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<document id="5E6055E10FE37981F16DCA680914E371" ID-CLB-Dataset="4031" ID-DOI="10.5852/ejt.2021.750.1365" ID-GBIF-Dataset="818d0bc4-31a6-4319-86a3-6e6b16ab72df" ID-ISSN="2118-9773" ID-Zenodo-Dep="5451747" ID-ZooBank="CE5D8517-AC04-4AF9-BD78-A2D6E21D33A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630516667691" checkinUser="felipe" docAuthor="Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z. &amp; Ahrens, Dirk" docDate="2021" docId="038B976CFF88FFEBCA28CCA5149DCB12" docLanguage="en" docName="ejt-750_94-123.pdf" docOrigin="European Journal of Taxonomy 750 (750)" docSource="http://dx.doi.org/10.5852/ejt.2021.750.1365" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Astaena biciliata L. W. Saylor 1946" docType="treatment" docVersion="9" lastPageNumber="99" masterDocId="FFB2EF14FF8BFFEEC84EC925155FCE04" masterDocTitle="Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor" masterLastPageNumber="123" masterPageNumber="94" pageNumber="97" updateTime="1699290068827" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4D82D2BA8C723946FCB56D92E60B6461">Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor</mods:title>
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<mods:namePart id="42D04E13D7DCE9C3D48F21C39C0F3B82">Pacheco, Thaynara L.</mods:namePart>
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<mods:affiliation id="68A1441A389B23F82F9CDD17DA86E1FC">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="38550693E061C29C1BC38B36C9AF5A2E" type="email">thay.lpacheco@gmail.com</mods:nameIdentifier>
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<mods:namePart id="DE258E415216069D3168AEB8639C30ED">Monné, Marcela L.</mods:namePart>
<mods:nameIdentifier id="A41EB81CB3130D23D0D4B9160D0A3B33" type="LSID">86B186B4-4297-4380-A98D-FC813B5D53A6&amp;2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="67B408E4CF97ECADD866ECBC4B76C141">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="7C2AAF0EE2220DB11FC656C9CD7D20DA" type="email">mlmonne2@gmail.com&amp;vazdemello@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="CBC065014C26F902F2D0D6E0C2850AA7">Author</mods:roleTerm>
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<mods:namePart id="F02D550FCC33EE3C511733B0E28E76B6">Vaz-De, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="AB3F3761CD84E91E7456504D7C670357" type="LSID">2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="4E2C4C05ADDA43839D01B6C1B222F476">Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="76DF71CA7C312584E1C31877BEEEAB19" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AEE22EF61484EFFF86CC09FB799CA258">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="E93408E6DE19BED93336C8490CCF7412">Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="95079CFD637F6F6064D2577DFE02B44B" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:title id="BDAA4F085952CBA45F6F8E209073D0F4">European Journal of Taxonomy</mods:title>
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<mods:date id="498D6A80EDD90C4193433D13C206C4FD">2021</mods:date>
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<treatment id="038B976CFF88FFEBCA28CCA5149DCB12" ID-DOI="http://doi.org/10.5281/zenodo.5572724" ID-GBIF-Taxon="188647623" ID-Zenodo-Dep="5572724" LSID="urn:lsid:plazi:treatment:038B976CFF88FFEBCA28CCA5149DCB12" httpUri="http://treatment.plazi.org/id/038B976CFF88FFEBCA28CCA5149DCB12" lastPageId="5" lastPageNumber="99" pageId="3" pageNumber="97">
