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<document id="3B4B68126E103B4201BD5019B24AC8DA" ID-CLB-Dataset="30367" ID-DOI="10.5852/ejt.2018.410" ID-GBIF-Dataset="6ec71c9e-1dcc-4135-a1e3-7389872abdcd" ID-Zenodo-Dep="1211198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1522682334120" checkinUser="plazi" docAuthor="Skelton, Paul H., Swartz, Ernst R. &amp; Vreven, Emmanuel J." docDate="2018" docId="038DB87EFFC8FFC07A57FA87FBF527DE" docLanguage="en" docName="ejt-410_skelton_swartz_vreven.pdf.imf" docOrigin="European Journal of Taxonomy 410" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pseudobarbus Smith 1841" docType="treatment" docVersion="13" lastPageNumber="17" masterDocId="FFB4C006FFC7FFD07807FFB6FFF8227E" masterDocTitle="The identity of Barbus capensis Smith, 1841 and the generic status of southern African tetraploid cyprinids (Teleostei, Cyprinidae)" masterLastPageNumber="29" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547732810861" originalUpdateUser="yanell" pageNumber="16" updateTime="1698535861530" updateUser="ExternalLinkService">
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<mods:title id="83D03276C142350AE8B2EA97E216FFEA">The identity of Barbus capensis Smith, 1841 and the generic status of southern African tetraploid cyprinids (Teleostei, Cyprinidae)</mods:title>
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<mods:namePart id="553BD79E89998A50023C621A5267ECFE">Swartz, Ernst R.</mods:namePart>
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<treatment id="038DB87EFFC8FFC07A57FA87FBF527DE" ID-DOI="http://doi.org/10.5281/zenodo.5687605" ID-GBIF-Taxon="143448362" ID-Zenodo-Dep="5687605" LSID="urn:lsid:plazi:treatment:038DB87EFFC8FFC07A57FA87FBF527DE" httpUri="http://treatment.plazi.org/id/038DB87EFFC8FFC07A57FA87FBF527DE" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C33E5AE3FFC8FFDF7A57FA87FC1C2735" box="[592,996,1328,1355]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B9B0968FFC8FFDF7A57FA87FC1C2735" blockId="15.[592,996,1328,1355]" box="[592,996,1328,1355]" pageId="15" pageNumber="16">
<heading id="D0D3BE04FFC8FFDF7A57FA87FC1C2735" box="[592,996,1328,1355]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
Genus
<taxonomicName id="4C2472EBFFC8FFDF7AA5FA86FC1C2735" ID-CoL="6Z9R" ID-ENA="100124" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[674,996,1328,1355]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFC8FFDF7AA5FA86FCB52735" bold="true" box="[674,845,1328,1355]" italics="true" pageId="15" pageNumber="16">Pseudobarbus</emphasis>
<bibRefCitation id="EFB57499FFC8FFDF7B53FA86FC1C2735" author="Smith A." box="[852,996,1328,1355]" journalOrPublisher="London" pageId="15" pageNumber="16" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
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<subSubSection id="C33E5AE3FFC8FFDF78BAFAC4FB1227B1" pageId="15" pageNumber="16" type="synonymic_list">
<paragraph id="8B9B0968FFC8FFDF78BAFAC4FAEE27F2" blockId="15.[189,1302,1394,1421]" box="[189,1302,1394,1421]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB342E46FFC8FFDF78BAFAC4FAEE27F2" box="[189,1302,1394,1421]" pageId="15" pageNumber="16">
<taxonomicName id="4C2472EBFFC8FFDF78BAFAC4FE1D27F3" ID-CoL="382L" authority="Daudin, 1805: 58" authorityName="Daudin" authorityPageNumber="58" authorityYear="1805" box="[189,485,1394,1421]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFC8FFDF78BAFAC4FEEA27F2" box="[189,274,1394,1420]" italics="true" pageId="15" pageNumber="16">Barbus</emphasis>
<treatmentCitation id="0A852F79FFC8FFDF791EFAC4FE1D27F3" author="" box="[281,485,1394,1421]" page="58" pageId="15" pageNumber="16">
