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<mods:title id="C5A1D2E5C9F482AB677F7752FE21D82B">Notes on Southeast Asian Stingless Bees of the Genus Tetragonula (Hymenoptera: Apidae), with the Description of a New Species from Thailand</mods:title>
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<taxonomicName id="4C204D694D721453FEE57B81FCE6E58A" ID-CoL="55NCS" authority="Engel &amp; Michener &amp; Boontop, 2017" authorityName="Engel &amp; Michener &amp; Boontop" authorityYear="2017" box="[347,820,395,422]" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="malaipanae" status="sp. nov." subGenus="Tetragonula">
<emphasis id="B954EAF84D721453FEE57B81FE3FE58A" bold="true" box="[347,493,395,422]" italics="true" pageId="4" pageNumber="5">Tetragonula</emphasis>
(
<emphasis id="B954EAF84D721453FE417B81FD43E58A" bold="true" box="[511,657,395,422]" italics="true" pageId="4" pageNumber="5">Tetragonula</emphasis>
)
<emphasis id="B954EAF84D721453FD1D7B81FCE6E58A" bold="true" box="[675,820,395,422]" italics="true" pageId="4" pageNumber="5">malaipanae</emphasis>
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<taxonomicNameLabel id="A26757834D721453FCFC7B86FC1EE589" box="[834,972,395,422]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
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<heading id="D0D781864D721453FDF17BC9FD0AE5F1" box="[591,728,451,477]" centered="true" fontSize="36" level="3" pageId="4" pageNumber="5" reason="7">
<figureCitation id="131B2A6F4D721453FDF17BC9FD0AE5F1" box="[591,728,451,477]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="5.[110,196,1510,1531]" captionStartId-1="6.[108,195,1528,1549]" captionStartId-2="7.[109,195,1559,1580]" captionTargetBox-0="[127,1200,230,1474]" captionTargetBox-1="[114,1200,235,1492]" captionTargetBox-2="[117,1206,225,1530]" captionTargetId-0="figure-6@5.[110,1210,224,1484]" captionTargetId-1="figure-9@6.[108,1212,224,1501]" captionTargetId-2="figure-6@7.[108,1212,224,1533]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 1. Worker of Tetragonula malaipanae, new species. A. Lateral habitus. B. Dorsal habitus. C. Facial view." captionText-1="FIGURE 2. Male of Tetragonula malaipanae, new species (specimen with metasoma dissected). A. Lateral habitus. B. Dorsal habitus. C. Facial view." captionText-2="FIGURE 3. Male of Tetragonula malaipanae, new species. A. Outer view of metatibia and metabasitarsis. B. Dorsal view of T7 (arrow indicates the position of the spiracle on the side of the sclerite). C. Ventral view of S5. D. Ventral view of S6. E. Ventral view of S7. F. Male genitalia (dorsal view at left, ventral view at right), distal part of membranous aedeagus omitted. G. Midventral membranous process. H. Male genitalia in right lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.5062282" figureDoi-1="http://doi.org/10.5281/zenodo.5062284" figureDoi-2="http://doi.org/10.5281/zenodo.5062286" httpUri-0="https://zenodo.org/record/5062282/files/figure.png" httpUri-1="https://zenodo.org/record/5062284/files/figure.png" httpUri-2="https://zenodo.org/record/5062286/files/figure.png" pageId="4" pageNumber="5">Figures 13</figureCitation>
</heading>
</paragraph>
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<subSubSection id="C33A65614D721453FF227BF1FDDFE7D7" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8B9F36EA4D721453FF227BF1FDDFE7D7" blockId="4.[108,1214,507,1766]" pageId="4" pageNumber="5">
DIAGNOSIS: The species generally agrees with the subgeneric characterization as presented by
<bibRefCitation id="EFB14B1B4D721453FF31782BFE86E617" author="Sakagami, S. F." box="[143,340,545,571]" pageId="4" pageNumber="5" pagination="165 - 247" refId="ref8490" refString="Sakagami, S. F. 1978. Tetragonula stingless bees of the continental Asia and Sri Lanka (Hymenoptera, Apidae). Journal of the Faculty of Science, Hokkaido University, Series 6, Zoology 21 (2): 165 - 247, + 1 pl. [pl. v]." type="journal article" year="1978">Sakagami (1978)</bibRefCitation>
. Worker dark but paler than
<taxonomicName id="4C204D694D721453FD1E782BFBEBE617" authority="(Smith, 1857)" baseAuthorityName="Smith" baseAuthorityYear="1857" box="[672,1081,544,571]" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="laeviceps">
<emphasis id="B954EAF84D721453FD1E782BFC40E617" box="[672,914,545,571]" italics="true" pageId="4" pageNumber="5">Tetragonula laeviceps</emphasis>
(
<bibRefCitation id="EFB14B1B4D721453FC1F782BFBE3E617" author="Smith, F." box="[929,1073,544,571]" pageId="4" pageNumber="5" pagination="42 - 88" refId="ref8775" refString="Smith, F. 1857. Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace. Journal of the Proceedings of the Linnean Society, Zoology 2 (6): 42 - 88, + 2 pls." type="book chapter" year="1857">Smith, 1857</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C204D694D721453FBFA782BFEF9E64C" authority="(Schwarz, 1939)" baseAuthorityName="Schwarz" baseAuthorityYear="1939" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pagdeni">
<emphasis id="B954EAF84D721453FBFA782BFB84E616" box="[1092,1110,545,570]" italics="true" pageId="4" pageNumber="5">T</emphasis>
.
<emphasis id="B954EAF84D721453FBDD782BFB6EE617" box="[1123,1212,545,571]" italics="true" pageId="4" pageNumber="5">pagdeni</emphasis>
(
<bibRefCitation id="EFB14B1B4D721453FFCA784CFEF1E64C" author="Schwarz, H. F." box="[116,291,582,608]" pageId="4" pageNumber="5" pagination="83 - 141" refId="ref8637" refString="Schwarz, H. F. 1939. The Indo-Malayan species of Trigona. Bulletin of the American Museum of Natural History 76 (3): 83 - 141." type="journal article" year="1939">Schwarz, 1939</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C204D694D721453FED0784DFCD6E64C" authority="(Sakagami, 1978)" baseAuthorityName="Sakagami" baseAuthorityYear="1978" box="[366,772,582,609]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pagdeniformis">
<emphasis id="B954EAF84D721453FED0784DFEACE64C" box="[366,382,583,608]" italics="true" pageId="4" pageNumber="5">T</emphasis>
.
<emphasis id="B954EAF84D721453FE30784CFDE2E64C" box="[398,560,582,608]" italics="true" pageId="4" pageNumber="5">pagdeniformis</emphasis>
(
<bibRefCitation id="EFB14B1B4D721453FDFE784CFD2EE64C" author="Sakagami, S. F." box="[576,764,582,608]" pageId="4" pageNumber="5" pagination="165 - 247" refId="ref8490" refString="Sakagami, S. F. 1978. Tetragonula stingless bees of the continental Asia and Sri Lanka (Hymenoptera, Apidae). Journal of the Faculty of Science, Hokkaido University, Series 6, Zoology 21 (2): 165 - 247, + 1 pl. [pl. v]." type="journal article" year="1978">Sakagami, 1978</bibRefCitation>
)
</taxonomicName>
(figs. 1A, 1B), clypeus yellowish (fig. 1C), legs partly or wholly red brown or testaceous (fig. 1A). Size similar to that of
<taxonomicName id="4C204D694D721453FB897866FEA1E687" authority="(Cameron, 1901)" baseAuthorityName="Cameron" baseAuthorityYear="1901" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="testaceitarsis">
<emphasis id="B954EAF84D721453FB897866FB9AE6A9" box="[1079,1096,620,645]" italics="true" pageId="4" pageNumber="5">T</emphasis>
.
<emphasis id="B954EAF84D721453FBE87866FF72E686" italics="true" pageId="4" pageNumber="5">testaceitarsis</emphasis>
(
<bibRefCitation id="EFB14B1B4D721453FF0F789BFEB9E687" author="Cameron, P." box="[177,363,657,683]" pageId="4" pageNumber="5" pagination="16 - 44" refId="ref7054" refString="Cameron, P. 1901. On the Hymenoptera collected during the &quot; Skeat Expedition &quot; to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London 2 (1): 16 - 44." type="journal article" year="1901">Cameron, 1901</bibRefCitation>
)
</taxonomicName>
(likely senior synonym of
<taxonomicName id="4C204D694D721453FD08789BFCFBE687" authorityName="Cockerell" authorityYear="1918" box="[694,809,657,683]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="valdezi">
<emphasis id="B954EAF84D721453FD08789BFD1AE686" box="[694,712,657,682]" italics="true" pageId="4" pageNumber="5">T</emphasis>
.