<subSubSection id="C33875F1FF88FFEDCA28CCA51691CB9E" box="[614,974,1407,1434]" pageId="3" pageNumber="97" type="nomenclature">
<paragraph id="8B9D267AFF88FFEDCA28CCA51691CB9E" blockId="3.[614,974,1407,1472]" box="[614,974,1407,1434]" pageId="3" pageNumber="97">
<heading id="D0D59116FF88FFEDCA28CCA51691CB9E" box="[614,974,1407,1434]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="97" reason="2">
<taxonomicName id="4C225DF9FF88FFEDCA28CCA51691CB9E" authority="Saylor, 1946" authorityName="L. W. Saylor" authorityYear="1946" box="[614,974,1407,1434]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="biciliata">
<emphasis id="B956FA68FF88FFEDCA28CCA5166ECB9E" bold="true" box="[614,817,1407,1434]" italics="true" pageId="3" pageNumber="97">Astaena biciliata</emphasis>
<bibRefCitation id="EFB35B8BFF88FFEDCB76CCA51691CB9E" author="Saylor L. W." box="[824,974,1408,1434]" pageId="3" pageNumber="97" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C33875F1FF88FFEDCA86CC831632CBC4" box="[712,877,1446,1472]" pageId="3" pageNumber="97" type="description">
<paragraph id="8B9D267AFF88FFEDCA86CC831632CBC4" blockId="3.[614,974,1407,1472]" box="[712,877,1446,1472]" pageId="3" pageNumber="97">
<figureCitation id="13193AFFFF88FFEDCA86CC831662CBC4" box="[712,829,1446,1472]" captionStart="Fig" captionStartId="4.[189,232,1856,1882]" captionTargetBox="[244,1343,265,1819]" captionTargetId="figure-11@4.[244,1343,265,1819]" captionTargetPageId="4" captionText="Fig. 1. A–D. Astaena abaca Saylor, 1946, holotype, ♀ (Type No. 7904; CAS). – E–I. A. biciliata Saylor, 1946, holotype, ♂ (Type No. 7907; CAS). – J–K. A. bogotana Saylor, 1946, paratype, ♂ (CAS). – L–N. A. bogotana, holotype, ♂ (Type No. 7909; CAS). A. Head and pronotum, dorsal view. B, G, L. Labels. C, H, M. Habitus, dorsal view. D, I, N. Habitus, lateral view. E, J. Parameres, dorsal view. F, K. Aedeagus, lateral view. Scale bars: A, E–F, J–K = 0.5 mm; C–D, H–I, M–N = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5451749" httpUri="https://zenodo.org/record/5451749/files/figure.png" pageId="3" pageNumber="97">Figs 1E–I</figureCitation>
,
<figureCitation id="13193AFFFF88FFEDCB07CC831632CBC4" box="[841,877,1446,1472]" captionStart="Fig" captionStartId="26.[189,232,1822,1848]" captionTargetBox="[261,1326,295,1750]" captionTargetId="figure-12@26.[241,1347,265,1784]" captionTargetPageId="26" captionText="Fig. 7. Distribution of type localities of the South American Sericini species described by Saylor. A. Astaena abaca Saylor, 1946, A. biciliata Saylor, 1946, A. bogotana Saylor, 1946, A. excisipes Saylor, 1947, A. explaniceps Saylor, 1947, A. incachaca Saylor, 1946, A. maqueta Saylor, 1947, and A. nigrona Saylor, 1947. B. A. fusagona Saylor, 1946, A. pygidia Saylor, 1946, A. rugithorax Saylor, 1946, A. salta Saylor, 1946, A. yungasa Saylor, 1946, and A. zyrota Saylor, 1946. C. Sayloria abcora (Saylor, 1946) comb. nov., S. bicoloripes (Saylor, 1946) comb. nov., and S. pottsi (Saylor, 1946) comb. nov.." figureDoi="http://doi.org/10.5281/zenodo.5451761" httpUri="https://zenodo.org/record/5451761/files/figure.png" pageId="3" pageNumber="97">7A</figureCitation>