<bibRefCitation id="EFB57499FFC8FFDF791EFAC4FE4527F3" author="Daudin F. M." box="[281,445,1394,1421]" editor="Cuvier F." journalOrPublisher="Serie" pageId="15" pageNumber="16" pagination="58" part="1" refId="ref12911" refString="Daudin F. M. 1805. Barbus. In: Cuvier F. (ed.) Dictionnaire des Sciences naturelles. BAN - BLU. Serie 1. Tome quatrieme: 58. Levrault, Schoell &amp; Cie, Paris." title="Barbus" type="journal article" volumeTitle="Dictionnaire des Sciences naturelles. BAN - BLU" year="1805">Daudin, 1805</bibRefCitation>
: 58
</treatmentCitation>
</taxonomicName>
(in part, non-Cuvier:
<taxonomicName id="4C2472EBFFC8FFDF7AE1FAC4FAF127F2" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[742,1289,1394,1420]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="burchelli" subGenus="Pseudobarbus">
<emphasis id="B950D57AFFC8FFDF7AE1FAC4FB9627F2" box="[742,1134,1394,1420]" italics="true" pageId="15" pageNumber="16">Barbus (Pseudobarbus) burchelli</emphasis>
<bibRefCitation id="EFB57499FFC8FFDF7C72FAC4FAF127F2" author="Smith A." box="[1141,1289,1394,1420]" journalOrPublisher="London" pageId="15" pageNumber="16" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B0968FFC8FFDF78BAFA02FB1227B1" blockId="15.[189,1258,1460,1487]" box="[189,1258,1460,1487]" pageId="15" pageNumber="16">
<taxonomicName id="4C2472EBFFC8FFDF78BAFA02FE9D27B0" authorityName="Smith" authorityYear="1841" box="[189,357,1460,1486]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFC8FFDF78BAFA02FE9D27B0" box="[189,357,1460,1486]" italics="true" pageId="15" pageNumber="16">Pseudobarbus</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFB57499FFC8FFDF7985FA02FDE627B0" author="Skelton P. H." box="[386,542,1460,1486]" journalOrPublisher="Annals of the Cape Provincial Museums (Natural History)" pageId="15" pageNumber="16" pagination="201 - 307" part="1 6" refId="ref14443" refString="Skelton P. H. 1988. A taxonomic revision of the redfin minnows (Pisces, Cyprinidae) from southern Africa. Annals of the Cape Provincial Museums (Natural History) 1 6 (10): 201 - 307." title="A taxonomic revision of the redfin minnows (Pisces, Cyprinidae) from southern Africa" type="journal article" year="1988">Skelton 1988</bibRefCitation>
: 263 (raised the subgenus of
<bibRefCitation id="EFB57499FFC8FFDF7B70FA02FBEB27B0" author="Smith A." box="[887,1043,1460,1486]" journalOrPublisher="London" pageId="15" pageNumber="16" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith (1841)</bibRefCitation>
to generic status).
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFC8FFDF78BAFA40FAEB254F" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8B9B0968FFC8FFDF78BAFA40FEA0246E" blockId="15.[189,344,1526,1552]" box="[189,344,1526,1552]" pageId="15" pageNumber="16">
<heading id="D0D3BE04FFC8FFDF78BAFA40FEA0246E" bold="true" box="[189,344,1526,1552]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B950D57AFFC8FFDF78BAFA40FEA0246E" bold="true" box="[189,344,1526,1552]" pageId="15" pageNumber="16">Type species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFC8FFDF78BAF993FD8B2441" blockId="15.[189,627,1573,1599]" box="[189,627,1573,1599]" pageId="15" pageNumber="16">
<taxonomicName id="4C2472EBFFC8FFDF78BAF993FD972441" ID-CoL="4NPJ8" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[189,623,1573,1599]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="burchelli">
<emphasis id="B950D57AFFC8FFDF78BAF993FE2D2441" box="[189,469,1573,1599]" italics="true" pageId="15" pageNumber="16">Pseudobarbus burchelli</emphasis>
<bibRefCitation id="EFB57499FFC8FFDF79DBF993FD972441" author="Smith A." box="[476,623,1573,1599]" journalOrPublisher="London" pageId="15" pageNumber="16" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9B0968FFC8FFDF78BAF9D2FE712400" blockId="15.[189,393,1636,1662]" box="[189,393,1636,1662]" pageId="15" pageNumber="16">
<heading id="D0D3BE04FFC8FFDF78BAF9D2FE712400" bold="true" box="[189,393,1636,1662]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B950D57AFFC8FFDF78BAF9D2FE712400" bold="true" box="[189,393,1636,1662]" pageId="15" pageNumber="16">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFC8FFDF78BAF926FAEB254F" blockId="15.[189,1399,1680,1841]" pageId="15" pageNumber="16">