<emphasis id="B954EAF84D721453FD66789BFCFBE687" box="[728,809,657,683]" italics="true" pageId="4" pageNumber="5">valdezi</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFB14B1B4D721453FC87789BFC29E687" author="Cockerell, T. D. A." box="[825,1019,656,683]" pageId="4" pageNumber="5" pagination="384 - 390" refId="ref7192" refString="Cockerell, T. D. A. 1918. Descriptions and records of bees - LXXX. Annals and Magazine of Natural History (9) 2 (10): 384 - 390." type="journal article" year="1918">Cockerell, 1918</bibRefCitation>
]), larger than
<taxonomicName id="4C204D694D721453FB1A789BFF03E6FC" baseAuthorityName="Smith" baseAuthorityYear="1857" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="laeviceps">
<emphasis id="B954EAF84D721453FB1A789BFB64E686" box="[1188,1206,657,682]" italics="true" pageId="4" pageNumber="5">T</emphasis>
.
<emphasis id="B954EAF84D721453FFD278BCFF03E6FC" box="[108,209,694,720]" italics="true" pageId="4" pageNumber="5">laeviceps</emphasis>
</taxonomicName>
(as restricted by
<bibRefCitation id="EFB14B1B4D721453FE1D78BDFCF0E6FC" author="Rasmussen, C. &amp; C. D. Michener" box="[419,802,694,721]" pageId="4" pageNumber="5" pagination="129 - 133" refId="ref8384" refString="Rasmussen, C., and C. D. Michener. 2010. The identity and neotype of Trigona laeviceps Smith (Hymenoptera: Apidae). Journal of the Kansas Entomological Society 83 (2): 129 - 133." type="journal article" year="2010">Rasmussen and Michener, 2010</bibRefCitation>
),
<taxonomicName id="4C204D694D721453FC8978BDFC61E6FC" box="[823,947,694,721]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pagdeni">
<emphasis id="B954EAF84D721453FC8978BDFC9BE6FC" box="[823,841,695,720]" italics="true" pageId="4" pageNumber="5">T</emphasis>
.
<emphasis id="B954EAF84D721453FCE778BCFC61E6FC" box="[857,947,694,720]" italics="true" pageId="4" pageNumber="5">pagdeni</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C204D694D721453FC4678BDFB6EE6FC" baseAuthorityName="Sakagami" baseAuthorityYear="1978" box="[1016,1212,694,721]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pagdeniformis">
<emphasis id="B954EAF84D721453FC4678BDFBD8E6FC" box="[1016,1034,695,720]" italics="true" pageId="4" pageNumber="5">T</emphasis>
.
<emphasis id="B954EAF84D721453FBA478BCFB6EE6FC" box="[1050,1212,694,720]" italics="true" pageId="4" pageNumber="5">pagdeniformis</emphasis>
</taxonomicName>
(see Discussion, below). Mesoscutum with pale pubescence not or weakly banded (fig. 1B), sublateral bare bands not or scarcely recognizable. Setae of body largely pale, but those of vertex, especially behind ocelli, blackish or sometimes dusky. Mean metatibial length
<quantity id="4CD89B0F4D721453FBF0792CFB65E76D" box="[1102,1207,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="4" pageNumber="5" unit="mm" value="1.81">1.81 mm</quantity>
, mean forewing length including tegula
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, mean head width
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. Male generally colored as in worker (fig. 2), with S5 lacking gradulus (fig. 3C), apical process of S6 long and parallel sided (fig. 3D), S7 and T7 sharply pointed apically (figs. 3B, 3E); genitalia with penis valves and gonostyli directed posteriorly (figs. 3F, 3H), sclerotization probably representing gonobase across base (figs. 3F, 3H).
</paragraph>
</subSubSection>
<subSubSection id="C33A65614D72145EFF227E0CFE08E03C" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5" type="description">
<paragraph id="8B9F36EA4D721453FF227E0CFE5DE229" blockId="4.[108,1214,507,1766]" pageId="4" pageNumber="5">
DESCRIPTION:
<emphasis id="B954EAF84D721453FEF47E0CFE7EE00C" bold="true" box="[330,428,1030,1056]" pageId="4" pageNumber="5">Worker.</emphasis>
Body length
<quantity id="4CD89B0F4D721453FDF47E0CFD61E00D" box="[586,691,1030,1057]" metricMagnitude="-3" metricUnit="m" metricValue="3.92" pageId="4" pageNumber="5" unit="mm" value="3.92">3.92 mm</quantity>
(
<quantity id="4CD89B0F4D721453FD7C7E0CFCA3E00D" box="[706,881,1030,1057]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.7" metricValueMin="3.5" pageId="4" pageNumber="5" unit="mm" value="4.1" valueMax="4.7" valueMin="3.5">3.504.70 mm</quantity>
); forewing length, including tegula,
<quantity id="4CD89B0F4D721453FF7D7E21FEFFE06A" box="[195,301,1067,1094]" metricMagnitude="-3" metricUnit="m" metricValue="4.58" pageId="4" pageNumber="5" unit="mm" value="4.58">4.58 mm</quantity>
(
<quantity id="4CD89B0F4D721453FE807E21FE3DE06A" box="[318,495,1067,1094]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.7" metricValueMin="4.4" pageId="4" pageNumber="5" unit="mm" value="4.55" valueMax="4.7" valueMin="4.4">4.404.70 mm</quantity>
); head length from anterior margin of clypeus to summit of vertex, in facial view
<quantity id="4CD89B0F4D721453FEDF7E5BFE1AE047" box="[353,456,1105,1131]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="4" pageNumber="5" unit="mm" value="1.42">1.42 mm</quantity>
(
<quantity id="4CD89B0F4D721453FE687E5AFD53E047" box="[470,641,1104,1131]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="4" pageNumber="5" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.401.50 mm</quantity>
); head width
<quantity id="4CD89B0F4D721453FCA37E5BFC56E047" box="[797,900,1105,1131]" metricMagnitude="-3" metricUnit="m" metricValue="1.91" pageId="4" pageNumber="5" unit="mm" value="1.91">1.91 mm</quantity>
(1.752.00 mm); metatibia length
<quantity id="4CD89B0F4D721453FF007E7CFEFAE0BD" box="[190,296,1142,1169]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="4" pageNumber="5" unit="mm" value="1.82">1.82 mm</quantity>
(
<quantity id="4CD89B0F4D721453FE867E7CFE35E0BD" box="[312,487,1142,1169]" metricMagnitude="-3" metricUnit="m" metricValue="1.755" metricValueMax="1.88" metricValueMin="1.63" pageId="4" pageNumber="5" unit="mm" value="1.755" valueMax="1.88" valueMin="1.63">1.631.88 mm</quantity>
); forewing diagonal from base of vein M to base of crossvein r-rs at margin of pterostigma
<quantity id="4CD89B0F4D721453FE6B7E91FDEDE09A" box="[469,575,1179,1206]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="4" pageNumber="5" unit="mm" value="1.31">1.31 mm</quantity>
(
<quantity id="4CD89B0F4D721453FDEF7E91FCD3E09A" box="[593,769,1179,1206]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.38" metricValueMin="1.23" pageId="4" pageNumber="5" unit="mm" value="1.305" valueMax="1.38" valueMin="1.23">1.231.38 mm</quantity>
). Scape (excluding basal bulb) more than half as long as compound eye; flagellomere IV slightly broader than long. Ocelloccipital distance over half OD, ocellocular distance less than interocellar distance. Genal area widest at about upper third of compound eye, and narrower than compound eye in profile. Metatibia and metabasitarsus shaped as usual in this group (see
<bibRefCitation id="EFB14B1B4D721453FD4F7F3BFC79E167" author="Sakagami, S. F." box="[753,939,1329,1355]" pageId="4" pageNumber="5" pagination="165 - 247" refId="ref8490" refString="Sakagami, S. F. 1978. Tetragonula stingless bees of the continental Asia and Sri Lanka (Hymenoptera, Apidae). Journal of the Faculty of Science, Hokkaido University, Series 6, Zoology 21 (2): 165 - 247, + 1 pl. [pl. v]." type="journal article" year="1978">Sakagami, 1978</bibRefCitation>
: fig. 2); basal sericeous area at base of inner surface of metabasitarsus well defined, reaching to middle of metabasitarsus [variable, not attaining middle or extending beyond middle]. Metasoma short, about as wide as head, widest at T2 and T3, remaining terga progressively much narrower and largely telescoped into preceding segments. Cuticular surface polished and shining with few minute punctures or setal bases.