</paragraph>
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<subSubSection id="C33875F1FF88FFEDC8F3CCCE16ADC874" pageId="3" pageNumber="97" type="reference_group">
<paragraph id="8B9D267AFF88FFEDC8F3CCCE173BC801" blockId="3.[189,612,1514,1541]" box="[189,612,1514,1541]" pageId="3" pageNumber="97">
<treatmentCitationGroup id="AB320154FF88FFEDC8F3CCCE173BC801" box="[189,612,1514,1541]" pageId="3" pageNumber="97">
<treatmentCitation id="0A83006BFF88FFEDC8F3CCCE173FC801" author="Saylor L. W." box="[189,608,1514,1541]" page="219" pageId="3" pageNumber="97" year="1946">
<taxonomicName id="4C225DF9FF88FFEDC8F3CCCE173FC801" authority="Saylor, 1946: 219" authorityName="L. W. Saylor" authorityPageNumber="219" authorityYear="1946" box="[189,608,1514,1541]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="biciliata">
<emphasis id="B956FA68FF88FFEDC8F3CCCE14DAC800" box="[189,389,1514,1541]" italics="true" pageId="3" pageNumber="97">Astaena biciliata</emphasis>
<bibRefCitation id="EFB35B8BFF88FFEDC9C2CCCF173FC801" author="Saylor L. W." box="[396,608,1514,1541]" pageId="3" pageNumber="97" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946: 219</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9D267AFF88FFEDC8F3CF0A16ADC874" blockId="3.[189,1399,1583,1648]" pageId="3" pageNumber="97">
<treatmentCitationGroup id="AB320154FF88FFEDC8F3CF0A16ADC874" pageId="3" pageNumber="97">
<taxonomicName id="4C225DF9FF88FFEDC8F3CF0A14DBC84D" authorityName="L. W. Saylor" authorityYear="1946" box="[189,388,1583,1609]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="biciliata">
<emphasis id="B956FA68FF88FFEDC8F3CF0A14DBC84D" box="[189,388,1583,1609]" italics="true" pageId="3" pageNumber="97">Astaena biciliata</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A83006BFF88FFEDC9EECF0A1709C84E" author="Frey G." box="[416,598,1583,1610]" page="335" pageId="3" pageNumber="97" year="1973">
<bibRefCitation id="EFB35B8BFF88FFEDC9EECF0A1709C84E" author="Frey G." box="[416,598,1583,1610]" pageId="3" pageNumber="97" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey 1973: 335</bibRefCitation>
</treatmentCitation>
, 347 (identification key). —
<treatmentCitation id="0A83006BFF88FFEDCBE8CF0A1104C84D" author="Evans A. V." box="[934,1115,1583,1609]" page="19" pageId="3" pageNumber="97" year="2003">
<bibRefCitation id="EFB35B8BFF88FFEDCBE8CF0A1104C84D" author="Evans A. V." box="[934,1115,1583,1609]" pageId="3" pageNumber="97" pagination="1 - 458" refId="ref13003" refString="Evans A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 19</bibRefCitation>
</treatmentCitation>
(checklist). — Evans &amp;
<treatmentCitation id="0A83006BFF88FFEDC8A4CF7014CEC874" author="Smith A. B. T. &amp; Evans A. V." box="[234,401,1621,1648]" page="5" pageId="3" pageNumber="97" year="2005">
<bibRefCitation id="EFB35B8BFF88FFEDC8A4CF7014CEC874" author="Smith A. B. T. &amp; Evans A. V." box="[234,401,1621,1648]" pageId="3" pageNumber="97" pagination="29 - 60" refId="ref14061" refString="Smith A. B. T. &amp; Evans A. V. 2005. A supplement to the checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) with notes on their tribal classification. Zootaxa 1032: 29 - 60. https: // doi. org / 10.11646 / zootaxa. 1032.1.2" type="journal article" year="2005">Smith 2005: 5</bibRefCitation>