<taxonomicName id="4C2472EBFFC8FFDF78BAF926FD8724D4" ID-CoL="4NPJ8" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[189,639,1680,1706]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="burchelli">
<emphasis id="B950D57AFFC8FFDF78BAF926FE2324D4" box="[189,475,1680,1706]" italics="true" pageId="15" pageNumber="16">Pseudobarbus burchelli</emphasis>
<bibRefCitation id="EFB57499FFC8FFDF79E0F926FD8724D4" author="Smith A." box="[487,639,1680,1706]" journalOrPublisher="London" pageId="15" pageNumber="16" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF7A96F926FBD324D5" ID-CoL="4NPJ5" authority="(Peters, 1864)" baseAuthorityName="Peters" baseAuthorityYear="1864" box="[657,1067,1680,1707]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="afer">
<emphasis id="B950D57AFFC8FFDF7A96F926FC8D24D4" box="[657,885,1680,1706]" italics="true" pageId="15" pageNumber="16">Pseudobarbus afer</emphasis>
(Peters, 1864)
</taxonomicName>
(
<figureCitation id="131F15EDFFC8FFDF7C46F926FB4C24D4" box="[1089,1204,1680,1706]" captionStart="Fig. 10" captionStartId="17.[189,232,1860,1886]" captionTargetBox="[189,1399,970,1817]" captionTargetId="figure@17.[189,1399,967,1818]" captionTargetPageId="17" captionText="Fig. 10. Species representative of southernAfrican tetraploid cyprinine genera. A. Pseudobarbus afer (Peters, 1864). B. Cheilobarbus capensis Smith, 1841. C. Amatolacypris trevelyani (Günther, 1877) gen.et comb.nov. D. Sedercypris calidus(Barnard, 1938) gen.et comb.nov. E. Namaquacyprishospes (Barnard, 1938) gen. et comb. nov. Photograph credits:A–D by P.H. Skelton (SAIAB); E by R.I. Bills (SAIAB)." httpUri="https://zenodo.org/record/1211218/files/figure.png" pageId="15" pageNumber="16">Fig. 10A</figureCitation>
);
<taxonomicName id="4C2472EBFFC8FFDF7CC8F926FE1324B5" ID-CoL="4NPJ7" authority="(Boulenger, 1911)" baseAuthorityName="Boulenger" baseAuthorityYear="1911" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="asper">
<emphasis id="B950D57AFFC8FFDF7CC8F926FF0624B5" italics="true" pageId="15" pageNumber="16">Pseudobarbus asper</emphasis>
(
<bibRefCitation id="EFB57499FFC8FFDF7913F907FE1A24B5" author="Boulenger G. A." box="[276,482,1713,1739]" journalOrPublisher="Trustees of the British Museum of Natural History, London" pageId="15" pageNumber="16" publicationUrl="https://doi.org/10.5962/bhl.title.8869" refId="ref12730" refString="Boulenger G. A. 1911. Catalogue of the Fresh-water Fishes of Africa in the British Museum (Natural History). Volume 2. Trustees of the British Museum of Natural History, London. https: // doi. org / 10.5962 / bhl. title. 8869" title="Catalogue of the Fresh-water Fishes of Africa in the British Museum (Natural History). Volume 2" type="book" year="1911">Boulenger, 1911</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF79FAF907FC2624B5" ID-CoL="4NPJ9" authority="(Boulenger, 1911)" baseAuthorityName="Boulenger" baseAuthorityYear="1911" box="[509,990,1713,1739]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="burgi">
<emphasis id="B950D57AFFC8FFDF79FAF907FD0924B5" box="[509,753,1713,1739]" italics="true" pageId="15" pageNumber="16">Pseudobarbus burgi</emphasis>
(
<bibRefCitation id="EFB57499FFC8FFDF7B00F907FC2D24B5" author="Boulenger G. A." box="[775,981,1713,1739]" journalOrPublisher="Trustees of the British Museum of Natural History, London" pageId="15" pageNumber="16" publicationUrl="https://doi.org/10.5962/bhl.title.8869" refId="ref12730" refString="Boulenger G. A. 1911. Catalogue of the Fresh-water Fishes of Africa in the British Museum (Natural History). Volume 2. Trustees of the British Museum of Natural History, London. https: // doi. org / 10.5962 / bhl. title. 8869" title="Catalogue of the Fresh-water Fishes of Africa in the British Museum (Natural History). Volume 2" type="book" year="1911">Boulenger, 1911</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF7BF7F907FEFD2493" ID-CoL="4NPJH" authority="(Smith, 1936)" baseAuthorityName="Smith" baseAuthorityYear="1936" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="senticeps">