</paragraph>
<paragraph id="8B9F36EA4D72145FFF227C1BFCF5E6BA" blockId="4.[108,1214,507,1766]" lastBlockId="8.[108,1214,225,1708]" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">Head black, the following parts yellow testaceous [to reddish testaceous or pale yellowish brown]: clypeus except dusky upper and upper lateral margins [entire clypeus commonly yellowish], supraclypeal area [sometimes dusky or blackish], mandible except blackish apex and extreme base, and labrum. [Lower genal area and hypostomal area behind mandibular base sometimes weakly brownish, grading into black background]. Antenna yellowish [to reddish] testaceous, flagellum blackish [or dusky] on upper surface, on lower surface part of each flagellomere dusky [or not]. Thorax black except pronotum dark brown [to blackish], pronotal lobe dusky testaceous [to black], mesoscutellum brownish [or black], [mesepisternum sometimes brownish black], anterolateral part of mesepisternum brown [to black]. Legs (including coxae and trochanters) reddish brown, meso- and metatibiae and tarsi dark reddish brown [or rarely legs testaceous or legs blackish brown, sometimes only mid and hind legs dark, inner surface of metatibia and metatarsus and often basal part of metafemur reddish brown]. Tegula dusky testaceous. Wings clear, faintly dusky, veins and pterostigma blackish. Metasoma with T1 dusky brown [brown], anterior surface largely testaceous; T5 and T6 [sometimes on T6] testaceous [brown or dusky]; T2 and T4 [sometimes also T5] brownish black, posterior margins paler translucent [posterior part of T1 sometimes similarly dark]. [Sometimes, especially specimens from nests 6 and 9, with all terga testaceous, posterior ones darker than anterior ones]. Metasomal sterna testaceous [brown, sometimes slightly dusky].</paragraph>
<caption id="DF5F66624D731452FFD07FECFF76E236" ID-DOI="http://doi.org/10.5281/zenodo.5062282" ID-Zenodo-Dep="5062282" httpUri="https://zenodo.org/record/5062282/files/figure.png" pageId="5" pageNumber="6" startId="5.[110,196,1510,1531]" targetBox="[127,1200,230,1474]" targetPageId="5">
<paragraph id="8B9F36EA4D731452FFD07FECFF76E236" blockId="5.[110,1214,1508,1562]" pageId="5" pageNumber="6">
FIGURE 1. Worker of
<taxonomicName id="4C204D694D731452FEE67FEEFD98E1D0" authority="Engel &amp; Michener &amp; Boontop, 2017" authorityName="Engel &amp; Michener &amp; Boontop" authorityYear="2017" box="[344,586,1508,1532]" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="malaipanae" status="sp. nov.">
<emphasis id="B954EAF84D731452FEE67FEEFD98E1D0" box="[344,586,1508,1532]" italics="true" pageId="5" pageNumber="6">Tetragonula malaipanae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26757834D731452FDE97FEFFD01E1D0" box="[599,723,1508,1532]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="B954EAF84D731452FD617FEEFD2AE1D0" bold="true" box="[735,760,1508,1532]" pageId="5" pageNumber="6">A.</emphasis>
Lateral habitus.
<emphasis id="B954EAF84D731452FC1A7FEFFC69E1D1" bold="true" box="[932,955,1509,1533]" pageId="5" pageNumber="6">B.</emphasis>
Dorsal habitus.
<emphasis id="B954EAF84D731452FBDB7FEEFBAFE1D0" bold="true" box="[1125,1149,1508,1532]" pageId="5" pageNumber="6">C.</emphasis>
Facial view.
</paragraph>
</caption>
<caption id="DF5F66624D701451FFD27FF2FDC7E207" ID-DOI="http://doi.org/10.5281/zenodo.5062284" ID-Zenodo-Dep="5062284" httpUri="https://zenodo.org/record/5062284/files/figure.png" pageId="6" pageNumber="7" startId="6.[108,195,1528,1549]" targetBox="[114,1200,235,1492]" targetPageId="6">
<paragraph id="8B9F36EA4D701451FFD27FF2FDC7E207" blockId="6.[108,1212,1526,1579]" pageId="6" pageNumber="7">
FIGURE 2. Male of
<taxonomicName id="4C204D694D701451FE817FFCFDE6E222" authority="Engel &amp; Michener &amp; Boontop, 2017" authorityName="Engel &amp; Michener &amp; Boontop" authorityYear="2017" box="[319,564,1526,1550]" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="malaipanae" status="sp. nov.">
<emphasis id="B954EAF84D701451FE817FFCFDE6E222" box="[319,564,1526,1550]" italics="true" pageId="6" pageNumber="7">Tetragonula malaipanae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26757834D701451FDFD7FFCFD6CE222" box="[579,702,1526,1550]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
(specimen with metasoma dissected).
<emphasis id="B954EAF84D701451FBEA7FFCFBBFE222" bold="true" box="[1108,1133,1526,1550]" pageId="6" pageNumber="7">A.</emphasis>
Lateral habitus.
<emphasis id="B954EAF84D701451FF7C7C19FF0BE207" bold="true" box="[194,217,1555,1579]" pageId="6" pageNumber="7">B.</emphasis>
Dorsal habitus.
<emphasis id="B954EAF84D701451FE3F7C19FE4BE207" bold="true" box="[385,409,1555,1579]" pageId="6" pageNumber="7">C.</emphasis>
Facial view.
</paragraph>
</caption>
<caption id="DF5F66624D711450FFD37C1DFF35E28F" ID-DOI="http://doi.org/10.5281/zenodo.5062286" ID-Zenodo-Dep="5062286" httpUri="https://zenodo.org/record/5062286/files/figure.png" pageId="7" pageNumber="8" startId="7.[109,195,1559,1580]" targetBox="[117,1206,225,1530]" targetPageId="7">
<paragraph id="8B9F36EA4D711450FFD37C1DFF35E28F" blockId="7.[109,1215,1557,1699]" pageId="7" pageNumber="8">
FIGURE 3. Male of
<taxonomicName id="4C204D694D711450FE827C1FFDFFE201" authority="Engel &amp; Michener &amp; Boontop, 2017" authorityName="Engel &amp; Michener &amp; Boontop" authorityYear="2017" box="[316,557,1557,1581]" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="malaipanae" status="sp. nov.">
<emphasis id="B954EAF84D711450FE827C1FFDFFE201" box="[316,557,1557,1581]" italics="true" pageId="7" pageNumber="8">Tetragonula malaipanae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26757834D711450FD847C1CFD64E201" box="[570,694,1557,1581]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="B954EAF84D711450FD7C7C1FFD09E201" bold="true" box="[706,731,1557,1581]" pageId="7" pageNumber="8">A.</emphasis>
Outer view of metatibia and metabasitarsis.
<emphasis id="B954EAF84D711450FB167C1FFB6DE201" bold="true" box="[1192,1215,1557,1581]" pageId="7" pageNumber="8">B.</emphasis>
Dorsal view of T7 (arrow indicates the position of the spiracle on the side of the sclerite).
<emphasis id="B954EAF84D711450FBBA7C38FBCEE266" bold="true" box="[1028,1052,1586,1610]" pageId="7" pageNumber="8">C.</emphasis>
Ventral view of S5.
<emphasis id="B954EAF84D711450FF2C7C5AFF79E244" bold="true" box="[146,171,1616,1640]" pageId="7" pageNumber="8">D.</emphasis>
Ventral view of S6.
<emphasis id="B954EAF84D711450FEC07C5AFE40E244" bold="true" box="[382,402,1616,1640]" pageId="7" pageNumber="8">E.</emphasis>
Ventral view of S7.
<emphasis id="B954EAF84D711450FDDD7C5AFDA7E244" bold="true" box="[611,629,1616,1640]" pageId="7" pageNumber="8">F.</emphasis>
Male genitalia (dorsal view at left, ventral view at right), distal part of membranous aedeagus omitted.
<emphasis id="B954EAF84D711450FDFD7C67FD8EE2A9" bold="true" box="[579,604,1645,1669]" pageId="7" pageNumber="8">G.</emphasis>
Midventral membranous process.
<emphasis id="B954EAF84D711450FC027C67FC04E2A9" bold="true" box="[956,982,1645,1669]" pageId="7" pageNumber="8">H.</emphasis>
Male genitalia in right lateral view.