</treatmentCitation>
(checklist); 2007: 5 (checklist); 2009: 5 (checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF88FFEBC8F3CFBF1483CF26" lastPageId="5" lastPageNumber="99" pageId="3" pageNumber="97" type="materials_examined">
<paragraph id="8B9D267AFF88FFEDC8F3CFBF14B3C8B7" blockId="3.[189,492,1689,1716]" box="[189,492,1689,1716]" pageId="3" pageNumber="97">
<heading id="D0D59116FF88FFEDC8F3CFBF14B3C8B7" bold="true" box="[189,492,1689,1716]" fontSize="11" level="3" pageId="3" pageNumber="97" reason="3">
<emphasis id="B956FA68FF88FFEDC8F3CFBF14B3C8B7" bold="true" box="[189,492,1689,1716]" pageId="3" pageNumber="97">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF88FFEDC8A4CFEE1154C935" blockId="3.[189,1398,1739,1842]" pageId="3" pageNumber="97">
<materialsCitation id="3B4A2C27FF88FFEDC8A4CFEE1154C935" ID-GBIF-Occurrence="3383726308" collectingDate="1935-09-20" collectionCode="CAS" collectorName="L. W. Saylor Collection" country="Peru" location="PERU" pageId="3" pageNumber="97" specimenCount="1" specimenCount-male="1" stateProvince="Lima" typeStatus="holotype">
<emphasis id="B956FA68FF88FFEDC8A4CFEE1403C8E1" bold="true" box="[234,348,1739,1765]" pageId="3" pageNumber="97">
<typeStatus id="549998D8FF88FFEDC8A4CFEE1403C8E1" box="[234,348,1739,1765]" pageId="3" pageNumber="97" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F33566EAFF88FFEDC8F3CFD71457C908" box="[189,264,1778,1804]" name="Peru" pageId="3" pageNumber="97">PERU</collectingCountry>
•
<specimenCount id="9D24EDF3FF88FFEDC96BCFD71461C908" box="[293,318,1778,1804]" pageId="3" pageNumber="97" type="male">♂</specimenCount>
; “
<date id="FF9C00BAFF88FFEDC919CFD414E0C908" box="[343,447,1777,1804]" pageId="3" pageNumber="97" value="1935-09-20">
<collectingDate id="EFD8F952FF88FFEDC919CFD414E0C908" box="[343,447,1777,1804]" pageId="3" pageNumber="97" value="1935-09-20">ix-20-35</collectingDate>
</date>
M. Sani Beni Lima Peru, 35 / L.W. Saylor Collection / California Academy of Sciences Type nº 7907 / Holotype
<taxonomicName id="4C225DF9FF88FFEDCA1FCE3D16EBC935" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1946" box="[593,948,1815,1842]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="biciliata">
<emphasis id="B956FA68FF88FFEDCA1FCE3D1646C935" box="[593,793,1815,1842]" italics="true" pageId="3" pageNumber="97">Astaena biciliata</emphasis>
L.W. Saylor
</taxonomicName>
”;
<collectionCode id="ED33BEBFFF88FFEDCB84CE321158C935" box="[970,1031,1815,1841]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="97" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9D267AFF88FFEDC8A4CE7914DCC9EC" blockId="3.[189,1399,1884,2024]" pageId="3" pageNumber="97">
<materialsCitation id="3B4A2C27FF88FFEDC8A4CE7914DCC9EC" ID-GBIF-Occurrence="3383726329" collectingDate="1935-09-20" collectionCode="CAS" collectorName="Sani Beni &amp; Peru &amp; L. W. Saylor Collection &amp; Collection &amp; San Francisco &amp; California" country="Peru" location="PERU" pageId="3" pageNumber="97" specimenCount="1" specimenCount-female="1" stateProvince="Lima" typeStatus="paratype">
<emphasis id="B956FA68FF88FFEDC8A4CE791403C972" bold="true" box="[234,348,1884,1910]" pageId="3" pageNumber="97">