<emphasis id="B950D57AFFC8FFDF7BF7F907FAEA24B5" box="[1008,1298,1713,1739]" italics="true" pageId="15" pageNumber="16">Pseudobarbus senticeps</emphasis>
(Smith, 1936)
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF791CF964FCC82492" ID-CoL="4NPJF" authority="(Barnard, 1938)" baseAuthorityName="Barnard" baseAuthorityYear="1938" box="[283,816,1746,1772]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="phlegethon">
<emphasis id="B950D57AFFC8FFDF791CF964FDAE2492" box="[283,598,1746,1772]" italics="true" pageId="15" pageNumber="16">Pseudobarbus phlegethon</emphasis>
(
<bibRefCitation id="EFB57499FFC8FFDF7A68F964FCDF2492" author="Barnard K. H." box="[623,807,1746,1772]" journalOrPublisher="Annals and Magazine of Natural History Ser. 11" pageId="15" pageNumber="16" pagination="80 - 88" part="2" refId="ref12509" refString="Barnard K. H. 1938. Notes on the species of Barbus from the Cape Province, with descriptions of new species. Annals and Magazine of Natural History Ser. 11, 2: 80 - 88." title="Notes on the species of Barbus from the Cape Province, with descriptions of new species" type="journal article" year="1938">Barnard, 1938</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF7B41F964FA8A2492" ID-CoL="4NPJG" authority="(Barnard, 1938)" baseAuthorityName="Barnard" baseAuthorityYear="1938" box="[838,1394,1746,1772]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="quathlambae">
<emphasis id="B950D57AFFC8FFDF7B41F964FB602492" box="[838,1176,1746,1772]" italics="true" pageId="15" pageNumber="16">Pseudobarbus quathlambae</emphasis>
(
<bibRefCitation id="EFB57499FFC8FFDF7CB6F964FA912492" author="Barnard K. H." box="[1201,1385,1746,1772]" journalOrPublisher="Annals and Magazine of Natural History Ser. 11" pageId="15" pageNumber="16" pagination="80 - 88" part="2" refId="ref12509" refString="Barnard K. H. 1938. Notes on the species of Barbus from the Cape Province, with descriptions of new species. Annals and Magazine of Natural History Ser. 11, 2: 80 - 88." title="Notes on the species of Barbus from the Cape Province, with descriptions of new species" type="journal article" year="1938">Barnard, 1938</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF78BAF945FD802573" ID-CoL="4NPJM" authority="(Barnard, 1938)" baseAuthorityName="Barnard" baseAuthorityYear="1938" box="[189,632,1779,1805]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="tenuis">
<emphasis id="B950D57AFFC8FFDF78BAF945FE4A2573" box="[189,434,1779,1805]" italics="true" pageId="15" pageNumber="16">Pseudobarbus tenuis</emphasis>
(
<bibRefCitation id="EFB57499FFC8FFDF79C6F945FD972573" author="Barnard K. H." box="[449,623,1779,1805]" journalOrPublisher="Annals and Magazine of Natural History Ser. 11" pageId="15" pageNumber="16" pagination="80 - 88" part="2" refId="ref12509" refString="Barnard K. H. 1938. Notes on the species of Barbus from the Cape Province, with descriptions of new species. Annals and Magazine of Natural History Ser. 11, 2: 80 - 88." title="Notes on the species of Barbus from the Cape Province, with descriptions of new species" type="journal article" year="1938">Barnard, 1938</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF7A83F945FB462573" ID-CoL="4NPJK" authority="Chakona &amp; Swartz, 2013" authorityName="Chakona &amp; Swartz" authorityYear="2013" box="[644,1214,1779,1805]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="skeltoni">
<emphasis id="B950D57AFFC8FFDF7A83F945FC762573" box="[644,910,1779,1805]" italics="true" pageId="15" pageNumber="16">Pseudobarbus skeltoni</emphasis>
Chakona &amp; Swartz, 2013
