119 lines
14 KiB
XML
119 lines
14 KiB
XML
<document id="BD84DABD8E858F924356554D70587BC3" ID="10.11646/zootaxa.4268.4.9SLASH11076" ID-CLB-Dataset="33020" ID-DOI="10.11646/zootaxa.4268.4.9" ID-GBIF-Dataset="287e7161-50ad-4ba2-83bb-4a458f5e0514" ID-ISSN="1175-5326" ID-Zenodo-Dep="580997" ID-ZooBank="FC8A0118-20FD-4829-BEAC-D1FAF107D310" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495098681352" checkinUser="plazi" docAuthor="Makarkin, Vladimir N." docDate="2017" docId="038AC01AFFC2EF0560818B8A81E988AA" docLanguage="en" docName="zootaxa.4268.4.9SLASH11076.pdf" docOrigin="Zootaxa 4268 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudosmylidia relicta Makarkin, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="583" masterDocId="FFB3B862FFC3EF0760168F48853A895E" masterDocTitle="A remarkable new genus of Protosmylinae (Neuroptera: Osmylidae) from late Eocene Florissant, Colorado" masterLastPageNumber="587" masterPageNumber="581" pageNumber="582" updateTime="1698439719004" updateUser="ExternalLinkService">
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<mods:mods id="70D8D105A21E72F887B22479F3C4A522" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="331C7D7743F4F7D632B7F4CCCF9756FF">
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<mods:title id="6D7C14CFE839DC97ACD760122F3C24E1">A remarkable new genus of Protosmylinae (Neuroptera: Osmylidae) from late Eocene Florissant, Colorado</mods:title>
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</mods:titleInfo>
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<mods:name id="48F5C170A4216B2834FA6682DF0E8B30" type="personal">
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<mods:role id="E00DE8E9B3977DF20F8F9CF7D597F2B1">
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<mods:roleTerm id="3B6C1439788EE8FD9F9B4BB0EB82A9DC">Author</mods:roleTerm>
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<mods:namePart id="79AFF731F6B5BFE91F67F8B9EF26FC42">Makarkin, Vladimir N.</mods:namePart>
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<mods:typeOfResource id="5F3A5AECA47BAA7C0E9354803AC79D74">text</mods:typeOfResource>
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<mods:titleInfo id="191F167A82B3C272FF53805D3C879FE8">
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<mods:title id="41B5DA1E076411165866EB955C878F64">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5C15661B9783E8A1567B6BC5AD683DD4">
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<mods:date id="1DD51DD12D11E9C27B48A5A660BF8B27">2017</mods:date>
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<mods:detail id="5702DD07E7D2ED5E095AF08B0D8FB337" type="volume">
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<mods:number id="522AB269CD284C7670002648CE4A83C2">4268</mods:number>
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<mods:detail id="3AF96050865B927E642F9D8A0B942EC5" type="issue">
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<mods:number id="2E4B70EA669517404673A2374F33ADDE">4</mods:number>
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<mods:start id="CE79EBDE7E2A0E0DAAFBBD3560465614">581</mods:start>
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<mods:classification id="9E7D5D91318E3FC9C9F3764923530191">journal article</mods:classification>
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<mods:identifier id="5D39C057E7F8B28AA4081D04A79CA3C0" type="CLB-Dataset">33020</mods:identifier>
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<mods:identifier id="9C91F010B3DD3E616F88D4E6915D3ACC" type="DOI">10.11646/zootaxa.4268.4.9</mods:identifier>
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<mods:identifier id="E5DB52734A72219AFE1247FC77F6E95E" type="GBIF-Dataset">287e7161-50ad-4ba2-83bb-4a458f5e0514</mods:identifier>
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<mods:identifier id="654D33D564E38688FCCD7C12CDA6374E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="692B606F235805341C2A3F01376B7D9D" type="Zenodo-Dep">580997</mods:identifier>
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<mods:identifier id="5195DA227401B45A0B1588789AA1C390" type="ZooBank">FC8A0118-20FD-4829-BEAC-D1FAF107D310</mods:identifier>