</paragraph>
</caption>
<paragraph id="8B9F36EA4D7E145FFF2278ABFD4FE1E7" blockId="8.[108,1214,225,1708]" pageId="8" pageNumber="9">
Pubescence decumbent and pallid except as otherwise noted, mostly ochraceous; clypeus without erect simple setae, even lower margin without such setae. Setae of face short (0.5 OD), plumose, rather dense but not hiding surface, particularly sparse and thin on lower paraocular area, whitish, those of frons [or upper half of frons] brownish to yellowish and hence inconspicuous in certain views; erect setae of scape absent except for exceedingly short ones; longer setae (1 OD) of vertex, especially behind ocelli, blackish or dusky; short pale setae of mesoscutum weakly [or not recognizably] banded. When recognizable, median asetose band (“G1” of
<bibRefCitation id="EFB14B1B4D7E145FFF3579ADFE91E7ED" author="Sakagami, S. F." box="[139,323,935,961]" pageId="8" pageNumber="9" pagination="165 - 247" refId="ref8490" refString="Sakagami, S. F. 1978. Tetragonula stingless bees of the continental Asia and Sri Lanka (Hymenoptera, Apidae). Journal of the Faculty of Science, Hokkaido University, Series 6, Zoology 21 (2): 165 - 247, + 1 pl. [pl. v]." type="journal article" year="1978">Sakagami, 1978</bibRefCitation>
) narrow, extending full length of mesoscutum except anterior extremity, G2 similarly narrow anteriorly but broadening and diverging posteriorly, G3 (lateral asetose band) poorly defined, especially on anterior half of mesoscutum [not recognizable except in some fresh, clean specimens]; longer, erect setae of mesoscutum (2 OD) pale or in certain illuminations dusky; fringe of long setae (up to 3 OD) on posterior margin of mesoscutellum ochraceous, with longest setae (34 OD) sometimes appearing dusky; metepisternum, side of propodeum, and anterior half of mesepisternum covered with dense white setae hiding surface [those of mesepisternum sometimes ochraceous]; some setae of outer surfaces of meso- and metatibiae and basitarsi dusky [commonly most setae of these parts, including large simple discal setae of corbicula, black or blackish]; anterior fringe of corbicula with a few [nearly all] setae dusky or blackish. T1 smooth and asetose; T2 to T5 smooth and asetose except for preapical very sparse bands, that of T2 consisting of only a few minute ochraceous setae, those of following terga with progressively more and longer setae, a few of those of T4 reaching beyond apical margin of tergum, those of T5 and T6 still longer, but all small and not conspicuous; setae of sterna denser and longer, almost white.
</paragraph>
<paragraph id="8B9F36EA4D7E145FFF227FDDFDD4E280" blockId="8.[108,1214,225,1708]" pageId="8" pageNumber="9">
<emphasis id="B954EAF84D7E145FFF227FDDFF32E1DD" bold="true" box="[156,224,1495,1521]" pageId="8" pageNumber="9">Male.</emphasis>
Agrees with description of worker except for variation noted above in brackets, in measurements provided, and as follows (except for measurements, description based on male collected from nest 4 on
<date id="FF9E102A4D7E145FFE327C2BFDA2E217" box="[396,624,1569,1595]" pageId="8" pageNumber="9" value="2007-09-17">17 September 2007</date>
): body length
<quantity id="4CD89B0F4D7E145FFCAA7C2BFCAFE210" box="[788,893,1569,1596]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="9" unit="mm" value="4.5">4.50 mm</quantity>
; forewing length, including tegula,
<quantity id="4CD89B0F4D7E145FFF7F7C4CFEF8E24D" box="[193,298,1606,1633]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" pageId="8" pageNumber="9" unit="mm" value="4.45">4.45 mm</quantity>
(
<quantity id="4CD89B0F4D7E145FFE877C4CFE3BE24D" box="[313,489,1606,1633]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="8" pageNumber="9" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.404.50 mm</quantity>
); head length (measured as for worker)
<quantity id="4CD89B0F4D7E145FFC7F7C4DFBF9E24D" box="[961,1067,1607,1633]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="9" unit="mm" value="1.4">1.40 mm</quantity>
; head width
<quantity id="4CD89B0F4D7E145FFFD37C66FF0FE2AB" box="[109,221,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="8" pageNumber="9" unit="mm" value="1.78">1.78 mm</quantity>
; metatibia length
<quantity id="4CD89B0F4D7E145FFE037C66FDFBE2AB" box="[445,553,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" pageId="8" pageNumber="9" unit="mm" value="1.46">1.46 mm</quantity>
(
<quantity id="4CD89B0F4D7E145FFD837C66FD23E2AB" box="[573,753,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="1.455" metricValueMax="1.48" metricValueMin="1.43" pageId="8" pageNumber="9" unit="mm" value="1.455" valueMax="1.48" valueMin="1.43">1.431.48 mm</quantity>
); forewing diagonal (measured as for worker)
<quantity id="4CD89B0F4D7E145FFF6F7C9BFEE8E280" box="[209,314,1681,1708]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="8" pageNumber="9" unit="mm" value="1.26">1.26 mm</quantity>
(
<quantity id="4CD89B0F4D7E145FFEF47C9BFE2BE280" box="[330,505,1681,1708]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" metricValueMax="1.27" metricValueMin="1.25" pageId="8" pageNumber="9" unit="mm" value="1.26" valueMax="1.27" valueMin="1.25">1.251.27 mm</quantity>
).
</paragraph>
<paragraph id="8B9F36EA4D7F145EFF227AEBFC4BE65F" blockId="9.[108,1213,225,1754]" pageId="9" pageNumber="10">Scape (excluding basal bulb) slightly less than half as long as compound eye. Flagellomere 4 (also 210) longer than broad. Ocelloccipital distance less than half OD; ocellocular distance about half of interocellar distance. Metatibia widest at about apical third, narrowed at apex, without plumose setae (fig. 3A). Metasomal T7 without Y-shaped ridge, apex broadly rounded (fig. 3B); S5 short, without noticeable gradulus, with shallow, broad, midapical emargination (fig. 3C); apical process of S6 slender, parallel sided (fig. 3D); terminalia as illustrated in figures 3D3H; unbroken, weakly sclerotized band in position of gonobase (fig. 3F); long membranous midventral genital process present (figs. 3G, 3H), presumably fragmentum from base of penis; penis valves and gonostyli directed posteriorly (figs. 3F, 3H); gonostylus arising near outer distal margin of gonocoxite (figs. 3F, 3H), flattened, so that in lateral view it is rather broad and subapically constricted (fig. 3H), in dorsal view very slender apically.</paragraph>
<paragraph id="8B9F36EA4D7F145EFF227874FCA6E779" blockId="9.[108,1213,225,1754]" pageId="9" pageNumber="10">Pale areas of head yellowish brown, upper median part of supraclypeal area black. Genal and hypostomal areas black. Flagellum black. Pronotum including pronotal lobe, mesoscutellum, and mesepisternum black. Legs yellowish brown, distal parts of femora, basal parts of tibiae, and distal parts of meso- and metatibia infuscated. Metasoma with T1 testaceous, with broad preapical dusky band; remaining terga cleared for study but apparently dusky testaceous (note that when dissected the metasoma appears pale testaceous).</paragraph>
<paragraph id="8B9F36EA4D7F145EFF22796AFE08E03C" blockId="9.[108,1213,225,1754]" pageId="9" pageNumber="10">Pubescence generally thinner and sparser than in female, particularly on clypeus, mesoscutellum, and mesepisternum; pubescence of head, mesosoma, and legs largely ochraceous. Mesoscutum almost without banding of setae; setae of mesepisternum and sides of propodeum white, somewhat less dense than in worker; dense setae of anterior part of mesepisternum dusky.</paragraph>
</subSubSection>
<subSubSection id="C33A65614D7F145DFF227E16FB6EE6CD" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B9F36EA4D7F145EFF227E16FBF6E0AD" blockId="9.[108,1213,225,1754]" pageId="9" pageNumber="10">
<materialsCitation id="3B483CB74D7F145EFF227E16FBF6E0AD" ID-GBIF-Occurrence="3312970301" collectingDate="2007-09-17" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="2" stateProvince="Kanchanaburi" typeStatus="holotype">
<typeStatus id="549B88484D7F145EFF227E16FECEE01A" box="[156,284,1052,1078]" pageId="9" pageNumber="10" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="F337767A4D7F145EFEF57E11FE67E019" box="[331,437,1051,1077]" name="Thailand" pageId="9" pageNumber="10">Thailand</collectingCountry>
,
<collectingRegion id="49E4F8084D7F145EFE7E7E11FDB4E019" box="[448,614,1051,1077]" country="Thailand" name="Kanchanaburi" pageId="9" pageNumber="10">Kanchanaburi</collectingRegion>
[Province],
<location id="8EFF60314D7F145EFD487E11FC10E019" LSID="urn:lsid:plazi:treatment:038987FC4D72145DFEE57B81FED4E177:8EFF60314D7F145EFD487E11FC10E019" box="[758,962,1051,1077]" country="Thailand" latitude="14.42" longLatPrecision="773" longitude="98.28" name="Thong Pha Phum" pageId="9" pageNumber="10" stateProvince="Kanchanaburi">Thong Pha Phum</location>
[District],
<date id="FF9E102A4D7F145EFBFA7E11FB6EE019" box="[1092,1212,1051,1077]" pageId="9" pageNumber="10" value="2007-09-17">
<collectingDate id="EFDAE9C24D7F145EFBFA7E11FB6EE019" box="[1092,1212,1051,1077]" pageId="9" pageNumber="10" value="2007-09-17">17.ix.2007</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFFCA7E4BFEB4E077" box="[116,358,1088,1115]" pageId="9" pageNumber="10" value="2007-09-17">
<collectingDate id="EFDAE9C24D7F145EFFCA7E4BFEB4E077" box="[116,358,1088,1115]" pageId="9" pageNumber="10" value="2007-09-17">17 September 2007</collectingDate>
</date>
],
<collectorName id="26D5533C4D7F145EFEC27E4BFDD4E077" box="[380,518,1089,1115]" pageId="9" pageNumber="10">Y. Boontop</collectorName>
, nest 2 (
<collectionCode id="ED31AE2F4D7F145EFDCF7E4BFD17E077" box="[625,709,1089,1115]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
).