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF7CC8F945FD3B254F" ID-CoL="4NPJP" authority="Chakona, Swartz &amp; Skelton, 2014" authorityName="Chakona, Swartz &amp; Skelton" authorityYear="2014" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="verloreni">
<emphasis id="B950D57AFFC8FFDF7CC8F945FEDF254F" italics="true" pageId="15" pageNumber="16">Pseudobarbus verloreni</emphasis>
Chakona, Swartz &amp; Skelton, 2014
</taxonomicName>
;
<taxonomicName id="4C2472EBFFC8FFDF7AC8F8A1FAF6254F" ID-CoL="4NPJL" authority="Chakona &amp; Skelton, 2017" authorityName="Chakona &amp; Skelton" authorityYear="2017" box="[719,1294,1815,1841]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="swartzi">
<emphasis id="B950D57AFFC8FFDF7AC8F8A1FC2B254F" box="[719,979,1815,1841]" italics="true" pageId="15" pageNumber="16">Pseudobarbus swartzi</emphasis>
Chakona &amp; Skelton, 2017
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFC8FFC078BAF8EFFE842168" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B9B0968FFC8FFDF78BAF8EFFECE250D" blockId="15.[189,310,1881,1907]" box="[189,310,1881,1907]" pageId="15" pageNumber="16">
<heading id="D0D3BE04FFC8FFDF78BAF8EFFECE250D" bold="true" box="[189,310,1881,1907]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B950D57AFFC8FFDF78BAF8EFFECE250D" bold="true" box="[189,310,1881,1907]" pageId="15" pageNumber="16">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFC8FFC078BAF83FFE842168" blockId="15.[189,1398,1928,2027]" lastBlockId="16.[189,1399,267,790]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<taxonomicName id="4C2472EBFFC8FFDF78BAF83FFE9D25DD" authorityName="Smith" authorityYear="1841" box="[189,357,1929,1955]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFC8FFDF78BAF83FFE9D25DD" box="[189,357,1929,1955]" italics="true" pageId="15" pageNumber="16">Pseudobarbus</emphasis>
</taxonomicName>
is distinct from all other southern African tetraploid cyprinine genera in having a flexible, non-serrated dorsal-fin unbranched ray (vs serrated), in adults sexual dimorphism expressed in nuptial tubercle development and fin size and shape (
<taxonomicName id="4C2472EBFFC8FFDF7ACCF867FC8B2595" authorityName="Smith" authorityYear="1841" box="[715,883,2001,2027]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFC8FFDF7ACCF867FC8B2595" box="[715,883,2001,2027]" italics="true" pageId="15" pageNumber="16">Pseudobarbus</emphasis>
</taxonomicName>
males develop conical tubercles on the head in a distinct pattern, on the scales in a row along the free edge, and in bands on the pectoral fins vs adults of both sexes having small erupted tubercles scattered over the head and scales; males of
<taxonomicName id="4C2472EBFFD7FFC07CC8FE99FA8F2337" authorityName="Smith" authorityYear="1841" box="[1231,1399,303,329]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC07CC8FE99FA8F2337" box="[1231,1399,303,329]" italics="true" pageId="16" pageNumber="17">Pseudobarbus</emphasis>
</taxonomicName>
have longer and more expansive fins than females vs similar finnage in both sexes), a reduced or absent mandibular lateral line canal (vs normally developed mandibular canal), and in having weakly ossified characteristics of the skeleton, especially the supraneural and intra-muscular bones (vs regularly ossified supraneural and intramuscular bones). In addition to these characters
<taxonomicName id="4C2472EBFFD7FFC07C0BFE0BFB4C23A9" authorityName="Smith" authorityYear="1841" box="[1036,1204,445,471]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC07C0BFE0BFB4C23A9" box="[1036,1204,445,471]" italics="true" pageId="16" pageNumber="17">Pseudobarbus</emphasis>
</taxonomicName>
differs from all these genera except
<taxonomicName id="4C2472EBFFD7FFC079AAFE56FDCE2384" authority="Skelton &amp; Swartz &amp; Vreven, 2018" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" box="[429,566,480,506]" class="Actinopterygii" family="Cyprinidae" genus="Sedercypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B950D57AFFD7FFC079AAFE56FDCE2384" box="[429,566,480,506]" italics="true" pageId="16" pageNumber="17">Sedercypris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD7FFC07A39FE57FD522385" box="[574,682,481,507]" pageId="16" pageNumber="17" rank="genus">gen. nov.</taxonomicNameLabel>
in having red pigmentation at the base of the fins (vs no red pigmentation); from
<taxonomicName id="4C2472EBFFD7FFC079B4FDB5FDAB2063" authorityName="Smith" authorityYear="1841" box="[435,595,515,541]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC079B4FDB5FDAB2063" box="[435,595,515,541]" italics="true" pageId="16" pageNumber="17">Cheilobarbus</emphasis>
</taxonomicName>
in adult size (&lt;
<quantity id="4CDCA48DFFD7FFC07B12FDB5FC802060" box="[789,888,515,542]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="mm" value="150.0">150 mm</quantity>
SL vs&gt;
<quantity id="4CDCA48DFFD7FFC07BE3FDB5FBBE2060" box="[996,1094,515,542]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="mm" value="150.0">150 mm</quantity>
SL); from
<taxonomicName id="4C2472EBFFD7FFC07CCCFDB5FA8E2063" authority="Skelton &amp; Swartz &amp; Vreven, 2018" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" box="[1227,1398,515,541]" class="Actinopterygii" family="Cyprinidae" genus="Amatolacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B950D57AFFD7FFC07CCCFDB5FA8E2063" box="[1227,1398,515,541]" italics="true" pageId="16" pageNumber="17">Amatolacypris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD7FFC078BAFD91FED1203F" box="[189,297,551,577]" pageId="16" pageNumber="17" rank="genus">gen. nov.</taxonomicNameLabel>
in having slender third and fourth infraorbital bones (vs broad third and fourth infraorbitals); and from
<taxonomicName id="4C2472EBFFD7FFC07934FDFDFE15201A" authority="Skelton &amp; Swartz &amp; Vreven, 2018" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" box="[307,493,587,612]" class="Actinopterygii" family="Cyprinidae" genus="Namaquacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B950D57AFFD7FFC07934FDFDFE15201A" box="[307,493,587,612]" italics="true" pageId="16" pageNumber="17">Namaquacypris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD7FFC079F1FDFDFD9B201B" box="[502,611,587,613]" pageId="16" pageNumber="17" rank="genus">gen. nov.</taxonomicNameLabel>
in the position of the dorsal fin (
<taxonomicName id="4C2472EBFFD7FFC07BF3FDFCFB64201A" authorityName="Smith" authorityYear="1841" box="[1012,1180,586,612]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC07BF3FDFCFB64201A" box="[1012,1180,586,612]" italics="true" pageId="16" pageNumber="17">Pseudobarbus</emphasis>
</taxonomicName>
dorsal fin in midbody, origin over or just behind the origin of the pelvic fins vs posteriorly, origin behind the pelvic fin), and in not having a membrane connecting the inner pelvic fin rays to the body (vs a membrane connecting the inner pelvic rays to the body).
<taxonomicName id="4C2472EBFFD7FFC07ADAFD03FC7D20B1" authorityName="Smith" authorityYear="1841" box="[733,901,693,719]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC07ADAFD03FC7D20B1" box="[733,901,693,719]" italics="true" pageId="16" pageNumber="17">Pseudobarbus</emphasis>
</taxonomicName>
differs further from
<taxonomicName id="4C2472EBFFD7FFC07C7DFD03FAFB20B1" authority="Skelton &amp; Swartz &amp; Vreven, 2018" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" box="[1146,1283,693,719]" class="Actinopterygii" family="Cyprinidae" genus="Sedercypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B950D57AFFD7FFC07C7DFD03FAFB20B1" box="[1146,1283,693,719]" italics="true" pageId="16" pageNumber="17">Sedercypris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD7FFC07D0CFD03FA8F20B1" box="[1291,1399,693,719]" pageId="16" pageNumber="17" rank="genus">gen. nov.</taxonomicNameLabel>
in the position of the mouth (subterminal vs terminal), and the number of anal-fin branched rays (five vs six or seven).