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</mods:mods>
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<treatment id="038AC01AFFC2EF0560818B8A81E988AA" ID-DOI="http://doi.org/10.5281/zenodo.6041691" ID-GBIF-Taxon="130810328" ID-Zenodo-Dep="6041691" LSID="urn:lsid:plazi:treatment:038AC01AFFC2EF0560818B8A81E988AA" httpUri="http://treatment.plazi.org/id/038AC01AFFC2EF0560818B8A81E988AA" lastPageId="2" lastPageNumber="583" pageId="1" pageNumber="582">
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<subSubSection id="C3392287FFC2EF0660818B8A85C38DA0" pageId="1" pageNumber="582" type="nomenclature">
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<paragraph id="8B9C710CFFC2EF0660818B8A87368D83" blockId="1.[151,524,1218,1278]" box="[151,524,1218,1245]" pageId="1" pageNumber="582">
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<heading id="D0D4C660FFC2EF0660818B8A87368D83" bold="true" box="[151,524,1218,1245]" fontSize="11" level="1" pageId="1" pageNumber="582" reason="1">
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<emphasis id="B957AD1EFFC2EF0660818B8A87368D83" bold="true" box="[151,524,1218,1245]" pageId="1" pageNumber="582">
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<taxonomicName id="4C230A8FFFC2EF0660818B8A849D8D82" ID-CoL="4PBP5" box="[151,423,1218,1244]" class="Insecta" family="Osmylidae" genus="Pseudosmylidia" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="relicta" status="sp. nov.">
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<emphasis id="B957AD1EFFC2EF0660818B8A849D8D82" bold="true" box="[151,423,1218,1244]" italics="true" pageId="1" pageNumber="582">Pseudosmylidia relicta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2641065FFC2EF0661B88B8B87368D83" box="[430,524,1219,1245]" pageId="1" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9C710CFFC2EF0660818BAD85C38DA0" blockId="1.[151,524,1218,1278]" box="[151,249,1253,1278]" pageId="1" pageNumber="582">
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<figureCitation id="13186D89FFC2EF0660818BAD85E68DA0" box="[151,220,1253,1278]" captionStart="FIGURE 1" captionStartId="2.[151,249,1091,1113]" captionTargetBox="[151,1436,559,1052]" captionTargetId="figure@2.[151,1436,559,1052]" captionTargetPageId="2" captionText="FIGURE 1. The holotype UCM 85899 of Pseudosmylidia relicta gen. et sp. nov. Scale bar = 2 mm." httpUri="https://zenodo.org/record/580998/files/figure.png" pageId="1" pageNumber="582">Figs 1</figureCitation>
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,
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<figureCitation id="13186D89FFC2EF0660FC8BAE85C38DA0" box="[234,249,1254,1278]" captionStart="FIGURE 2" captionStartId="3.[151,249,1675,1697]" captionTargetBox="[172,1416,193,1643]" captionTargetId="figure@3.[172,1416,193,1643]" captionTargetPageId="3" captionText="FIGURE 2. The holotype UCM 85899 of Pseudosmylidia relicta gen. et sp. nov. A, photograph of wings taken with a polarized filter. B, forewing venation. C, portion of wings showing the hind wing CuP and AA 1 (hm, hind margin of hind wing). All converted to standard view, with apex to the right. Scale bars = 1 mm (A, B), 0.5 mm (C)." httpUri="https://zenodo.org/record/580999/files/figure.png" pageId="1" pageNumber="582">2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3392287FFC2EF0560818A6584E189A6" lastPageId="2" lastPageNumber="583" pageId="1" pageNumber="582" type="description">
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<paragraph id="8B9C710CFFC2EF0660818A6584D18CD0" blockId="1.[151,1437,1325,1962]" pageId="1" pageNumber="582">
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<emphasis id="B957AD1EFFC2EF0660818A6584138C18" bold="true" box="[151,297,1325,1350]" pageId="1" pageNumber="582">Description.</emphasis>
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Head visible in lateral view, with vertex slightly raised, eyes relatively small; antennae probably incompletely preserved, with scapus not enlarged. Prothorax relatively short. Other parts of body, legs poorly preserved (details not visible).