<typeStatus id="549998D8FF88FFEDC8A4CE791403C972" box="[234,348,1884,1910]" pageId="3" pageNumber="97" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F33566EAFF88FFEDC8F3CEA71457C998" box="[189,264,1922,1948]" name="Peru" pageId="3" pageNumber="97">PERU</collectingCountry>
•
<specimenCount id="9D24EDF3FF88FFEDC953CEA71418C998" box="[285,327,1922,1948]" pageId="3" pageNumber="97" type="female">1 ♀</specimenCount>
; “
<date id="FF9C00BAFF88FFEDC910CEA71499C999" box="[350,454,1922,1949]" pageId="3" pageNumber="97" value="1935-09-20">
<collectingDate id="EFD8F952FF88FFEDC910CEA71499C999" box="[350,454,1922,1949]" pageId="3" pageNumber="97" value="1935-09-20">ix-20-35</collectingDate>
</date>
M. Sani Beni Lima Peru, 35 / L.W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype
<taxonomicName id="4C225DF9FF88FFEDCBC4CE8D11B5C9C6" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1946" box="[906,1258,1960,1987]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="biciliata">
<emphasis id="B956FA68FF88FFEDCBC4CE8D110BC9C6" box="[906,1108,1960,1986]" italics="true" pageId="3" pageNumber="97">Astaena biciliata</emphasis>
L.W. Saylor
</taxonomicName>
/ CASENT 8438128”;
<collectionCode id="ED33BEBFFF88FFEDC90CCEEB1420C9EC" box="[322,383,1998,2024]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="97" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="DF5D76F2FF8FFFEAC8F3CE65119CC9EC" ID-DOI="http://doi.org/10.5281/zenodo.5451749" ID-Zenodo-Dep="5451749" httpUri="https://zenodo.org/record/5451749/files/figure.png" pageId="4" pageNumber="98" startId="4.[189,232,1856,1882]" targetBox="[244,1343,265,1819]" targetPageId="4">
<paragraph id="8B9D267AFF8FFFEAC8F3CE65119CC9EC" blockId="4.[189,1400,1856,2024]" pageId="4" pageNumber="98">
<emphasis id="B956FA68FF8FFFEAC8F3CE65140CC95E" bold="true" box="[189,339,1856,1882]" pageId="4" pageNumber="98">Fig. 1. A–D</emphasis>
.
<taxonomicName id="4C225DF9FF8FFFEAC928CE6417E0C95F" authority="Saylor, 1946" authorityName="L. W. Saylor" authorityYear="1946" box="[358,703,1857,1883]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="98" phylum="Arthropoda" rank="species" species="abaca">
<emphasis id="B956FA68FF8FFFEAC928CE641747C95F" box="[358,536,1857,1883]" italics="true" pageId="4" pageNumber="98">Astaena abaca</emphasis>
<bibRefCitation id="EFB35B8BFF8FFFEACA6DCE6417E0C95F" author="Saylor L. W." box="[547,703,1857,1883]" pageId="4" pageNumber="98" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
</taxonomicName>
, holotype, ♀ (Type No. 7904; CAS). –
<emphasis id="B956FA68FF8FFFEACCE1CE641182C95F" bold="true" box="[1199,1245,1857,1883]" pageId="4" pageNumber="98">E–I</emphasis>
.
<taxonomicName id="4C225DF9FF8FFFEACCBECE64140CC97A" authority="Saylor, 1946" authorityName="L. W. Saylor" authorityYear="1946" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="98" phylum="Arthropoda" rank="species" species="biciliata">
<emphasis id="B956FA68FF8FFFEACCBECE641028C95F" box="[1264,1399,1857,1883]" italics="true" pageId="4" pageNumber="98">A. biciliata</emphasis>
<bibRefCitation id="EFB35B8BFF8FFFEAC8F3CE41140CC97A" author="Saylor L. W." box="[189,339,1892,1918]" pageId="4" pageNumber="98" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
</taxonomicName>
, holotype, ♂ (Type No. 7907; CAS). –
<emphasis id="B956FA68FF8FFFEACB55CE411611C97A" bold="true" box="[795,846,1892,1918]" pageId="4" pageNumber="98">J–K</emphasis>
.