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFD7FFC078BAFC88FBE521D4" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8B9B0968FFD7FFC078BAFC88FEBC2126" blockId="16.[189,324,830,856]" box="[189,324,830,856]" pageId="16" pageNumber="17">
<heading id="D0D3BE04FFD7FFC078BAFC88FEBC2126" bold="true" box="[189,324,830,856]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B950D57AFFD7FFC078BAFC88FEBC2126" bold="true" box="[189,324,830,856]" pageId="16" pageNumber="17">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD7FFC078BAFCDAFBE521D4" blockId="16.[189,1398,876,938]" pageId="16" pageNumber="17">
<taxonomicName id="4C2472EBFFD7FFC078BAFCDAFE9D21F8" authorityName="Smith" authorityYear="1841" box="[189,357,876,902]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC078BAFCDAFE9D21F8" box="[189,357,876,902]" italics="true" pageId="16" pageNumber="17">Pseudobarbus</emphasis>
</taxonomicName>
is derived from ‘
<emphasis id="B950D57AFFD7FFC07A37FCDAFD7521F8" box="[560,653,876,902]" italics="true" pageId="16" pageNumber="17">pseudes</emphasis>
’ (Greek) meaning false, deceptive, and ‘
<emphasis id="B950D57AFFD7FFC07C6CFCDAFB4921F8" box="[1131,1201,876,902]" italics="true" pageId="16" pageNumber="17">barba</emphasis>
’ (Latin) a beard; in reference to a deceptive similarity with the genus
<taxonomicName id="4C2472EBFFD7FFC07B23FC26FC8121D4" authorityName="Daudin" authorityYear="1805" box="[804,889,912,938]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC07B23FC26FC8121D4" box="[804,889,912,938]" italics="true" pageId="16" pageNumber="17">Barbus</emphasis>
</taxonomicName>
from Europe.
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFD7FFC078BAFC67FBF527DE" pageId="16" pageNumber="17" type="description">
<paragraph id="8B9B0968FFD7FFC078BAFC67FEB62195" blockId="16.[189,334,977,1003]" box="[189,334,977,1003]" pageId="16" pageNumber="17">
<heading id="D0D3BE04FFD7FFC078BAFC67FEB62195" bold="true" box="[189,334,977,1003]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B950D57AFFD7FFC078BAFC67FEB62195" bold="true" box="[189,334,977,1003]" pageId="16" pageNumber="17">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD7FFC078BAFBB6FBF527DE" blockId="16.[189,1399,1024,1441]" pageId="16" pageNumber="17">
Species of
<taxonomicName id="4C2472EBFFD7FFC0793AFBB6FE1D2664" authorityName="Smith" authorityYear="1841" box="[317,485,1024,1050]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC0793AFBB6FE1D2664" box="[317,485,1024,1050]" italics="true" pageId="16" pageNumber="17">Pseudobarbus</emphasis>
</taxonomicName>
are moderate-sized (&lt;
<quantity id="4CDCA48DFFD7FFC07AF6FBB6FCAA2665" box="[753,850,1024,1051]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="mm" value="150.0">150 mm</quantity>
SL) fusiform or terete, tetraploid smiliogastrin minnows, with one or two pairs of simple circum-oral barbels; lips variably developed, pharyngeal teeth in two or three rows; tooth formulae 0-2,3,3,4+5-5-4,3,0-2; pharyngeal tooth crowns variable with offset major cusp; intestine variable in length from 1:
<quantity id="4CDCA48DFFD7FFC07B19FBDDFCB626FA" box="[798,846,1130,1157]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0">1 in</quantity>
SL to 3–4:1 SL; scales radially striated, from small to moderate in size, nape and breast scales reduced or embedded; no pectoral or pelvic axil scale; lateral line interrupted or complete, in mid-body; cephalic lateral line system with pre-opercular branch disconnected and reduced or absent on the mandible. Dorsal fin with simple flexible unbranched ray and normally seven branched rays. Anal fin with five branched rays. Pectoral fins sexually dimorphic in mature adults, males with longer more expansive pectorals. Adults with bright red patches at the base of the fins. Mature males develop conical tubercles on the head, in a distinctive pattern as illustrated in
<bibRefCitation id="EFB57499FFD7FFC078BAFAD5FE332703" author="Skelton P. H." box="[189,459,1379,1405]" journalOrPublisher="Annals of the Cape Provincial Museums (Natural History)" pageId="16" pageNumber="17" pagination="201 - 307" part="1 6" refId="ref14443" refString="Skelton P. H. 1988. A taxonomic revision of the redfin minnows (Pisces, Cyprinidae) from southern Africa. Annals of the Cape Provincial Museums (Natural History) 1 6 (10): 201 - 307." title="A taxonomic revision of the redfin minnows (Pisces, Cyprinidae) from southern Africa" type="journal article" year="1988">Skelton (1988: fig. 31)</bibRefCitation>
, in single rows along the free edge of scales and in bands over anterior pectoral fins. Axial skeleton without ossified supraneural or intramuscular bones.
</paragraph>
</subSubSection>
</treatment>
</document>
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