<collectorName id="26D5533C4D7F145EFD657E4BFCD5E077" box="[731,775,1089,1115]" pageId="9" pageNumber="10">The</collectorName>
<typeStatus id="549B88484D7F145EFCAC7E4BFCA8E077" box="[786,890,1089,1115]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
is from nest number
<locationDeviation id="9A2D2FC84D7F145EFB327E4BFF0CE0AD" location="Boontop's" pageId="9" pageNumber="10">
<quantity id="4CD89B0F4D7F145EFB327E4BFB6EE077" box="[1164,1212,1089,1115]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="10" unit="in" value="2.0">2 in</quantity>
Boontops
</locationDeviation>
records, a site
<elevation id="000DD1D94D7F145EFE2E7E6CFD42E0AC" box="[400,656,1126,1153]" metricMagnitude="2" metricUnit="m" metricValue="5.49" pageId="9" pageNumber="10" unit="m" value="549.0">
<quantity id="4CD89B0F4D7F145EFE2E7E6CFE08E0AD" box="[400,474,1126,1153]" metricMagnitude="2" metricUnit="m" metricValue="5.49" pageId="9" pageNumber="10" unit="m" value="549.0">549 m</quantity>
above sea level
</elevation>
at coordinates
<geoCoordinate id="EE14502D4D7F145EFCF87E6DFC7DE0AD" box="[838,943,1127,1153]" degrees="14.42" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="555" value="14.42">14.42° N</geoCoordinate>
,
<geoCoordinate id="EE14502D4D7F145EFC057E6DFBF2E0AD" box="[955,1056,1127,1153]" degrees="98.28" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="555" value="98.28">98.28° E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9F36EA4D7F145DFF237E86FB6EE6CD" blockId="9.[108,1213,225,1754]" lastBlockId="10.[108,1214,225,1371]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<typeStatus id="549B88484D7F145EFF237E86FEF2E08A" box="[157,288,1164,1190]" pageId="9" pageNumber="10" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="9D26FD634D7F145EFE8F7E86FEBDE08B" box="[305,367,1164,1191]" pageId="9" pageNumber="10" type="male">2♂♂</specimenCount>
, 340⚲⚲):
<materialsCitation id="3B483CB74D7F145EFE527E86FBEFE08A" ID-GBIF-Occurrence="3312970323" box="[492,1085,1164,1191]" collectingDate="2007-09-17" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="paratype">
<specimenCount id="9D26FD634D7F145EFE527E86FDC3E08B" box="[492,529,1164,1191]" pageId="9" pageNumber="10" type="male">1♂</specimenCount>
, same data as holotype except nest 4 (
<collectionCode id="ED31AE2F4D7F145EFC5C7E86FBE6E08A" box="[994,1076,1164,1190]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFBF47E86FC15E0E0" ID-GBIF-Occurrence="3312970314" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="paratype">
<specimenCount id="9D26FD634D7F145EFBF47E86FBA2E08B" box="[1098,1136,1164,1191]" pageId="9" pageNumber="10" type="male">1♂</specimenCount>
, same data as holotype except
<date id="FF9E102A4D7F145EFE397EB8FDD2E0E0" box="[391,512,1202,1228]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFE397EB8FDD2E0E0" box="[391,512,1202,1228]" pageId="9" pageNumber="10" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFDAE7EB8FD15E0E0" box="[528,711,1201,1228]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFDAE7EB8FD15E0E0" box="[528,711,1201,1228]" pageId="9" pageNumber="10" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 11 (
<collectionCode id="ED31AE2F4D7F145EFCD37EB8FC6DE0E0" box="[877,959,1202,1228]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFC6A7EB8FF04E13B" ID-GBIF-Occurrence="3312970334" collectingDate="2008-03-12" collectionCode="SEMC, CSCA, DZUP, EMBT, ICNC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
24⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFE937EDDFE74E0DD" box="[301,422,1239,1265]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFE937EDDFE74E0DD" box="[301,422,1239,1265]" pageId="9" pageNumber="10" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFE0B7EDDFDB9E0DE" box="[437,619,1239,1266]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFE0B7EDDFDB9E0DE" box="[437,619,1239,1266]" pageId="9" pageNumber="10" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 1 (16
<collectionCode id="ED31AE2F4D7F145EFC9B7EDDFCA1E0DD" box="[805,883,1239,1265]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFC2D7EDDFC0DE0DD" box="[915,991,1239,1265]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="9" pageNumber="10" type="Museum">CSCA</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFC417ED2FB99E0DE" box="[1023,1099,1240,1266]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="9" pageNumber="10" type="University or college">DZUP</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFBD27ED2FB68E0DE" box="[1132,1210,1240,1266]" country="Thailand" httpUri="http://grbio.org/cool/01w0-hzbr" name="Department of Agriculture, Thailand" pageId="9" pageNumber="10">EMBT</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFF3D7EF7FF1CE13B" box="[131,206,1277,1303]" pageId="9" pageNumber="10">ICNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFF5D7EF7FB6AE13B" ID-GBIF-Occurrence="3312970320" box="[227,1208,1276,1303]" collectingDate="2009-07-27" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
11⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFD277EF7FCCAE13B" box="[665,792,1277,1303]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFD277EF7FCCAE13B" box="[665,792,1277,1303]" pageId="9" pageNumber="10" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFC967EF7FC10E13B" box="[808,962,1276,1303]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFC967EF7FC10E13B" box="[808,962,1276,1303]" pageId="9" pageNumber="10" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 1 (
<collectionCode id="ED31AE2F4D7F145EFBE37EF7FB7DE13B" box="[1117,1199,1277,1303]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFFD07F28FD02E111" ID-GBIF-Occurrence="3312970336" box="[110,720,1314,1341]" collectingDate="2007-09-17" collectionCode="SEMC, EMBT" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
18⚲⚲, same data as holotype (16
<collectionCode id="ED31AE2F4D7F145EFDBD7F28FD83E110" box="[515,593,1314,1340]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFDC87F29FD1BE111" box="[630,713,1315,1341]" country="Thailand" httpUri="http://grbio.org/cool/01w0-hzbr" name="Department of Agriculture, Thailand" pageId="9" pageNumber="10">EMBT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFD607F28FC89E14E" ID-GBIF-Occurrence="3312970312" collectingDate="2008-03-12" collectionCode="SEMC, AMNH, BBSL, CNIN" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
17⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFB277F28FF11E14E" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFB277F28FF11E14E" pageId="9" pageNumber="10" value="2008-03-12">12. iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFF6F7F42FE5AE14E" box="[209,392,1351,1378]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFF6F7F42FE5AE14E" box="[209,392,1351,1378]" pageId="9" pageNumber="10" value="2008-03-12">12 March 2008</collectingDate>
</date>
] (9
<collectionCode id="ED31AE2F4D7F145EFE0B7F42FDD1E14E" box="[437,515,1352,1378]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 4
<collectionCode id="ED31AE2F4D7F145EFD9A7F42FD53E14E" box="[548,641,1352,1378]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="10">AMNH</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFD1C7F42FD37E14E" box="[674,741,1352,1378]" country="USA" httpUri="http://grbio.org/cool/q49p-qw3v" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="9" pageNumber="10" type="Museum">BBSL</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFCB87F42FC81E14E" box="[774,851,1352,1378]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="9" pageNumber="10">CNIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFCD97F42FBE1E1A4" ID-GBIF-Occurrence="3312970339" collectingDate="2009-07-27" collectionCode="SEMC, NHML, SANC, USNM, EMBT" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
24⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFF037F67FEE9E1AB" box="[189,315,1389,1415]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFF037F67FEE9E1AB" box="[189,315,1389,1415]" pageId="9" pageNumber="10" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFEF67F67FE0EE1A4" box="[328,476,1389,1416]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFEF67F67FE0EE1A4" box="[328,476,1389,1416]" pageId="9" pageNumber="10" value="2009-07-27">27 July 2009</collectingDate>
</date>
] (16
<collectionCode id="ED31AE2F4D7F145EFDAB7F67FDB1E1AB" box="[533,611,1389,1415]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFD3C7F64FD08E1A4" box="[642,730,1390,1416]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="9" pageNumber="10">NHML</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFD477F67FC94E1AB" box="[761,838,1389,1415]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="9" pageNumber="10" type="Museum">SANC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFCDB7F67FC68E1AB" box="[869,954,1389,1415]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFC677F64FBF9E1A4" box="[985,1067,1390,1416]" country="Thailand" httpUri="http://grbio.org/cool/01w0-hzbr" name="Department of Agriculture, Thailand" pageId="9" pageNumber="10">EMBT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFB807F67FD96E181" ID-GBIF-Occurrence="3312970324" collectingDate="2007-09-17" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
8⚲⚲, same data as holotype except nest 3 (
<collectionCode id="ED31AE2F4D7F145EFE547F99FDEEE181" box="[490,572,1427,1453]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFDED7F99FE37E1FF" ID-GBIF-Occurrence="3312970331" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
13⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFBAC7F99FB59E181" box="[1042,1163,1427,1453]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFBAC7F99FB59E181" box="[1042,1163,1427,1453]" pageId="9" pageNumber="10" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFB237F99FF28E1FF" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFB237F99FF28E1FF" pageId="9" pageNumber="10" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 3 (
<collectionCode id="ED31AE2F4D7F145EFE327FB3FE0CE1FF" box="[396,478,1465,1491]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFE4E7FB3FD82E1D4" ID-GBIF-Occurrence="3312970313" collectingDate="2009-07-27" collectionCode="SEMC, ZMHB, ZISP" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
14⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFC287FB2FBC6E1FE" box="[918,1044,1464,1490]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFC287FB2FBC6E1FE" box="[918,1044,1464,1490]" pageId="9" pageNumber="10" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFB9C7FB3FB64E1FF" box="[1058,1206,1464,1491]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFB9C7FB3FB64E1FF" box="[1058,1206,1464,1491]" pageId="9" pageNumber="10" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 3 (10
<collectionCode id="ED31AE2F4D7F145EFEA17FD4FEBFE1D4" box="[287,365,1502,1528]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFE2F7FD5FE34E1D5" box="[401,486,1503,1529]" pageId="9" pageNumber="10">ZMHB</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFDB47FD4FD9AE1D4" box="[522,584,1502,1528]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="9" pageNumber="10">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFDE37FD4FB6AE1D4" ID-GBIF-Occurrence="3312970326" box="[605,1208,1502,1529]" collectingDate="2007-09-17" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
2⚲⚲, same data as holotype except nest 4 (
<collectionCode id="ED31AE2F4D7F145EFBE37FD4FB7DE1D4" box="[1117,1199,1502,1528]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFFD07C0EFF1BE26F" ID-GBIF-Occurrence="3312970322" collectingDate="2008-03-12" collectionCode="SEMC, KSMA" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
12⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFD8B7C09FD7DE231" box="[565,687,1539,1565]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFD8B7C09FD7DE231" box="[565,687,1539,1565]" pageId="9" pageNumber="10" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFD7C7C0EFC52E232" box="[706,896,1539,1566]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFD7C7C0EFC52E232" box="[706,896,1539,1566]" pageId="9" pageNumber="10" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 4 (10
<collectionCode id="ED31AE2F4D7F145EFBEE7C0EFB4DE232" box="[1104,1183,1540,1566]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFFD27C23FF13E26F" box="[108,193,1577,1603]" pageId="9" pageNumber="10">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFF687C23FB6AE26F" ID-GBIF-Occurrence="3312970325" box="[214,1208,1576,1604]" collectingDate="2009-07-27" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
11⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFD2D7C23FCC0E26F" box="[659,786,1577,1603]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFD2D7C23FCC0E26F" box="[659,786,1577,1603]" pageId="9" pageNumber="10" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFC9D7C23FC6CE26F" box="[803,958,1576,1603]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFC9D7C23FC6CE26F" box="[803,958,1576,1603]" pageId="9" pageNumber="10" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 4 (
<collectionCode id="ED31AE2F4D7F145EFBE07C23FB62E26F" box="[1118,1200,1577,1603]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFFD27C45FD1CE245" ID-GBIF-Occurrence="3312970318" box="[108,718,1614,1641]" collectingDate="2007-09-17" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
2⚲⚲, same data as holotype except nest 5 (
<collectionCode id="ED31AE2F4D7F145EFDCA7C45FD14E245" box="[628,710,1615,1641]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFD627C45FCD9E2A3" ID-GBIF-Occurrence="3312970315" collectingDate="2008-03-12" collectionCode="SEMC, EMBT" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
14⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFB277C45FF11E2A2" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFB277C45FF11E2A2" pageId="9" pageNumber="10" value="2008-03-12">12. iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFF6F7C7EFE55E2A3" box="[209,391,1652,1679]" pageId="9" pageNumber="10" value="2008-03-12">
<collectingDate id="EFDAE9C24D7F145EFF6F7C7EFE55E2A3" box="[209,391,1652,1679]" pageId="9" pageNumber="10" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 5 (11
<collectionCode id="ED31AE2F4D7F145EFDFF7C7EFD5DE2A2" box="[577,655,1652,1678]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 3
<collectionCode id="ED31AE2F4D7F145EFD0E7C7FFCD1E2A3" box="[688,771,1653,1679]" country="Thailand" httpUri="http://grbio.org/cool/01w0-hzbr" name="Department of Agriculture, Thailand" pageId="9" pageNumber="10">EMBT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFCA87C7EFC5DE298" ID-GBIF-Occurrence="3312970303" collectingDate="2009-07-27" collectionCode="SEMC, KUEC, EMBT" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
15⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFFD27C90FF39E298" box="[108,235,1690,1716]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFFD27C90FF39E298" box="[108,235,1690,1716]" pageId="9" pageNumber="10" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFF457C90FE41E298" box="[251,403,1689,1716]" pageId="9" pageNumber="10" value="2009-07-27">
<collectingDate id="EFDAE9C24D7F145EFF457C90FE41E298" box="[251,403,1689,1716]" pageId="9" pageNumber="10" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 5 (11
<collectionCode id="ED31AE2F4D7F145EFDEC7C90FD72E298" box="[594,672,1690,1716]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFD7C7C90FCC0E298" box="[706,786,1690,1716]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="9" pageNumber="10">KUEC</collectionCode>
, 2
<collectionCode id="ED31AE2F4D7F145EFC8A7C90FC55E298" box="[820,903,1690,1716]" country="Thailand" httpUri="http://grbio.org/cool/01w0-hzbr" name="Department of Agriculture, Thailand" pageId="9" pageNumber="10">EMBT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145EFC257C93FCC1E2F5" ID-GBIF-Occurrence="3312970316" collectingDate="2006-07-31" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
5⚲⚲, same data as holotype except
<date id="FF9E102A4D7F145EFF447CB5FEAAE2F5" box="[250,376,1727,1753]" pageId="9" pageNumber="10" value="2006-07-31">
<collectingDate id="EFDAE9C24D7F145EFF447CB5FEAAE2F5" box="[250,376,1727,1753]" pageId="9" pageNumber="10" value="2006-07-31">31.vii.2006</collectingDate>
</date>
[
<date id="FF9E102A4D7F145EFE367CB5FDF2E2F6" box="[392,544,1727,1754]" pageId="9" pageNumber="10" value="2006-07-31">
<collectingDate id="EFDAE9C24D7F145EFE367CB5FDF2E2F6" box="[392,544,1727,1754]" pageId="9" pageNumber="10" value="2006-07-31">31 July 2006</collectingDate>
</date>
] and nest 6 (
<collectionCode id="ED31AE2F4D7F145EFD077CB5FCD9E2F5" box="[697,779,1727,1753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="10" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7F145DFCA17CB5FEC8E4D7" ID-GBIF-Occurrence="3312970332" collectingDate="2007-09-17" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" lastPageId="10" lastPageNumber="11" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
9⚲⚲, same data as holotype except nest 6 (
<collectionCode id="ED31AE2F4D7C145DFF017AEBFEC3E4D7" box="[191,273,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFE9B7AEBFE5FE50D" ID-GBIF-Occurrence="3312970335" collectingDate="2008-03-12" collectionCode="SEMC, FSCA, LIPI" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
15⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFD717AEBFC9AE4D7" box="[719,840,225,251]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFD717AEBFC9AE4D7" box="[719,840,225,251]" pageId="10" pageNumber="11" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFCE87AEBFBD9E4D0" box="[854,1035,225,252]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFCE87AEBFBD9E4D0" box="[854,1035,225,252]" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 6 (13
<collectionCode id="ED31AE2F4D7C145DFFD27B0DFF68E50D" box="[108,186,263,289]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
, 1
<collectionCode id="ED31AE2F4D7C145DFF617B0DFEF5E50D" box="[223,295,263,289]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="10" pageNumber="11" type="Museum">FSCA</collectionCode>
, 1
<collectionCode id="ED31AE2F4D7C145DFEF27B0DFE57E50D" box="[332,389,263,289]" pageId="10" pageNumber="11">LIPI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFE247B0DFDDAE56B" ID-GBIF-Occurrence="3312970337" collectingDate="2009-07-27" collectionCode="SEMC, FSCA, LIPI" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
21⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFCEB7B0DFC06E50D" box="[853,980,263,289]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFCEB7B0DFC06E50D" box="[853,980,263,289]" pageId="10" pageNumber="11" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFC5B7B0DFB52E50D" box="[997,1152,262,289]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFC5B7B0DFB52E50D" box="[997,1152,262,289]" pageId="10" pageNumber="11" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 6 (19
<collectionCode id="ED31AE2F4D7C145DFF557B26FEEBE56A" box="[235,313,300,326]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
, 1
<collectionCode id="ED31AE2F4D7C145DFEE27B26FE76E56A" box="[348,420,300,326]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="10" pageNumber="11" type="Museum">FSCA</collectionCode>
, 1
<collectionCode id="ED31AE2F4D7C145DFE767B27FDD3E56B" box="[456,513,301,327]" pageId="10" pageNumber="11">LIPI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFDA87B26FE4FE547" ID-GBIF-Occurrence="3312970327" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
5⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFC007B26FBE5E56A" box="[958,1079,300,326]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFC007B26FBE5E56A" box="[958,1079,300,326]" pageId="10" pageNumber="11" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFBF67B26FF78E540" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFBF67B26FF78E540" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 7 (
<collectionCode id="ED31AE2F4D7C145DFEFD7B5BFE47E547" box="[323,405,337,363]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFE147B5BFEF1E5BD" ID-GBIF-Occurrence="3312970333" collectingDate="2009-07-27" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
12⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFCE57B5BFC0BE547" box="[859,985,337,363]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFCE57B5BFC0BE547" box="[859,985,337,363]" pageId="10" pageNumber="11" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFC577B5BFB53E540" box="[1001,1153,337,364]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFC577B5BFB53E540" box="[1001,1153,337,364]" pageId="10" pageNumber="11" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 7 (
<collectionCode id="ED31AE2F4D7C145DFF767B7DFEC9E5BD" box="[200,283,375,401]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFE8F7B7CFF1CE59A" ID-GBIF-Occurrence="3312970304" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
5⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFD5A7B7DFC8CE5BD" box="[740,862,375,401]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFD5A7B7DFC8CE5BD" box="[740,862,375,401]" pageId="10" pageNumber="11" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFCCE7B7DFBFEE5BD" box="[880,1068,374,401]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFCCE7B7DFBFEE5BD" box="[880,1068,374,401]" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 8 (
<collectionCode id="ED31AE2F4D7C145DFFCA7B96FF14E59A" box="[116,198,412,438]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFF637B96FB65E59A" ID-GBIF-Occurrence="3312970309" box="[221,1207,411,439]" collectingDate="2009-07-27" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
9⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFD317B96FCDCE59A" box="[655,782,412,438]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFD317B96FCDCE59A" box="[655,782,412,438]" pageId="10" pageNumber="11" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFCA17B96FC6EE59A" box="[799,956,411,438]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFCA17B96FC6EE59A" box="[799,956,411,438]" pageId="10" pageNumber="11" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 9 (
<collectionCode id="ED31AE2F4D7C145DFBE37B96FB7DE59A" box="[1117,1199,412,438]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFFD27BCBFBA2E5F7" ID-GBIF-Occurrence="3312970302" box="[108,1136,449,476]" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
5⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFDA07BCBFD4BE5F7" box="[542,665,449,475]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFDA07BCBFD4BE5F7" box="[542,665,449,475]" pageId="10" pageNumber="11" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFD147BCBFCB4E5F0" box="[682,870,449,476]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFD147BCBFCB4E5F0" box="[682,870,449,476]" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 10 (
<collectionCode id="ED31AE2F4D7C145DFBA87BCBFBBBE5F7" box="[1046,1129,449,475]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFBC17BCBFC3BE62D" ID-GBIF-Occurrence="3312970310" collectingDate="2009-07-27" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
4⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFE797BEDFD94E62D" box="[455,582,487,513]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFE797BEDFD94E62D" box="[455,582,487,513]" pageId="10" pageNumber="11" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFDEB7BEDFD39E62D" box="[597,747,486,513]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFDEB7BEDFD39E62D" box="[597,747,486,513]" pageId="10" pageNumber="11" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 10 (
<collectionCode id="ED31AE2F4D7C145DFC317BEDFC33E62D" box="[911,993,487,513]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFC4B7BEDFC5DE60A" ID-GBIF-Occurrence="3312970319" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
14⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFEF07806FE15E60A" box="[334,455,524,550]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFEF07806FE15E60A" box="[334,455,524,550]" pageId="10" pageNumber="11" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFE697806FD5CE60A" box="[471,654,523,550]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFE697806FD5CE60A" box="[471,654,523,550]" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 11 (
<collectionCode id="ED31AE2F4D7C145DFC8B7806FC55E60A" box="[821,903,524,550]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFC257806FCCBE667" ID-GBIF-Occurrence="3312970328" collectingDate="2009-07-27" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
3⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFF46783BFEA4E667" box="[248,374,561,587]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFF46783BFEA4E667" box="[248,374,561,587]" pageId="10" pageNumber="11" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFE3B783BFDCEE660" box="[389,540,561,588]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFE3B783BFDCEE660" box="[389,540,561,588]" pageId="10" pageNumber="11" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 11 (
<collectionCode id="ED31AE2F4D7C145DFD00783BFCC2E667" box="[702,784,561,587]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFC9A783BFD4FE65D" ID-GBIF-Occurrence="3312970307" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
6⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFFD2785DFF36E65D" box="[108,228,599,625]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFFD2785DFF36E65D" box="[108,228,599,625]" pageId="10" pageNumber="11" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFF4C785DFE76E65D" box="[242,420,598,625]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFF4C785DFE76E65D" box="[242,420,598,625]" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 12 (
<collectionCode id="ED31AE2F4D7C145DFDFA785DFD44E65D" box="[580,662,599,625]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFD16785DFDDFE6BA" ID-GBIF-Occurrence="3312970306" collectingDate="2009-07-27" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
3⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFB80785DFB6EE65D" box="[1086,1212,599,625]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFB80785DFB6EE65D" box="[1086,1212,599,625]" pageId="10" pageNumber="11" value="2009-07-27">27.vii.2009</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFFCD7876FEDEE6BA" box="[115,268,635,662]" pageId="10" pageNumber="11" value="2009-07-27">
<collectingDate id="EFDAE9C24D7C145DFFCD7876FEDEE6BA" box="[115,268,635,662]" pageId="10" pageNumber="11" value="2009-07-27">27 July 2009</collectingDate>
</date>
] and nest 12 (
<collectionCode id="ED31AE2F4D7C145DFE0D7876FDD7E6BA" box="[435,517,636,662]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFDA47876FE79E697" ID-GBIF-Occurrence="3312970338" collectingDate="2008-03-12" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
8⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFC017876FBEAE6BA" box="[959,1080,636,662]" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFC017876FBEAE6BA" box="[959,1080,636,662]" pageId="10" pageNumber="11" value="2008-03-12">12.iii.2008</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFBF67876FF78E690" pageId="10" pageNumber="11" value="2008-03-12">
<collectingDate id="EFDAE9C24D7C145DFBF67876FF78E690" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</collectingDate>
</date>
] and nest 13 (
<collectionCode id="ED31AE2F4D7C145DFEEE78ABFE70E697" box="[336,418,673,699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFE0978ABFEF9E6CD" ID-GBIF-Occurrence="3312970329" collectingDate="2006-07-31" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
2⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFCE578ABFC0BE697" box="[859,985,673,699]" pageId="10" pageNumber="11" value="2006-07-31">
<collectingDate id="EFDAE9C24D7C145DFCE578ABFC0BE697" box="[859,985,673,699]" pageId="10" pageNumber="11" value="2006-07-31">31.vii.2006</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFC5778ABFB53E690" box="[1001,1153,673,700]" pageId="10" pageNumber="11" value="2006-07-31">
<collectingDate id="EFDAE9C24D7C145DFC5778ABFB53E690" box="[1001,1153,673,700]" pageId="10" pageNumber="11" value="2006-07-31">31 July 2006</collectingDate>
</date>
] and nest 19 (
<collectionCode id="ED31AE2F4D7C145DFF6F78CDFEF1E6CD" box="[209,291,711,737]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CB74D7C145DFE8978CDFB6AE6CD" ID-GBIF-Occurrence="3312970311" box="[311,1208,710,737]" collectingDate="2007-09-17" collectionCode="SEMC" collectorName="Y. Boontop &amp; The" country="Thailand" elevation="549" latitude="14.42" location="Thong Pha Phum" longLatPrecision="773" longitude="98.28" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
2⚲⚲, same data as holotype except
<date id="FF9E102A4D7C145DFD6778CDFC83E6CD" box="[729,849,711,737]" pageId="10" pageNumber="11" value="2007-09-17">
<collectingDate id="EFDAE9C24D7C145DFD6778CDFC83E6CD" box="[729,849,711,737]" pageId="10" pageNumber="11" value="2007-09-17">17.ix.2007</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFCDE78CDFB95E6CD" box="[864,1095,710,737]" pageId="10" pageNumber="11" value="2007-09-17">
<collectingDate id="EFDAE9C24D7C145DFCDE78CDFB95E6CD" box="[864,1095,710,737]" pageId="10" pageNumber="11" value="2007-09-17">17 September 2007</collectingDate>
</date>
] (
<collectionCode id="ED31AE2F4D7C145DFBE378CDFB62E6CD" box="[1117,1200,711,737]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A65614D7C145DFF2278E6FEA0E7CA" pageId="10" pageNumber="11" type="description">
<paragraph id="8B9F36EA4D7C145DFF2278E6FEA0E7CA" blockId="10.[108,1214,225,1371]" pageId="10" pageNumber="11">
<materialsCitation id="3B483CB74D7C145DFF2278E6FD36E7B7" ID-GBIF-Occurrence="3312970330" collectingDate="2007-05-03" collectionCode="SEMC" collectorName="Y. Boontop" country="Thailand" elevation="359" latitude="8.91" location="Nakhonsithammarat" longLatPrecision="780" longitude="98.52" municipality="Khao Laung" pageId="10" pageNumber="11" specimenCount="1">
ADDITIONAL MATERIAL: A further six workers are remarkably similar to
<taxonomicName id="4C204D694D7C145DFBAD78E7FB6EE72A" authorityName="Engel &amp; Michener &amp; Boontop" authorityYear="2017" box="[1043,1212,748,775]" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="malaipanae">
<emphasis id="B954EAF84D7C145DFBAD78E7FBF1E72A" box="[1043,1059,749,774]" italics="true" pageId="10" pageNumber="11">T</emphasis>
.