<taxonomicName id="4C225DF9FF8FFFEACB13CE4111D7C97A" authority="Saylor, 1946" authorityName="L. W. Saylor" authorityYear="1946" box="[861,1160,1892,1918]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="98" phylum="Arthropoda" rank="species" species="bogotana">
<emphasis id="B956FA68FF8FFFEACB13CE4116B4C97A" box="[861,1003,1892,1918]" italics="true" pageId="4" pageNumber="98">A. bogotana</emphasis>
<bibRefCitation id="EFB35B8BFF8FFFEACBBCCE4111D7C97A" author="Saylor L. W." box="[1010,1160,1892,1918]" pageId="4" pageNumber="98" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
</taxonomicName>
, paratype, ♂ (CAS). –
<emphasis id="B956FA68FF8FFFEAC89BCEA21453C9A5" bold="true" box="[213,268,1927,1953]" pageId="4" pageNumber="98">L–N</emphasis>
.
<taxonomicName id="4C225DF9FF8FFFEAC953CEA214F0C9A5" authorityName="L. W. Saylor" authorityYear="1946" box="[285,431,1927,1953]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="98" phylum="Arthropoda" rank="species" species="bogotana">
<emphasis id="B956FA68FF8FFFEAC953CEA214F0C9A5" box="[285,431,1927,1953]" italics="true" pageId="4" pageNumber="98">A. bogotana</emphasis>
</taxonomicName>
, holotype, ♂ (Type No. 7909; CAS).
<emphasis id="B956FA68FF8FFFEACB36CEA216D2C9A5" bold="true" box="[888,909,1927,1953]" pageId="4" pageNumber="98">A</emphasis>
. Head and pronotum, dorsal view.
<emphasis id="B956FA68FF8FFFEACD78CEA21017C9A5" bold="true" box="[1334,1352,1927,1953]" pageId="4" pageNumber="98">B</emphasis>
,
<emphasis id="B956FA68FF8FFFEACD16CEA21030C9A5" bold="true" box="[1368,1391,1927,1953]" pageId="4" pageNumber="98">G</emphasis>
,
<emphasis id="B956FA68FF8FFFEAC8F3CE8F158EC9C0" bold="true" box="[189,209,1962,1988]" pageId="4" pageNumber="98">L</emphasis>
. Labels.
<emphasis id="B956FA68FF8FFFEAC974CE8F14C1C9C0" bold="true" box="[314,414,1962,1988]" pageId="4" pageNumber="98">C, H, M</emphasis>
. Habitus, dorsal view.
<emphasis id="B956FA68FF8FFFEACAE6CE8F17A6C9C0" bold="true" box="[680,761,1962,1988]" pageId="4" pageNumber="98">D, I, N</emphasis>
. Habitus, lateral view.
<emphasis id="B956FA68FF8FFFEACC48CE8F1169C9C0" bold="true" box="[1030,1078,1962,1988]" pageId="4" pageNumber="98">E, J</emphasis>
. Parameres, dorsal view.
<emphasis id="B956FA68FF8FFFEACD11CE8F158BC9EC" bold="true" pageId="4" pageNumber="98">F, K</emphasis>
. Aedeagus, lateral view. Scale bars: A, E–F, J–K = 0.5 mm; C–D, H–I, M–N = 1 mm.