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</paragraph>
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<paragraph id="8B9C710CFFC2EF0660D18AD184D68EF4" blockId="1.[151,1437,1325,1962]" pageId="1" pageNumber="582">
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Forewing oval,
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<quantity id="4CDBDCE9FFC2EF0661688AD284DC8CEF" box="[382,486,1434,1458]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="582" unit="mm" value="12.1">12.1 mm</quantity>
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long,
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<quantity id="4CDBDCE9FFC2EF0662228AD287B78CEF" box="[564,653,1434,1458]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="582" unit="mm" value="4.7">4.7 mm</quantity>
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wide. Costal space broad, broadest approximately at proximal 1/5; rather abruptly narrowed basad, radually narrowed distad. Subcostal veinlets moderately widely spaced; simple proximally, shallowly forked distally (some veinlets poorly preserved). ScP, RA fused far from wing apex. ScP+RA entering margin before from wing apex. Veinlets of ScP+RA poorly preserved; at least forked once. Subcostal space poorly preserved, narrow. RA space moderately broad, with five detected crossveins (more crossveins possible). RP with nine branches. RP1 originated rather close to origin of RP. All branches of RP shallowly forked (far distad outer gradate series of crossveins), mostly dichotomous. M forked at level of origin of RP2. MA, MP dichotomously forked slightly distad outer gradate series of crossveins. Cu rather long, dividing into CuA, CuP relatively far from wing base. Anterior trace of CuA shallowly trident-like forked, with four shallowlyforked pectinate branches (termination of two proximal-most branches poorly visible). CuP few pectinate, with two shallowly forked branches (terminations poorly preserved). AA1 pectinate, with four oblique branched; all probably shallowly forked. AA2 poorly preserved, probably few branched. AA3 not detected. All preserved crossveins in radial to mediocubital spaces arranged in four gradate series. Three crossveins detected between CuA, CuP. Trichosors prominent along outer wing margin; basally indistinct or absent. Nygmata not detected. Color pattern probably absent.
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</paragraph>
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<paragraph id="8B9C710CFFC1EF0560D18FDF84E189A6" blockId="2.[151,1437,151,501]" pageId="2" pageNumber="583">
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Hind wing very poorly preserved, ca.
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<quantity id="4CDBDCE9FFC1EF0562838FDF861289F1" box="[661,808,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="2" pageNumber="583" unit="mm" value="10.75" valueMax="11.0" valueMin="10.5">10.5–11 mm</quantity>
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long, 4.0 mm wide. RP with nine branches. Radial crossveins mostly arranged in three gradate series. CuP short, simple or forked once (poorly preserved). AA1 probably pectinate (
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<figureCitation id="13186D89FFC1EF0561608F97848389A6" box="[374,441,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,249,1675,1697]" captionTargetBox="[172,1416,193,1643]" captionTargetId="figure@3.[172,1416,193,1643]" captionTargetPageId="3" captionText="FIGURE 2. The holotype UCM 85899 of Pseudosmylidia relicta gen. et sp. nov. A, photograph of wings taken with a polarized filter. B, forewing venation. C, portion of wings showing the hind wing CuP and AA 1 (hm, hind margin of hind wing). All converted to standard view, with apex to the right. Scale bars = 1 mm (A, B), 0.5 mm (C)." httpUri="https://zenodo.org/record/580999/files/figure.png" pageId="2" pageNumber="583">Fig. 2</figureCitation>
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C).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3392287FFC1EF0560D18E4C802A88D9" pageId="2" pageNumber="583" type="materials_examined">
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<paragraph id="8B9C710CFFC1EF0560D18E4C8188883A" blockId="2.[151,1437,151,501]" pageId="2" pageNumber="583">
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<emphasis id="B957AD1EFFC1EF0560D18E4C84438843" bold="true" box="[199,377,260,285]" pageId="2" pageNumber="583">Type material.</emphasis>
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Holotype UCM 85899 (only part), likely collected by T.D.A. Cockerell, and deposited in UCM. A nearly complete, rather poorly preserved specimen with all four wings overlapping. Written on the rock is “W14”, indicating one of Cockerell’s collecting localities at Florissant (T. Karim, pers. com.).
