.</emphasis>
The specific name is derived from the Biokovo Mountain, where the species was discovered.
</paragraph>
</subSubSection>
</treatment>
</document>