<emphasis id="B954EAF84D7C145DFB8C78E6FB6EE72A" box="[1074,1212,748,774]" italics="true" pageId="10" pageNumber="11">malaipanae</emphasis>
</taxonomicName>
and we tentatively include them here, pending further work on these more southern populations, particularly utilizing morphometrics, males, and DNA data: 4⚲⚲,
<collectingCountry id="F337767A4D7C145DFC66793DFB96E77D" box="[984,1092,823,849]" name="Thailand" pageId="10" pageNumber="11">Thailand</collectingCountry>
,
<location id="8EFF60314D7C145DFBEF793DFED4E75A" LSID="urn:lsid:plazi:treatment:038987FC4D72145DFEE57B81FED4E177:8EFF60314D7C145DFBEF793DFED4E75A" country="Thailand" latitude="8.91" longLatPrecision="780" longitude="98.52" municipality="Khao Laung" name="Nakhonsithammarat" pageId="10" pageNumber="11">Nakhonsithammarat</location>
,
<collectingMunicipality id="6BFBAC904D7C145DFEAA7956FE79E75B" box="[276,427,860,887]" pageId="10" pageNumber="11">Khao Laung</collectingMunicipality>
,
<date id="FF9E102A4D7C145DFE077956FDC8E75A" box="[441,538,860,886]" pageId="10" pageNumber="11" value="2007-05-03">
<collectingDate id="EFDAE9C24D7C145DFE077956FDC8E75A" box="[441,538,860,886]" pageId="10" pageNumber="11" value="2007-05-03">3.v.2007</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFD927956FD10E75A" box="[556,706,859,887]" pageId="10" pageNumber="11" value="2007-05-03">
<collectingDate id="EFDAE9C24D7C145DFD927956FD10E75A" box="[556,706,859,887]" pageId="10" pageNumber="11" value="2007-05-03">3 May 2007</collectingDate>
</date>
],
<collectorName id="26D5533C4D7C145DFD697957FCB2E75B" box="[727,864,861,887]" pageId="10" pageNumber="11">Y. Boontop</collectorName>
, nest 11 [a site
<elevation id="000DD1D94D7C145DFB9F7956FF00E7B7" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="10" pageNumber="11" unit="m" value="359.0">
<quantity id="4CD89B0F4D7C145DFB9F7956FBBFE75B" box="[1057,1133,860,887]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="10" pageNumber="11" unit="m" value="359.0">359 m</quantity>
above sea level
</elevation>
at coordinates
<geoCoordinate id="EE14502D4D7C145DFE2D798BFE20E7B0" box="[403,498,897,924]" degrees="8.49" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="555" value="8.49">8.49° N</geoCoordinate>
,
<geoCoordinate id="EE14502D4D7C145DFDBF798BFDB9E7B0" box="[513,619,897,924]" degrees="99.78" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="555" value="99.78">99.78° E</geoCoordinate>
] (
<collectionCode id="ED31AE2F4D7C145DFD3B798BFD0BE7B7" box="[645,729,897,923]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B483CB74D7C145DFD51798BFEBCE7CA" ID-GBIF-Occurrence="3312970321" collectingDate="2007-05-03" collectionCode="SEMC" collectorName="Y. Boontop" country="Thailand" elevation="359" latitude="8.91" location="Nakhonsithammarat" longLatPrecision="780" longitude="98.52" municipality="Khao Laung" pageId="10" pageNumber="11" specimenCount="1">
2⚲⚲,
<collectingCountry id="F337767A4D7C145DFC86798BFC76E7B7" box="[824,932,897,923]" name="Thailand" pageId="10" pageNumber="11">Thailand</collectingCountry>
, Suratthani, Khao sok,
<date id="FF9E102A4D7C145DFFD279ADFF1FE7ED" box="[108,205,935,961]" pageId="10" pageNumber="11" value="2007-05-03">
<collectingDate id="EFDAE9C24D7C145DFFD279ADFF1FE7ED" box="[108,205,935,961]" pageId="10" pageNumber="11" value="2007-05-03">3.v.2007</collectingDate>
</date>
[
<date id="FF9E102A4D7C145DFF6079ADFEA2E7ED" box="[222,368,934,961]" pageId="10" pageNumber="11" value="2007-05-03">
<collectingDate id="EFDAE9C24D7C145DFF6079ADFEA2E7ED" box="[222,368,934,961]" pageId="10" pageNumber="11" value="2007-05-03">3 May 2007</collectingDate>
</date>
],
<collectorName id="26D5533C4D7C145DFE3A79ADFDD9E7ED" box="[388,523,935,961]" pageId="10" pageNumber="11">Y. Boontop</collectorName>
, nest 15 [a site
<elevation id="000DD1D94D7C145DFD7A79ADFC15E7ED" box="[708,967,935,961]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="10" pageNumber="11" unit="m" value="421.0">
<quantity id="4CD89B0F4D7C145DFD7A79ADFCDDE7ED" box="[708,783,935,961]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="10" pageNumber="11" unit="m" value="421.0">421 m</quantity>
above sea level
</elevation>
at coordinates
<geoCoordinate id="EE14502D4D7C145DFB3E79ADFF57E7CB" degrees="8.91" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="555" value="8.91">8.91° N</geoCoordinate>
,
<geoCoordinate id="EE14502D4D7C145DFF2C79C6FF2BE7CA" box="[146,249,971,999]" degrees="98.52" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="555" value="98.52">98.52° E</geoCoordinate>
] (
<collectionCode id="ED31AE2F4D7C145DFEAF79C6FEB7E7CA" box="[273,357,972,998]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A65614D7C145DFF2279FBFBC5E0C0" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8B9F36EA4D7C145DFF2279FBFBC5E0C0" blockId="10.[108,1214,225,1371]" pageId="10" pageNumber="11">
COMMENTS: Probably variation in coloration of males is more or less comparable to that of workers. The male taken
<date id="FF9E102A4D7C145DFE377E1DFDE6E01D" box="[393,564,1047,1073]" pageId="10" pageNumber="11" value="2008-03-12">12 March 2008</date>
at nest number 11 differs from the male described above as follows: integument of head and mesosoma (including clypeus and tegula) black; scape and pedicel reddish brown; legs dusky pale brown; meso- and metatarsi and distal parts of tibiae blackish, palest areas on outer and posterior surfaces of protibia and basal parts of meso- and metatibiae; metasoma reddish brown with vague transverse dusky areas, apical terga darker; membranous midventral genital process not evident (likely a result of damage during dissection).
</paragraph>
</subSubSection>
<subSubSection id="C33A65614D7C145DFF227EFDFED4E177" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8B9F36EA4D7C145DFF227EFDFED4E177" blockId="10.[108,1214,225,1371]" pageId="10" pageNumber="11">
ETYMOLOGY: The specific epithet honors Savitree Malaipan of the Department of Agriculture, Kasetsart University,
<collectingRegion id="49E4F8084D7C145DFE737F16FDE9E11A" box="[461,571,1308,1334]" country="Thailand" name="Krung Thep Mahanakhon" pageId="10" pageNumber="11">Bangkok</collectingRegion>
, beloved mentor of Y.B. and a prominent Thai melittologist.
</paragraph>
</subSubSection>
</treatment>
</document>