</paragraph>
</caption>
<paragraph id="8B9D267AFF8EFFEBC8F3C82C1483CF26" blockId="5.[189,476,264,291]" box="[189,476,264,291]" pageId="5" pageNumber="99">
<heading id="D0D59116FF8EFFEBC8F3C82C1483CF26" bold="true" box="[189,476,264,291]" fontSize="11" level="3" pageId="5" pageNumber="99" reason="3">
<emphasis id="B956FA68FF8EFFEBC8F3C82C1483CF26" bold="true" box="[189,476,264,291]" pageId="5" pageNumber="99">
<typeStatus id="549998D8FF8EFFEBC8F3C82C1470CF27" box="[189,303,265,291]" pageId="5" pageNumber="99" type="holotype">Holotype</typeStatus>
redescription
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF8EFFEBC8F3C812149DCB12" pageId="5" pageNumber="99" type="description">
<paragraph id="8B9D267AFF8EFFEBC8F3C81217A1CF56" blockId="5.[189,766,311,338]" box="[189,766,311,338]" pageId="5" pageNumber="99">
MEASUREMENTS. Length:
<quantity id="4CDA8B9FFF8EFFEBC9A8C8121765CF56" box="[486,570,311,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="99" unit="mm" value="11.0">11 mm</quantity>
. Width:
<quantity id="4CDA8B9FFF8EFFEBCAD3C81217A5CF56" box="[669,762,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="99" unit="mm" value="5.6">5.6 mm</quantity>
.
</paragraph>
<paragraph id="8B9D267AFF8EFFEBC8F3C85C11C9CC25" blockId="5.[189,1399,377,545]" pageId="5" pageNumber="99">HEAD. Reddish-brown, surface with upright setae. Labroclypeus, shape of anterior margin convex, posteriorly with sparse punctation. Ocular canthus with terminal setae, ratio to eye: 1/4. Eyes, ratio diameter/interocular width: 0.55. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club/remaining antennomeres: 1/1; ratio third vs fourth antennomere: 1/4; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1/5.</paragraph>
<paragraph id="8B9D267AFF8EFFEBC8F3CB6D162CCD32" blockId="5.[189,1399,584,822]" pageId="5" pageNumber="99">PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, reddishbrown, widest at base, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and smooth (not serrated), antero-lateral angle dorso-ventrally not flattened, posterolateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddishbrown, surface opaque, microsculpture absent, punctation on intervals present and dense, diameter of strial and interval punctation equal, posterior edge straight.</paragraph>
<paragraph id="8B9D267AFF8EFFEBC8F3CA7B17BECD9F" blockId="5.[189,1398,862,924]" pageId="5" pageNumber="99">ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite.</paragraph>
<paragraph id="8B9D267AFF8EFFEBC8F3CAE61172CAD1" blockId="5.[189,1399,963,1237]" pageId="5" pageNumber="99">LEGS. Ratio of length of metepisternum/metacoxa: 1/0.77. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, ratio of width/length: 1/2.69, dorsal margin not carinated, ventral margin not serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur 1/2. Dorsal surface of tarsi glabrous and without punctures. Metatarsi, length of first protarsomere smaller than second and third combined, ventral margin not serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute.</paragraph>
<paragraph id="8B9D267AFF8EFFEBC8F3CDD9149DCB12" blockId="5.[189,450,1276,1302]" box="[189,450,1276,1302]" pageId="5" pageNumber="99">
AEDEAGUS.
<figureCitation id="13193AFFFF8EFFEBC906CDD914E1CB12" box="[328,446,1276,1302]" captionStart="Fig" captionStartId="4.[189,232,1856,1882]" captionTargetBox="[244,1343,265,1819]" captionTargetId="figure-11@4.[244,1343,265,1819]" captionTargetPageId="4" captionText="Fig. 1. A–D. Astaena abaca Saylor, 1946, holotype, ♀ (Type No. 7904; CAS). – E–I. A. biciliata Saylor, 1946, holotype, ♂ (Type No. 7907; CAS). – J–K. A. bogotana Saylor, 1946, paratype, ♂ (CAS). – L–N. A. bogotana, holotype, ♂ (Type No. 7909; CAS). A. Head and pronotum, dorsal view. B, G, L. Labels. C, H, M. Habitus, dorsal view. D, I, N. Habitus, lateral view. E, J. Parameres, dorsal view. F, K. Aedeagus, lateral view. Scale bars: A, E–F, J–K = 0.5 mm; C–D, H–I, M–N = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5451749" httpUri="https://zenodo.org/record/5451749/files/figure.png" pageId="5" pageNumber="99">Fig. 1E–F</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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