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</paragraph>
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<paragraph id="8B9C710CFFC1EF0560D18E27802A88D9" blockId="2.[151,1437,151,501]" box="[199,1296,367,392]" pageId="2" pageNumber="583">
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<materialsCitation id="3B4B7B51FFC1EF0560D18E27802A88D9" ID-GBIF-Occurrence="1503373118" box="[199,1296,367,392]" collectionCode="UCM" country="United States of America" location="Florissant" pageId="2" pageNumber="583" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
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<emphasis id="B957AD1EFFC1EF0560D18E2784CD88D6" bold="true" box="[199,503,367,392]" pageId="2" pageNumber="583">
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<typeStatus id="5498CFAEFFC1EF0560D18E27843888D6" box="[199,258,367,392]" pageId="2" pageNumber="583">Type</typeStatus>
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locality and horizon
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</emphasis>
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.
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<collectingCountry id="F334319CFFC1EF0562128E38876F88D6" box="[516,597,368,392]" name="United States of America" pageId="2" pageNumber="583">U.S.A.</collectingCountry>
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:
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<collectingRegion id="49E7BFEEFFC1EF0562768E2787F788D6" box="[608,717,367,392]" country="United States of America" name="Colorado" pageId="2" pageNumber="583">Colorado</collectingRegion>
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:
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<location id="8EFC27D7FFC1EF0562C18E27867F88D6" LSID="urn:lsid:plazi:treatment:038AC01AFFC2EF0560818B8A81E988AA:8EFC27D7FFC1EF0562C18E27867F88D6" box="[727,837,367,392]" country="United States of America" name="Florissant" pageId="2" pageNumber="583" stateProvince="Colorado">Florissant</location>
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(
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<collectionCode id="ED32E9C9FFC1EF0563428E3886AC88D6" box="[852,918,368,392]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="2" pageNumber="583">UCM</collectionCode>
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locality No. 83014); late Eocene.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3392287FFC1EF0560D18EDC803788F3" box="[199,1293,404,429]" pageId="2" pageNumber="583" type="etymology">
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<paragraph id="8B9C710CFFC1EF0560D18EDC803788F3" blockId="2.[151,1437,151,501]" box="[199,1293,404,429]" pageId="2" pageNumber="583">
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<emphasis id="B957AD1EFFC1EF0560D18EDC847488F3" bold="true" box="[199,334,404,429]" pageId="2" pageNumber="583">Etymology.</emphasis>
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From the Latin
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<emphasis id="B957AD1EFFC1EF0562158EDC876F88F2" box="[515,597,404,428]" italics="true" pageId="2" pageNumber="583">relictus</emphasis>
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, -
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<emphasis id="B957AD1EFFC1EF05627B8EDD874188F2" box="[621,635,405,428]" italics="true" pageId="2" pageNumber="583">a</emphasis>
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, -
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<emphasis id="B957AD1EFFC1EF0562858EDD878F88F2" box="[659,693,405,428]" italics="true" pageId="2" pageNumber="583">um</emphasis>
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- abandoned, referring to relict nature of this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3392287FFC1EF0560D18EFF81E988AA" pageId="2" pageNumber="583" type="discussion">
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<paragraph id="8B9C710CFFC1EF0560D18EFF81E988AA" blockId="2.[151,1437,151,501]" pageId="2" pageNumber="583">
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<emphasis id="B957AD1EFFC1EF0560D18EFF8401888E" bold="true" box="[199,315,439,464]" pageId="2" pageNumber="583">Remarks.</emphasis>
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Only one forewing is rather distinctly preserved and allows a detailed drawing. The other forewing and the hind wings are poorly preserved. Many details of the hind wings are difficult to separate.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |