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<mods:title id="C3C28D3B005375234964C40BDCA1C2F0">Five species of the family Cyproideidae (Crustacea: Amphipoda) from Japan, with the description of a new genus and two new species</mods:title>
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<treatment id="0388D03FFFDBFFE882DFF8A7FE5AF97C" ID-DOI="http://doi.org/10.5281/zenodo.6056311" ID-GBIF-Taxon="119330788" ID-Zenodo-Dep="6056311" LSID="urn:lsid:plazi:treatment:0388D03FFFDBFFE882DFF8A7FE5AF97C" httpUri="http://treatment.plazi.org/id/0388D03FFFDBFFE882DFF8A7FE5AF97C" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302">
<subSubSection id="C33B32A2FFDBFFEB82DFF8A7FE79FD98" pageId="1" pageNumber="302" type="reference_group">
<paragraph id="8B9E6129FFDBFFEB82DFF8A7FDC4FD47" blockId="1.[151,567,1812,1872]" box="[151,542,1812,1838]" pageId="1" pageNumber="302">
<heading id="D0D6D645FFDBFFEB82DFF8A7FDC4FD47" bold="true" box="[151,542,1812,1838]" fontSize="11" level="1" pageId="1" pageNumber="302" reason="1">
<emphasis id="B955BD3BFFDBFFEB82DFF8A7FDC4FD47" bold="true" box="[151,542,1812,1838]" pageId="1" pageNumber="302">
Genus
<taxonomicName id="4C211AAAFFDBFFEB82A6F8A7FDC4FD47" ID-CoL="7NV8P" authority="Haswell, 1880" authorityName="Haswell" authorityYear="1880" box="[238,542,1812,1838]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B955BD3BFFDBFFEB82A6F8A7FEB0FD47" bold="true" box="[238,362,1812,1838]" italics="true" pageId="1" pageNumber="302">Cyproidea</emphasis>
<bibRefCitation id="EFB01CD8FFDBFFEB8339F8A7FDC4FD47" author="Haswell" box="[369,542,1812,1838]" pageId="1" pageNumber="331" refString="Haswell, W. A. (1880) Preliminary report on the Australian Amphipoda. Annals and Magazine of Natural History, Series 5, 5, 30 - 34. http: // dx. doi. org / 10.1080 / 00222938009459377" type="journal article" year="1880">Haswell, 1880</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9E6129FFDBFFEB82DFF88BFE53FDDA" blockId="1.[151,567,1812,1872]" lastBlockId="1.[151,1436,1919,2033]" pageId="1" pageNumber="302">
[Japanese name: Hoteiyokoebi-zoku]
<taxonomicName id="4C211AAAFFDBFFEB82DFF8CCFE0FFDFC" ID-CoL="7NV8P" authority="Haswell, 1880: 223" authorityName="Haswell" authorityPageNumber="223" authorityYear="1880" box="[151,469,1919,1941]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B955BD3BFFDBFFEB82DFF8CCFEDBFDFC" box="[151,257,1919,1941]" italics="true" pageId="1" pageNumber="302">Cyproidea</emphasis>
Haswell, 1880: 223
</taxonomicName>
.—Stebbing, 1906: 157.—Barnard, 1969: 139.—Barnard &amp; Karaman, 1991: 251.—Lowry &amp; Stoddart, 2003: 95.
</paragraph>
<paragraph id="8B9E6129FFDBFFEB82DFF80FFDD9FDBB" blockId="1.[151,1436,1919,2033]" box="[151,515,1980,2002]" pageId="1" pageNumber="302">
<taxonomicName id="4C211AAAFFDBFFEB82DFF80FFF27FDBB" ID-CoL="e0161d0d-be6b-483c-8294-f618f2c6fcd3" box="[151,253,1980,2002]" class="Malacostraca" family="Cyproideidae" genus="Cyproidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B955BD3BFFDBFFEB82DFF80FFF27FDBB" box="[151,253,1980,2002]" italics="true" pageId="1" pageNumber="302">Cyproidia</emphasis>
</taxonomicName>
[sic] Haswell, 1879: 320.
</paragraph>
<paragraph id="8B9E6129FFDBFFEB82DFF868FE79FD98" blockId="1.[151,1436,1919,2033]" box="[151,419,2011,2033]" pageId="1" pageNumber="302">
<taxonomicName id="4C211AAAFFDBFFEB82DFF868FE45FD98" authority="Walker, 1904: 256" authorityName="Walker" authorityPageNumber="256" authorityYear="1904" box="[151,415,2011,2033]" class="Malacostraca" family="Cyproideidae" genus="Gallea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B955BD3BFFDBFFEB82DFF868FF06FD98" box="[151,220,2011,2033]" italics="true" pageId="1" pageNumber="302">Gallea</emphasis>
Walker, 1904: 256
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B32A2FFD8FFE882DFFF24FB1FFAD9" box="[151,1221,151,176]" pageId="2" pageNumber="303" type="nomenclature">
<paragraph id="8B9E6129FFD8FFE882DFFF24FB1FFAD9" blockId="2.[151,1437,151,789]" box="[151,1221,151,176]" pageId="2" pageNumber="303">
<emphasis id="B955BD3BFFD8FFE882DFFF24FEE8FAD9" bold="true" box="[151,306,151,176]" pageId="2" pageNumber="303">
<typeStatus id="549ADF8BFFD8FFE882DFFF24FF08FAD9" box="[151,210,151,176]" pageId="2" pageNumber="303">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C211AAAFFD8FFE88371FF24FE70FAD9" ID-CoL="e0161d0d-be6b-483c-8294-f618f2c6fcd3" box="[313,426,151,176]" class="Malacostraca" family="Cyproideidae" genus="Cyproidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B955BD3BFFD8FFE88371FF24FE70FAD9" box="[313,426,151,176]" italics="true" pageId="2" pageNumber="303">Cyproidia</emphasis>
</taxonomicName>
[sic]
<emphasis id="B955BD3BFFD8FFE883A1FF2AFDEEFAD9" box="[489,564,153,176]" italics="true" pageId="2" pageNumber="303">ornata</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE88073FF24FD06FAD9" author="Haswell" box="[571,732,151,176]" pageId="2" pageNumber="331" refString="Haswell, W. A. (1879) On some additional new genera and species of amphipodous crustaceans. Proceedings of the Linnean Society of New South Wales, 4, 319 - 350, pls. 18 - 24. http: // dx. doi. org / 10.5962 / bhl. part. 22854" type="journal article" year="1879">Haswell, 1879</bibRefCitation>
by subsequent designation (
<bibRefCitation id="EFB01CD8FFD8FFE88651FF24FB62FAD9" author="Barnard" box="[1049,1208,151,176]" pageId="2" pageNumber="330" refString="Barnard, J. L. (1969) The families and genera of marine gammaridean Amphipoda. Bulletin of the United States National Museum, 271, 1 - 535." type="journal article" year="1969">Barnard 1969</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33B32A2FFD8FFE8828FFF0FFE5AF97C" pageId="2" pageNumber="303" type="description">
<paragraph id="8B9E6129FFD8FFE8828FFF0FFBB9F8C1" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="303">
<emphasis id="B955BD3BFFD8FFE8828FFF0FFE6FFABC" bold="true" box="[199,437,188,213]" pageId="2" pageNumber="303">Emended diagnosis.</emphasis>
<emphasis id="B955BD3BFFD8FFE883F3FF0FFE2FFABC" box="[443,501,188,213]" italics="true" pageId="2" pageNumber="303">Body</emphasis>
subglobular in lateral view but flattened laterally.
<emphasis id="B955BD3BFFD8FFE88660FF0EFAE1FABC" box="[1064,1339,188,213]" italics="true" pageId="2" pageNumber="303">Pereonites and pleonites</emphasis>
smooth, with pereonites 34 enlarged, pleonite 3 unproduced posterodorsally.
<emphasis id="B955BD3BFFD8FFE88186FF52FB94FA91" box="[974,1102,225,248]" italics="true" pageId="2" pageNumber="303">Urosomites</emphasis>
free, urosomite 1 unkeeled, posterodorsal end of urosomite 3 strongly projected.
<emphasis id="B955BD3BFFD8FFE880B9FEB7FCBFFB75" box="[753,869,260,284]" italics="true" pageId="2" pageNumber="303">Antenna 1</emphasis>
stout; peduncular article 2 dorsally crested, distal end distinctly toothed; primary flagellum short, with 69 articles.
<emphasis id="B955BD3BFFD8FFE881DDFE9BFBD1FB29" box="[917,1035,296,320]" italics="true" pageId="2" pageNumber="303">Antenna 2</emphasis>
slender; flagellum short, with 34 articles.
<emphasis id="B955BD3BFFD8FFE882BFFEFEFEB9FB0C" box="[247,355,332,357]" italics="true" pageId="2" pageNumber="303">Upper lip</emphasis>
incised, asymmetrical.
<emphasis id="B955BD3BFFD8FFE88039FEFFFD06FB0C" box="[625,732,332,357]" italics="true" pageId="2" pageNumber="303">Mandible</emphasis>
, molar reduced; palp slender, with 3 articles.
<emphasis id="B955BD3BFFD8FFE886AEFEFFFA89FB0D" box="[1254,1363,332,357]" italics="true" pageId="2" pageNumber="303">Maxilla 1</emphasis>
, inner plate medium-sized to small; outer plate with 68 distal robust setae; palp 1-articulated, with several robust setae apically.
<emphasis id="B955BD3BFFD8FFE882B4FE27FEAFFBC4" box="[252,373,404,429]" italics="true" pageId="2" pageNumber="303">Maxilliped</emphasis>
, outer plate feeble; palp stout, consisting of 4 articles, article 3 with small apicodistal process.
<emphasis id="B955BD3BFFD8FFE882DFFE0BFF05FBA6" box="[151,223,440,463]" italics="true" pageId="2" pageNumber="303">Coxae</emphasis>
12 very small, hidden by coxa 3, coxae 34 immensely broadened, coxa 5 short and wide, coxae 67 also very small; gills present on coxae 26, oostegites of female present on coxae 25.
<emphasis id="B955BD3BFFD8FFE88675FE6FFB61FB9C" box="[1085,1211,476,501]" italics="true" pageId="2" pageNumber="303">Gnathopod</emphasis>
1 large, subchelate, propodus large, palm very oblique, dactylus narrow; gnathopod 2 large, subchelate (or carpochelate), posterodistal corner of carpus strongly protruded, palm almost transverse, dactylus narrow.
<emphasis id="B955BD3BFFD8FFE88668FD97FB4DF854" box="[1056,1175,548,573]" italics="true" pageId="2" pageNumber="303">Pereopods</emphasis>
37 elongate, slender, bases of pereopods 57 rectolinear.
<emphasis id="B955BD3BFFD8FFE88064FDF4FD1CF809" box="[556,710,583,608]" italics="true" pageId="2" pageNumber="303">Pleopods 13</emphasis>
, peduncles long, each with 2 coupling hooks; inner rami almost same length as outer.
<emphasis id="B955BD3BFFD8FFE883C5FDDFFE34F8EC" box="[397,494,620,645]" italics="true" pageId="2" pageNumber="303">Uropods</emphasis>
biramous, slender; both rami of uropods 12 extending subequally, inner ramus of uropod 3 longer than outer.
<emphasis id="B955BD3BFFD8FFE88386FD3CFDCEF8C1" box="[462,532,655,680]" italics="true" pageId="2" pageNumber="303">Telson</emphasis>
entire, short, reaching middle of uropod 3 peduncle.
</paragraph>
<paragraph id="8B9E6129FFD8FFE8828FFD07FE5AF97C" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="303">
<emphasis id="B955BD3BFFD8FFE8828FFD07FEA8F8A4" bold="true" box="[199,370,692,717]" pageId="2" pageNumber="303">Included taxa.</emphasis>
<taxonomicName id="4C211AAAFFD8FFE88330FD07FD33F8A5" authority="Haswell, 1879" authorityName="Haswell" authorityYear="1879" box="[376,745,692,717]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="ornate">
<emphasis id="B955BD3BFFD8FFE88330FD07FDE7F8A5" box="[376,573,692,717]" italics="true" pageId="2" pageNumber="303">Cyproidea ornate</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE8800BFD07FD33F8A5" author="Haswell" box="[579,745,692,717]" pageId="2" pageNumber="331" refString="Haswell, W. A. (1879) On some additional new genera and species of amphipodous crustaceans. Proceedings of the Linnean Society of New South Wales, 4, 319 - 350, pls. 18 - 24. http: // dx. doi. org / 10.5962 / bhl. part. 22854" type="journal article" year="1879">Haswell, 1879</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C211AAAFFD8FFE880BBFD06FB5FF8A5" authority="Schellenberg, 1938" authorityName="Schellenberg" authorityYear="1938" box="[755,1157,692,717]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B955BD3BFFD8FFE880BBFD06FC7EF8A4" box="[755,932,692,717]" italics="true" pageId="2" pageNumber="303">C. serratipalma</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE881E2FD07FB5FF8A5" author="Schellenberg" box="[938,1157,692,717]" pageId="2" pageNumber="331" refString="Schellenberg, A. (1938) Litorale Amphipoden des tropischen Pazifiks nach Sammlungen von Prof. Bock (Stockholm), Prof. Dahl + (Berlin) und Prof. Pietschmann (Wien). Kunglia Svenska Vetenskapsakademiens Handlingar, Series 3, 16 (6), 1 - 105." type="journal article" year="1938">Schellenberg, 1938</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C211AAAFFD8FFE886D8FD06FF09F899" authority="Hirayama, 1978" authorityName="Hirayama" authorityYear="1978" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="liodactyla">
<emphasis id="B955BD3BFFD8FFE886D8FD06FAFAF8A4" box="[1168,1312,692,717]" italics="true" pageId="2" pageNumber="303">C. liodactyla</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE8876EFD07FF09F899" author="Hirayama" pageId="2" pageNumber="331" refString="Hirayama, A. (1978) A new species of the amphipod genus Cyproides [sic] from Amakusa, Kyushu. Publications from the Amakusa Marine Biological Laboratory, 4, 245 - 251." type="journal article" year="1978">Hirayama, 1978</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C211AAAFFD8FFE882A8FD6BFDF8F899" authority="Moore, 1981" authorityName="Moore" authorityYear="1981" box="[224,546,728,752]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis id="B955BD3BFFD8FFE882A8FD6BFE58F899" box="[224,386,728,752]" italics="true" pageId="2" pageNumber="303">C. marmorata</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE883C4FD6BFDF8F899" author="Moore" box="[396,546,728,752]" pageId="2" pageNumber="331" refString="Moore, P. G. (1981) Marine Amphipoda (Crustacea) new to science from the Tasmanian phytal fauna. Journal of Natural History, 15, 939 - 964. http: // dx. doi. org / 10.1080 / 00222938100770701" type="journal article" year="1981">Moore, 1981</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C211AAAFFD8FFE88078FD6BFCF6F899" authority="Ren, 2006" authorityName="Ren" authorityYear="2006" box="[560,812,727,752]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B955BD3BFFD8FFE88078FD6BFD72F899" box="[560,680,727,752]" italics="true" pageId="2" pageNumber="303">C. robusta</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE880FAFD6BFCF6F899" author="Ren" box="[690,812,727,752]" pageId="2" pageNumber="331" refString="Ren, X. (2006) Fauna Sinica, Invertebrata Vol. 41. Crustacea: Amphipoda: Gammaridea (I). Science Press, Beijing, 588 pp. [in Chinese with English abstract and descriptions of new species]" type="book" year="2006">Ren, 2006</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C211AAAFFD8FFE88171FD6BFB9BF899" authority="Azman, 2009" authorityName="Azman" authorityYear="2009" box="[825,1089,727,752]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="cobia">
<emphasis id="B955BD3BFFD8FFE88171FD6BFC41F899" box="[825,923,727,752]" italics="true" pageId="2" pageNumber="303">C. cobia</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE881EDFD6BFB9BF899" author="Azman" box="[933,1089,728,752]" pageId="2" pageNumber="330" refString="Azman, B. A. R. (2009) Cyproideidae. In: Lowry, J. K. &amp; Myers, A. A. (Eds.), Benthic Amphipoda of the Great Barrier Reef, Australia. Zootaxa, 2260, 380 - 392." type="journal article" year="2009">Azman, 2009</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C211AAAFFD8FFE88618FD6BFAAEF899" authority="Myers, 2012" authorityName="Myers" authorityYear="2012" box="[1104,1396,728,752]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="excavata">
<emphasis id="B955BD3BFFD8FFE88618FD6BFB0DF899" box="[1104,1239,728,752]" italics="true" pageId="2" pageNumber="303">C. excavata</emphasis>
<bibRefCitation id="EFB01CD8FFD8FFE886A8FD6BFAAEF899" author="Myers" box="[1248,1396,728,752]" pageId="2" pageNumber="331" refString="Myers, A. A. (2012) Amphipoda (Crustacea) from Palau, Micronesia: families Ampeliscidae, Ampithoidae, Aoridae, Colomastigidae and Cyproideidae. ZooKeys, 193, 1 - 25. http: // dx. doi. org / 10.3897 / zookeys. 193.3109" type="journal article" year="2012">Myers, 2012</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C211AAAFFD8FFE887CAFD6BFEFAF97C" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="okinawensis" status="sp. nov.">
<emphasis id="B955BD3BFFD8FFE887CAFD6BFEFAF97C" italics="true" pageId="2" pageNumber="303">C. okinawensis</emphasis>
</taxonomicName>
<emphasis id="B955BD3BFFD8FFE8836FFD4FFE5AF97C" bold="true" box="[295,384,764,789]" pageId="2" pageNumber="303">
<taxonomicNameLabel id="A2660040FFD8FFE8836FFD4FFE5AF97C" box="[295,384,764,789]" pageId="2" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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</treatment>
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