treatments-xml/data/03/8C/87/038C87C4FFC2FFE4BDAEFF08FD116088.xml
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<document id="166D546A80C8DD2D276A0C4A454E9AD2" ID-DOI="10.11646/zootaxa.5363.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146851" ID-ZooBank="B8979382-355B-410A-B1C9-20AD910233E6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1700196986855" checkinUser="plazi" docAuthor="Riedel, Matthias" docDate="2023" docId="038C87C4FFC2FFE4BDAEFF08FD116088" docLanguage="en" docName="zootaxa.5363.1.1.pdf" docOrigin="Zootaxa 5363 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eccoptosage sabahensis Riedel 2023, nov. sp." docType="treatment" docVersion="4" lastPageNumber="56" masterDocId="FFB5FFBCFFF5FFD3BD39FF9FFF806516" masterDocTitle="Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia" masterLastPageNumber="94" masterPageNumber="1" pageNumber="56" updateTime="1700571439667" updateUser="diego">
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<mods:title id="3CE0A1417D472DCA62CC01D54A9BCACA">Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia</mods:title>
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<mods:date id="8890E9F8415941B2CD211346C0EA744D">2023</mods:date>
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<treatment id="038C87C4FFC2FFE4BDAEFF08FD116088" ID-DOI="http://doi.org/10.5281/zenodo.10167804" ID-Zenodo-Dep="10167804" LSID="urn:lsid:plazi:treatment:038C87C4FFC2FFE4BDAEFF08FD116088" httpUri="http://treatment.plazi.org/id/038C87C4FFC2FFE4BDAEFF08FD116088" lastPageNumber="56" pageId="55" pageNumber="56">
<subSubSection id="C33F6559FFC2FFE4BDAEFF08FDA065A7" box="[151,544,151,178]" pageId="55" pageNumber="56" type="nomenclature">
<paragraph id="8B9A36D2FFC2FFE4BDAEFF08FDA065A7" blockId="55.[151,544,151,213]" box="[151,544,151,178]" pageId="55" pageNumber="56">
<heading id="D0D281BEFFC2FFE4BDAEFF08FDA065A7" bold="true" box="[151,544,151,178]" fontSize="11" level="2" pageId="55" pageNumber="56" reason="4">
<emphasis id="B951EAC0FFC2FFE4BDAEFF08FDA065A7" bold="true" box="[151,544,151,178]" pageId="55" pageNumber="56">
<taxonomicName id="4C254D51FFC2FFE4BDAEFF08FE3A65A4" authority="Riedel, 2023" authorityName="Riedel" authorityYear="2023" box="[151,442,151,178]" class="Insecta" family="Ichneumonidae" genus="Eccoptosage" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B951EAC0FFC2FFE4BDAEFF08FE3A65A4" bold="true" box="[151,442,151,178]" italics="true" pageId="55" pageNumber="56">Eccoptosage sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257BBFFC2FFE4BCF8FF08FDA065A7" box="[449,544,151,177]" pageId="55" pageNumber="56" rank="species">nov. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6559FFC2FFE4BDAEFF24FEB265C3" box="[151,306,187,213]" pageId="55" pageNumber="56" type="description">
<paragraph id="8B9A36D2FFC2FFE4BDAEFF24FEB265C3" blockId="55.[151,544,151,213]" box="[151,306,187,213]" pageId="55" pageNumber="56">
(
<figureCitation id="131E2A57FFC2FFE4BDA6FF24FEAA65C3" box="[159,298,187,213]" captionStart="FIGURE 23" captionStartId="54.[151,250,1868,1892]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure-23@54.[151,1436,181,1844]" captionTargetPageId="54" captionText="FIGURE 23. Eccoptosage sabahensis nov. sp. HT ♂. A face, frontal view, B propodeum, dorsal view, C basal tergites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10146898" httpUri="https://zenodo.org/record/10146898/files/figure.png" pageId="55" pageNumber="56">Figs 23AC</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F6559FFC2FFE4BDAEFE9CFD8B649F" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="8B9A36D2FFC2FFE4BDAEFE9CFD8B6457" blockId="55.[151,1437,259,1438]" pageId="55" pageNumber="56">
<materialsCitation id="3B4D3C8FFFC2FFE4BDAEFE9CFD8B6457" collectingDate="1987-10-20" collectingDateMax="1987-12-22" collectingDateMin="1987-10-20" collectorName="Danum Valley Field C &amp; C. van Achterberg" country="Malaysia" elevation="150" location="Valley Field" municipality="Danum Valley" pageId="55" pageNumber="56" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B951EAC0FFC2FFE4BDAEFE9CFE84640B" bold="true" box="[151,260,259,285]" pageId="55" pageNumber="56">
<typeStatus id="549E8870FFC2FFE4BDAEFE9CFE84640B" box="[151,260,259,285]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
</emphasis>
. (♂)
<collectingCountry id="F3327642FFC2FFE4BC78FE9CFE2C640B" box="[321,428,259,285]" name="Malaysia" pageId="55" pageNumber="56">Malaysia</collectingCountry>
: SE
<collectingRegion id="49E1F830FFC2FFE4BCE6FE9CFDA6640B" box="[479,550,259,285]" country="Malaysia" name="Sabah" pageId="55" pageNumber="56">Sabah</collectingRegion>
, nr
<collectorName id="26D05304FFC2FFE4BF69FE9BFCCE640B" box="[592,846,259,286]" pageId="55" pageNumber="56">
<collectingMunicipality id="6BFEACA8FFC2FFE4BF69FE9BFD71640B" box="[592,753,259,286]" pageId="55" pageNumber="56">
Danum
<location id="8EFA6009FFC2FFE4BF90FE9CFD71640B" LSID="urn:lsid:plazi:treatment:038C87C4FFC2FFE4BDAEFF08FD116088:8EFA6009FFC2FFE4BF90FE9CFD71640B" box="[681,753,259,285]" country="Malaysia" municipality="Danum Valley" name="Valley Field" pageId="55" pageNumber="56" stateProvince="Sabah">Valley</location>
</collectingMunicipality>
Field C
</collectorName>
,
<collectingMethod id="52644EC5FFC2FFE4BE60FE9CFC656408" box="[857,997,259,286]" pageId="55" pageNumber="56">Malaise trap</collectingMethod>
15, c
<quantity id="4CDD9B37FFC2FFE4B913FE9CFBF36408" box="[1066,1139,259,286]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="55" pageNumber="56" unit="m" value="150.0">
<elevation id="0008D1E1FFC2FFE4B913FE9CFBF36408" box="[1066,1139,259,286]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="55" pageNumber="56" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="FF9B1012FFC2FFE4B946FE9CFAC7640B" box="[1151,1351,259,285]" pageId="55" pageNumber="56" value="1987-10-20" valueMax="1987-12-22" valueMin="1987-10-20">
<collectingDate id="EFDFE9FAFFC2FFE4B946FE9CFAC7640B" box="[1151,1351,259,285]" pageId="55" pageNumber="56" value="1987-10-20" valueMax="1987-12-22" valueMin="1987-10-20">20.x.22.xii.1987</collectingDate>
</date>
,
<collectorName id="26D05304FFC2FFE4B86AFE9CFE976457" pageId="55" pageNumber="56">C. van Achterberg</collectorName>
, RMNH´87 (Leiden).
</materialsCitation>
</paragraph>
<paragraph id="8B9A36D2FFC2FFE4BDFEFED4FD8B649F" blockId="55.[151,1437,259,1438]" pageId="55" pageNumber="56">
<materialsCitation id="3B4D3C8FFFC2FFE4BDFEFED4FD87649F" collectingDate="1987-04-01" collectingDateMax="1987-04-14" collectingDateMin="1987-04-01" collectorName="Mal. &amp; C. van Achterberg" country="Malaysia" elevation="1050" location="Long Pa Sia" pageId="55" pageNumber="56" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B951EAC0FFC2FFE4BDFEFED4FEB46473" bold="true" box="[199,308,331,357]" pageId="55" pageNumber="56">
<typeStatus id="549E8870FFC2FFE4BDFEFED4FEB46473" box="[199,308,331,357]" pageId="55" pageNumber="56" type="paratype">Paratype</typeStatus>
</emphasis>
. (♂)
<collectingCountry id="F3327642FFC2FFE4BC4EFED4FE626473" box="[375,482,331,357]" name="Malaysia" pageId="55" pageNumber="56">Malaysia</collectingCountry>
: SW
<collectingRegion id="49E1F830FFC2FFE4BF1DFED4FDEB6473" box="[548,619,331,357]" country="Malaysia" name="Sabah" pageId="55" pageNumber="56">Sabah</collectingRegion>
, nr
<location id="8EFA6009FFC2FFE4BFA5FED3FCAC6473" LSID="urn:lsid:plazi:treatment:038C87C4FFC2FFE4BDAEFF08FD116088:8EFA6009FFC2FFE4BFA5FED3FCAC6473" box="[668,812,331,358]" country="Malaysia" name="Long Pa Sia" pageId="55" pageNumber="56" stateProvince="Sabah">Long Pa Sia</location>
(West), c.
<quantity id="4CDD9B37FFC2FFE4BE96FED4FB896470" box="[943,1033,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="55" pageNumber="56" unit="m" value="1050.0">
<elevation id="0008D1E1FFC2FFE4BE96FED4FB896470" box="[943,1033,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="55" pageNumber="56" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="FF9B1012FFC2FFE4B921FED4FB2D6473" box="[1048,1197,331,357]" pageId="55" pageNumber="56" value="1987-04-01" valueMax="1987-04-14" valueMin="1987-04-01">
<collectingDate id="EFDFE9FAFFC2FFE4B921FED4FB2D6473" box="[1048,1197,331,357]" pageId="55" pageNumber="56" value="1987-04-01" valueMax="1987-04-14" valueMin="1987-04-01">114.iv.1987</collectingDate>
</date>
,
<collectorName id="26D05304FFC2FFE4B985FED4FB706473" box="[1212,1264,331,357]" pageId="55" pageNumber="56">Mal.</collectorName>
trap 3,
<collectorName id="26D05304FFC2FFE4B869FED4FE97649F" pageId="55" pageNumber="56">C. van Achterberg</collectorName>
, RMNH´87 (Leiden)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6559FFC2FFE4BDFEFE0CFB9361FF" pageId="55" pageNumber="56" type="description">
<paragraph id="8B9A36D2FFC2FFE4BDFEFE0CFD3D64B8" blockId="55.[151,1437,259,1438]" box="[199,701,403,432]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4BDFEFE0CFED164BB" bold="true" box="[199,337,403,429]" pageId="55" pageNumber="56">Description</emphasis>
. ♂. Body length 10.0
<quantity id="4CDD9B37FFC2FFE4BF6AFE0CFD3964B8" box="[595,697,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="55" pageNumber="56" unit="mm" value="10.5">10.5 mm</quantity>
.
</paragraph>
<paragraph id="8B9A36D2FFC2FFE4BDFEFE28FE3E67BF" blockId="55.[151,1437,259,1438]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4BDFEFE28FE8464C7" box="[199,260,439,465]" italics="true" pageId="55" pageNumber="56">Head</emphasis>
. Flagellum with 35 flagellomeres, bristle-shaped; 1
<superScript id="7C509B9AFFC2FFE4BE63FE28FCE564D3" attach="left" box="[858,869,439,453]" fontSize="6" pageId="55" pageNumber="56">st</superScript>
flagellomere 2.8 × longer than wide. Tyloids on flagellomeres 10/1113/14, long-oval, maximally 0.5 × as long as their flagellomeres. Temple short, strongly and roundly narrowed behind eye. OED 1.6 × and OOD 0.4 × ocellar diameter. Frons smooth, with some sparse punctures dorsally. Face densely punctate. Clypeus large, flat, with rather dense punctuation; apical margin sharp, slightly rounded (fig. 23A). Mandible seen from frontal sickle-shaped, ventral tooth small and strongly turned inwards (fig. 23A). Malar space 0.6 × as long as width of mandibular base. Gena densely punctate. Genal carina reaching mandibular base.
</paragraph>
<paragraph id="8B9A36D2FFC2FFE4BDFEFD2BFBA56697" blockId="55.[151,1437,259,1438]" pageId="55" pageNumber="56">
<taxonomicName id="4C254D51FFC2FFE4BDFEFD2BFEBF67DB" box="[199,319,692,717]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAC0FFC2FFE4BDFEFD2BFEBF67DB" box="[199,319,692,717]" italics="true" pageId="55" pageNumber="56">Mesosoma</emphasis>
</taxonomicName>
. Mesoscutum finely and densely punctate, dull. Mesopleuron with rather dense superficial punctures, shining; speculum with fine punctures. Metapleuron punctate (in HT) or ± striate (in PT). Scutellum pyramidal, with strong lateral carina forming a distinct lamelliform extension which is slightly bent outwards. Propodeum with strong apophysis (fig. 23B). Area superomedia hexagonal, slightly wider than long (confluent with area basalis in HT); costula reaching its basal 0.4. Hind femur 4.7 × longer than wide. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 1.0 × their width. Vein 1cu-a postfurcal by 2.0 × its width.
</paragraph>
<paragraph id="8B9A36D2FFC2FFE4BDFEFC13FB456107" blockId="55.[151,1437,259,1438]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4BDFEFC13FEBC66B3" box="[199,316,908,933]" italics="true" pageId="55" pageNumber="56">Metasoma</emphasis>
. Postpetiole slightly widened, with microsculpture and few fine punctures, shining. 2
<superScript id="7C509B9AFFC2FFE4B9C5FC14FA8C668F" attach="left" box="[1276,1292,907,921]" fontSize="6" pageId="55" pageNumber="56">nd</superScript>
tergite 1.1 × longer than wide. Gastrocoelus slightly impressed. Thyridium small, transverse, 0.3 × as wide as interval between thyridia (fig. 23C). 2
<superScript id="7C509B9AFFC2FFE4BCB2FC4CFE1B66F7" attach="left" box="[395,411,979,993]" fontSize="6" pageId="55" pageNumber="56">nd</superScript>
tergite finely and densely punctate, finely rugose and aciculate basally, dull. Following tergites with superficial punctures, ± shining. Hypopygium unmodified, apical margin rounded.
</paragraph>
<paragraph id="8B9A36D2FFC2FFE4BDFEFB84FB9361FF" blockId="55.[151,1437,259,1438]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4BDFEFB84FE966123" box="[199,278,1051,1077]" italics="true" pageId="55" pageNumber="56">Colour</emphasis>
. Flagellum black, flagellomeres 8/916 ivory dorsally. Head and mesosoma red. Palps, mandible except teeth, clypeus, face and frontal orbit ivory; gena yellowish. Frons and temples black. Scape red ventrally. 1
<superScript id="7C509B9AFFC2FFE4B87AFBA0FACE615B" attach="left" box="[1347,1358,1087,1101]" fontSize="6" pageId="55" pageNumber="56">st</superScript>
tergite reddish. 2
<superScript id="7C509B9AFFC2FFE4BC3DFBFCFE946167" attach="left" box="[260,276,1123,1137]" fontSize="6" pageId="55" pageNumber="56">nd</superScript>
and 3
<superScript id="7C509B9AFFC2FFE4BC6FFBFCFEE46167" attach="left" box="[342,356,1123,1137]" fontSize="6" pageId="55" pageNumber="56">rd</superScript>
tergites mainly black, yellowish laterally and apically (in PT) or yellowish, with black basal spots (in HT) (fig. 23C). 4
<superScript id="7C509B9AFFC2FFE4BC46FB18FE0C6183" attach="left" box="[383,396,1159,1173]" fontSize="6" pageId="55" pageNumber="56">th</superScript>
to 6
<superScript id="7C509B9AFFC2FFE4BC86FB18FE4C6183" attach="left" box="[447,460,1159,1173]" fontSize="6" pageId="55" pageNumber="56">th</superScript>
tergites blackish or brownish, yellowish laterally and apically. 7
<superScript id="7C509B9AFFC2FFE4B998FB18FB2E6183" attach="left" box="[1185,1198,1159,1173]" fontSize="6" pageId="55" pageNumber="56">th</superScript>
tergite and paramere yellowish. Fore and mid legs including coxae reddish-yellow; mid coxa basally and mid tarsus mainly brown; hind leg black, hind coxa red dorsally. Wings slightly yellowish; pterostigma yellow.
</paragraph>
</subSubSection>
<subSubSection id="C33F6559FFC2FFE4BDFEFB6CFBAE606C" pageId="55" pageNumber="56" type="discussion">
<paragraph id="8B9A36D2FFC2FFE4BDFEFB6CFBAE606C" blockId="55.[151,1437,259,1438]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4BDFEFB6CFEB4601B" bold="true" box="[199,308,1267,1293]" pageId="55" pageNumber="56">Remarks</emphasis>
. I was not able to correlate these ♂♂ with any
<specimenCount id="9D23FD5BFFC2FFE4BE52FB6BFC156018" box="[875,917,1268,1294]" count="1" pageId="55" pageNumber="56" type="female">♀♀</specimenCount>
of the known
<taxonomicName id="4C254D51FFC2FFE4B97DFB6BFAAF601B" box="[1092,1327,1267,1293]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4B97DFB6BFB53601B" box="[1092,1235,1268,1293]" italics="true" pageId="55" pageNumber="56">Eccoptosage</emphasis>
species
</taxonomicName>
from the Oriental region and describe them as a new species here, but the taxonomy of the Oriental
<taxonomicName id="4C254D51FFC2FFE4B9A3FA87FA026027" box="[1178,1410,1303,1329]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4B9A3FA87FAA96027" box="[1178,1321,1304,1329]" italics="true" pageId="55" pageNumber="56">Eccoptosage</emphasis>
species
</taxonomicName>
is fragmentary to date. More studies of the available
<typeStatus id="549E8870FFC2FFE4BFFCFAA3FD756040" box="[709,757,1340,1366]" pageId="55" pageNumber="56">type</typeStatus>
material, additional collection activities and genomic studies are necessary in the future for a comprehensive revision of this rather large genus.
</paragraph>
</subSubSection>
<subSubSection id="C33F6559FFC2FFE4BDFEFA1CFD116088" box="[199,657,1411,1438]" pageId="55" pageNumber="56" type="distribution">
<paragraph id="8B9A36D2FFC2FFE4BDFEFA1CFD116088" blockId="55.[151,1437,259,1438]" box="[199,657,1411,1438]" pageId="55" pageNumber="56">
<emphasis id="B951EAC0FFC2FFE4BDFEFA1CFED9608B" bold="true" box="[199,345,1411,1437]" pageId="55" pageNumber="56">Distribution</emphasis>
. Only known from Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>